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<document id="568C045AD59CD53CCB6DA6DB243803C7" ID-CLB-Dataset="9157" ID-DOI="10.11646/zootaxa.4838.1.8" ID-GBIF-Dataset="84867756-fb8e-44e0-9dc2-0472215c147f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403757" ID-ZooBank="8ED941B5-EB3D-45BB-8307-E5D6836DFF95" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609352119797" checkinUser="plazi" docAuthor="Agnihotri, Priya, Chandra, Kajal, Shukla, Anumeha, Singh, Hukam &amp; Mehrotra, Rakesh C." docDate="2020" docId="713D87D9FFC0FFACFF24F954061CFDFD" docLanguage="en" docName="zootaxa.4838.1.8.pdf" docOrigin="Zootaxa 4838 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Teloganella gurhaensis Agnihotri &amp; Chandra &amp; Shukla &amp; Singh &amp; Mehrotra 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="141" masterDocId="8D04FFA1FFC1FFA8FFB3FF84055BFF9C" masterDocTitle="First fossil record of a nymph (Ephemeroptera, Teloganellidae) from the Indian subcontinent" masterLastPageNumber="142" masterPageNumber="137" pageNumber="138" updateTime="1698850127703" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="636780E1D5F44B8DBFD58301F1F2909D">First fossil record of a nymph (Ephemeroptera, Teloganellidae) from the Indian subcontinent</mods:title>
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<mods:namePart id="6B9B9C81FBE3AD463EF6F8BC7E1B8AAD">Agnihotri, Priya</mods:namePart>
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<mods:affiliation id="FB3032B486564224AF421814D4F5E8AE">Birbal Sahni Institute of Palaeosciences, 53 University Road, Lucknow- 226007, India &amp; agnihotripriya 18 @ yahoo. com; https: // orcid. org / 0000 - 0002 - 2207 - 5578</mods:affiliation>
<mods:nameIdentifier id="2269F0D0EA9E707DEA42FD5843F96549" type="email">agnihotripriya18@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="0525C6B924757E1E52C660FD4AE3FFAE">Chandra, Kajal</mods:namePart>
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<mods:affiliation id="993C62074F6E775A0B331BA8C5B292CB">Birbal Sahni Institute of Palaeosciences, 53 University Road, Lucknow- 226007, India &amp; kajalchandra 74 @ gmail. com; https: // orcid. org / 0000 - 0002 - 7137 - 4371</mods:affiliation>
<mods:nameIdentifier id="5A9C1A41E120ACA92FA5A3261B67E28E" type="email">kajalchandra74@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5E8F45C826616A2D4D58A1A52879481D">Shukla, Anumeha</mods:namePart>
<mods:nameIdentifier id="E7759951F833451BBFE69379820AC389" type="ORCID">0000-0001-5424-0032</mods:nameIdentifier>
<mods:affiliation id="0B8AA31D59E50A3E8B4C7C64FD002B26">Birbal Sahni Institute of Palaeosciences, 53 University Road, Lucknow- 226007, India &amp; anu _ bsip @ yahoo. co. in; https: // orcid. org / 0000 - 0001 - 5424 - 0032</mods:affiliation>
<mods:nameIdentifier id="6AF8E73288DE4EBA24D61F6FCDAE7815" type="email">anu_bsip@yahoo.co.in</mods:nameIdentifier>
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<mods:namePart id="B9D94951B28B0C7BA6624CFCBEE55F68">Singh, Hukam</mods:namePart>
<mods:nameIdentifier id="B329CF5D1BF031BF76EB8764F2F7CFB7" type="ORCID">0000-0002-5426-4891</mods:nameIdentifier>
<mods:affiliation id="79D0D51EF07ED04E0165A8E48F9C1CF1">Birbal Sahni Institute of Palaeosciences, 53 University Road, Lucknow- 226007, India &amp; hukams @ gmail. com; https: // orcid. org / 0000 - 0002 - 5426 - 4891</mods:affiliation>
<mods:nameIdentifier id="EAF7A18A19DD0A58553F861DAA0B69E7" type="email">hukams@gmail.com</mods:nameIdentifier>
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<mods:namePart id="EA2B37D39A35E43A2219FFFA0BECF9F9">Mehrotra, Rakesh C.</mods:namePart>
<mods:nameIdentifier id="50D0FEBBD1595757D5DB9D6A1F46EADC" type="ORCID">0000-0002-6654-8621</mods:nameIdentifier>
<mods:affiliation id="365E2CAD68879E08344DF14EAD68AFB1">Birbal Sahni Institute of Palaeosciences, 53 University Road, Lucknow- 226007, India &amp; rcmehrotra @ yahoo. com; https: // orcid. org / 0000 - 0002 - 6654 - 8621</mods:affiliation>
<mods:nameIdentifier id="3A92FBD77ABA21907E79329362000CE3" type="email">rcmehrotra@yahoo.com</mods:nameIdentifier>
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<subSubSection id="B18E6544FFC0FFA9FF24F954075EF977" box="[151,517,1744,1771]" pageId="1" pageNumber="138" type="nomenclature">
<paragraph id="F92B36CFFFC0FFA9FF24F954075EF977" blockId="1.[151,517,1744,1807]" box="[151,517,1744,1771]" pageId="1" pageNumber="138">
<heading id="A26381A3FFC0FFA9FF24F954075EF977" bold="true" box="[151,517,1744,1771]" fontSize="11" level="1" pageId="1" pageNumber="138" reason="1">
<emphasis id="CBE0EADDFFC0FFA9FF24F954075EF977" bold="true" box="[151,517,1744,1771]" pageId="1" pageNumber="138">
<taxonomicName id="3E944D4CFFC0FFA9FF24F95404FFF977" authority="Agnihotri &amp; Chandra &amp; Shukla &amp; Singh &amp; Mehrotra, 2020" authorityName="Agnihotri &amp; Chandra &amp; Shukla &amp; Singh &amp; Mehrotra" authorityYear="2020" box="[151,420,1744,1771]" class="Insecta" family="Teloganellidae" genus="Teloganella" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="gurhaensis" status="sp. nov.">
<emphasis id="CBE0EADDFFC0FFA9FF24F95404FFF977" bold="true" box="[151,420,1744,1771]" italics="true" pageId="1" pageNumber="138">Teloganella gurhaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0D357A6FFC0FFA9FE18F955075EF977" box="[427,517,1745,1771]" pageId="1" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="B18E6544FFC0FFA9FF24F9710450F893" box="[151,267,1781,1807]" pageId="1" pageNumber="138" type="description">
<paragraph id="F92B36CFFFC0FFA9FF24F9710450F893" blockId="1.[151,517,1744,1807]" box="[151,267,1781,1807]" pageId="1" pageNumber="138">
(
<figureCitation id="61AF2A4AFFC0FFA9FF2CF97105B3F893" box="[159,232,1781,1807]" captionStart="FIGURE 2" captionStartId="2.[151,250,1718,1742]" captionTargetBox="[151,1436,181,1694]" captionTargetId="figure-26@2.[151,1436,181,1694]" captionTargetPageId="2" captionText="FIGURE 2. AD. Teloganella gurhaensis Agnihotri et al., sp. nov. A. Fossil insect in reflected light. B. Enlarged view of the upper half showing fore-femora, mid-femora and mid-tibia. C. Enlarged head portion with setae (red arrows). D. Enlarged lower half with abdominal segments (red arrows) and terminal filaments (black arrows)." figureDoi="http://doi.org/10.5281/zenodo.4403763" httpUri="https://zenodo.org/record/4403763/files/figure.png" pageId="1" pageNumber="138">Figs 2</figureCitation>
,
<figureCitation id="61AF2A4AFFC0FFA9FF47F9710459F893" box="[244,258,1781,1807]" captionStart="FIGURE 3" captionStartId="3.[151,250,1982,2006]" captionTargetBox="[151,1436,273,1958]" captionTargetId="figure-71@3.[151,1436,273,1958]" captionTargetPageId="3" captionText="FIGURE 3. Numbered abdominal segments of Teloganella gurhaensis Agnihotri et al., sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4403765" httpUri="https://zenodo.org/record/4403765/files/figure.png" pageId="1" pageNumber="138">3</figureCitation>
)
</paragraph>
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<subSubSection id="B18E6544FFC0FFA9FF24F8B90750F803" pageId="1" pageNumber="138" type="diagnosis">
<paragraph id="F92B36CFFFC0FFA9FF24F8B90750F803" blockId="1.[151,1437,1853,2023]" pageId="1" pageNumber="138">
<emphasis id="CBE0EADDFFC0FFA9FF24F8B90449F8CB" bold="true" box="[151,274,1853,1879]" pageId="1" pageNumber="138">Diagnosis.</emphasis>
Body with defined proportions, flattened pronotum and mesonotum, eyes widely separated, fore-femora greatly expanded, claws without denticles, fore-femora margined with bristle-like setae, elongated abdomen, three caudal filaments well-developed.
</paragraph>
</subSubSection>
<subSubSection id="B18E6544FFC0FFA9FF74F82D0492F87B" pageId="1" pageNumber="138" type="materials_examined">
<paragraph id="F92B36CFFFC0FFA9FF74F82D0492F87B" blockId="1.[151,1437,1853,2023]" pageId="1" pageNumber="138">
<materialsCitation id="49FC3C92FFC0FFA9FF74F82D0492F87B" ID-GBIF-Occurrence="3006040301" county="Late Paleocene-early Eocene" location="Gurha" municipality="Palana Formation" pageId="1" pageNumber="138" specimenCount="1" stateProvince="Rajasthan" typeStatus="holotype">
<emphasis id="CBE0EADDFFC0FFA9FF74F82D0460F85F" bold="true" box="[199,315,1961,1987]" pageId="1" pageNumber="138">
<typeStatus id="262F886DFFC0FFA9FF74F82D046DF85F" box="[199,310,1961,1987]" pageId="1" pageNumber="138" type="holotype">Holotype</typeStatus>
.
</emphasis>
BSIP specimen no. 41813,
<collectingCounty id="104A4E43FFC0FFA9FD30F82E068AF858" box="[643,977,1961,1988]" pageId="1" pageNumber="138">Late Paleocene-early Eocene</collectingCounty>
,
<collectingMunicipality id="194FACB5FFC0FFA9FC53F82D01F6F85F" box="[992,1197,1961,1987]" pageId="1" pageNumber="138">Palana Formation</collectingMunicipality>
,
<location id="FC4B6014FFC0FFA9FB08F82D005AF85F" LSID="urn:lsid:plazi:treatment:713D87D9FFC0FFACFF24F954061CFDFD:FC4B6014FFC0FFA9FB08F82D005AF85F" box="[1211,1281,1961,1987]" county="Late Paleocene-early Eocene" municipality="Palana Formation" name="Gurha" pageId="1" pageNumber="138" stateProvince="Rajasthan">Gurha</location>
lignite mine,
<location id="FC4B6014FFC0FFA9FF24F8490413F87B" LSID="urn:lsid:plazi:treatment:713D87D9FFC0FFACFF24F954061CFDFD:FC4B6014FFC0FFA9FF24F8490413F87B" box="[151,328,1997,2023]" county="Late Paleocene-early Eocene" municipality="Palana Formation" name="Bikaner district" pageId="1" pageNumber="138" stateProvince="Rajasthan">Bikaner district</location>
,
<collectingRegion id="3B50F82DFFC0FFA9FEE0F849049EF87B" box="[339,453,1997,2023]" country="India" name="Rajasthan" pageId="1" pageNumber="138">Rajasthan</collectingRegion>
.
</materialsCitation>
</paragraph>
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<caption id="ADEB6647FFC3FFAAFF24F932068DF88B" ID-DOI="http://doi.org/10.5281/zenodo.4403763" ID-Zenodo-Dep="4403763" httpUri="https://zenodo.org/record/4403763/files/figure.png" pageId="2" pageNumber="139" startId="2.[151,250,1718,1742]" targetBox="[151,1436,181,1694]" targetPageId="2">
<paragraph id="F92B36CFFFC3FFAAFF24F932068DF88B" blockId="2.[151,1437,1718,1815]" pageId="2" pageNumber="139">
<emphasis id="CBE0EADDFFC3FFAAFF24F9320408F952" bold="true" box="[151,339,1718,1742]" pageId="2" pageNumber="139">FIGURE 2. AD.</emphasis>
<taxonomicName id="3E944D4CFFC3FFAAFEEAF93207B0F952" authority="Agnihotri et al." authorityName="Agnihotri" authorityYear="2020" box="[345,747,1718,1742]" class="Insecta" family="Teloganellidae" genus="Teloganella" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="gurhaensis" status="sp. nov.">
<emphasis id="CBE0EADDFFC3FFAAFEEAF932071DF952" box="[345,582,1718,1742]" italics="true" pageId="2" pageNumber="139">Teloganella gurhaensis</emphasis>
Agnihotri
<emphasis id="CBE0EADDFFC3FFAAFD05F93307B0F952" box="[694,747,1718,1742]" italics="true" pageId="2" pageNumber="139">et al.</emphasis>
</taxonomicName>
<emphasis id="CBE0EADDFFC3FFAAFD58F93207B4F952" box="[747,751,1718,1742]" italics="true" pageId="2" pageNumber="139">,</emphasis>
<emphasis id="CBE0EADDFFC3FFAAFD45F933063CF952" bold="true" box="[758,871,1718,1743]" pageId="2" pageNumber="139">
<taxonomicNameLabel id="D0D357A6FFC3FFAAFD45F933061CF953" box="[758,839,1719,1743]" pageId="2" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Fossil insect in reflected light.
<emphasis id="CBE0EADDFFC3FFAAFB1AF933019BF953" bold="true" box="[1193,1216,1719,1743]" pageId="2" pageNumber="139">B.</emphasis>
Enlarged view of the upper half showing fore-femora, mid-femora and mid-tibia.
<emphasis id="CBE0EADDFFC3FFAAFD41F95E0651F96E" bold="true" box="[754,778,1754,1778]" pageId="2" pageNumber="139">C.</emphasis>
Enlarged head portion with setae (red arrows).
<emphasis id="CBE0EADDFFC3FFAAFB57F95F01A6F96F" bold="true" box="[1252,1277,1755,1779]" pageId="2" pageNumber="139">D.</emphasis>
Enlarged lower half with abdominal segments (red arrows) and terminal filaments (black arrows).
</paragraph>
</caption>
<subSubSection id="B18E6544FFC3FFAAFF74F8C1005CF8C3" box="[199,1287,1861,1887]" pageId="2" pageNumber="139" type="etymology">
<paragraph id="F92B36CFFFC3FFAAFF74F8C1005CF8C3" blockId="2.[151,1436,1861,2037]" box="[199,1287,1861,1887]" pageId="2" pageNumber="139">
<emphasis id="CBE0EADDFFC3FFAAFF74F8C10413F8C3" bold="true" box="[199,328,1861,1887]" pageId="2" pageNumber="139">Etymology</emphasis>
. The specific name is given after the locality Gurha, where the fossil was discovered.
</paragraph>
</subSubSection>
<subSubSection id="B18E6544FFC3FFACFF74F8EC061CFDFD" lastPageId="4" lastPageNumber="141" pageId="2" pageNumber="139" type="description">
<paragraph id="F92B36CFFFC3FFABFF74F8EC018DFF65" blockId="2.[151,1436,1861,2037]" lastBlockId="3.[151,1436,151,249]" lastPageId="3" lastPageNumber="140" pageId="2" pageNumber="139">
<emphasis id="CBE0EADDFFC3FFAAFF74F8EC0402F81E" bold="true" box="[199,345,1896,1922]" pageId="2" pageNumber="139">Description.</emphasis>
Nymph impression in dorsal position; satisfactory preservation quality of the specimen; body
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in length; head
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long with lateral location of eyes and fringe of setae over the head capsule, antennae visible; prothorax
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divided into two lobes, hard sclerite impression over the dorsal surface; fore-femora wider than long (FF
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; FF
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), setae on the inner and outer margins of femur and tibia, single tarsal claw without denticles; mesothorax
<quantity id="3E6C9B2AFFC2FFABFE43FF130713FF2D" box="[496,584,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="140" unit="mm" value="1.4">1.4 mm</quantity>
, forewing pads slightly visible, left and right mid-tibia margined with setae; abdomen
<quantity id="3E6C9B2AFFC2FFABFEB6FF3F0406FF49" box="[261,349,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="140" unit="mm" value="3.4">3.4 mm</quantity>
, segments I to X visible,
<superScript id="0EE19B87FFC2FFABFDC9FF3F07C0FF54" attach="left" box="[634,667,186,213]" fontSize="6" pageId="3" pageNumber="140">Xth</superScript>
segment bearing 3 caudal filaments (cerci-
<quantity id="3E6C9B2AFFC2FFABFB35FF3F0190FF49" box="[1158,1227,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="140" unit="mm" value="2.0">2 mm</quantity>
), filaments 1/4
<superScript id="0EE19B87FFC2FFABFAC2FF3E0025FF54" attach="left" box="[1393,1406,186,200]" fontSize="6" pageId="3" pageNumber="140">th</superScript>
of the body length, median filament visible, slender with no spines; location of gills not discernible.
</paragraph>
<caption id="ADEB6647FFC2FFABFF24F83A013CF84A" ID-DOI="http://doi.org/10.5281/zenodo.4403765" ID-Zenodo-Dep="4403765" box="[151,1127,1982,2007]" httpUri="https://zenodo.org/record/4403765/files/figure.png" pageId="3" pageNumber="140" startId="3.[151,250,1982,2006]" targetBox="[151,1436,273,1958]" targetPageId="3">
<paragraph id="F92B36CFFFC2FFABFF24F83A013CF84A" blockId="3.[151,1127,1982,2007]" box="[151,1127,1982,2007]" pageId="3" pageNumber="140">
<emphasis id="CBE0EADDFFC2FFABFF24F83A0456F84A" bold="true" box="[151,269,1982,2006]" pageId="3" pageNumber="140">FIGURE 3</emphasis>
. Numbered abdominal segments of
<taxonomicName id="3E944D4CFFC2FFABFDCFF83A0151F84A" authority="Agnihotri et al." authorityName="Agnihotri" authorityYear="2020" box="[636,1034,1982,2006]" class="Insecta" family="Teloganellidae" genus="Teloganella" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="gurhaensis" status="sp. nov.">
<emphasis id="CBE0EADDFFC2FFABFDCFF83A0633F84A" box="[636,872,1982,2006]" italics="true" pageId="3" pageNumber="140">Teloganella gurhaensis</emphasis>
Agnihotri
<emphasis id="CBE0EADDFFC2FFABFC64F83B0151F84A" box="[983,1034,1982,2006]" italics="true" pageId="3" pageNumber="140">et al.</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D0D357A6FFC2FFABFBA5F83A013CF84A" box="[1046,1127,1982,2006]" pageId="3" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="F92B36CFFFC5FFACFF74FF120684FE4D" blockId="4.[151,1437,150,609]" pageId="4" pageNumber="141">
<emphasis id="CBE0EADDFFC5FFACFF74FF120466FF2C" bold="true" box="[199,317,150,176]" pageId="4" pageNumber="141">Affinities.</emphasis>
The fossil exhibits morphological similarities with modern
<taxonomicName id="3E944D4CFFC5FFACFC68FF130105FF2D" authorityName="Ulmer" authorityYear="1939" box="[987,1118,151,177]" class="Insecta" family="Teloganellidae" genus="Teloganella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="CBE0EADDFFC5FFACFC68FF130105FF2D" box="[987,1118,151,177]" italics="true" pageId="4" pageNumber="141">Teloganella</emphasis>
</taxonomicName>
studied from
<collectingCountry id="8183765FFFC5FFACFB4FFF13006DFF2D" box="[1276,1334,151,177]" name="India" pageId="4" pageNumber="141">India</collectingCountry>
and
<collectingCountry id="8183765FFFC5FFACFADDFF130587FF49" name="Malaysia" pageId="4" pageNumber="141">Malaysia</collectingCountry>
. Expanded fore-femora are considered a convergence trait that has evolved through diverse taxa of
<taxonomicName id="3E944D4CFFC5FFACFAF1FF3F00C7FF49" authorityName="McCafferty &amp; Edmunds" authorityYear="1979" box="[1346,1436,187,213]" class="Insecta" higherTaxonomySource="GBIF" infraOrder="Pannota" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="infraOrder">Pannota</taxonomicName>
(
<bibRefCitation id="9D054B3EFFC5FFACFF2DFF5B0497FF65" author="McCafferty, W. P. &amp; Wang, T. Q." box="[158,460,223,249]" pageId="4" pageNumber="141" pagination="9 - 101" refId="ref2412" refString="McCafferty, W. P. &amp; Wang, T. Q. (2000) Phylogenetic systematics of the major lineages of Pannote mayflies (Ephemeroptera, Pannota). Transactions of the American Entomological Society, 126, 9 - 101." type="journal article" year="2000">McCafferty &amp; Wang 2000</bibRefCitation>
) and also justifies its convergence with
<taxonomicName id="3E944D4CFFC5FFACFC25FF5B0161FF65" authorityName="Allen" authorityYear="1965" box="[918,1082,223,249]" class="Insecta" family="Teloganodidae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Teloganodidae</taxonomicName>
. Row of well-developed broad setae on the larval fore-femora distinguishes the
<taxonomicName id="3E944D4CFFC5FFACFD72FE87063EFE81" box="[705,869,259,285]" class="Insecta" family="Teloganellidae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Teloganellidae</taxonomicName>
from the other families of
<taxonomicName id="3E944D4CFFC5FFACFB28FE870587FEDD" authority="(Kluge 2004)" baseAuthorityName="Kluge" baseAuthorityYear="2004" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="order">
Ephemeroptera (
<bibRefCitation id="9D054B3EFFC5FFACFAE5FE87058FFEDD" author="Kluge, N. J." pageId="4" pageNumber="141" refId="ref2084" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, Dordrecht, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004</bibRefCitation>
)
</taxonomicName>
. This trait also suggests co-relation with the
<taxonomicName id="3E944D4CFFC5FFACFD52FEA30623FEDD" authorityName="Lestage" authorityYear="1942" box="[737,888,295,321]" class="Insecta" family="Tricorythidae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Tricorythidae</taxonomicName>
and is also considered as a homophyly (resemblance due to common ancestry) both in
<taxonomicName id="3E944D4CFFC5FFACFDD2FECF07E0FEF9" authorityName="McCafferty &amp; Edmunds" authorityYear="1979" box="[609,699,331,357]" class="Insecta" higherTaxonomySource="GBIF" infraOrder="Pannota" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="infraOrder">Pannota</taxonomicName>
and other mayflies under similar environmental conditions.
<taxonomicName id="3E944D4CFFC5FFACFAD1FECF0451FE15" class="Insecta" family="Teloganellidae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Teloganellidae</taxonomicName>
belongs to the superfamily
<taxonomicName id="3E944D4CFFC5FFACFDE0FEEB064AFE15" authorityName="Klapalek" authorityYear="1909" box="[595,785,367,393]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="superFamily" superFamily="Ephemerelloidea">Ephemerelloidea</taxonomicName>
which also encloses
<taxonomicName id="3E944D4CFFC5FFACFBB9FEEB01E6FE15" authorityName="Klapalek" authorityYear="1909" box="[1034,1213,367,393]" class="Insecta" family="Ephemerellidae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Ephemerellidae</taxonomicName>
,
<taxonomicName id="3E944D4CFFC5FFACFB78FEEB00C3FE15" box="[1227,1432,367,393]" class="Insecta" family="Melanemerellidae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Melanemerellidae</taxonomicName>
,
<taxonomicName id="3E944D4CFFC5FFACFF24FE170460FE31" authorityName="Allen" authorityYear="1965" box="[151,315,403,429]" class="Insecta" family="Teloganodidae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Teloganodidae</taxonomicName>
and
<taxonomicName id="3E944D4CFFC5FFACFEC1FE170756FE31" authorityName="Lestage" authorityYear="1942" box="[370,525,403,429]" class="Insecta" family="Tricorythidae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="family">Tricorythidae</taxonomicName>
; however, the affinities and the position of
<taxonomicName id="3E944D4CFFC5FFACFC42FE17012FFE31" authorityName="Ulmer" authorityYear="1939" box="[1009,1140,403,429]" class="Insecta" family="Teloganellidae" genus="Teloganella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="CBE0EADDFFC5FFACFC42FE17012FFE31" box="[1009,1140,403,429]" italics="true" pageId="4" pageNumber="141">Teloganella</emphasis>
</taxonomicName>
are still unsolved (
<bibRefCitation id="9D054B3EFFC5FFACFAFAFE1705ADFE4D" author="Sites, R. W. &amp; Wang, T. &amp; Permkam, S. &amp; Hubbard, M. D." pageId="4" pageNumber="141" pagination="243 - 268" refId="ref2858" refString="Sites, R. W., Wang, T., Permkam, S. &amp; Hubbard, M. D. (2001) The mayfly genera (Ephemeroptera) of southern Thailand. Natural History Bulletin of the Siam Society, 49, 243 - 268." type="journal article" year="2001">
Sites
<emphasis id="CBE0EADDFFC5FFACFA3BFE1705EFFE4D" italics="true" pageId="4" pageNumber="141">et al.</emphasis>
2001
</bibRefCitation>
,
<bibRefCitation id="9D054B3EFFC5FFACFEB2FE3304CAFE4D" author="Soldan, T." box="[257,401,439,465]" pageId="4" pageNumber="141" pagination="53 - 65" refId="ref2908" refString="Soldan, T. (2001) Status of the systematic knowledge and priorities in Ephemeroptera studies: the Oriental Region. In: Dominguez, E. (Ed.), Trends in Research in Ephemeroptera &amp; Plecoptera. Kluwer Academic / Plenum Publishers, New York, pp. 53 - 65. https: // doi. org / 10.1007 / 978 - 1 - 4615 - 1257 - 8 _ 9" type="book chapter" year="2001">Soldan 2001</bibRefCitation>
,
<bibRefCitation id="9D054B3EFFC5FFACFE2FFE330732FE4D" author="Sartori, M. &amp; Derleth, P. &amp; Gattolliat, J. L." box="[412,617,439,465]" pageId="4" pageNumber="141" pagination="403 - 406" refId="ref2617" refString="Sartori, M., Derleth, P. &amp; Gattolliat, J. L. (2003) New data about the mayflies (Ephemeroptera) from Borneo. in: Gaino, E. (Ed.), Research update on Ephemeroptera &amp; Plecoptera. Universita di Perugia, Perugia, pp. 403 - 406." type="book chapter" year="2003">
Sartori
<emphasis id="CBE0EADDFFC5FFACFE5CFE33077CFE4D" box="[495,551,439,465]" italics="true" pageId="4" pageNumber="141">et al.</emphasis>
2003
</bibRefCitation>
,
<bibRefCitation id="9D054B3EFFC5FFACFDC7FE3307A1FE4D" author="Kluge, N. J." box="[628,762,439,465]" pageId="4" pageNumber="141" refId="ref2084" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, Dordrecht, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004</bibRefCitation>
,
<bibRefCitation id="9D054B3EFFC5FFACFCB6FE330688FE4D" author="Ogden, T. H. &amp; Gattolliat, J. L. &amp; Sartori, M. &amp; Staniczek, A. H. &amp; Soldan, T. &amp; Whiting, M. F." box="[773,979,439,465]" pageId="4" pageNumber="141" pagination="616 - 634" refId="ref2456" refString="Ogden, T. H., Gattolliat, J. L., Sartori, M., Staniczek, A. H., Soldan, T. &amp; Whiting, M. F. (2009) Towards a new paradigm in mayfly phylogeny (Ephemeroptera): combined analysis of morphological and molecular data. Systematic Entomology, 34, 616 - 634. https: // doi. org / 10.1111 / j. 1365 - 3113.2009.00488. x" type="journal article" year="2009">
Ogden
<emphasis id="CBE0EADDFFC5FFACFCE4FE3306D4FE4D" box="[855,911,439,465]" italics="true" pageId="4" pageNumber="141">et al.</emphasis>
2009
</bibRefCitation>
).
</paragraph>
<paragraph id="F92B36CFFFC5FFACFF74FE5F061CFDFD" blockId="4.[151,1437,150,609]" pageId="4" pageNumber="141">
A number of fossil
<taxonomicName id="3E944D4CFFC5FFACFE14FE5F04A8FE69" box="[423,499,475,501]" class="Insects" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="142" phylum="Arthropoda" rank="class">insects</taxonomicName>
from the early Eocene amber belonging to different families have been described from the Indian subcontinent (
<bibRefCitation id="9D054B3EFFC5FFACFE41FE7B065EFD85" author="Grimaldi, D. A. &amp; Singh, H." box="[498,773,511,537]" pageId="4" pageNumber="141" pagination="17 - 28" refId="ref1766" refString="Grimaldi, D. A. &amp; Singh, H. (2012) The extinct genus Pareuthychaeta in Eocene ambers (Diptera: Schizophora: Ephydroidea). The Canadian Entomologist, 144, 17 - 28. https: // doi. org / 10.4039 / tce. 2012.5" type="journal article" year="2012">Grimaldi &amp; Singh 2012</bibRefCitation>
,
<bibRefCitation id="9D054B3EFFC5FFACFCA1FE7B06B4FD85" author="Stebner, F. &amp; Szadziewski, R. &amp; Singh, H. &amp; Gunkel, S. &amp; Rust, J." box="[786,1007,511,537]" pageId="4" pageNumber="141" pagination="0169144" refId="ref2984" refString="Stebner, F., Szadziewski, R., Singh, H., Gunkel, S. &amp; Rust, J. (2017) Biting midges (Diptera: Ceratopogonidae) from Cambay amber indicate that the Eocene fauna of the Indian subcontinent was not isolated. PLoS ONE, 12 (1), e 0169144. https: // doi. org / 10.1371 / journal. pone. 0169144" type="journal article" year="2017">
Stebner
<emphasis id="CBE0EADDFFC5FFACFCC2FE7B06F0FD85" box="[881,939,511,537]" italics="true" pageId="4" pageNumber="141">et al.</emphasis>
2017
</bibRefCitation>
,
<bibRefCitation id="9D054B3EFFC5FFACFC4FFE7B0051FD85" author="Zakrzewska, M. &amp; Stebner, F. &amp; Mateusz, P. &amp; Singh, H. &amp; Gilka, W." box="[1020,1290,511,537]" pageId="4" pageNumber="141" pagination="255 - 261" refId="ref3226" refString="Zakrzewska, M., Stebner, F., Mateusz, P., Singh, H. &amp; Gilka, W. (2017) A peculiar leg structure in the first non-biting midge de- scribed from Cambay amber, India (Diptera: Chironomidae). Earth and Environmental Science, Transactions of the Royal Society of Edinburgh, 107 (2 - 3), 255 - 261. https: // doi. org / 10.1017 / S 1755691017000421" type="journal article" year="2017">
Zakrzewska
<emphasis id="CBE0EADDFFC5FFACFB3FFE7B019DFD85" box="[1164,1222,511,537]" italics="true" pageId="4" pageNumber="141">et al.</emphasis>
2017
</bibRefCitation>
,
<bibRefCitation id="9D054B3EFFC5FFACFAA4FE7B0589FDA1" author="Kania, I. &amp; Krzeminski, W. &amp; Stebner, F. &amp; Singh, H." pageId="4" pageNumber="141" pagination="263 - 269" refId="ref1985" refString="Kania, I., Krzeminski, W., Stebner, F. &amp; Singh, H. (2018) The first representative of Tipulomorpha (Diptera) from Early Eocene Cambay amber (India). Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 107, 263 - 269. https: // doi. org / 10.1017 / S 1755691017000433" type="journal article" year="2018">
Kania
<emphasis id="CBE0EADDFFC5FFACFAD1FE7B00C6FD85" box="[1378,1437,511,537]" italics="true" pageId="4" pageNumber="141">et al.</emphasis>
2018
</bibRefCitation>
, and reference therein). The present fossil is the first record of a mayfly from the Indian subcontinent, and is described as
<taxonomicName id="3E944D4CFFC5FFACFE9BFDC307BAFDFD" authority="Agnihotri et al." authorityName="Agnihotri" authorityYear="2020" box="[296,737,583,609]" class="Insecta" family="Teloganellidae" genus="Teloganella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="gurhaensis" status="sp. nov.">
<emphasis id="CBE0EADDFFC5FFACFE9BFDC30777FDFD" box="[296,556,583,609]" italics="true" pageId="4" pageNumber="141">Teloganella gurhaensis</emphasis>
Agnihotri
<emphasis id="CBE0EADDFFC5FFACFD14FDC307BAFDFD" box="[679,737,583,609]" italics="true" pageId="4" pageNumber="141">et al.</emphasis>
</taxonomicName>
<emphasis id="CBE0EADDFFC5FFACFD52FDC307BDFDFD" box="[737,742,583,609]" italics="true" pageId="4" pageNumber="141">,</emphasis>
<emphasis id="CBE0EADDFFC5FFACFD5EFDC3061CFDFD" bold="true" box="[749,839,583,609]" pageId="4" pageNumber="141">
<taxonomicNameLabel id="D0D357A6FFC5FFACFD5EFDC3061CFDFD" box="[749,839,583,609]" pageId="4" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>