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<document ID-DOI="10.15560/16.2.361" ID-ISSN="1809-127X" approvalRequired="256" approvalRequired_for_taxonomicNames="5" approvalRequired_for_textStreams="149" approvalRequired_for_treatments="102" checkinTime="1630319602342" checkinUser="marcus" docAuthor="Kharel, Bhim Prasad, Schoolmeester, Paul &amp; Sarkar, Subhankar Kumar" docDate="2020" docId="6D0F879DFFC8C824F1DF89791AA3C14A" docLanguage="en" docName="CheckList.2020.16.2.361-381.pdf" docOrigin="Check List 16 (2)" docSource="http://dx.doi.org/10.15560/16.2.361" docStyle="DocumentStyle:BB30DA0EB5D11E1D6FE264CD32192DE2.1:CheckList.2017-.journal_article.annotatedlist" docStyleId="BB30DA0EB5D11E1D6FE264CD32192DE2" docStyleName="CheckList.2017-.journal_article.annotatedlist" docStyleVersion="1" docTitle="Onthophagus (Colobonthophagus) dama" docType="treatment" docVersion="1" lastPageNumber="366" masterDocId="9136FFE5FFCDC821F1778A37190CC21D" masterDocTitle="A first faunistic account on the Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae, Scarabaeinae) of the Nadia district, West Bengal, with a preliminary checklist from India" masterLastPageNumber="381" masterPageNumber="361" pageId="5" pageNumber="366" updateTime="1630319606135" updateUser="marcus" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A first faunistic account on the Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae, Scarabaeinae) of the Nadia district, West Bengal, with a preliminary checklist from India</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kharel, Bhim Prasad</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schoolmeester, Paul</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sarkar, Subhankar Kumar</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Check List</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-04-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>16</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>361</mods:start>
<mods:end>381</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.15560/16.2.361</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.15560/16.2.361</mods:identifier>
<mods:identifier type="ISSN">1809-127X</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:6D0F879DFFC8C824F1DF89791AA3C14A" httpUri="http://treatment.plazi.org/id/6D0F879DFFC8C824F1DF89791AA3C14A" lastPageNumber="366" pageId="5" pageNumber="366">
<subSubSection pageId="5" pageNumber="366" type="nomenclature">
<paragraph blockId="5.[168,753,846,903]" pageId="5" pageNumber="366">
<heading centered="true" fontSize="10" level="2" pageId="5" pageNumber="366" reason="2">
<taxonomicName authority="(Fabricius, 1798)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="dama" subGenus="Colobonthophagus">
<emphasis bold="true" pageId="5" pageNumber="366">
<emphasis box="[168,753,846,870]" inLineHeading="true" pageId="5" pageNumber="366" reason="1">
<emphasis bold="true" box="[168,319,846,870]" italics="true" pageId="5" pageNumber="366">Onthophagus</emphasis>
(
<emphasis bold="true" box="[334,545,846,870]" italics="true" pageId="5" pageNumber="366">Colobonthophagus</emphasis>
)
<emphasis bold="true" box="[561,623,846,870]" italics="true" pageId="5" pageNumber="366">dama</emphasis>
(Fabricius,
</emphasis>
1798)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[168,308,913,937]" pageId="5" pageNumber="366" type="description">
<paragraph blockId="5.[168,308,913,937]" box="[168,308,913,937]" pageId="5" pageNumber="366">
<figureCitation box="[168,308,913,937]" captionStart="Figure 3" captionStartId="6.[168,231,1953,1973]" captionTargetBox="[209,1380,238,1888]" captionTargetId="figure-120@6.[191,1394,209,1897]" captionTargetPageId="6" captionText="Figure 3. Dorsal habitus of male/female of Onthophagus Latreille, 1802 species recorded from Nadia district of West Bengal, India. A. Onthophagus (Colobonthophagus) dama (Fabricius,1798)♂. B. Onthophagus (Colobonthophagus) dama (Fabricius, 1798) ♀. C. Onthophagus (Colobonthophagus) hindu Arrow, 1931 ♀. D. Onthophagus (Onthophagus) mauritii Boucomont, 1919 ♀. E. Onthophagus (Indonthophagus) mopsus (Fabricius, 1792) ♀. F. Onthophagus (Colobonthophagus) paliceps Arrow, 1931 ♂. G. Onthophagus (Colobonthophagus) pardalis Boucomont, 1914 ♀. H. Onthophagus (Onthophagus) porcus Arrow, 1931 ♂. Scale bars = 2 mm." pageId="5" pageNumber="366">Figure 3A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="366" type="materials_examined">
<paragraph blockId="5.[168,775,954,1815]" pageId="5" pageNumber="366">
<emphasis bold="true" box="[168,385,954,978]" pageId="5" pageNumber="366">Material examined</emphasis>
(
<specimenCount box="[397,445,955,979]" pageId="5" pageNumber="366" type="male">26♂</specimenCount>
,
<specimenCount box="[454,500,955,979]" pageId="5" pageNumber="366" type="female">39♀</specimenCount>
).
<collectingCountry box="[519,578,955,979]" name="India" pageId="5" pageNumber="366">India</collectingCountry>
,
<collectingRegion box="[588,726,955,979]" country="India" name="West Bengal" pageId="5" pageNumber="366">West Bengal</collectingRegion>
, Nadia, Karimpur (
<geoCoordinate box="[349,476,987,1012]" degrees="23" direction="north" minutes="58" orientation="latitude" pageId="5" pageNumber="366" precision="15" seconds="12" value="23.970001">23°5812″N</geoCoordinate>
,
<geoCoordinate box="[493,631,987,1012]" degrees="088" direction="east" minutes="37" orientation="longitude" pageId="5" pageNumber="366" precision="15" seconds="12" value="88.62">088°3712″E</geoCoordinate>
), S.K.Sarkar leg.,
<date box="[221,326,1022,1046]" pageId="5" pageNumber="366" value="2017-05-08">
08.
<collectionCode box="[250,269,1022,1046]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="366" type="Museum">V</collectionCode>
.2017
</date>
,
<collectionCode box="[337,428,1022,1045]" pageId="5" pageNumber="366">KUZEN</collectionCode>
S0138S0141 (
<specimenCount box="[596,630,1022,1046]" pageId="5" pageNumber="366" type="male">4♂</specimenCount>
).
<collectingCountry box="[650,709,1022,1046]" name="India" pageId="5" pageNumber="366">India</collectingCountry>
,
<collectingRegion country="India" name="West Bengal" pageId="5" pageNumber="366">West Bengal</collectingRegion>
, Nadia, Kaliganj (
<geoCoordinate box="[482,609,1054,1079]" degrees="23" direction="north" minutes="44" orientation="latitude" pageId="5" pageNumber="366" precision="15" seconds="03" value="23.734167">23°4403″N</geoCoordinate>
,
<geoCoordinate box="[633,767,1054,1079]" degrees="088" direction="east" minutes="13" orientation="longitude" pageId="5" pageNumber="366" precision="15" seconds="57" value="88.2325">088°1357″E</geoCoordinate>
) (
<geoCoordinate box="[176,303,1087,1112]" degrees="23" direction="north" minutes="44" orientation="latitude" pageId="5" pageNumber="366" precision="15" seconds="03" value="23.734167">23°4403″N</geoCoordinate>
,
<geoCoordinate box="[320,456,1087,1112]" degrees="088" direction="east" minutes="13" orientation="longitude" pageId="5" pageNumber="366" precision="15" seconds="57" value="88.2325">088°1357″E</geoCoordinate>
), B.P.Kharel leg.,
<date box="[670,771,1089,1113]" pageId="5" pageNumber="366" value="2017-05-11">
11.
<collectionCode box="[695,714,1089,1113]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="366" type="Museum">V</collectionCode>
.2017
</date>
,
<collectionCode box="[168,259,1122,1145]" pageId="5" pageNumber="366">KUZEN</collectionCode>
S0142-S0143 (
<specimenCount box="[424,455,1122,1146]" pageId="5" pageNumber="366" type="female">2♀</specimenCount>
).
<collectingCountry box="[477,536,1122,1146]" name="India" pageId="5" pageNumber="366">India</collectingCountry>
,
<collectingRegion box="[549,691,1122,1146]" country="India" name="West Bengal" pageId="5" pageNumber="366">West Bengal</collectingRegion>
, Nadia, Krishnanagar (
<geoCoordinate box="[341,469,1154,1179]" degrees="23" direction="north" minutes="23" orientation="latitude" pageId="5" pageNumber="366" precision="15" seconds="59" value="23.399721">23°2359″N</geoCoordinate>
,
<geoCoordinate box="[490,629,1154,1179]" degrees="088" direction="east" minutes="30" orientation="longitude" pageId="5" pageNumber="366" precision="15" seconds="37" value="88.51028">088°3037″E</geoCoordinate>
), B.P.Kharel leg.,
<date box="[225,340,1189,1213]" pageId="5" pageNumber="366" value="2018-06-01">
01.
<collectionCode box="[253,282,1189,1213]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="366">VI</collectionCode>
.2018
</date>
,
<collectionCode box="[357,448,1189,1212]" pageId="5" pageNumber="366">KUZEN</collectionCode>
S0144S0146 (
<specimenCount box="[627,658,1189,1213]" pageId="5" pageNumber="366" type="female">3♀</specimenCount>
),
<collectionCode box="[683,774,1189,1212]" pageId="5" pageNumber="366">KUZEN</collectionCode>
S0147-S0151 (
<specimenCount box="[319,353,1222,1246]" pageId="5" pageNumber="366" type="male">5♂</specimenCount>
).
<collectingCountry box="[371,430,1222,1246]" name="India" pageId="5" pageNumber="366">India</collectingCountry>
,
<collectingRegion box="[439,578,1222,1246]" country="India" name="West Bengal" pageId="5" pageNumber="366">West Bengal</collectingRegion>
, Nadia, Nabadwip (
<geoCoordinate box="[176,301,1254,1279]" degrees="23" direction="north" minutes="24" orientation="latitude" pageId="5" pageNumber="366" precision="15" seconds="11" value="23.403055">23°2411″N</geoCoordinate>
,
<geoCoordinate box="[316,442,1254,1279]" degrees="088" direction="east" minutes="22" orientation="longitude" pageId="5" pageNumber="366" precision="15" seconds="5" value="88.36806">088°225″E</geoCoordinate>
), S.K.Sarkar leg.,
<date box="[653,771,1256,1280]" pageId="5" pageNumber="366" value="2018-06-03">
03.
<collectionCode box="[683,712,1256,1280]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="366">VI</collectionCode>
.2018
</date>
,
<collectionCode box="[168,259,1290,1313]" pageId="5" pageNumber="366">KUZEN</collectionCode>
S0152S0154 (
<specimenCount box="[454,484,1289,1313]" pageId="5" pageNumber="366" type="female">3♀</specimenCount>
),
<collectionCode box="[519,610,1290,1313]" pageId="5" pageNumber="366">KUZEN</collectionCode>
S0155S0157 (
<specimenCount box="[177,211,1323,1347]" pageId="5" pageNumber="366" type="male">3♂</specimenCount>
).
<collectingCountry box="[232,291,1323,1347]" name="India" pageId="5" pageNumber="366">India</collectingCountry>
,
<collectingRegion box="[304,445,1323,1347]" country="India" name="West Bengal" pageId="5" pageNumber="366">West Bengal</collectingRegion>
, Nadia, Santipur (
<geoCoordinate box="[646,770,1321,1346]" degrees="23" direction="north" minutes="15" orientation="latitude" pageId="5" pageNumber="366" precision="15" seconds="56" value="23.265556">23°1556″N</geoCoordinate>
,
<geoCoordinate box="[168,306,1355,1380]" degrees="088" direction="east" minutes="26" orientation="longitude" pageId="5" pageNumber="366" precision="15" seconds="10" value="88.43611">088°2610″E</geoCoordinate>
), S.K.Sarkar leg.,
<date box="[534,662,1356,1380]" pageId="5" pageNumber="366" value="2018-07-08">08.VII.2018</date>
,
<collectionCode box="[683,774,1357,1380]" pageId="5" pageNumber="366">KUZEN</collectionCode>
S0158S0163 (
<specimenCount box="[339,369,1390,1414]" pageId="5" pageNumber="366" type="female">6♀</specimenCount>
),
<collectionCode box="[400,491,1390,1413]" pageId="5" pageNumber="366">KUZEN</collectionCode>
S0164S0171 (
<specimenCount box="[679,713,1390,1414]" pageId="5" pageNumber="366" type="male">8♂</specimenCount>
).
<collectingCountry name="India" pageId="5" pageNumber="366">India</collectingCountry>
,
<collectingRegion box="[228,380,1423,1447]" country="India" name="West Bengal" pageId="5" pageNumber="366">West Bengal</collectingRegion>
, Nadia, Haringhata (
<geoCoordinate box="[643,771,1422,1447]" degrees="22" direction="north" minutes="57" orientation="latitude" pageId="5" pageNumber="366" precision="15" seconds="49" value="22.963612">22°5749″N</geoCoordinate>
,
<geoCoordinate box="[168,305,1455,1480]" degrees="088" direction="east" minutes="33" orientation="longitude" pageId="5" pageNumber="366" precision="15" seconds="36" value="88.560005">088°3336″E</geoCoordinate>
), S.K.Sarkar leg.,
<date box="[535,662,1457,1481]" pageId="5" pageNumber="366" value="2018-07-10">10.VII.2018</date>
,
<collectionCode box="[684,775,1457,1480]" pageId="5" pageNumber="366">KUZEN</collectionCode>
S0172S0179 (
<specimenCount box="[329,359,1490,1514]" pageId="5" pageNumber="366" type="female">8♀</specimenCount>
).
<collectingCountry box="[380,439,1490,1514]" name="India" pageId="5" pageNumber="366">India</collectingCountry>
,
<collectingRegion box="[451,591,1490,1514]" country="India" name="West Bengal" pageId="5" pageNumber="366">West Bengal</collectingRegion>
, Nadia, Hanskhali (
<geoCoordinate box="[235,360,1522,1547]" degrees="23" direction="north" minutes="21" orientation="latitude" pageId="5" pageNumber="366" precision="15" seconds="56" value="23.365557">23°2156″N</geoCoordinate>
,
<geoCoordinate box="[397,536,1522,1547]" degrees="088" direction="east" minutes="36" orientation="longitude" pageId="5" pageNumber="366" precision="15" seconds="07" value="88.601944">088°3607″E</geoCoordinate>
), B.P.Kharel leg.,
<date box="[168,305,1557,1581]" pageId="5" pageNumber="366" value="2018-08-05">05.VIII.2018</date>
,
<collectionCode box="[321,412,1557,1580]" pageId="5" pageNumber="366">KUZEN</collectionCode>
S0180S0185 (
<specimenCount box="[590,621,1557,1581]" pageId="5" pageNumber="366" type="female">6♀</specimenCount>
).
<collectingCountry box="[646,705,1557,1581]" name="India" pageId="5" pageNumber="366">India</collectingCountry>
,
<collectingRegion country="India" name="West Bengal" pageId="5" pageNumber="366">West Bengal</collectingRegion>
, Nadia, Ranaghat (
<geoCoordinate box="[477,606,1589,1614]" degrees="23" direction="north" minutes="09" orientation="latitude" pageId="5" pageNumber="366" precision="15" seconds="53" value="23.164722">23°0953″N</geoCoordinate>
,
<geoCoordinate box="[624,763,1589,1614]" degrees="088" direction="east" minutes="33" orientation="longitude" pageId="5" pageNumber="366" precision="15" seconds="27" value="88.5575">088°3327″E</geoCoordinate>
), B.P.Kharel leg.,
<date box="[373,491,1624,1648]" pageId="5" pageNumber="366" value="2019-03-09">09.III.2019</date>
,
<collectionCode box="[516,607,1624,1647]" pageId="5" pageNumber="366">KUZEN</collectionCode>
S0186S0192 (
<specimenCount box="[177,209,1657,1681]" pageId="5" pageNumber="366" type="female">7♀</specimenCount>
).
<collectingCountry box="[235,294,1657,1681]" name="India" pageId="5" pageNumber="366">India</collectingCountry>
,
<collectingRegion box="[312,458,1657,1681]" country="India" name="West Bengal" pageId="5" pageNumber="366">West Bengal</collectingRegion>
, Nadia,Tehatta (
<geoCoordinate box="[646,771,1656,1681]" degrees="23" direction="north" minutes="43" orientation="latitude" pageId="5" pageNumber="366" precision="15" seconds="53" value="23.73139">23°4353″N</geoCoordinate>
,
<geoCoordinate box="[168,304,1689,1714]" degrees="088" direction="east" minutes="31" orientation="longitude" pageId="5" pageNumber="366" precision="15" seconds="44" value="88.52889">088°3144″E</geoCoordinate>
), S.K.Sarkar leg.,
<date box="[526,664,1691,1715]" pageId="5" pageNumber="366" value="2018-08-22">22.VIII.2018</date>
,
<collectionCode box="[684,775,1691,1714]" pageId="5" pageNumber="366">KUZEN</collectionCode>
S0193S0198 (
<specimenCount box="[331,366,1724,1748]" pageId="5" pageNumber="366" type="male">6♂</specimenCount>
).
<collectingCountry box="[388,447,1724,1748]" name="India" pageId="5" pageNumber="366">India</collectingCountry>
,
<collectingRegion box="[460,602,1724,1748]" country="India" name="West Bengal" pageId="5" pageNumber="366">West Bengal</collectingRegion>
, Nadia, Chapra (
<geoCoordinate box="[176,305,1756,1781]" degrees="23" direction="north" minutes="30" orientation="latitude" pageId="5" pageNumber="366" precision="15" seconds="50" value="23.51389">23°3050″N</geoCoordinate>
,
<geoCoordinate box="[316,454,1756,1781]" degrees="088" direction="east" minutes="32" orientation="longitude" pageId="5" pageNumber="366" precision="15" seconds="56" value="88.54889">088°3256″E</geoCoordinate>
), B.P.Kharel leg.,
<date box="[650,770,1758,1782]" pageId="5" pageNumber="366" value="2018-09-23">23.IX.2018</date>
,
<collectionCode box="[168,259,1792,1815]" pageId="5" pageNumber="366">KUZEN</collectionCode>
S0199S0202 (
<specimenCount box="[430,461,1791,1815]" pageId="5" pageNumber="366" type="female">4♀</specimenCount>
).
</paragraph>
<paragraph blockId="5.[168,775,1832,2091]" lastBlockId="5.[813,1420,160,253]" pageId="5" pageNumber="366">
<emphasis bold="true" box="[168,323,1832,1856]" pageId="5" pageNumber="366">Identification</emphasis>
(
<specimenCount box="[343,364,1833,1857]" pageId="5" pageNumber="366" type="male"></specimenCount>
). This species can be distinguished from other
<taxonomicName box="[299,534,1866,1890]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[299,445,1866,1890]" italics="true" pageId="5" pageNumber="366">Onthophagus</emphasis>
species
</taxonomicName>
by the body entirely glabrous dorsally; head short and broad, with a pair of long, backwardly inclined, outwardly curved, and toothed horns on the vertex; clypeus strongly punctate; clypeal margin semi-circular and medially slightly projected; pronotum finely and sparsely punctate, with the sides nearly rounded and basal margins slightly angulate medially; elytra distinctly striate, with interstria enearly flat and finely and scarcely punctate; pygidium moderately punctate.
</paragraph>
<paragraph blockId="5.[812,1420,271,777]" pageId="5" pageNumber="366">
<emphasis bold="true" box="[813,1122,271,295]" pageId="5" pageNumber="366">Geographical distribution.</emphasis>
<collectingCountry box="[1130,1211,272,296]" name="Bhutan" pageId="5" pageNumber="366">Bhutan</collectingCountry>
,
<collectingCountry box="[1223,1291,272,296]" name="China" pageId="5" pageNumber="366">China</collectingCountry>
,
<collectingCountry box="[1303,1370,272,296]" name="Nepal" pageId="5" pageNumber="366">Nepal</collectingCountry>
, and
<collectingCountry box="[813,926,306,330]" name="Sri Lanka" pageId="5" pageNumber="366">Sri Lanka</collectingCountry>
(
<bibRefCitation author="Arrow GJ" box="[945,1067,306,330]" pageId="5" pageNumber="366" refId="ref9072" refString="Arrow GJ (1931) The fauna of British India including Ceylon and Burma, part III, Coleoptera: Lamellicornia (Coprinae). Taylor and Francis, London, 353 pp." type="book" year="1931">Arrow1931</bibRefCitation>
;
<bibRefCitation author="Biswas S &amp; Chatterjee SK" box="[1083,1415,306,330]" pageId="5" pageNumber="366" pagination="57 - 67" refId="ref9435" refString="Biswas S, Chatterjee SK (1986 a) Dung beetles fauna (Coleoptera: Scarabaeidae: Scarabainae) of Palamou Tiger Reserve, Bihar, with description of a new species. Records of the Zoological Survey of India 83 (3 - 4): 57 - 67." type="journal article" year="1986">Biswas and Chatterjee 1986a</bibRefCitation>
; Singh et al. 2017).
<collectingCountry box="[1041,1101,340,364]" name="India" pageId="5" pageNumber="366">India</collectingCountry>
:
<collectingRegion box="[1118,1335,340,364]" country="India" name="Arunachal Pradesh" pageId="5" pageNumber="366">Arunachal Pradesh</collectingRegion>
,
<collectingRegion box="[1353,1415,340,364]" country="India" name="Bihar" pageId="5" pageNumber="366">Bihar</collectingRegion>
,
<collectingRegion box="[813,956,375,399]" country="India" name="Chhattisgarh" pageId="5" pageNumber="366">Chhattisgarh</collectingRegion>
,
<collectingRegion box="[969,1065,375,399]" country="India" name="Haryana" pageId="5" pageNumber="366">Haryana</collectingRegion>
,
<collectingRegion box="[1080,1284,375,399]" country="India" name="Himachal Pradesh" pageId="5" pageNumber="366">Himachal Pradesh</collectingRegion>
,
<collectingRegion box="[1299,1416,375,399]" country="India" name="Karnataka" pageId="5" pageNumber="366">Karnataka</collectingRegion>
,
<collectingRegion box="[813,888,409,433]" country="India" name="Kerala" pageId="5" pageNumber="366">Kerala</collectingRegion>
,
<collectingRegion box="[898,1082,409,433]" country="India" name="Madhya Pradesh" pageId="5" pageNumber="366">Madhya Pradesh</collectingRegion>
,
<collectingRegion box="[1092,1232,409,433]" country="India" name="Maharashtra" pageId="5" pageNumber="366">Maharashtra</collectingRegion>
,
<collectingRegion box="[1242,1322,409,433]" country="India" name="Odisha" pageId="5" pageNumber="366">Odisha</collectingRegion>
,
<collectingRegion box="[1333,1416,409,433]" country="India" name="Sikkim" pageId="5" pageNumber="366">Sikkim</collectingRegion>
,
<collectingRegion box="[812,943,444,468]" country="India" name="Tamil Nadu" pageId="5" pageNumber="366">Tamil Nadu</collectingRegion>
,
<collectingRegion box="[952,1092,444,468]" country="India" name="Uttarakhand" pageId="5" pageNumber="366">Uttarakhand</collectingRegion>
,
<collectingRegion box="[1102,1254,444,468]" country="India" name="Uttar Pradesh" pageId="5" pageNumber="366">Uttar Pradesh</collectingRegion>
, and
<collectingRegion country="India" name="West Bengal" pageId="5" pageNumber="366">West Bengal</collectingRegion>
(
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;
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,
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).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="366" type="discussion">
<paragraph blockId="5.[813,1418,796,855]" pageId="5" pageNumber="366">
<emphasis bold="true" box="[813,924,796,820]" pageId="5" pageNumber="366">Remarks.</emphasis>
This species is widely distributed throughout the country.
</paragraph>
</subSubSection>
</treatment>
</document>