treatments-xml/data/60/F0/14/60F01412C606574AA4C48CA3F3E05F6C.xml
2024-06-21 12:38:05 +02:00

198 lines
20 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.884.32550" ID-GBIF-Dataset="b07c4790-847b-4e8c-9461-b4063d8a9a0e" ID-PMC="PMC6834725" ID-Pensoft-Pub="1313-2970-884-81" ID-Pensoft-UUID="8DBA9A85768052D8B3603886F5DA2BE8" ID-PubMed="31723328" ID-ZooBank="AC4289EF56024BB9807D578B858BB0B1" ModsDocID="1313-2970-884-81" checkinTime="1572509056192" checkinUser="pensoft" docAuthor="Yang, Yuxia, Xi, Huacong, Yang, Xingke &amp; Liu, Haoyu" docDate="2019" docId="60F01412C606574AA4C48CA3F3E05F6C" docLanguage="en" docName="ZooKeys 884: 81-106" docOrigin="ZooKeys 884" docSource="http://dx.doi.org/10.3897/zookeys.884.32550" docTitle="Themus (Telephorops) crassimargo Champion 1926" docType="treatment" docVersion="4" id="8DBA9A85768052D8B3603886F5DA2BE8" lastPageNumber="81" masterDocId="8DBA9A85768052D8B3603886F5DA2BE8" masterDocTitle="Taxonomic review of the Themus (Telephorops) nepalensis species-group (Coleoptera, Cantharidae)" masterLastPageNumber="106" masterPageNumber="81" pageNumber="81" updateTime="1668167998012" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic review of the Themus (Telephorops) nepalensis species-group (Coleoptera, Cantharidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Yuxia</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xi, Huacong</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Xingke</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Haoyu</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>884</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>81</mods:start>
<mods:end>106</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.884.32550</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.884.32550</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-884-81</mods:identifier>
<mods:identifier type="ZooBank">AC4289EF56024BB9807D578B858BB0B1</mods:identifier>
<mods:identifier type="Pensoft-UUID">8DBA9A85768052D8B3603886F5DA2BE8</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="159932939" LSID="urn:lsid:plazi:treatment:60F01412C606574AA4C48CA3F3E05F6C" httpUri="http://treatment.plazi.org/id/60F01412C606574AA4C48CA3F3E05F6C" lastPageNumber="81" pageId="0" pageNumber="81">
<subSubSection pageId="0" pageNumber="81" type="nomenclature">
<paragraph pageId="0" pageNumber="81">
<taxonomicName LSID="60F01412-C606-574A-A4C4-8CA3F3E05F6C" authority="Champion, 1926" authorityName="Champion" authorityYear="1926" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus (Telephorops) crassimargo" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="crassimargo" subGenus="Telephorops">Themus (Telephorops) crassimargo Champion, 1926</taxonomicName>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male habitus, dorsal view A Themus cavipennis Champion, 1926 (the specimen of Xizang) B T. crassimargo Champion, 1926 (the specimen of Sichuan). Scale bars: 5.0 mm." figureDoi="10.3897/zookeys.884.32550.figure3" httpUri="https://binary.pensoft.net/fig/351092" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.884.32550.figure3">Figs 3B</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus, dorsal view A Themus crassimargo Champion, 1926 (lectotype) B Themus (Telephorops) subcaeruleiformis Wittmer, 1983 (holotype) C Triblius laboissierei Pic, 1929 (lectotype) D Themus (Tryblius) separandus Wittmer, 1975 (holotype)." figureDoi="10.3897/zookeys.884.32550.figure4" httpUri="https://binary.pensoft.net/fig/351093" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.884.32550.figure4">4A, B</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female abdominal sternite VIII, ventral view A Themus bicoloricornis Wittmer, 1983 B T. cavipennis Champion, 1926 C T. coelestis (Gorham, 1889) D T. crassimargo Champion, 1926 E T. crassipes Pic, 1929 F T. impressipennis (Fairmaire, 1886) G T. laboissierei (Pic, 1921) H T. masatakai Okushima, 2003 I T. sauteri (Pic, 1912) J T. uncinatus Wittmer, 1983. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.884.32550.figure9" httpUri="https://binary.pensoft.net/fig/351098" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.884.32550.figure9">9D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="reference_group">
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus crassimargo" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="crassimargo">Themus crassimargo</taxonomicName>
Champion, 1926: 127.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus (Tryblius) crassimargo" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="crassimargo" subGenus="Tryblius">Themus (Tryblius) crassimargo</taxonomicName>
:
<bibRefCitation author="Wittmer, W" journalOrPublisher="Entomologica Basiliensia" pageId="0" pageNumber="81" pagination="249 - 278" refId="B22" refString="Wittmer, W, 1975. Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: lam. Cantharidae (2.Teil). Entomologica Basiliensia 1: 249 - 278" title="Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: lam. Cantharidae (2. Teil)." volume="1" year="1975">Wittmer 1975</bibRefCitation>
: 251, fig. 2 (aedeagus illustration).
</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus (Telephorops) subcaeruleiformis" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="subcaeruleiformis" subGenus="Telephorops">Themus (Telephorops) subcaeruleiformis</taxonomicName>
Wittmer, 1983b: 199, fig. 3, 3a (aedeagus illustration), syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus (Telephorops) crassimargo" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="crassimargo" subGenus="Telephorops">Themus (Telephorops) crassimargo</taxonomicName>
:
<bibRefCitation author="Okushima, Y" journalOrPublisher="Insecta Matsumurana" pageId="0" pageNumber="81" pagination="51 - 68" publicationUrl="http://hdl.handle.net/" refId="B12" refString="Okushima, Y, 1999. Cantharidae collected by the Hokkaido University Expeditions to Nepal Himalaya (Coleoptera). Insecta Matsumurana 56: 51 - 68" title="Cantharidae collected by the Hokkaido University Expeditions to Nepal Himalaya (Coleoptera)." url="http://hdl.handle.net/" volume="56" year="1999">Okushima 1999</bibRefCitation>
: 59, figs 11 (habitus photo), 35 (female abdominal sternite VIII illustration).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="type material">
<paragraph pageId="0" pageNumber="81">Type material examined.</paragraph>
<paragraph pageId="0" pageNumber="81">
1♂ (BMNH,
<emphasis bold="true" pageId="0" pageNumber="81">lectotype</emphasis>
of
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus crassimargo" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="crassimargo">
<emphasis italics="true" pageId="0" pageNumber="81">Themus crassimargo</emphasis>
</taxonomicName>
), [p]&quot;Gopaldhara, Sikkim, vii.1924, H. Stevens&quot;, [p]
<normalizedToken originalValue="“LECTOTYPUS”">&quot;LECTOTYPUS&quot;</normalizedToken>
, [p] &quot;Themus \ crassimargo \ Champ.&quot;. The lectotype was designated by
<bibRefCitation author="Wittmer, W" journalOrPublisher="Entomologica Basiliensia" pageId="0" pageNumber="81" pagination="249 - 278" refId="B22" refString="Wittmer, W, 1975. Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: lam. Cantharidae (2.Teil). Entomologica Basiliensia 1: 249 - 278" title="Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: lam. Cantharidae (2. Teil)." volume="1" year="1975">Wittmer (1975)</bibRefCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="81">
1♂ (NHMB, holotype of
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus (Telephorops) subcaeruleiformis" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="subcaeruleiformis" subGenus="Telephorops">Themus (Telephorops) subcaeruleiformis</taxonomicName>
), [h] &quot;Kuanshien (Guanxian, now is Dujiangyan) \ Umg. VIII.1934&quot;,
<normalizedToken originalValue="“600">&quot;600-</normalizedToken>
1300m \ Szechwan (Sichuan), China&quot;, [p]
<normalizedToken originalValue="“HOLOTYPUS”">&quot;HOLOTYPUS&quot;</normalizedToken>
, [h] &quot;T. (Telephorops) subcaerulei- \ formis Wittm. \ det. W. Wittmer&quot;, [p] &quot;Naturhist. \ Museum Basel \ coll. W. Wittmer&quot;, [p] &quot;CANTHARIDAE \ CANTH00001233&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="materials_examined">
<paragraph pageId="0" pageNumber="81">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">CHINA</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="81">Xizang</emphasis>
: 1♂ (CAUB),
<normalizedToken originalValue="Zayü">Zayue</normalizedToken>
, Shajiong, 1700 m, 1978.VI.26, leg. F.S. Li.
<emphasis bold="true" pageId="0" pageNumber="81">Sichuan</emphasis>
: 1♀ (IZAS), 70 km West Chengdu, Qingcheng, Hou Shan mts., 1360 m,
<geoCoordinate degrees="30" direction="north" minutes="44" orientation="latitude" precision="925" value="30.733334">30°44'N</geoCoordinate>
,
<geoCoordinate degrees="103" direction="east" minutes="08" orientation="longitude" precision="925" value="103.13333">103°08'E</geoCoordinate>
, 2004.VIII.28, S. Murzin.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="distribution">
<paragraph pageId="0" pageNumber="81">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="81">China (Xizang, Sichuan), N. India, Bhutan, Nepal.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="description">
<paragraph pageId="0" pageNumber="81">Supplementary description.</paragraph>
<paragraph pageId="0" pageNumber="81">
Female (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male habitus, dorsal view A Themus cavipennis Champion, 1926 (the specimen of Xizang) B T. crassimargo Champion, 1926 (the specimen of Sichuan). Scale bars: 5.0 mm." figureDoi="10.3897/zookeys.884.32550.figure3" httpUri="https://binary.pensoft.net/fig/351092" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.884.32550.figure3">Fig. 3B</figureCitation>
). Like male, but antennomeres
<normalizedToken originalValue="VXI">V-XI</normalizedToken>
without impressions along outer edges (while present with smooth narrow longitudinal or oblong impressions in male), terminal abdominal ventrite wide (while narrow and triangular in male) (
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female abdominal sternite VIII, ventral view A Themus bicoloricornis Wittmer, 1983 B T. cavipennis Champion, 1926 C T. coelestis (Gorham, 1889) D T. crassimargo Champion, 1926 E T. crassipes Pic, 1929 F T. impressipennis (Fairmaire, 1886) G T. laboissierei (Pic, 1921) H T. masatakai Okushima, 2003 I T. sauteri (Pic, 1912) J T. uncinatus Wittmer, 1983. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.884.32550.figure9" httpUri="https://binary.pensoft.net/fig/351098" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.884.32550.figure9">Fig. 9D</figureCitation>
) with posterior edge triangularly protuberant on each side, space between lateral protuberances about twice as wide as each width.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="remarks">
<paragraph pageId="0" pageNumber="81">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus (T.) subcaeruleiformis" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="subcaeruleiformis" subGenus="T.">Themus (T.) subcaeruleiformis</taxonomicName>
Wittmer, 1983 was originally described based on a single male type, from China, Szechwan, Kuanshien Umg. (now in Sichuan, Dujiangyan). Here a female (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male habitus, dorsal view A Themus cavipennis Champion, 1926 (the specimen of Xizang) B T. crassimargo Champion, 1926 (the specimen of Sichuan). Scale bars: 5.0 mm." figureDoi="10.3897/zookeys.884.32550.figure3" httpUri="https://binary.pensoft.net/fig/351092" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.884.32550.figure3">Fig. 3B</figureCitation>
) collected from Qingcheng, which is near the type locality, is discovered for the first time. The structure of its abdominal sternite VIII (
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female abdominal sternite VIII, ventral view A Themus bicoloricornis Wittmer, 1983 B T. cavipennis Champion, 1926 C T. coelestis (Gorham, 1889) D T. crassimargo Champion, 1926 E T. crassipes Pic, 1929 F T. impressipennis (Fairmaire, 1886) G T. laboissierei (Pic, 1921) H T. masatakai Okushima, 2003 I T. sauteri (Pic, 1912) J T. uncinatus Wittmer, 1983. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.884.32550.figure9" httpUri="https://binary.pensoft.net/fig/351098" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.884.32550.figure9">Fig. 9D</figureCitation>
) is like that of
<taxonomicName lsidName="T. crassimargo" pageId="0" pageNumber="81" rank="species" species="crassimargo">
<emphasis italics="true" pageId="0" pageNumber="81">T. crassimargo</emphasis>
</taxonomicName>
Champion, 1926, illustrated by
<bibRefCitation author="Okushima, Y" journalOrPublisher="Insecta Matsumurana" pageId="0" pageNumber="81" pagination="51 - 68" publicationUrl="http://hdl.handle.net/" refId="B12" refString="Okushima, Y, 1999. Cantharidae collected by the Hokkaido University Expeditions to Nepal Himalaya (Coleoptera). Insecta Matsumurana 56: 51 - 68" title="Cantharidae collected by the Hokkaido University Expeditions to Nepal Himalaya (Coleoptera)." url="http://hdl.handle.net/" volume="56" year="1999">Okushima (1999</bibRefCitation>
: fig. 35). Furthermore,
<taxonomicName lsidName="T. subcaeruleiformis" pageId="0" pageNumber="81" rank="species" species="subcaeruleiformis">
<emphasis italics="true" pageId="0" pageNumber="81">T. subcaeruleiformis</emphasis>
</taxonomicName>
was only compared with
<taxonomicName lsidName="T. subcaeruleus" pageId="0" pageNumber="81" rank="species" species="subcaeruleus">
<emphasis italics="true" pageId="0" pageNumber="81">T. subcaeruleus</emphasis>
</taxonomicName>
located in Yunnan, China in the original publication (
<bibRefCitation author="Wittmer, W" journalOrPublisher="Entomological Review of Japan" pageId="0" pageNumber="81" refId="B24" refString="Wittmer, W, 1983b. Beitrag zur einer Revision der Gattung ThemusMotsch. Coleoptera: Cantharidae. Entomologischen Arbeiten aus dem Museum G. Frey 31/32: 189-239." title="Beitrag zur einer Revision der Gattung ThemusMotsch. Coleoptera: Cantharidae. Entomologischen Arbeiten aus dem Museum G. Frey 31 / 32: 189 - 239." year="1983 b">Wittmer 1983b</bibRefCitation>
), but not with species from the Himalayas (
<bibRefCitation author="Wittmer, W" journalOrPublisher="Entomologica Basiliensia" pageId="0" pageNumber="81" pagination="249 - 278" refId="B22" refString="Wittmer, W, 1975. Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: lam. Cantharidae (2.Teil). Entomologica Basiliensia 1: 249 - 278" title="Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: lam. Cantharidae (2. Teil)." volume="1" year="1975">Wittmer 1975</bibRefCitation>
).Moreover, the types of
<taxonomicName lsidName="T. subcaeruleiformis" pageId="0" pageNumber="81" rank="species" species="subcaeruleiformis">
<emphasis italics="true" pageId="0" pageNumber="81">T. subcaeruleiformis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="T. crassimargo" pageId="0" pageNumber="81" rank="species" species="crassimargo">
<emphasis italics="true" pageId="0" pageNumber="81">T. crassimargo</emphasis>
</taxonomicName>
were compared, but no differences between them were found in external morphology (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus, dorsal view A Themus crassimargo Champion, 1926 (lectotype) B Themus (Telephorops) subcaeruleiformis Wittmer, 1983 (holotype) C Triblius laboissierei Pic, 1929 (lectotype) D Themus (Tryblius) separandus Wittmer, 1975 (holotype)." figureDoi="10.3897/zookeys.884.32550.figure4" httpUri="https://binary.pensoft.net/fig/351093" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.884.32550.figure4">Fig. 4A, B</figureCitation>
) and aedeagi illustrated by
<bibRefCitation author="Wittmer, W" journalOrPublisher="Entomologica Basiliensia" pageId="0" pageNumber="81" pagination="249 - 278" refId="B22" refString="Wittmer, W, 1975. Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: lam. Cantharidae (2.Teil). Entomologica Basiliensia 1: 249 - 278" title="Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: lam. Cantharidae (2. Teil)." volume="1" year="1975">Wittmer (1975</bibRefCitation>
: fig. 2; 1983b: fig. 3, 3a).Therefore,
<taxonomicName lsidName="T. subcaeruleiformis" pageId="0" pageNumber="81" rank="species" species="subcaeruleiformis">
<emphasis italics="true" pageId="0" pageNumber="81">T. subcaeruleiformis</emphasis>
</taxonomicName>
is proposed here to be junior synonym of
<taxonomicName lsidName="T. crassimargo" pageId="0" pageNumber="81" rank="species" species="crassimargo">
<emphasis italics="true" pageId="0" pageNumber="81">T. crassimargo</emphasis>
</taxonomicName>
, according to the Principle of Priority (
<bibRefCitation author="Hope, FW" journalOrPublisher=") Zoologica Miscellanea" pageId="0" pageNumber="81" refId="B7" refString="ICZN (1999) International Code of Zoological Nomenclature (Fourth Edition)." year="1831">ICZN 1999</bibRefCitation>
, Article 23.1).
</paragraph>
</subSubSection>
</treatment>
</document>