treatments-xml/data/03/A0/87/03A087C1FFE8FF98FF717957FD175200.xml

<document id="CBA8CD6E278F8DD2EF281ACA50EEBDB8" ID-CLB-Dataset="72883" ID-DOI="10.11646/zootaxa.5150.3.7" ID-GBIF-Dataset="6ca82f5e-b51d-4beb-b33f-00a76c6a7bc8" ID-ISSN="1175-5326" ID-Zenodo-Dep="6623165" ID-ZooBank="A379F26C-DCAB-43A8-B075-ED4ED10D8476" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" checkinTime="1654677040107" checkinUser="plazi" docAuthor="Tang, William &amp; Dominguez, Hector Gomez" docDate="2022" docId="03A087C1FFE8FF98FF717957FD175200" docLanguage="en" docName="zootaxa.5150.3.7.pdf" docOrigin="Zootaxa 5150 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceratophila (Vovidesa) chipjonesi Tang &amp; Gomez Dominguez 2022, new species" docType="treatment" docVersion="9" lastPageNumber="434" masterDocId="FF99FFB9FFECFF9EFFE67A00FFEE5019" masterDocTitle="Two new species of Pharaxonothinae beetles (Coleoptera: Erotylidae) inhabiting cones of the cycad Ceratozamia santillanii Pérez-Farr. &amp; Vovides (Cycadales: Zamiaceae) in Mexico" masterLastPageNumber="442" masterPageNumber="428" pageNumber="432" updateTime="1728482944120" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="58A7C6A799AD0E28AF7083F640F93B0B">Two new species of Pharaxonothinae beetles (Coleoptera: Erotylidae) inhabiting cones of the cycad Ceratozamia santillanii Pérez-Farr. &amp; Vovides (Cycadales: Zamiaceae) in Mexico</mods:title>
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<mods:namePart id="EC83E955D7A9210990B9C307F6F87DA6">Tang, William</mods:namePart>
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<mods:affiliation id="D2F7D8E49B1E603A3848273ADE4845AD">Florida State Collection of Arthropods, Florida Department of Agriculture - DPI, P. O. Box 147100, Gainesville, FL 32614 - 7100, USA. wlmtang 60 @ gmail. com; https: // orcid. org / 0000 - 0001 - 9009 - 3113</mods:affiliation>
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<mods:namePart id="0B062DACF010F306EF8B84B2DC77C18F">Dominguez, Hector Gomez</mods:namePart>
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<mods:affiliation id="960EB46D5A810CFDBFDE4FE237B2015D">Senda Sustentable A. C., Calle Cerro Azul, 208 B, Fraccionamiento Lomas del Pedregal, Berriozabal, CP 29130, Chiapas, Mexico hectorgomezdominguez @ hotmail. com; https: // orcid. org / 0000 - 0001 - 7125 - 4639</mods:affiliation>
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<subSubSection id="C313655CFFE8FF9AFF717957FBC9536B" box="[151,1063,855,882]" pageId="4" pageNumber="432" type="nomenclature">
<paragraph id="8BB636D7FFE8FF9AFF717957FBC9536B" blockId="4.[151,1063,855,919]" box="[151,1063,855,882]" pageId="4" pageNumber="432">
<heading id="D0FE81BBFFE8FF9AFF717957FBC9536B" bold="true" box="[151,1063,855,882]" fontSize="11" level="1" pageId="4" pageNumber="432" reason="1">
<emphasis id="B97DEAC5FFE8FF9AFF717957FBC9536B" bold="true" box="[151,1063,855,882]" pageId="4" pageNumber="432">
<taxonomicName id="4C094D54FFE8FF9AFF717957FC64536B" authority="Tang &amp; Gomez Dominguez" authorityName="Tang &amp; Gomez Dominguez" authorityYear="2022" box="[151,906,855,882]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="chipjonesi" status="sp. nov." subGenus="Vovidesa">
<emphasis id="B97DEAC5FFE8FF9AFF717957FEC8536B" bold="true" box="[151,294,855,882]" italics="true" pageId="4" pageNumber="432">Ceratophila</emphasis>
(
<emphasis id="B97DEAC5FFE8FF9AFEDE7957FE4C536B" bold="true" box="[312,418,855,882]" italics="true" pageId="4" pageNumber="432">Vovidesa</emphasis>
) 
<emphasis id="B97DEAC5FFE8FF9AFE557957FDC1536B" bold="true" box="[435,559,855,882]" italics="true" pageId="4" pageNumber="432">chipjonesi</emphasis>
Tang &amp; Gomez Dominguez
</taxonomicName>
, 
<taxonomicNameLabel id="A24E57BEFFE8FF9AFC717958FBC9536B" box="[919,1063,856,882]" pageId="4" pageNumber="432" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
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<subSubSection id="C313655CFFE8FF9AFF71797CFE98538E" box="[151,374,892,919]" pageId="4" pageNumber="432" type="description">
<paragraph id="8BB636D7FFE8FF9AFF71797CFE98538E" blockId="4.[151,1063,855,919]" box="[151,374,892,919]" pageId="4" pageNumber="432">
(
<figureCitation id="13322A52FFE8FF9AFF79797CFEF3538F" box="[159,285,892,918]" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="4" pageNumber="432">Figs. 4A–J</figureCitation>
, 
<figureCitation id="13322A52FFE8FF9AFECE797CFE80538E" box="[296,366,892,919]" captionStart="FIGURE 5" captionStartId="8.[151,250,1834,1858]" captionTargetBox="[151,1436,181,1810]" captionTargetId="figure-21@8.[151,1436,181,1810]" captionTargetPageId="8" captionText="FIGURE 5. Ceratophila chipjonesi female: A–C) allotype dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views; G) female genitalia, ventral view; H) spermatheca. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G; black = 0.1 mm for H." figureDoi="http://doi.org/10.5281/zenodo.6623181" httpUri="https://zenodo.org/record/6623181/files/figure.png" pageId="4" pageNumber="432">5A–H</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C313655CFFE8FF9AFF7179C4FD82555F" pageId="4" pageNumber="432" type="diagnosis">
<paragraph id="8BB636D7FFE8FF9AFF7179C4FD625477" blockId="4.[151,1437,964,2034]" pageId="4" pageNumber="432">
<emphasis id="B97DEAC5FFE8FF9AFF7179C4FEB953C7" bold="true" box="[151,343,964,990]" pageId="4" pageNumber="432">Adult diagnosis.</emphasis>
Distinguished from other 
<taxonomicName id="4C094D54FFE8FF9AFD9F79C4FC9253C7" box="[633,892,964,990]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="432" phylum="Arthropoda" rank="subGenus" subGenus="Vovidesa">
<emphasis id="B97DEAC5FFE8FF9AFD9F79C4FCEE53C7" box="[633,768,964,990]" italics="true" pageId="4" pageNumber="432">Ceratophila</emphasis>
(
<emphasis id="B97DEAC5FFE8FF9AFCE979C4FC9D53C7" box="[783,883,964,990]" italics="true" pageId="4" pageNumber="432">Vovidesa</emphasis>
)
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species by male with subapical emarginations on tibiae, submentum of both sexes with dense punctures and setae, male submental setae projecting laterally, pronotum relatively long and narrow with pronotal width to pronotal width = 0.78–0.86; male genitalia with median lobe nearly laterally flattened and curved, parameres triangular in lateral view, and with a known distribution in 
<collectingCountry id="F31E7647FFE8FF9AFAA67E30FA765453" box="[1344,1432,1072,1098]" name="Mexico" pageId="4" pageNumber="432">Mexico</collectingCountry>
, state of 
<collectingRegion id="49CDF835FFE8FF9AFF167E54FEA35477" box="[240,333,1108,1134]" country="Mexico" name="Chiapas" pageId="4" pageNumber="432">Chiapas</collectingRegion>
, on 
<emphasis id="B97DEAC5FFE8FF9AFE9D7E54FD625477" box="[379,652,1108,1134]" italics="true" pageId="4" pageNumber="432">
<taxonomicName id="4C094D54FFE8FF9AFE9D7E54FD665477" box="[379,648,1108,1134]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="432" phylum="Arthropoda" rank="species" species="santillanii">Ceratozamia santillanii</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BB636D7FFE8FF9AFF217E78FD82555F" blockId="4.[151,1437,964,2034]" pageId="4" pageNumber="432">
<emphasis id="B97DEAC5FFE8FF9AFF217E78FE0F548B" bold="true" box="[199,481,1144,1170]" pageId="4" pageNumber="432">Adult male description.</emphasis>
Length 
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, width 
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(n = 10). Body in dorsal view elongateoval, greatest width at middle of elytra; in lateral view convex dorsally. General body color brown (
<figureCitation id="13322A52FFE8FF9AFAED7E9CFA7E54AF" box="[1291,1424,1180,1206]" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="4" pageNumber="432">Figs. 4A–F</figureCitation>
), head, pronotum and elytra concolorous; dorsal surface punctate, shining, short procumbent hairs associated with punctation; ventrally shining except meso- and metaventrites and oval areas on the lateral portions of the abdominal ventrites subshining to alutaceous, appearing glabrous except meso- and metaventrites and shining portions of abdomen covered with long procumbent setae.
</paragraph>
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<subSubSection id="C313655CFFE8FF9BFF217F50FA7B5680" lastPageId="5" lastPageNumber="433" pageId="4" pageNumber="432" type="description">
<paragraph id="8BB636D7FFE8FF9AFF217F50FB5C56CB" blockId="4.[151,1437,964,2034]" pageId="4" pageNumber="432">
<emphasis id="B97DEAC5FFE8FF9AFF217F50FEE95573" bold="true" box="[199,263,1360,1386]" pageId="4" pageNumber="432">Head</emphasis>
in dorsal view conical, gradually narrowed anteriorly, surface flat to slightly convex, finely, moderately punctured (
<figureCitation id="13322A52FFE8FF9AFEFE7F74FE4F5597" box="[280,417,1396,1422]" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="4" pageNumber="432">Figs. 4A, D</figureCitation>
), average distance between closest punctures 2× width of puncture; width 
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; dorsal interocular distance 
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, head width/dorsal interocular distance ratio 1.55–1.63, ventral interocular distance 
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, head width/ventral interocular distance ratio 2.07–2.24. Eye with large black facets. Antennal length approximately equal to pronotal width, 1.5× head width; antennomere I (scape) fairly large, slightly elongate, antennomere II subequal to III; IV–VIII small, width equals length; club fairly large, IX and X similar in length, XI slightly longer, globular with acuminate apex. Clypeus truncate anteriorly, moderately punctate, with narrow margin. Mentum with moderate punctation, distance between nearest punctures ~ 1× own width; submentum finely, densely punctate, each puncture with a long seta ~ 2–3× width of eye facet, setae projecting laterally (
<figureCitation id="13322A52FFE8FF9AFA827C70FF3756B7" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="4" pageNumber="432">Figs. 4E–F</figureCitation>
). Gular area smooth, without punctation or setae, border with submentum usually marked with a shallow depression without punctation. Mandibles arcuate-triangular, apically with 3 teeth (
<figureCitation id="13322A52FFE8FF9AFBC67CB8FB4B56CB" box="[1056,1189,1720,1746]" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="4" pageNumber="432">Figs. 4D–E</figureCitation>
).
</paragraph>
<paragraph id="8BB636D7FFE8FF9BFF217CDCFCBC52D4" blockId="4.[151,1437,964,2034]" lastBlockId="5.[151,1437,151,2014]" lastPageId="5" lastPageNumber="433" pageId="4" pageNumber="432">
<emphasis id="B97DEAC5FFE8FF9AFF217CDCFECE56EF" bold="true" box="[199,288,1756,1782]" pageId="4" pageNumber="432">Thorax</emphasis>
with pronotum moderately transverse in dorsal view with marginal beads basally and laterally, anterior margin with weak bead, reduced to absent medially; length/width ratio (PL/PW) = 0.81–0.86, rectangular; anterior and posterior angles present, anterior angles projecting; lateral carinae expanding in basal 1/10 to parallel sides, sometimes slightly emarginated, parallel for 50% of length, then gradually converging to anterior angles; posterior margin slightly projecting medially, projection beginning approximately by pair of small, dark pores located 1/5 length of posterior margin from posterior corners and touching and just anterior to marginal bead, pores mark base of longitudinal furrows onto disc. Prosternum in ventral view convex; anterior margin slightly emarginate, finely denticulate with row of long, anteriorly directed setae, longest setae ~ 1/3 length of eye. Hypomeron laterally apparently lacking punctures; medially with longitudinal striations. Elytra in dorsal view elongate-oval, convex; length/width 1.74–1.90, greatest width near midlength; without marginal line basally; 10 complete striae of moderate puncture size, scutellary striole extending ¼ elytral length, with 9–14 punctures; intervals of striae with fine, shallow punctures of similar size to strial punctures. All punctures of elytra bearing a single fine, procumbent seta; seta often only visible in profile (often abraded), due to poor preservation unclear whether interval puncture setae are longer than strial puncture setae as in other members of the subgenus. Punctation on meso- and metaventrite moderate to dense, distance between nearest punctures 1–2× width of punctures, puncture depth shallow. Metaventrite long, convex laterally, slightly impressed medially, metathoracic discrimen extending slightly over 3/4 metaventrite length. Legs with procoxa oval; mesocoxa globular; metacoxa transversely elongate-oval; trochanters obliquely truncate apically; in males all femora swollen, stout, dorso-ventral width greater than eye length (
<figureCitation id="13322A52FFE9FF9BFA827BDBFF305200" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="5" pageNumber="433">Figs. 4A–C</figureCitation>
); protibia with shallow medial subapical emargination, meso- and metatibia with a distinct medial subapical emargination 
<figureCitation id="13322A52FFE9FF9BFED57823FE555224" box="[307,443,547,573]" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="5" pageNumber="433">Figs. 4A, C</figureCitation>
); all tibiae obliquely truncate at apex, with complete fringe of fine spinules on anterior and posterior margins; protibia swollen, not flattened in cross section notably larger than other tibiae, triangularly dilated to apex; mesotibia weakly swollen, moderately triangularly dilated to apex; metatibia narrowed, weakly dilating to midlength then slightly narrowing to apex; pro- and mesotarsi greatly enlarged, corresponding tarsomere I width ~ 3× own length, metatarsomere I width ~ own length.
</paragraph>
<paragraph id="8BB636D7FFE9FF9BFF2178D6FA9154DC" blockId="5.[151,1437,151,2014]" pageId="5" pageNumber="433">
<emphasis id="B97DEAC5FFE9FF9BFF2178D6FEAD52E9" bold="true" box="[199,323,726,752]" pageId="5" pageNumber="433">Abdomen.</emphasis>
Ventrite I with intercoxal process narrow, with triangular point anteromedially, lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I longer medially than II; II–IV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex; apical margin bearing short, dense setae; all ventrites strongly alutaceous laterally, less so and glossy medially, bearing dense, moderate, shallow punctation, distance to nearest puncture ~ 1× width of puncture, punctures bearing mostly reclining setae; ventrites I–V with setae length ~ 2–3× width of puncture; I–V each with 2 or more median pairs of longer, semi-erect sensory hairs obscured in dense setation, V with additional 4–5 pairs of long semi-erect hairs located postero-laterally, but not at submargin. Male genitalia (n=3) with genital capsule fringed with fine setae (
<figureCitation id="13322A52FFE9FF9BFCDC79F7FC7B5408" box="[826,917,1015,1041]" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="5" pageNumber="433">Fig. 4H</figureCitation>
); as in all other known members of the genus, tegmen and median lobe held upside down in body (
<bibRefCitation id="EF984B26FFE9FF9BFD3D7E1BFC4D542F" author="Tang, W. &amp; Perez-Farrera, M. A." box="[731,931,1051,1078]" pageId="5" pageNumber="433" pagination="151 - 178" refId="ref10958" refString="Tang, W., Skelley &amp; P. &amp; Perez-Farrera, M. A. (2018 a) Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae). Zootaxa, 4508 (2), 151 - 178. https: // doi. org / 10.11646 / zootaxa. 4508.2.1" type="journal article" year="2018">
Tang 
<emphasis id="B97DEAC5FFE9FF9BFCFD7E1BFCA2542C" box="[795,844,1051,1077]" italics="true" pageId="5" pageNumber="433">et al</emphasis>
. 2018a
</bibRefCitation>
,b), tegmen sclerotized, triangular, (
<figureCitation id="13322A52FFE9FF9BFACC7E1BFA61542C" box="[1322,1423,1051,1077]" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="5" pageNumber="433">Figs. 4G</figureCitation>
); parameres in dorso-ventral view approximately parallel-sided, length/width ratio = 2.3–2.7, rounded apex with 6+ setae, 1 or 2 setae also visibly projecting just posteriad of center of inner margins, length of all setae &lt;width of paramere, in lateral view parameres triangular (
<figureCitation id="13322A52FFE9FF9BFD957E87FD0854B8" box="[627,742,1159,1185]" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="5" pageNumber="433">Figs. 4I–J</figureCitation>
); median lobe laterally compressed, sclerotized, in lateral view strongly curved, apex tapering to a point. Ratio of penile strut length to median lobe length 1.65–1.82 (
<figureCitation id="13322A52FFE9FF9BFAFE7EABFA9C54DC" box="[1304,1394,1195,1221]" captionStart="FIGURE 4" captionStartId="7.[151,250,1782,1806]" captionTargetBox="[266,1322,181,1757]" captionTargetId="figure-26@7.[266,1322,181,1758]" captionTargetPageId="7" captionText="FIGURE 4. Ceratophila chipjonesi male holotype: A–F: A–C) dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views, arrow indicates tuff of setae projecting laterally from submentum; G) tegmen attached to median lobe, (top to bottom) ventral and lateral views; H) male genital capsule, ventral view; I) parameres of tegmen, ventral view; J) paramere, lateral view. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G &amp; H; black = 0.1 mm for I &amp; J." figureDoi="http://doi.org/10.5281/zenodo.6623177" httpUri="https://zenodo.org/record/6623177/files/figure.png" pageId="5" pageNumber="433">Fig. 4G</figureCitation>
).
</paragraph>
<paragraph id="8BB636D7FFE9FF9BFF217ECFFA7B5680" blockId="5.[151,1437,151,2014]" pageId="5" pageNumber="433">
<emphasis id="B97DEAC5FFE9FF9BFF217ECFFECB54F0" bold="true" box="[199,293,1231,1257]" pageId="5" pageNumber="433">Female.</emphasis>
Generally similar to male, except for sexual dimorphism (
<figureCitation id="13322A52FFE9FF9BFC5D7ECFFBAE54F0" box="[955,1088,1230,1257]" captionStart="FIGURE 5" captionStartId="8.[151,250,1834,1858]" captionTargetBox="[151,1436,181,1810]" captionTargetId="figure-21@8.[151,1436,181,1810]" captionTargetPageId="8" captionText="FIGURE 5. Ceratophila chipjonesi female: A–C) allotype dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views; G) female genitalia, ventral view; H) spermatheca. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G; black = 0.1 mm for H." figureDoi="http://doi.org/10.5281/zenodo.6623181" httpUri="https://zenodo.org/record/6623181/files/figure.png" pageId="5" pageNumber="433">Figs. 5A–F</figureCitation>
). Length 
<quantity id="4CF19B32FFE9FF9BFB4C7ECFFABE54F0" box="[1194,1360,1231,1257]" metricMagnitude="-3" metricUnit="m" metricValue="4.265" metricValueMax="4.84" metricValueMin="3.69" pageId="5" pageNumber="433" unit="mm" value="4.265" valueMax="4.84" valueMin="3.69">3.69–4.84 mm</quantity>
, width 
<quantity id="4CF19B32FFE9FF9BFF717EF3FED35514" box="[151,317,1267,1293]" metricMagnitude="-3" metricUnit="m" metricValue="1.595" metricValueMax="1.77" metricValueMin="1.42" pageId="5" pageNumber="433" unit="mm" value="1.595" valueMax="1.77" valueMin="1.42">1.42–1.77 mm</quantity>
(n = 13). Head width 
<quantity id="4CF19B32FFE9FF9BFDA37EF3FD005514" box="[581,750,1267,1293]" metricMagnitude="-4" metricUnit="m" metricValue="7.949999999999999" metricValueMax="8.7" metricValueMin="7.199999999999999" pageId="5" pageNumber="433" unit="mm" value="0.7949999999999999" valueMax="0.87" valueMin="0.72">0.72–0.87 mm</quantity>
; dorsal interocular distance 
<quantity id="4CF19B32FFE9FF9BFBD47EF2FB345514" box="[1074,1242,1266,1293]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.4" metricValueMin="4.6" pageId="5" pageNumber="433" unit="mm" value="0.5" valueMax="0.54" valueMin="0.46">0.46–0.54 mm</quantity>
, head width/dorsal interocular distance ratio 1.55–1.63, ventral interocular distance 
<quantity id="4CF19B32FFE9FF9BFC717F17FBD35528" box="[919,1085,1303,1329]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.9000000000000004" metricValueMin="3.3000000000000003" pageId="5" pageNumber="433" unit="mm" value="0.36" valueMax="0.39" valueMin="0.33">0.33–0.39 mm</quantity>
, head width/ventral interocular distance ratio 2.09–2.33; submentum densely punctate, but unlike male setae short, not projecting (
<figureCitation id="13322A52FFE9FF9BFAEB7F3BFA7E554C" box="[1293,1424,1338,1365]" captionStart="FIGURE 5" captionStartId="8.[151,250,1834,1858]" captionTargetBox="[151,1436,181,1810]" captionTargetId="figure-21@8.[151,1436,181,1810]" captionTargetPageId="8" captionText="FIGURE 5. Ceratophila chipjonesi female: A–C) allotype dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views; G) female genitalia, ventral view; H) spermatheca. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G; black = 0.1 mm for H." figureDoi="http://doi.org/10.5281/zenodo.6623181" httpUri="https://zenodo.org/record/6623181/files/figure.png" pageId="5" pageNumber="433">Figs. 5E–F</figureCitation>
). Pronotum on average slightly wider than in male, PL/PW = 0.78–0.82; elytra length/width 1.70–1.80, on average shorter than male; legs unmodified, fore and middle legs not swollen, femora dorso-ventral width about equal to eye length, all tibia gradually dilating to narrowly triangular apex; tibia lacking median subapical emarginations; all legs with tarsomere I width ~ own length; metaventrite flattened medially, not impressed; metaventrite and abdomen less densely punctate and setose than in male. Genitalia (n = 3): gonostylus set apically on gonocoxite, gonostylus cylindrical, slightly tapering toward apex, length = 2.6–3.3× width (
<figureCitation id="13322A52FFE9FF9BFC637C13FC335634" box="[901,989,1554,1581]" captionStart="FIGURE 5" captionStartId="8.[151,250,1834,1858]" captionTargetBox="[151,1436,181,1810]" captionTargetId="figure-21@8.[151,1436,181,1810]" captionTargetPageId="8" captionText="FIGURE 5. Ceratophila chipjonesi female: A–C) allotype dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views; G) female genitalia, ventral view; H) spermatheca. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G; black = 0.1 mm for H." figureDoi="http://doi.org/10.5281/zenodo.6623181" httpUri="https://zenodo.org/record/6623181/files/figure.png" pageId="5" pageNumber="433">Fig. 5G</figureCitation>
); spermatheca C-shaped, surface texture smooth throughout, apex rounded, arc of unsclerotized tissue appearing very small or absent in central third, spermathecal duct located at tip of base, glandular duct attached dorsally on basal third midway between apex and arc of unsclerotized tissue, short and connected to a spherical gland with diameter &lt;width of spermatheca base (
<figureCitation id="13322A52FFE9FF9BFAC97C7FFA665680" box="[1327,1416,1662,1689]" captionStart="FIGURE 5" captionStartId="8.[151,250,1834,1858]" captionTargetBox="[151,1436,181,1810]" captionTargetId="figure-21@8.[151,1436,181,1810]" captionTargetPageId="8" captionText="FIGURE 5. Ceratophila chipjonesi female: A–C) allotype dorsal, lateral and ventral habitus; D–F) head dorsal, lateral and ventral views; G) female genitalia, ventral view; H) spermatheca. Scale bars: white = 1 mm for A, B &amp; C; brown = 0.5 mm for D, E &amp; F; gray = 0.5 mm for G; black = 0.1 mm for H." figureDoi="http://doi.org/10.5281/zenodo.6623181" httpUri="https://zenodo.org/record/6623181/files/figure.png" pageId="5" pageNumber="433">Fig. 5H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C313655CFFE9FF9BFF217CA3FB5757C4" pageId="5" pageNumber="433" type="materials_examined">
<paragraph id="8BB636D7FFE9FF9BFF217CA3FD8F56F8" blockId="5.[151,1437,151,2014]" pageId="5" pageNumber="433">
<materialsCitation id="3B613C8AFFE9FF9BFF217CA3FDB056F8" ID-GBIF-Occurrence="3808181305" collectorName="Reserva de Biosfera Selva el Ocote" country="Mexico" location="Reserva de Biosfera Selva" pageId="5" pageNumber="433" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<emphasis id="B97DEAC5FFE9FF9BFF217CA3FE8B56A5" bold="true" box="[199,357,1698,1725]" pageId="5" pageNumber="433">
<typeStatus id="54B28875FFE9FF9BFF217CA3FEEC56A4" box="[199,258,1699,1725]" pageId="5" pageNumber="433">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31E7647FFE9FF9BFE8D7CA3FE2D56A4" box="[363,451,1699,1725]" name="Mexico" pageId="5" pageNumber="433">Mexico</collectingCountry>
, 
<collectingRegion id="49CDF835FFE9FF9BFE287CA3FDC556A4" box="[462,555,1699,1725]" country="Mexico" name="Chiapas" pageId="5" pageNumber="433">Chiapas</collectingRegion>
, near the border of 
<collectorName id="26FC5301FFE9FF9BFCE57CA3FB6356A4" box="[771,1165,1699,1725]" pageId="5" pageNumber="433">
<location id="8ED6600CFFE9FF9BFCE57CA3FBC656A4" LSID="urn:lsid:plazi:treatment:03A087C1FFE8FF98FF717957FD175200:8ED6600CFFE9FF9BFCE57CA3FBC656A4" box="[771,1064,1699,1725]" country="Mexico" name="Reserva de Biosfera Selva" pageId="5" pageNumber="433" stateProvince="Chiapas">Reserva de Biosfera Selva</location>
el Ocote
</collectorName>
[GPS and locality name omitted], forest understory on limestone
</materialsCitation>
.
</paragraph>
<paragraph id="8BB636D7FFE9FF9BFF217CEBFAA05730" blockId="5.[151,1437,151,2014]" pageId="5" pageNumber="433">
<emphasis id="B97DEAC5FFE9FF9BFF217CEBFEF4571C" bold="true" box="[199,282,1771,1797]" pageId="5" pageNumber="433">Range.</emphasis>
The original description of the host of this beetle species, 
<taxonomicName id="4C094D54FFE9FF9BFC467CEBFB47571C" box="[928,1193,1771,1797]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="santillanii">
<emphasis id="B97DEAC5FFE9FF9BFC467CEBFB47571C" box="[928,1193,1771,1797]" italics="true" pageId="5" pageNumber="433">Ceratozamia santillanii</emphasis>
</taxonomicName>
, indicates its range as the northern highlands of 
<collectingRegion id="49CDF835FFE9FF9BFE5E7D0FFDBB5730" box="[440,597,1807,1833]" country="Mexico" name="Chiapas" pageId="5" pageNumber="433">Chiapas State</collectingRegion>
, including the Ocote Biosphere Reserve (
<bibRefCitation id="EF984B26FFE9FF9BFBC37D0FFAAC5730" author="Perez-Farrera, M. A. &amp; Vovides, A. P. &amp; Martinez-Camilo, R. &amp; Martinez-Melendez, N. &amp; Iglesias, C." box="[1061,1346,1807,1833]" pageId="5" pageNumber="433" pagination="433 - 443" refId="ref10238" refString="Perez-Farrera, M. A., Vovides, A. P., Martinez-Camilo, R., Martinez-Melendez, N. &amp; Iglesias, C. (2009) A reassessment of the Ceratozamia miqueliana species complex (Zamiaceae) of southeastern Mexico, with comments on species relationships. Systematics and Biodiversity, 7, 433 - 443. https: // doi. org / 10.1017 / S 1477200009990211" type="journal article" year="2009">
Pérez-Farrera 
<emphasis id="B97DEAC5FFE9FF9BFB207D0FFB195730" box="[1222,1271,1807,1833]" italics="true" pageId="5" pageNumber="433">et al</emphasis>
. 2009
</bibRefCitation>
).
</paragraph>
<paragraph id="8BB636D7FFE9FF9BFF217D32FB5757C4" blockId="5.[151,1437,151,2014]" pageId="5" pageNumber="433">
<emphasis id="B97DEAC5FFE9FF9BFF217D32FE5F5755" bold="true" box="[199,433,1842,1868]" pageId="5" pageNumber="433">Material examined.</emphasis>
<materialsCitation id="3B613C8AFFE9FF9BFE5E7D33FBB7578C" ID-GBIF-Occurrence="3808181306" collectingDate="2021-05-16" collectorName="H. Gomez Dominguez &amp; M. Jones" country="Mexico" location="Chis" pageId="5" pageNumber="433" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B97DEAC5FFE9FF9BFE5E7D33FDCB5754" bold="true" box="[440,549,1843,1869]" pageId="5" pageNumber="433">
<typeStatus id="54B28875FFE9FF9BFE5E7D33FDCB5754" box="[440,549,1843,1869]" pageId="5" pageNumber="433" type="holotype">Holotype</typeStatus>
</emphasis>
(by designation) male of 
<taxonomicName id="4C094D54FFE9FF9BFCA17D33FC325754" authorityName="Tang &amp; Gomez Dominguez" authorityYear="2022" box="[839,988,1843,1869]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="chipjonesi">
<emphasis id="B97DEAC5FFE9FF9BFCA17D33FC325754" box="[839,988,1843,1869]" italics="true" pageId="5" pageNumber="433">C. chipjonesi</emphasis>
</taxonomicName>
with the following labels: 1) [rectangular; white; printed in black ink] “ 
<collectingCountry id="F31E7647FFE9FF9BFDEE7D57FD925768" box="[520,636,1879,1905]" name="Mexico" pageId="5" pageNumber="433">MEXICO</collectingCountry>
, 
<location id="8ED6600CFFE9FF9BFD617D57FD535768" LSID="urn:lsid:plazi:treatment:03A087C1FFE8FF98FF717957FD175200:8ED6600CFFE9FF9BFD617D57FD535768" box="[647,701,1879,1905]" country="Mexico" name="Chis" pageId="5" pageNumber="433">Chis</location>
, [GPS and locality omitted], cone 
<taxonomicName id="4C094D54FFE9FF9BFBA47D57FAA25768" box="[1090,1356,1879,1905]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="santillanii">
<emphasis id="B97DEAC5FFE9FF9BFBA47D57FAA25768" box="[1090,1356,1879,1905]" italics="true" pageId="5" pageNumber="433">Ceratozamia santillanii</emphasis>
</taxonomicName>
<specimenCount id="9D0FFD5EFFE9FF9BFAB57D57FA845768" box="[1363,1386,1879,1905]" pageId="5" pageNumber="433" type="male">♂</specimenCount>
, 
<date id="FFB71017FFE9FF9BFA917D57FF03578C" pageId="5" pageNumber="433" value="2021-05-16">
<collectingDate id="EFF3E9FFFFE9FF9BFA917D57FF03578C" pageId="5" pageNumber="433" value="2021-05-16">16- V-2021</collectingDate>
</date>
, 
<collectorName id="26FC5301FFE9FF9BFF117D7BFE1D578C" box="[247,499,1915,1941]" pageId="5" pageNumber="433">H. Gomez Dominguez</collectorName>
&amp; 
<collectorName id="26FC5301FFE9FF9BFDF37D7BFD91578C" box="[533,639,1915,1941]" pageId="5" pageNumber="433">M. Jones</collectorName>
”; 2) [rectangular; red; printed in black ink]
</materialsCitation>
<materialsCitation id="3B613C8AFFE9FF9BFB867D7BFC5F57A0" ID-GBIF-Occurrence="3808181308" collectionCode="FSCA" country="Mexico" pageId="5" pageNumber="433" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54B28875FFE9FF9BFB867D7BFB1A578C" box="[1120,1268,1915,1941]" pageId="5" pageNumber="433" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D0FFD5EFFE9FF9BFB1C7D7BFAE1578C" box="[1274,1295,1915,1941]" pageId="5" pageNumber="433" type="male">♂</specimenCount>
<taxonomicName id="4C094D54FFE9FF9BFAF37D7BFD7B57A0" authority="Tang &amp; Gomez Dominguez 2022" authorityName="Tang &amp; Gomez Dominguez" authorityYear="2022" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="chipjonesi">
<emphasis id="B97DEAC5FFE9FF9BFAF37D7BFEE457A0" italics="true" pageId="5" pageNumber="433">Ceratophila chipjonesi</emphasis>
Tang &amp; Gomez Dominguez 2022
</taxonomicName>
. Deposited in the 
<collectionCode id="ED18AE12FFE9FF9BFC817D9FFC5F57A0" box="[871,945,1951,1977]" country="USA" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="433" type="Museum">FSCA</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B613C8AFFE9FF9BFC5B7D9EFB5B57C4" ID-GBIF-Occurrence="3808181302" collectionCode="FSCA, ANIC, IEXA, NHMUK, NZAC, UNAM, USNM" country="Mexico" pageId="5" pageNumber="433" specimenCount="39" specimenCount-adult="38" typeStatus="allotype">
<emphasis id="B97DEAC5FFE9FF9BFC5B7D9EFBCC57A1" bold="true" box="[957,1058,1950,1976]" pageId="5" pageNumber="433">
<typeStatus id="54B28875FFE9FF9BFC5B7D9EFBCC57A1" box="[957,1058,1950,1976]" pageId="5" pageNumber="433" type="allotype">Allotype</typeStatus>
</emphasis>
(
<collectionCode id="ED18AE12FFE9FF9BFBD57D9FFB9257A0" box="[1075,1148,1951,1977]" country="USA" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="433" type="Museum">FSCA</collectionCode>
) and 
<specimenCount id="9D0FFD5EFFE9FF9BFB597D9FFAF357A0" box="[1215,1309,1951,1977]" pageId="5" pageNumber="433" type="adult">38 adult</specimenCount>
<emphasis id="B97DEAC5FFE9FF9BFAC07D9FFA7257A0" bold="true" box="[1318,1436,1951,1977]" pageId="5" pageNumber="433">
<typeStatus id="54B28875FFE9FF9BFAC07D9FFA7257A0" box="[1318,1436,1951,1977]" pageId="5" pageNumber="433" type="paratype">paratypes</typeStatus>
</emphasis>
(same data as holotype) deposited at 
<collectionCode id="ED18AE12FFE9FF9BFDD57DC3FD9457C4" box="[563,634,1987,2013]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="5" pageNumber="433">ANIC</collectionCode>
, 
<collectionCode id="ED18AE12FFE9FF9BFD607DC3FD2157C4" box="[646,719,1987,2013]" country="USA" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="433" type="Museum">FSCA</collectionCode>
, 
<collectionCode id="ED18AE12FFE9FF9BFD3C7DC3FCCE57C4" box="[730,800,1987,2013]" pageId="5" pageNumber="433">IEXA</collectionCode>
, 
<collectionCode id="ED18AE12FFE9FF9BFCCD7DC3FC7657C4" box="[811,920,1987,2013]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="5" pageNumber="433" type="Museum">NHMUK</collectionCode>
, 
<collectionCode id="ED18AE12FFE9FF9BFC447DC3FC1F57C4" box="[930,1009,1987,2013]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="433" type="Other institutional type">NZAC</collectionCode>
, 
<collectionCode id="ED18AE12FFE9FF9BFC1B7DC3FBBB57C4" box="[1021,1109,1987,2013]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="5" pageNumber="433">UNAM</collectionCode>
, 
<collectionCode id="ED18AE12FFE9FF9BFB867DC3FB5B57C4" box="[1120,1205,1987,2013]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="433" type="Museum">USNM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C313655CFFEAFF98FF217A96FE6A50CD" pageId="6" pageNumber="434" type="etymology">
<paragraph id="8BB636D7FFEAFF98FF217A96FE6A50CD" blockId="6.[151,1437,150,537]" pageId="6" pageNumber="434">
<emphasis id="B97DEAC5FFEAFF98FF217A96FEA050A9" bold="true" box="[199,334,150,176]" pageId="6" pageNumber="434">Etymology.</emphasis>
The species is named in honor of Malcolm “Chip” Jones for his contributions to cycad conservation and horticulture.
</paragraph>
</subSubSection>
<subSubSection id="C313655CFFEAFF98FF217ADEFD175200" pageId="6" pageNumber="434" type="discussion">
<paragraph id="8BB636D7FFEAFF98FF217ADEFD175200" blockId="6.[151,1437,150,537]" pageId="6" pageNumber="434">
<emphasis id="B97DEAC5FFEAFF98FF217ADEFED550E1" bold="true" box="[199,315,222,248]" pageId="6" pageNumber="434">Remarks.</emphasis>
This species is closely related to 
<taxonomicName id="4C094D54FFEAFF98FD207ADEFBD950E1" box="[710,1079,222,249]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="vazquezi" subGenus="Vovidesa">
<emphasis id="B97DEAC5FFEAFF98FD207ADEFCA350E1" box="[710,845,222,248]" italics="true" pageId="6" pageNumber="434">Ceratophila</emphasis>
(
<emphasis id="B97DEAC5FFEAFF98FC867ADEFC2A50E1" box="[864,964,222,248]" italics="true" pageId="6" pageNumber="434">Vovidesa</emphasis>
) 
<emphasis id="B97DEAC5FFEAFF98FC317ADFFBD950E1" box="[983,1079,223,248]" italics="true" pageId="6" pageNumber="434">vazquezi</emphasis>
</taxonomicName>
, which, based on the principal component analysis (
<figureCitation id="13322A52FFEAFF98FE647B02FE2C5105" box="[386,450,258,284]" captionStart="FIGURE 1" captionStartId="2.[151,250,1116,1140]" captionTargetBox="[239,1350,517,1076]" captionTargetId="figure-155@2.[216,1370,489,1092]" captionTargetPageId="2" captionText="FIGURE 1. Scatter plot score derived from the functions produced by stepwise discriminant analysis of 3 anatomical characters from five populations of Ceratophila: A: males; B: females. PC1 is related to the two characters of: ratio of pronotal length to pronotal width (PL/PW) and ratio of pronotal length to elytra length (PL/EL). PC2 is related to the character: ratio of head width to ventral interocular distance (HW/VI)" figureDoi="http://doi.org/10.5281/zenodo.6623169" httpUri="https://zenodo.org/record/6623169/files/figure.png" pageId="6" pageNumber="434">Fig. 1</figureCitation>
), appears widely distributed in central 
<collectingRegion id="49CDF835FFEAFF98FC947B03FC3B5104" box="[882,981,259,285]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="434">Veracruz</collectingRegion>
and 
<collectingRegion id="49CDF835FFEAFF98FBEC7B02FB865105" box="[1034,1128,258,284]" country="Mexico" name="Hidalgo" pageId="6" pageNumber="434">Hidalgo</collectingRegion>
. Both 
<taxonomicName id="4C094D54FFEAFF98FB487B03FA805105" authorityName="Tang &amp; Gomez Dominguez" authorityYear="2022" box="[1198,1390,258,285]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="chipjonesi" subGenus="Vovidesa">
<emphasis id="B97DEAC5FFEAFF98FB487B03FB265104" box="[1198,1224,259,285]" italics="true" pageId="6" pageNumber="434">C.</emphasis>
(
<emphasis id="B97DEAC5FFEAFF98FB317B03FB025105" box="[1239,1260,259,284]" italics="true" pageId="6" pageNumber="434">V.</emphasis>
) 
<emphasis id="B97DEAC5FFEAFF98FB1D7B02FA805105" box="[1275,1390,258,284]" italics="true" pageId="6" pageNumber="434">chipjonesi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C094D54FFEAFF98FF717B27FEA95159" box="[151,327,294,321]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="vazquezi" subGenus="Vovidesa">
<emphasis id="B97DEAC5FFEAFF98FF717B27FF5F5158" box="[151,177,295,321]" italics="true" pageId="6" pageNumber="434">C.</emphasis>
(
<emphasis id="B97DEAC5FFEAFF98FF277B27FF395159" box="[193,215,295,320]" italics="true" pageId="6" pageNumber="434">V.</emphasis>
) 
<emphasis id="B97DEAC5FFEAFF98FF017B27FEA95159" box="[231,327,295,320]" italics="true" pageId="6" pageNumber="434">vazquezi</emphasis>
</taxonomicName>
share the diagnostic characters of males with meso- and metatibia with a distinct medial subapical emargination and mentum with long setae projecting laterally. 
<taxonomicName id="4C094D54FFEAFF98FC817B4AFB75517D" authorityName="Tang &amp; Gomez Dominguez" authorityYear="2022" box="[871,1179,330,357]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="chipjonesi" subGenus="Vovidesa">
<emphasis id="B97DEAC5FFEAFF98FC817B4AFC00517D" box="[871,1006,330,356]" italics="true" pageId="6" pageNumber="434">Ceratophila</emphasis>
(
<emphasis id="B97DEAC5FFEAFF98FBE77B4BFBF8517D" box="[1025,1046,331,356]" italics="true" pageId="6" pageNumber="434">V.</emphasis>
) 
<emphasis id="B97DEAC5FFEAFF98FBCE7B4AFB75517D" box="[1064,1179,330,356]" italics="true" pageId="6" pageNumber="434">chipjonesi</emphasis>
</taxonomicName>
differs mainly in possessing a longer, narrower pronotum (see results of PCA above). On the 
<specimenCount id="9D0FFD5EFFEAFF98FC2F7B6FFBC15191" box="[969,1071,366,393]" pageId="6" pageNumber="434" type="male">one male</specimenCount>
cone of 
<taxonomicName id="4C094D54FFEAFF98FB747B6FFA735191" box="[1170,1437,366,393]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="santillanii">
<emphasis id="B97DEAC5FFEAFF98FB747B6FFA735191" box="[1170,1437,366,393]" italics="true" pageId="6" pageNumber="434">Ceratozamia santillanii</emphasis>
</taxonomicName>
sampled for this paper, 
<taxonomicName id="4C094D54FFEAFF98FE7B7B93FDB051B5" authorityName="Tang &amp; Gomez Dominguez" authorityYear="2022" box="[413,606,402,429]" class="Insecta" family="Erotylidae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="chipjonesi" subGenus="Vovidesa">
<emphasis id="B97DEAC5FFEAFF98FE7B7B93FE5951B4" box="[413,439,403,429]" italics="true" pageId="6" pageNumber="434">C.</emphasis>
(
<emphasis id="B97DEAC5FFEAFF98FE217B93FE3251B5" box="[455,476,403,428]" italics="true" pageId="6" pageNumber="434">V.</emphasis>
) 
<emphasis id="B97DEAC5FFEAFF98FE0D7B92FDB051B5" box="[491,606,402,428]" italics="true" pageId="6" pageNumber="434">chipjonesi</emphasis>
</taxonomicName>
accounted for 70% of adult 
<taxonomicName id="4C094D54FFEAFF98FC7A7B92FBB651B5" authorityName="Crowson" authorityYear="1952" box="[924,1112,402,428]" class="Insecta" family="Erotylidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
collected, while 
<taxonomicName id="4C094D54FFEAFF98FAF07B92FEEA51C9" authorityName="Tang &amp; Gomez Dominguez" authorityYear="2022" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="perezi">
<emphasis id="B97DEAC5FFEAFF98FAF07B92FEEA51C9" italics="true" pageId="6" pageNumber="434">Pharaxonotha perezi</emphasis>
</taxonomicName>
(see below) accounted for 25% of adult 
<taxonomicName id="4C094D54FFEAFF98FD2F7BB6FC6B51C9" authorityName="Crowson" authorityYear="1952" box="[713,901,438,464]" class="Insecta" family="Erotylidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
in the cone. A third undescribed 
<taxonomicName id="4C094D54FFEAFF98FB137BB6FA7B51C9" authorityName="Reitter" authorityYear="1875" box="[1269,1429,438,464]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAC5FFEAFF98FB137BB6FA7B51C9" box="[1269,1429,438,464]" italics="true" pageId="6" pageNumber="434">Pharaxonotha</emphasis>
</taxonomicName>
, with a distribution that appears to extend north into 
<collectingRegion id="49CDF835FFEAFF98FD327BDBFCD551EC" box="[724,827,475,501]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="434">Veracruz</collectingRegion>
, 
<collectingRegion id="49CDF835FFEAFF98FCA27BDAFC7051ED" box="[836,926,474,500]" country="Mexico" name="Hidalgo" pageId="6" pageNumber="434">Hidalgo</collectingRegion>
and 
<collectingRegion id="49CDF835FFEAFF98FC357BDAFB6F51ED" box="[979,1153,474,500]" country="Mexico" name="San Luis Potosi" pageId="6" pageNumber="434">San Luis Potosí</collectingRegion>
(Tang, unpublished data) was also collected on the cone in small numbers (5%).
</paragraph>
</subSubSection>
</treatment>
</document>