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<document ID-DOI="10.15560/16.2.361" ID-ISSN="1809-127X" approvalRequired="256" approvalRequired_for_taxonomicNames="5" approvalRequired_for_textStreams="149" approvalRequired_for_treatments="102" checkinTime="1630319602342" checkinUser="marcus" docAuthor="Kharel, Bhim Prasad, Schoolmeester, Paul &amp; Sarkar, Subhankar Kumar" docDate="2020" docId="6D0F879DFFC9C825F1DF8EC51A61C35A" docLanguage="en" docName="CheckList.2020.16.2.361-381.pdf" docOrigin="Check List 16 (2)" docSource="http://dx.doi.org/10.15560/16.2.361" docStyle="DocumentStyle:BB30DA0EB5D11E1D6FE264CD32192DE2.1:CheckList.2017-.journal_article.annotatedlist" docStyleId="BB30DA0EB5D11E1D6FE264CD32192DE2" docStyleName="CheckList.2017-.journal_article.annotatedlist" docStyleVersion="1" docTitle="Onthophagus (Onthophagus) cervus" docType="treatment" docVersion="1" lastPageNumber="365" masterDocId="9136FFE5FFCDC821F1778A37190CC21D" masterDocTitle="A first faunistic account on the Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae, Scarabaeinae) of the Nadia district, West Bengal, with a preliminary checklist from India" masterLastPageNumber="381" masterPageNumber="361" pageId="4" pageNumber="365" updateTime="1630319606135" updateUser="marcus" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A first faunistic account on the Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae, Scarabaeinae) of the Nadia district, West Bengal, with a preliminary checklist from India</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kharel, Bhim Prasad</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schoolmeester, Paul</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sarkar, Subhankar Kumar</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Check List</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-04-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>16</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>361</mods:start>
<mods:end>381</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.15560/16.2.361</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.15560/16.2.361</mods:identifier>
<mods:identifier type="ISSN">1809-127X</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:6D0F879DFFC9C825F1DF8EC51A61C35A" httpUri="http://treatment.plazi.org/id/6D0F879DFFC9C825F1DF8EC51A61C35A" lastPageNumber="365" pageId="4" pageNumber="365">
<subSubSection pageId="4" pageNumber="365" type="nomenclature">
<paragraph blockId="4.[168,772,1266,1324]" pageId="4" pageNumber="365">
<taxonomicName authority="(Fabricius, 1798)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[168,772,1266,1290]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="species" species="cervus" subGenus="Onthophagus">
<emphasis bold="true" box="[168,772,1266,1290]" inLineHeading="true" pageId="4" pageNumber="365" reason="1">
<emphasis bold="true" box="[168,319,1266,1290]" italics="true" pageId="4" pageNumber="365">Onthophagus</emphasis>
(
<emphasis bold="true" box="[334,485,1266,1290]" italics="true" pageId="4" pageNumber="365">Onthophagus</emphasis>
)
<emphasis bold="true" box="[501,574,1267,1290]" italics="true" pageId="4" pageNumber="365">cervus</emphasis>
(Fabricius, 1798)
</emphasis>
</taxonomicName>
<figureCitation box="[168,278,1300,1324]" captionStart="Figure 2" captionStartId="3.[168,231,1953,1973]" captionTargetBox="[213,1415,221,1931]" captionTargetId="figure-114@3.[205,1419,211,1941]" captionTargetPageId="3" captionText="Figure 2. Dorsal habitus of adults of Onthophagus Latreille, 1802 species recorded from Nadia district of West Bengal, India. A. Onthophagus (Colobonthophagus) agnus Gillet, 1925 ♀. B. Onthophagus (Onthophagus) andrewesi Arrow, 1931 ♂. C. Onthophagus (Colobonthophagus) bison Boucomont, 1919 ♂. D. Onthophagus (Onthophagus) cervus (Fabricius,1798)♂. E. Onthophagus (Onthophagus) ceylonicus Harold, 1859 ♀. F. Onthophagus (Onthophagus) circulifer Arrow, 1931 ♂. G. Onthophagus (Onthophagus) circulifer Arrow, 1931 ♀. H. Onthophagus (Onthophagiellus) crassicollis Boucomont, 1913 ♀. Scale bars = 2 mm." pageId="4" pageNumber="365">Figure 2D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="365" type="materials_examined">
<paragraph blockId="4.[168,775,1342,1434]" pageId="4" pageNumber="365">
<emphasis bold="true" box="[168,391,1342,1366]" pageId="4" pageNumber="365">Material examined</emphasis>
(
<specimenCount box="[412,444,1342,1366]" pageId="4" pageNumber="365" type="male">1♂</specimenCount>
).
<collectingCountry box="[469,528,1342,1366]" name="India" pageId="4" pageNumber="365">India</collectingCountry>
,
<collectingRegion box="[543,688,1342,1366]" country="India" name="West Bengal" pageId="4" pageNumber="365">West Bengal</collectingRegion>
, Nadia, Hanskhali (
<geoCoordinate box="[298,424,1374,1399]" degrees="23" direction="north" minutes="21" orientation="latitude" pageId="4" pageNumber="365" precision="15" seconds="40" value="23.36111">23°2140″N</geoCoordinate>
,
<geoCoordinate box="[440,577,1374,1399]" degrees="088" direction="east" minutes="35" orientation="longitude" pageId="4" pageNumber="365" precision="15" seconds="56" value="88.59889">088°3556″E</geoCoordinate>
), B.P.Kharel leg.,
<date box="[168,296,1409,1433]" pageId="4" pageNumber="365" value="2018-07-05">05.VII.2018</date>
,
<collectionCode box="[306,397,1410,1433]" pageId="4" pageNumber="365">KUZEN</collectionCode>
S0131 (
<specimenCount box="[483,515,1410,1434]" pageId="4" pageNumber="365" type="male">1♂</specimenCount>
).
</paragraph>
<paragraph blockId="4.[168,775,1451,1745]" pageId="4" pageNumber="365">
<emphasis bold="true" box="[168,323,1451,1475]" pageId="4" pageNumber="365">Identification</emphasis>
(
<specimenCount box="[343,364,1451,1475]" pageId="4" pageNumber="365" type="male"></specimenCount>
). This species can be distinguished from other
<taxonomicName box="[300,535,1485,1509]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[300,446,1485,1509]" italics="true" pageId="4" pageNumber="365">Onthophagus</emphasis>
species
</taxonomicName>
by the head narrow, with rounded sides and a pair of long, backwardly sloping, and internally curved horns on the vertex; clypeus truncate, little produced but not pointed, and separated from the forehead by a feeble carina; pronotum strongly, unevenly, and scantily punctate, with sharp front angles; elytra finely striate, with slightly convex and finely punctate interstriae; pygidium coarsely and finely punctate.
</paragraph>
<paragraph blockId="4.[168,775,1763,2091]" pageId="4" pageNumber="365">
<emphasis bold="true" box="[168,476,1763,1787]" pageId="4" pageNumber="365">Geographical distribution.</emphasis>
<collectingCountry box="[482,578,1763,1787]" name="Pakistan" pageId="4" pageNumber="365">Pakistan</collectingCountry>
,
<collectingCountry box="[588,732,1763,1787]" name="Saudi Arabia" pageId="4" pageNumber="365">Saudi Arabia</collectingCountry>
,
<collectingCountry name="Sri Lanka" pageId="4" pageNumber="365">Sri Lanka</collectingCountry>
, and
<collectingCountry box="[303,396,1797,1821]" name="Vietnam" pageId="4" pageNumber="365">Vietnam</collectingCountry>
(
<bibRefCitation author="Sewak R" box="[413,565,1797,1821]" pageId="4" pageNumber="365" pagination="1 - 48" refId="ref12194" refString="Sewak R (2009 a) Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Thar Desert of Gujrat. Records of the Zoological Survey of India, Occasional Paper 295: 1 - 48." type="journal article" year="2009">Sewak 2009a</bibRefCitation>
;
<bibRefCitation author="Biswas S &amp; Chatterjee SK" pageId="4" pageNumber="365" pagination="57 - 67" refId="ref9435" refString="Biswas S, Chatterjee SK (1986 a) Dung beetles fauna (Coleoptera: Scarabaeidae: Scarabainae) of Palamou Tiger Reserve, Bihar, with description of a new species. Records of the Zoological Survey of India 83 (3 - 4): 57 - 67." type="journal article" year="1986">Biswas and Chatterjee 1986a</bibRefCitation>
).
<collectingCountry box="[336,396,1831,1855]" name="India" pageId="4" pageNumber="365">India</collectingCountry>
:
<collectingRegion box="[415,771,1831,1855]" country="India" name="Andaman and Nicobar Islands" pageId="4" pageNumber="365">Andaman and Nicobar Islands</collectingRegion>
,
<collectingRegion box="[168,383,1864,1888]" country="India" name="Arunachal Pradesh" pageId="4" pageNumber="365">Arunachal Pradesh</collectingRegion>
,
<collectingRegion box="[398,541,1864,1888]" country="India" name="Chhattisgarh" pageId="4" pageNumber="365">Chhattisgarh</collectingRegion>
,
<collectingRegion box="[555,639,1864,1888]" country="India" name="Gujarat" pageId="4" pageNumber="365">Gujarat</collectingRegion>
,
<collectingRegion box="[654,771,1864,1888]" country="India" name="Karnataka" pageId="4" pageNumber="365">Karnataka</collectingRegion>
,
<collectingRegion box="[168,243,1898,1922]" country="India" name="Kerala" pageId="4" pageNumber="365">Kerala</collectingRegion>
,
<collectingRegion box="[253,437,1898,1922]" country="India" name="Madhya Pradesh" pageId="4" pageNumber="365">Madhya Pradesh</collectingRegion>
,
<collectingRegion box="[448,588,1898,1922]" country="India" name="Maharashtra" pageId="4" pageNumber="365">Maharashtra</collectingRegion>
,
<collectingCountry box="[598,726,1898,1922]" name="India" pageId="4" pageNumber="365">Puducherry</collectingCountry>
,
<collectingRegion country="India" name="Rajasthan" pageId="4" pageNumber="365">Rajasthan</collectingRegion>
,
<collectingRegion box="[263,397,1932,1956]" country="India" name="Tamil Nadu" pageId="4" pageNumber="365">Tamil Nadu</collectingRegion>
,
<collectingRegion box="[412,551,1932,1956]" country="India" name="Uttarakhand" pageId="4" pageNumber="365">Uttarakhand</collectingRegion>
,
<collectingRegion box="[566,721,1932,1956]" country="India" name="Uttar Pradesh" pageId="4" pageNumber="365">Uttar Pradesh</collectingRegion>
, and
<collectingRegion box="[168,304,1966,1990]" country="India" name="West Bengal" pageId="4" pageNumber="365">West Bengal</collectingRegion>
(
<bibRefCitation author="Arrow GJ" box="[317,447,1966,1990]" pageId="4" pageNumber="365" refId="ref9072" refString="Arrow GJ (1931) The fauna of British India including Ceylon and Burma, part III, Coleoptera: Lamellicornia (Coprinae). Taylor and Francis, London, 353 pp." type="book" year="1931">Arrow 1931</bibRefCitation>
;
<bibRefCitation author="Biswas S &amp; Chatterjee SK" box="[457,771,1966,1990]" pageId="4" pageNumber="365" pagination="57 - 67" refId="ref9435" refString="Biswas S, Chatterjee SK (1986 a) Dung beetles fauna (Coleoptera: Scarabaeidae: Scarabainae) of Palamou Tiger Reserve, Bihar, with description of a new species. Records of the Zoological Survey of India 83 (3 - 4): 57 - 67." type="journal article" year="1986">Biswas and Chatterjee 1986a</bibRefCitation>
;
<bibRefCitation author="Chandra K" box="[168,331,1999,2023]" pageId="4" pageNumber="365" pagination="359 - 362" refId="ref9673" refString="Chandra K (2000) Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh, India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000</bibRefCitation>
;
<bibRefCitation author="Chandra K &amp; Ahirwar SC" box="[350,683,1999,2023]" pageId="4" pageNumber="365" pagination="1961 - 1964" refId="ref9812" refString="Chandra K, Ahirwar SC (2005 a) Scarabaeid beetles of Bandhavgarh National Park, Madhya Pradesh. Zoos' Print Journal 20 (8): 1961 - 1964. https: // doi. org / 10.11609 / JoTT. ZPJ. 1281.1961 - 4" type="journal article" year="2005">Chandra and Ahirwar 2005a</bibRefCitation>
,
<bibRefCitation author="Chandra K &amp; Ahirwar SC" box="[701,771,1999,2023]" pageId="4" pageNumber="365" pagination="147 - 155" refId="ref9859" refString="Chandra K, Ahirwar SC (2005 b) Scarabaeid beetles (Coleoptera) of Kanha Tiger Reserve, Madhya Pradesh. Records of the Zoological Survey of India 105 (1 - 2): 147 - 155." type="journal article" year="2005">2005b</bibRefCitation>
,
<bibRefCitation author="Chandra K &amp; Ahirwar SC" box="[168,224,2033,2057]" pageId="4" pageNumber="365" pagination="273 - 300" refId="ref9899" refString="Chandra K, Ahirwar SC (2007) Insecta: Coleoptera: Scarabaeidae, fauna of Madhya Pradesh (including Chhattisgarh). Zoological Survey of India, State Fauna Series 15: 273 - 300." type="book chapter" year="2007">2007</bibRefCitation>
;
<bibRefCitation author="Sewak R" box="[242,384,2033,2057]" pageId="4" pageNumber="365" pagination="191 - 224" refId="ref12157" refString="Sewak R (2006) Coleoptera: Scarabaeidae: Coprinae (dung beetles). Zoological Survey of India, Fauna of Arunachal Pradesh, State Fauna Series 13 (2): 191 - 224." type="journal article" year="2006">Sewak 2006</bibRefCitation>
,
<bibRefCitation author="Sewak R" box="[402,471,2033,2057]" pageId="4" pageNumber="365" pagination="1 - 48" refId="ref12194" refString="Sewak R (2009 a) Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Thar Desert of Gujrat. Records of the Zoological Survey of India, Occasional Paper 295: 1 - 48." type="journal article" year="2009">2009a</bibRefCitation>
,
<bibRefCitation author="Sewak R" box="[488,557,2033,2057]" pageId="4" pageNumber="365" refId="ref12231" refString="Sewak R (2009 b) Dung beetles (Coleoptera: Scarabaeidae: Coprinae)" type="book" year="2009">2009b</bibRefCitation>
, 2010;
<bibRefCitation author="Jadhav MJ &amp; Sharma RM" pageId="4" pageNumber="365" pagination="489 - 494" refId="ref11030" refString="Jadhav MJ, Sharma RM (2012) Insecta: Coleoptera: Scarabaeidae (scarabaeid beetles). Fauna of Maharashtra, State Fauna Series, Zoological Survey of India 20 (2): 489 - 494." type="journal article" year="2012">Jadhav and Sharma 2012</bibRefCitation>
; Chandra et al. 2012b;
<bibRefCitation author="Chandra K &amp; Gupta D" box="[562,775,2067,2091]" pageId="4" pageNumber="365" pagination="886 - 891" refId="ref10103" refString="Chandra K, Gupta D (2012 c) An inventory of scarab beetles (Coleoptera: Scarabaeidae) of Achanakmar-Amarkantak Biosphere Reserve, Chhattisgarh, India. International Journal of Science and Nature 3 (4): 886 - 891." type="journal article" year="2012">Chandra and Gupta</bibRefCitation>
</paragraph>
<paragraph blockId="4.[813,1420,161,251]" box="[813,1420,160,184]" pageId="4" pageNumber="365">
2012c,
<bibRefCitation author="Chandra K &amp; Gupta D" box="[895,963,160,184]" pageId="4" pageNumber="365" pagination="99 - 106" refId="ref10146" refString="Chandra K, Gupta D (2012 d) First report on five species of genus Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae) from Madhya Pradesh, India and their description of external male genitalia. Biodiversity Journal 3 (2): 99 - 106." type="journal article" year="2012">2012d</bibRefCitation>
,
<bibRefCitation author="Chandra K &amp; Gupta D" box="[979,1032,160,184]" pageId="4" pageNumber="365" pagination="331 - 360" refId="ref10244" refString="Chandra K, Gupta D (2013) Taxonomic studies on dung beetles (Coleoptera: Scarabaeidae, Geotrupidae, Hybosoridae) of Chhattisgarh, India. Munis Entomology &amp; Zoology 8 (1): 331 - 360." type="journal article" year="2013">2013</bibRefCitation>
;
<bibRefCitation author="Anitha Rani A &amp; Sanjayan KP" box="[1049,1415,160,184]" pageId="4" pageNumber="365" pagination="241 - 246" refId="ref8966" refString="Anitha Rani A, Sanjayan KP (2013) Community composition of dung beetle and population dynamics of Tiniocellus spinipes in a scarab Jungle ecosystem. Asian Journal of Biological and Life Sciences 2 (3): 241 - 246." type="journal article" year="2013">Anitha Rani and Sanjayan 2013</bibRefCitation>
,
</paragraph>
<paragraph blockId="4.[813,1420,161,251]" pageId="4" pageNumber="365">
<bibRefCitation author="Karimbumkara SN &amp; Priyadarsanan DR" box="[813,1250,194,218]" pageId="4" pageNumber="365" pagination="173 - 178" refId="ref11154" refString="Karimbumkara SN, Priyadarsanan DR (2013) Fauna of Karnataka, State Fauna Series. Zoological Survey of India 21: 173 - 178." type="journal article" year="2013">Karimbumkara and Priyadarsanan 2013</bibRefCitation>
;
<bibRefCitation author="Sathiandran N &amp; Thomas KS &amp; Flemming AT" pageId="4" pageNumber="365" pagination="8250 - 8258" refId="ref11769" refString="Sathiandran N, Thomas KS, Flemming AT (2015) An illustrated checklist of dung beetles (Coleoptera: Scarabaeidae) from the PeriyarTiger Reserve, Kerala, India. Journal of Threatened Taxa 7 (15): 8250 - 8258. https: // doi. org / 10.11609 / jott. 2466.7.15.8250 - 8258" type="journal article" year="2015">Sathiandran et al. 2015</bibRefCitation>
;
<bibRefCitation author="Gajendra N &amp; Prasad SK" box="[909,1195,227,251]" pageId="4" pageNumber="365" pagination="710 - 714" refId="ref10831" refString="Gajendra N, Prasad SK (2016) A Review of Coleoptera diversity of Chhattisgarh: updated checklist 2015. International Journal of Science and Research 5 (4): 710 - 714." type="journal article" year="2016">Gajendra and Prasad 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="365" type="discussion">
<paragraph blockId="4.[813,1418,269,327]" pageId="4" pageNumber="365">
<emphasis bold="true" box="[813,924,269,293]" pageId="4" pageNumber="365">Remarks.</emphasis>
This species is widely distributed throughout
<collectingCountry box="[813,873,303,327]" name="India" pageId="4" pageNumber="365">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>