248 lines
32 KiB
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248 lines
32 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e81736" ID-Pensoft-Pub="1864-8312-80-411" ID-Pensoft-UUID="775020E4F468563291A208DB113C16F3" ID-ZooBank="48AB3E3B392849CAB26D4B22839FA7D5" ModsDocID="1864-8312-80-411" checkinTime="1660884374599" checkinUser="pensoft" docAuthor="Li, Yan-Da, Zhang, Yu-Bo, Szawaryn, Karol, Huang, Di-Ying & Cai, Chen-Yang" docDate="2022" docId="B26F4236273B5D53BF06D29EBBBFCB92" docLanguage="en" docName="ArthropodSystPhyl 80: 411-422" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-08-18" docSource="http://dx.doi.org/10.3897/asp.80.e81736" docTitle="Xenostanus jiangkuni Li, Szawaryn & Cai 2022, sp. nov." docType="treatment" docUuid="DF781A4B-9A46-460E-AD27-BDC09BCB1958" docUuidSource="ZooBank" docVersion="1" id="775020E4F468563291A208DB113C16F3" lastPageNumber="411" masterDocId="775020E4F468563291A208DB113C16F3" masterDocTitle="Earliest fossil record of Corylophidae from Burmese amber and phylogeny of Corylophidae (Coleoptera: Coccinelloidea)" masterLastPageNumber="422" masterPageNumber="411" pageNumber="411" updateTime="1660884374599" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Earliest fossil record of Corylophidae from Burmese amber and phylogeny of Corylophidae (Coleoptera: Coccinelloidea)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Yan-Da</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9439-202X</mods:nameIdentifier>
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<mods:affiliation>State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; Yan-Da Li [ydli @ pku. edu. cn]; Di-Ying Huang [dyhuang @ nigpas. ac. cn] & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol, BS 8 1 TQ, United Kingdom</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Yu-Bo</mods:namePart>
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<mods:affiliation>State Key Laboratory of Protein and Plant Gene Research, and Peking-Tsinghua Center for Life Sciences, Academy for Advanced Interdisciplinary Studies, Peking University, Beijing 100871, China; Yu-Bo Zhang [yubozhang @ pku. edu. cn]</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Szawaryn, Karol</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9329-4268</mods:nameIdentifier>
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<mods:affiliation>Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, Warsaw, Poland; Karol Szawaryn [k. szawaryn @ gmail. com]</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Huang, Di-Ying</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5637-4867</mods:nameIdentifier>
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<mods:affiliation>State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; Yan-Da Li [ydli @ pku. edu. cn]; Di-Ying Huang [dyhuang @ nigpas. ac. cn]</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cai, Chen-Yang</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9283-8323</mods:nameIdentifier>
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<mods:affiliation>State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; Yan-Da Li [ydli @ pku. edu. cn]; Di-Ying Huang [dyhuang @ nigpas. ac. cn] & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol, BS 8 1 TQ, United Kingdom</mods:affiliation>
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<mods:nameIdentifier type="email">cycai@nigpas.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-18</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>80</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>411</mods:start>
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<mods:end>422</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e81736</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e81736</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-80-411</mods:identifier>
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<mods:identifier type="ZooBank">48AB3E3B392849CAB26D4B22839FA7D5</mods:identifier>
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<mods:identifier type="Pensoft-UUID">775020E4F468563291A208DB113C16F3</mods:identifier>
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<subSection pageId="0" pageNumber="411" type="3. systematic paleontology">
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<treatment LSID="urn:lsid:zoobank.org:act:DF781A4B-9A46-460E-AD27-BDC09BCB1958" httpUri="http://treatment.plazi.org/id/B26F4236273B5D53BF06D29EBBBFCB92" lastPageNumber="411" pageId="0" pageNumber="411">
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<subSubSection pageId="0" pageNumber="411" type="nomenclature">
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<paragraph pageId="0" pageNumber="411">
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<taxonomicName LSID="http://zoobank.org/DF781A4B-9A46-460E-AD27-BDC09BCB1958" authority="Li, Szawaryn & Cai" authorityName="Li, Szawaryn & Cai" authorityYear="2022" family="Corylophidae" genus="Xenostanus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Xenostanus jiangkuni" order="Coleoptera" pageId="0" pageNumber="411" rank="species" species="jiangkuni" status="sp. nov.">Xenostanus jiangkuni Li, Szawaryn & Cai</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="411">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="411" type="description">
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<paragraph pageId="0" pageNumber="411">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under incident light. A Dorsal view. B Ventral view. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411">Figs 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. General habitus of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Dorsal view. B Ventral view, with arrowheads indicating the metaventral and abdominal postcoxal lines. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure2" httpUri="https://binary.pensoft.net/fig/730769" pageId="0" pageNumber="411">, 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782. A Dorsal view. B Ventral view. C Lateral view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure4" httpUri="https://binary.pensoft.net/fig/730771" pageId="0" pageNumber="411">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under " Sum along Ray " mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">, 5, S1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="411" type="etymology">
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<paragraph pageId="0" pageNumber="411">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="411">The species is named after Mr. Kun Jiang, who kindly donated many fossils for our research.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="411" type="type material">
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<paragraph pageId="0" pageNumber="411">Type materials.</paragraph>
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<paragraph pageId="0" pageNumber="411">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="411">Holotype</emphasis>
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: NIGP177782. Two
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<emphasis bold="true" italics="true" pageId="0" pageNumber="411">paratypes</emphasis>
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, GPIH no. 5058 (CCGG no. 11948), GPIH no. 5059 (CCGG no. 11105).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="411" type="type locality">
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<paragraph pageId="0" pageNumber="411">Type locality and horizon.</paragraph>
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<paragraph pageId="0" pageNumber="411">Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District, Kachin State, Myanmar; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="411" type="diagnosis">
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<paragraph pageId="0" pageNumber="411">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="411">As for the tribe.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="411" type="description">
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<paragraph pageId="0" pageNumber="411">Description.</paragraph>
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<paragraph pageId="0" pageNumber="411">
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Body elongate, widest at middle of elytra, very weakly convex. Surface with hair-like setae. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="411">Head</emphasis>
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partially exposed and visible from above (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under " Sum along Ray " mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5F</figureCitation>
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). Eyes coarsely faceted, without interfacetal setae (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3B,C</figureCitation>
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). Frontoclypeal suture absent (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3B</figureCitation>
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). Labrum free, transverse (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3B</figureCitation>
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). Subantennal grooves absent. Antennae (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782. A Dorsal view. B Ventral view. C Lateral view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure4" httpUri="https://binary.pensoft.net/fig/730771" pageId="0" pageNumber="411">4C,D</figureCitation>
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) with 10 antennomeres; scape distinctly longer and wider than pedicel; antennomeres 3-7 small, subquadrate to transverse; club (antennomeres 8-10) asymmetrical, as long as antennomeres 3-7 combined. Mandibles short (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3A</figureCitation>
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). Maxillary palps (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3A</figureCitation>
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) 3-segmented; apical palpomere about twice as long as penultimate one, conical. Labial palps (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3A</figureCitation>
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) 2-segmented; apical palpomere about as long as basal one. Ventral head surface seemingly with a pair of parallel subgenal ridges (Figs
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3A</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under " Sum along Ray " mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5E</figureCitation>
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). -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="411">Prothorax</emphasis>
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: Pronotal disc (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under " Sum along Ray " mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5F</figureCitation>
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) widest at base; anterior margin straight; lateral margins bordered in posterior part; posterior angles pointed; posterior margin bisinuate. Prosternum (Figs
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3A</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under " Sum along Ray " mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5E</figureCitation>
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) in front of coxae about as long as longitudinal coxal diameter, anteriorly not produced forward; prosternal carinae absent; prosternal process broad, widened beyond front coxae, meeting postcoxal hypomeral projections, truncate at apex. Procoxal cavities externally closed, ovaloid, without lateral slit (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under " Sum along Ray " mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5E</figureCitation>
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). -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="411">Meso- and metathorax</emphasis>
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: Scutellar shield strongly transverse. Elytra covering entire abdomen except for part of pygidium (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under incident light. A Dorsal view. B Ventral view. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411">1A</figureCitation>
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). Mesoventrite flat, without transverse carina (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under incident light. A Dorsal view. B Ventral view. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411">1B</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under " Sum along Ray " mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5A</figureCitation>
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). Mesocoxal cavities circular, outwardly closed (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under incident light. A Dorsal view. B Ventral view. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411">1B</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under " Sum along Ray " mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5A</figureCitation>
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). Meso-metaventral junction nearly straight (though with a small projection medially). Metaventrite with distinct postcoxal lines (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under incident light. A Dorsal view. B Ventral view. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411">1B</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. General habitus of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Dorsal view. B Ventral view, with arrowheads indicating the metaventral and abdominal postcoxal lines. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure2" httpUri="https://binary.pensoft.net/fig/730769" pageId="0" pageNumber="411">2B</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under " Sum along Ray " mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5A</figureCitation>
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); discrimen visible in posterior third of metaventrite. Metacoxae transverse, broadly separated. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="411">Legs</emphasis>
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: Femora flattened. Tibiae simple, not strongly widened, apically with small denticles; tibial spurs 2-2-2 (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3D</figureCitation>
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). Tarsi 4-4-4 (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Details of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Head and prothorax, ventral view. B Head, dorsal view. C Head and prothorax, lateral view. D Abdominal base, ventral view. Abbreviations: an, antenna; ey, compound eye; lb, labrum; lbp, labial palp; md, mandible; msf, mesofemur; msts, mesotarsus; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; pts, protarsus; v 1 - 3, ventrites 1 - 3. Scale bars: 200 μm." figureDoi="10.3897/asp.80.e81736.figure3" httpUri="https://binary.pensoft.net/fig/730770" pageId="0" pageNumber="411">3D</figureCitation>
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); tarsomeres 1 and 2 ventrally lobed; tarsomere 3 smaller and simple; tarsomere 4 elongate, as long as 1-3 combined. Pretarsal claws simple, with small basal angulation. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="411">Abdomen</emphasis>
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with six freely articulated ventrites. Ventrite 1 longer than 2-4 combined, with strongly arcuate postcoxal lines (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General habitus of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under incident light. A Dorsal view. B Ventral view. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411">1B</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. General habitus of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782, under confocal microscopy. A Dorsal view. B Ventral view, with arrowheads indicating the metaventral and abdominal postcoxal lines. Scale bars: 400 μm." figureDoi="10.3897/asp.80.e81736.figure2" httpUri="https://binary.pensoft.net/fig/730769" pageId="0" pageNumber="411">2B</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. X-ray microtomographic reconstruction of Xenostanus jiangkuni Li, Szawaryn & Cai gen. et sp. nov., holotype, NIGP 177782. A Ventral view, with legs removed. B Ventral view, rendered under " Sum along Ray " mode. C, D Antenna. E Head and prothorax, ventral view. F Head and prothorax, dorsal view. Scale bar: 400 μm." figureDoi="10.3897/asp.80.e81736.figure5" httpUri="https://binary.pensoft.net/fig/730772" pageId="0" pageNumber="411">5A</figureCitation>
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), anteriorly complete; intercoxal process very broad and truncate.
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</paragraph>
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<caption doi="10.3897/asp.80.e81736.figure1" httpUri="https://binary.pensoft.net/fig/730768" pageId="0" pageNumber="411" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="411">
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<emphasis bold="true" pageId="0" pageNumber="411">Figure 1.</emphasis>
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General habitus of
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<taxonomicName authorityName="Li, Szawaryn & Cai" authorityYear="2022" family="Corylophidae" genus="Xenostanus" kingdom="Animalia" lsidName="Xenostanus jiangkuni" order="Coleoptera" pageId="0" pageNumber="411" rank="species" species="jiangkuni">
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<emphasis italics="true" pageId="0" pageNumber="411">Xenostanus jiangkuni</emphasis>
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</taxonomicName>
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Li, Szawaryn & Cai
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|
<emphasis bold="true" pageId="0" pageNumber="411">gen. et sp. nov.</emphasis>
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, holotype, NIGP177782, under incident light.
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<emphasis bold="true" pageId="0" pageNumber="411">A</emphasis>
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Dorsal view.
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<emphasis bold="true" pageId="0" pageNumber="411">B</emphasis>
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Ventral view. Scale bars: 400
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<normalizedToken originalValue="μm">μm</normalizedToken>
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.
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</paragraph>
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</caption>
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<caption doi="10.3897/asp.80.e81736.figure2" httpUri="https://binary.pensoft.net/fig/730769" pageId="0" pageNumber="411" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="411">
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<emphasis bold="true" pageId="0" pageNumber="411">Figure 2.</emphasis>
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General habitus of
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<taxonomicName authorityName="Li, Szawaryn & Cai" authorityYear="2022" family="Corylophidae" genus="Xenostanus" kingdom="Animalia" lsidName="Xenostanus jiangkuni" order="Coleoptera" pageId="0" pageNumber="411" rank="species" species="jiangkuni">
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<emphasis italics="true" pageId="0" pageNumber="411">Xenostanus jiangkuni</emphasis>
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</taxonomicName>
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Li, Szawaryn & Cai
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|
<emphasis bold="true" pageId="0" pageNumber="411">gen. et sp. nov.</emphasis>
|
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, holotype, NIGP177782, under confocal microscopy.
|
|
<emphasis bold="true" pageId="0" pageNumber="411">A</emphasis>
|
|
Dorsal view.
|
|
<emphasis bold="true" pageId="0" pageNumber="411">B</emphasis>
|
|
Ventral view, with arrowheads indicating the metaventral and abdominal postcoxal lines. Scale bars: 400
|
|
<normalizedToken originalValue="μm">μm</normalizedToken>
|
|
.
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|
</paragraph>
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|
</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="411" type="description">
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<paragraph pageId="0" pageNumber="411">Measurements.</paragraph>
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<paragraph pageId="0" pageNumber="411">
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NIGP177782:
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<emphasis bold="true" pageId="0" pageNumber="411">BL</emphasis>
|
|
1.42 mm,
|
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<emphasis bold="true" pageId="0" pageNumber="411">BW</emphasis>
|
|
0.63 mm,
|
|
<emphasis bold="true" pageId="0" pageNumber="411">HL</emphasis>
|
|
0.27 mm,
|
|
<emphasis bold="true" pageId="0" pageNumber="411">HW</emphasis>
|
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0.31 mm,
|
|
<emphasis bold="true" pageId="0" pageNumber="411">PL</emphasis>
|
|
0.34 mm,
|
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<emphasis bold="true" pageId="0" pageNumber="411">PW</emphasis>
|
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0.51 mm,
|
|
<emphasis bold="true" pageId="0" pageNumber="411">EL</emphasis>
|
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0.96 mm. GPIH no. 5058:
|
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<emphasis bold="true" pageId="0" pageNumber="411">BL</emphasis>
|
|
1.88 mm,
|
|
<emphasis bold="true" pageId="0" pageNumber="411">BW</emphasis>
|
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0.74 mm. GPIH no. 5059:
|
|
<emphasis bold="true" pageId="0" pageNumber="411">BL</emphasis>
|
|
1.30 mm,
|
|
<emphasis bold="true" pageId="0" pageNumber="411">BW</emphasis>
|
|
0.62 mm
|
|
</paragraph>
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</subSubSection>
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</treatment>
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</subSection>
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</document> |