528 lines
61 KiB
XML
528 lines
61 KiB
XML
<document id="803B7F90C3DFD5AAE75211FC424C18C8" ID-DOI="10.5281/zenodo.194047" ID-GBIF-Dataset="18e7ebdc-9680-4301-9674-c14b5e87ecaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="194047" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460183944776" checkinUser="plazi" docAuthor="Samyn, Yves & Thandar, Ahmed S." docDate="2010" docId="03A2B434FFAD4B1AFF5F217DFBBAB08D" docLanguage="en" docName="zt02399p019.pdf" docOrigin="Zootaxa 2399" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Massinium magnum Ludwig 1882" docType="treatment" docVersion="9" lastPageNumber="8" masterDocId="FF9BCC4CFFAF4B1DFFC8237FFF9BB541" masterDocTitle="Two new species in the phyllophorid genus Massinium (Echinodermata: Holothuroidea) with redescription of Massinium magnum" masterLastPageNumber="19" masterPageNumber="1" pageNumber="3" updateTime="1698236274408" updateUser="plazi">
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<mods:title id="4878A57D1A8ABDBEB161687C6572D8DB">Two new species in the phyllophorid genus Massinium (Echinodermata: Holothuroidea) with redescription of Massinium magnum</mods:title>
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<mods:namePart id="B8EDBCCA9FE84F21FBEDFD234F6FBE25">Thandar, Ahmed S.</mods:namePart>
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<treatment id="03A2B434FFAD4B1AFF5F217DFBBAB08D" ID-DOI="http://doi.org/10.5281/zenodo.6196778" ID-GBIF-Taxon="119383632" ID-Zenodo-Dep="6196778" LSID="urn:lsid:plazi:treatment:03A2B434FFAD4B1AFF5F217DFBBAB08D" httpUri="http://treatment.plazi.org/id/03A2B434FFAD4B1AFF5F217DFBBAB08D" lastPageId="7" lastPageNumber="8" pageId="2" pageNumber="3">
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<subSubSection id="C31156A9FFAD4B1FFF5F217DFDB0B703" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8BB40522FFAD4B1FFF5F217DFDCEB75D" blockId="2.[151,597,514,578]" box="[151,597,514,540]" pageId="2" pageNumber="3">
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<heading id="D0FCB24EFFAD4B1FFF5F217DFDCEB75D" bold="true" box="[151,597,514,540]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<taxonomicName id="4C0B7EA1FFAD4B1FFF5F217DFDCEB75D" ID-CoL="3YCDB" authority="Ludwig, 1882" authorityName="Ludwig" authorityYear="1882" box="[151,597,514,540]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="magnum">
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<emphasis id="B97FD930FFAD4B1FFF5F217DFDCEB75D" bold="true" box="[151,597,514,540]" pageId="2" pageNumber="3">
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<emphasis id="B97FD930FFAD4B1FFF5F217DFE0AB75D" bold="true" box="[151,401,514,540]" italics="true" pageId="2" pageNumber="3">Massinium magnum</emphasis>
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(
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<bibRefCitation id="EF9A78D3FFAD4B1FFE68217DFDD6B75D" author="Ludwig" box="[416,589,514,540]" pageId="2" pageNumber="18" refString="Ludwig, H. (1882) List of the holothurians in the collection of the Leyden Museum. Notes Leyden Museum, 4 (10), 127 - 137." type="journal article" year="1882">Ludwig, 1882</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BB40522FFAD4B1FFF5F2157FDB0B703" blockId="2.[151,597,514,578]" box="[151,555,552,578]" pageId="2" pageNumber="3">(figs 1A–E, 2A–H, 3A–N, 4A&B)</paragraph>
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</subSubSection>
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||
<subSubSection id="C31156A9FFAD4B1FFF5F210DFC1CB688" pageId="2" pageNumber="3" type="reference_group">
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<paragraph id="8BB40522FFAD4B1FFF5F210DFC72B7C8" blockId="2.[151,1437,626,969]" box="[151,1001,626,649]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="AB1B220CFFAD4B1FFF5F210DFC72B7C8" box="[151,1001,626,649]" pageId="2" pageNumber="3">
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<taxonomicName id="4C0B7EA1FFAD4B1FFF5F210DFDD5B7C8" authority="Ludwig, 1882: 132" authorityName="Ludwig" authorityPageNumber="132" authorityYear="1882" box="[151,590,626,649]" class="Holothuroidea" family="Cucumariidae" genus="Thyonidium" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="magnum">
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<emphasis id="B97FD930FFAD4B1FFF5F210DFEE2B7C8" box="[151,377,626,649]" italics="true" pageId="2" pageNumber="3">Thyonidium magnum</emphasis>
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<treatmentCitation id="0AAA2333FFAD4B1FFEB7210DFDD5B7C8" author="Ludwig" box="[383,590,626,649]" page="132" pageId="2" pageNumber="3" year="1882">
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<bibRefCitation id="EF9A78D3FFAD4B1FFEB7210DFD83B7C8" author="Ludwig" box="[383,536,626,649]" pageId="2" pageNumber="18" refString="Ludwig, H. (1882) List of the holothurians in the collection of the Leyden Museum. Notes Leyden Museum, 4 (10), 127 - 137." type="journal article" year="1882">Ludwig, 1882</bibRefCitation>
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: 132
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="0AAA2333FFAD4B1FFD91210DFCB3B7C8" author="Lampert" box="[601,808,626,649]" page="170" pageId="2" pageNumber="3" year="1885">
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<bibRefCitation id="EF9A78D3FFAD4B1FFD91210DFD68B7C8" author="Lampert" box="[601,755,626,649]" pageId="2" pageNumber="18" refString="Lampert, K. (1885) Die Seewalzen. Eine systematische Monographie. In: Semper, C. (Ed.), Reisen im Archipel der Philippinen, Teil 2, Wissenschaftliche Resultate. Wiesbaden, 1 - 310, 1 pl." type="journal article" year="1885">Lampert 1885</bibRefCitation>
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: 170
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</treatmentCitation>
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;
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<treatmentCitation id="0AAA2333FFAD4B1FFCFB210DFC7EB7C8" author="Theel" box="[819,997,626,649]" page="146" pageId="2" pageNumber="3" year="1886">
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<bibRefCitation id="EF9A78D3FFAD4B1FFCFB210DFC2BB7C8" author="Theel" box="[819,944,626,649]" pageId="2" pageNumber="19" refString="Theel, Hj. (1886) Holothuroidea. Part 2. Results of the Voyage of the Challenger (Zoology) 39, 1 - 290, pls 1 - 16." type="journal article" year="1886">Théel 1886</bibRefCitation>
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: 146
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BB40522FFAD4B1FFF5F21EDFDA6B788" blockId="2.[151,1437,626,969]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="AB1B220CFFAD4B1FFF5F21EDFDA6B788" pageId="2" pageNumber="3">
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<taxonomicName id="4C0B7EA1FFAD4B1FFF5F21EDFE1BB7E8" box="[151,384,658,681]" class="Holothuroidea" family="Phyllophoridae" genus="Phyllophorus" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="magnus">
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<emphasis id="B97FD930FFAD4B1FFF5F21EDFE1BB7E8" box="[151,384,658,681]" italics="true" pageId="2" pageNumber="3">Phyllophorus magnus</emphasis>
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</taxonomicName>
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||
;
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<bibRefCitation id="EF9A78D3FFAD4B1FFE4621EDFD84B7E8" author="Ludwig" box="[398,543,658,681]" lastPageId="2" lastPageNumber="19" pageId="2" pageNumber="18" refString="Ludwig, H. (1889 - 1892) Echinodermen: Die Zeewalzen. In: Bronn, H. G. (Ed.), Bronn's Klassen un Ordnungen des Their-Reichs, wissenschaftlich dargestellt in Wort und Bild. CF. Winter'sche Verlagshandlung, Leipzig, 460 pp., 17 pls." type="book" year="1889">
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Ludwig
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<date id="FFB523E2FFAD4B1FFE2021EDFD84B7E8" box="[488,543,658,681]" pageId="2" pageNumber="3" value="1889" valueMax="1892">1889</date>
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</bibRefCitation>
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–92: 347;
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<treatmentCitation id="0AAA2333FFAD4B1FFD4221EDFCDEB7E8" author="Sluiter" box="[650,837,658,681]" page="112" pageId="2" pageNumber="3" year="1901">
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<bibRefCitation id="EF9A78D3FFAD4B1FFD4221EDFC8BB7E8" author="Sluiter" box="[650,784,658,681]" pageId="2" pageNumber="19" refString="Sluiter, C. P. (1901) Die Holothurien der Siboga Expedition. Siboga Expedition, 44, 1 - 142, 10 pls." type="journal article" year="1901">Sluiter 1901</bibRefCitation>
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: 112
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||
</treatmentCitation>
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; pl. IV, fig. 5, pl. VII, fig. 1a–e;
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<treatmentCitation id="0AAA2333FFAD4B1FFB5721EDFAEEB7E8" author="Domantay" box="[1183,1397,658,681]" page="38" pageId="2" pageNumber="3" year="1933">
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<bibRefCitation id="EF9A78D3FFAD4B1FFB5721EDFAD5B7E8" author="Domantay" box="[1183,1358,658,681]" pageId="2" pageNumber="18" refString="Domantay, J. S. (1933) Littoral Holothurioidea of Port Galera Bay and Adjacent Waters. Natural Applied Science Bulletin, University Philippines. 3, 41 - 101, pls 1 - 4." type="journal article" year="1933">Domantay 1933</bibRefCitation>
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: 38
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</treatmentCitation>
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||
, pl. 2, fig. 11a–j;
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<treatmentCitation id="0AAA2333FFAD4B1FFE9D21CDFDA2B788" author="Domantay" box="[341,569,690,713]" page="399" pageId="2" pageNumber="3" year="1936">
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||
<bibRefCitation id="EF9A78D3FFAD4B1FFE9D21CDFD98B788" author="Domantay" box="[341,515,690,713]" pageId="2" pageNumber="18" refString="Domantay, J. S. (1936) The ecological distribution of the echinoderm fauana of the Puerto Galera Marine Biological Station. Natural Applied Science Bulletin, University Philippines. 5: 385 - 403, pls 1 - 7." type="journal article" year="1936">Domantay 1936</bibRefCitation>
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||
: 399
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||
</treatmentCitation>
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||
.
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</treatmentCitationGroup>
|
||
</paragraph>
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||
<paragraph id="8BB40522FFAD4B1FFF5F21ADFD4EB608" blockId="2.[151,1437,626,969]" pageId="2" pageNumber="3">
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||
<treatmentCitationGroup id="AB1B220CFFAD4B1FFF5F21ADFD4EB608" pageId="2" pageNumber="3">
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||
<taxonomicName id="4C0B7EA1FFAD4B1FFF5F21ADFE04B7A8" ID-CoL="46V8J" box="[151,415,722,745]" class="Holothuroidea" family="Phyllophoridae" genus="Neothyonidium" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="magnum">
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<emphasis id="B97FD930FFAD4B1FFF5F21ADFE04B7A8" box="[151,415,722,745]" italics="true" pageId="2" pageNumber="3">Neothyonidium magnum</emphasis>
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||
</taxonomicName>
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||
;
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||
<treatmentCitation id="0AAA2333FFAD4B1FFE6721ADFD6EB7A8" author="Heding" box="[431,757,722,745]" page="197" pageId="2" pageNumber="3" year="1954">
|
||
<bibRefCitation id="EF9A78D3FFAD4B1FFE6721ADFD24B7A8" author="Heding" box="[431,703,722,745]" pageId="2" pageNumber="18" refString="Heding, S. G. & Panning, A. (1954) Phyllophoridae. Eine Bearbeitung der Polytentaculaten Dendrochiroten Holothurien des Zoologische Museums in Kopenhagen. Spolia Zoologica Musei Hauniensis, 13, 1 - 209, figs 1 - 102." type="journal article" year="1954">Heding & Panning 1954</bibRefCitation>
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||
: 197
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||
</treatmentCitation>
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||
, fig. 98;
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<treatmentCitation id="0AAA2333FFAD4B1FFC9421ADFBEFB7A8" author="Clark" box="[860,1140,722,745]" page="182" pageId="2" pageNumber="3" year="1971">
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||
<bibRefCitation id="EF9A78D3FFAD4B1FFC9421ADFBA4B7A8" author="Clark" box="[860,1087,722,745]" pageId="2" pageNumber="18" refString="Clark, A. M. & Rowe, F. W. E. (1971) Monograph of shallow-water Indo-West Pacific echinoderms. Trusties of the British Museum of Natural History London, 238 pp, 31 pls" type="book" year="1971">Clark & Rowe 1971</bibRefCitation>
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||
: 182
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||
</treatmentCitation>
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(distribution);?
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||
<treatmentCitation id="0AAA2333FFAD4B1FFAF821ADFE16B648" author="George" page="246" pageId="2" pageNumber="3" year="1987">
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<bibRefCitation id="EF9A78D3FFAD4B1FFAF821ADFECDB648" author="George" pageId="2" pageNumber="18" refString="George, J. D. & George, J. (1987) The corals of the Bodgaya Islands (Sabah: Malaysia) and Pulau Sipadan. Malayan Nature Journal 40, 225 - 260." type="journal article" year="1987">George & George 1987</bibRefCitation>
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||
: 246
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</treatmentCitation>
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||
;?
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||
<treatmentCitation id="0AAA2333FFAD4B1FFE65218DFD5AB648" author="Rowe" box="[429,705,754,777]" page="312" pageId="2" pageNumber="3" year="1995">
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||
<bibRefCitation id="EF9A78D3FFAD4B1FFE65218DFD16B648" author="Rowe" box="[429,653,754,777]" pageId="2" pageNumber="19" refString="Rowe, F. W. E. & Gates, J. (1995) Echinodermata. In: Wells, A. (Ed.), Zoological Catalogue of Australia. CSIRO Australia, Melbourne, vol. 33, i-xiii, 1 - 510," type="journal article" year="1995">Rowe & Gates 1995</bibRefCitation>
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||
: 312
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</treatmentCitation>
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(synonymy);
|
||
<treatmentCitation id="0AAA2333FFAD4B1FFC91218DFB10B648" author="Colin" box="[857,1163,754,777]" page="262" pageId="2" pageNumber="3" year="1995">
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||
<bibRefCitation id="EF9A78D3FFAD4B1FFC91218DFBCFB648" author="Colin" box="[857,1108,754,777]" pageId="2" pageNumber="18" refString="Colin, P. L. & Arnison, Ch. (1995) Tropical Pacific Invertebrates. A Field Guide to the Marine Invertebrates Occurring on Tropical Pacific Coral Reefs, Sea Grass Beds and Mangroves. Coral Reef Press, Beverly Hills, i-viii, 1 - 296." type="book chapter" year="1995">Colin & Arneson 1995</bibRefCitation>
|
||
: 262
|
||
</treatmentCitation>
|
||
, fig 1247;
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||
<treatmentCitation id="0AAA2333FFAD4B1FFAC9218DFEAFB668" author="Gosliner" page="283" pageId="2" pageNumber="3" year="1996">
|
||
<bibRefCitation id="EF9A78D3FFAD4B1FFAC9218DFE9BB668" author="Gosliner" pageId="2" pageNumber="18" refString="Gosliner, T. M., Behrens, D. W. & Williams, G. C. (1996) Coral Reef Animals of the Indo-Pacific: animal life from Africa to Hawaii exclusive of the vertebrates. Sea Challengers, Montery, 314 pp." type="book" year="1996">
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||
Gosliner
|
||
<emphasis id="B97FD930FFAD4B1FFAAD218DFA07B648" box="[1381,1436,754,777]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
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||
1996
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||
</bibRefCitation>
|
||
: 283
|
||
</treatmentCitation>
|
||
, fig. 1045;
|
||
<treatmentCitation id="0AAA2333FFAD4B1FFE65206DFCBEB668" author="Erhardt" box="[429,805,786,809]" page="1151" pageId="2" pageNumber="3" year="1997">
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||
<bibRefCitation id="EF9A78D3FFAD4B1FFE65206DFD7FB668" author="Erhardt" box="[429,740,786,809]" pageId="2" pageNumber="18" refString="Erhardt H. A., & Moosleitner, H. (1997) Meerwasser Atlas Bd 3 Wirbellose Tiere. Mergus Verlag, Melle, 738 - 1328." type="book chapter" year="1997">Erhardt & Moosleitner, 1997</bibRefCitation>
|
||
: 1151
|
||
</treatmentCitation>
|
||
, 1152 (colour pictures); Forbes (ed.) et al.1999: 52;
|
||
<treatmentCitation id="0AAA2333FFAD4B1FFA87206DFEBCB608" author="Massin" page="88" pageId="2" pageNumber="3" year="1999">
|
||
<bibRefCitation id="EF9A78D3FFAD4B1FFA87206DFE9BB608" author="Massin" pageId="2" pageNumber="19" refString="Massin, C. (1999) Reef-dwelling Holothuroidea (Echinodermata) of the Spermonde Archipelago (South-West Sulawesi, Indonesia). Zoologische Verhandelingen, 329, 1 - 144." type="journal article" year="1999">Massin 1999</bibRefCitation>
|
||
: 88
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||
</treatmentCitation>
|
||
, figs 72a–k, 73;?
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<treatmentCitation id="0AAA2333FFAD4B1FFE23204DFD4BB608" author="Lane" box="[491,720,818,841]" page="491" pageId="2" pageNumber="3" year="2000">
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<bibRefCitation id="EF9A78D3FFAD4B1FFE23204DFD00B608" author="Lane" box="[491,667,818,841]" pageId="2" pageNumber="18" refString="Lane, D. J., Marsh, L. M., VandenSpiegel D., Rowe, F. W. E. (2000) Echinoderm fauna of the South China Sea: an inventory and analysis of distribution patterns. The Raffles Bulletin of Zoology, Supplement 8, 459 - 493." type="book chapter" year="2000">
|
||
Lane
|
||
<emphasis id="B97FD930FFAD4B1FFDEE204DFDC7B608" box="[550,604,818,841]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
|
||
2000
|
||
</bibRefCitation>
|
||
: 491
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB40522FFAD4B1FFF5F202DFDA1B6C8" blockId="2.[151,1437,626,969]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="AB1B220CFFAD4B1FFF5F202DFDA1B6C8" pageId="2" pageNumber="3">
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<taxonomicName id="4C0B7EA1FFAD4B1FFF5F202DFE39B628" ID-CoL="46V8J" box="[151,418,850,873]" class="Holothuroidea" family="Phyllophoridae" genus="Neothyonidium" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="magnum">
|
||
<emphasis id="B97FD930FFAD4B1FFF5F202DFE39B628" box="[151,418,850,873]" italics="true" pageId="2" pageNumber="3">Neothyonidium magnum</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<treatmentCitation id="0AAA2333FFAD4B1FFE7E202DFD27B628" author="Cherbonnier" box="[438,700,850,873]" page="656" pageId="2" pageNumber="3" year="1980">
|
||
<bibRefCitation id="EF9A78D3FFAD4B1FFE7E202DFD1AB628" author="Cherbonnier" box="[438,641,850,873]" pageId="2" pageNumber="18" refString="Cherbonnier, G. (1980) Holothuries de Nouvelle-Caledonie. Bulletin du Museum National d'Histoire Naturelle de Paris 4 eme serie 2, section A (3), 615 - 667." type="journal article" year="1980">Cherbonnier 1980</bibRefCitation>
|
||
: 656
|
||
</treatmentCitation>
|
||
, fig. 20A–Q;
|
||
<treatmentCitation id="0AAA2333FFAD4B1FFC97202DFB53B628" author="Feral" box="[863,1224,850,873]" page="102" pageId="2" pageNumber="3" year="1986">
|
||
<bibRefCitation id="EF9A78D3FFAD4B1FFC97202DFB15B628" author="Feral" box="[863,1166,850,873]" pageId="2" pageNumber="18" refString="Feral, J. - P. & Cherbonnier, G. (1986) Les holothurides. In: Guille, Laboute & Menou (eds), Guide des etoiles de mer, oursins et autres echinodermes du lagon de Nouvelle-Caledonie, ORSTOM, Paris, 55 - 107." type="book chapter" year="1986">Féral & Cherbonnier 1986</bibRefCitation>
|
||
: 102
|
||
</treatmentCitation>
|
||
, fig. 40T (non
|
||
<taxonomicName id="4C0B7EA1FFAD4B1FFAB7202DFEB8B6C8" ID-CoL="46V8J" class="Holothuroidea" family="Phyllophoridae" genus="Neothyonidium" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="magnum">
|
||
<emphasis id="B97FD930FFAD4B1FFAB7202DFEB8B6C8" italics="true" pageId="2" pageNumber="3">M. magnum</emphasis>
|
||
</taxonomicName>
|
||
) (=
|
||
<taxonomicName id="4C0B7EA1FFAD4B1FFE99200DFE55B6C8" ID-CoL="3YCD4" box="[337,462,882,905]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="albicans" status="sp. nov.">
|
||
<emphasis id="B97FD930FFAD4B1FFE99200DFE55B6C8" box="[337,462,882,905]" italics="true" pageId="2" pageNumber="3">M. albicans</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24C644BFFAD4B1FFE1D200DFDB0B6CB" box="[469,555,882,906]" pageId="2" pageNumber="3" rank="species">
|
||
<emphasis id="B97FD930FFAD4B1FFE1D200DFDB0B6CB" bold="true" box="[469,555,882,906]" pageId="2" pageNumber="3">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB40522FFAD4B1FFF5F20EDFBB1B6E8" blockId="2.[151,1437,626,969]" box="[151,1066,914,937]" pageId="2" pageNumber="3">
|
||
<treatmentCitationGroup id="AB1B220CFFAD4B1FFF5F20EDFBB1B6E8" box="[151,1066,914,937]" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C0B7EA1FFAD4B1FFF5F20EDFDB6B6E8" ID-CoL="56RN9" authority="H.L. Clark, 1938: 471" authorityName="H.L. Clark" authorityPageNumber="471" authorityYear="1938" box="[151,557,914,937]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="perissa">
|
||
<emphasis id="B97FD930FFAD4B1FFF5F20EDFEA2B6E8" box="[151,313,914,937]" italics="true" pageId="2" pageNumber="3">Thyone perissa</emphasis>
|
||
<treatmentCitation id="0AAA2333FFAD4B1FFE8820EDFDB6B6E8" author="Clark" box="[320,557,914,937]" page="471" pageId="2" pageNumber="3" year="1938">
|
||
<bibRefCitation id="EF9A78D3FFAD4B1FFE8820EDFE62B6E8" author="Clark" box="[320,505,914,937]" pageId="2" pageNumber="18" refString="Clark, H. L. (1938) Echinoderms from Australia. Memoirs of the Museum of Comparative Zoology, Harvard 55 (8), i-viii, 1 - 596, pls 1 - 28." type="journal article" year="1938">H.L. Clark, 1938</bibRefCitation>
|
||
: 471
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, fig. 43;
|
||
<treatmentCitation id="0AAA2333FFAD4B1FFD4420EDFC00B6E8" author="Rowe" box="[652,923,914,937]" page="312" pageId="2" pageNumber="3" year="1995">
|
||
<bibRefCitation id="EF9A78D3FFAD4B1FFD4420EDFCF2B6E8" author="Rowe" box="[652,873,914,937]" pageId="2" pageNumber="19" refString="Rowe, F. W. E. & Gates, J. (1995) Echinodermata. In: Wells, A. (Ed.), Zoological Catalogue of Australia. CSIRO Australia, Melbourne, vol. 33, i-xiii, 1 - 510," type="journal article" year="1995">Rowe & Gates 1995</bibRefCitation>
|
||
: 312
|
||
</treatmentCitation>
|
||
(synonymy).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB40522FFAD4B1FFF5F20CDFC1CB688" blockId="2.[151,1437,626,969]" box="[151,903,946,969]" pageId="2" pageNumber="3">
|
||
<treatmentCitationGroup id="AB1B220CFFAD4B1FFF5F20CDFC1CB688" box="[151,903,946,969]" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C0B7EA1FFAD4B1FFF5F20CDFEF5B688" ID-CoL="3YCDB" box="[151,366,946,969]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="magnum">
|
||
<emphasis id="B97FD930FFAD4B1FFF5F20CDFEF5B688" box="[151,366,946,969]" italics="true" pageId="2" pageNumber="3">Massinium magnum</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<treatmentCitation id="0AAA2333FFAD4B1FFEB420CDFD2EB688" author="Samyn" box="[380,693,946,969]" page="135" pageId="2" pageNumber="3" year="2003">
|
||
<bibRefCitation id="EF9A78D3FFAD4B1FFEB420CDFD1AB688" author="Samyn" box="[380,641,946,969]" pageId="2" pageNumber="19" refString="Samyn, Y. & Thandar, A. S. (2003) Massinium, a new genus in the family Phyllophoridae (Echinodermata: Holothuroidea: Dendrochirotida) with description of a new south-west Indian Ocean species Massinium maculosum. Belgian Journal of Zoology 133, 132 - 142." type="journal article" year="2003">Samyn & Thandar 2003</bibRefCitation>
|
||
: 135
|
||
</treatmentCitation>
|
||
, 141, 142 (
|
||
<emphasis id="B97FD930FFAD4B1FFCE520CDFCECB688" box="[813,887,946,969]" italics="true" pageId="2" pageNumber="3">passim</emphasis>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31156A9FFAD4B1FFF5F2088FEE7B11D" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="8BB40522FFAD4B1FFF5F2088FC92B153" blockId="2.[151,1437,1015,2012]" box="[151,777,1015,1042]" pageId="2" pageNumber="3">
|
||
<emphasis id="B97FD930FFAD4B1FFF5F2088FECDB150" bold="true" box="[151,342,1015,1041]" pageId="2" pageNumber="3">Original name:</emphasis>
|
||
<taxonomicName id="4C0B7EA1FFAD4B1FFE952087FC9EB153" authority="Ludwig, 1882" authorityName="Ludwig" authorityYear="1882" box="[349,773,1016,1042]" class="Holothuroidea" family="Cucumariidae" genus="Thyonidium" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="magnum">
|
||
<emphasis id="B97FD930FFAD4B1FFE952087FDCDB150" box="[349,598,1016,1041]" italics="true" pageId="2" pageNumber="3">Thyonidium magnum</emphasis>
|
||
<bibRefCitation id="EF9A78D3FFAD4B1FFD952087FC9EB153" author="Ludwig" box="[605,773,1016,1042]" pageId="2" pageNumber="18" refString="Ludwig, H. (1882) List of the holothurians in the collection of the Leyden Museum. Notes Leyden Museum, 4 (10), 127 - 137." type="journal article" year="1882">Ludwig, 1882</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB40522FFAD4B1FFF0C2762FEE7B11D" blockId="2.[151,1437,1015,2012]" pageId="2" pageNumber="3">
|
||
<emphasis id="B97FD930FFAD4B1FFF0C2762FE5BB176" bold="true" box="[196,448,1053,1079]" pageId="2" pageNumber="3">
|
||
Name-bearing
|
||
<typeStatus id="54B0BB80FFAD4B1FFE482762FE21B176" box="[384,442,1053,1079]" pageId="2" pageNumber="3">type</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<typeStatus id="54B0BB80FFAD4B1FFE002762FDA0B176" box="[456,571,1053,1079]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
|
||
: NMNH-Leiden 1989 (calcareous ring with attached structures, introvert and tentacles only).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31156A9FFAD4B1EFF0C2718FCFAB2A7" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="materials_examined">
|
||
<paragraph id="8BB40522FFAD4B1FFF0C2718FD4EB1C3" blockId="2.[151,1437,1015,2012]" box="[196,725,1127,1154]" pageId="2" pageNumber="3">
|
||
<emphasis id="B97FD930FFAD4B1FFF0C2718FEF6B1C0" bold="true" box="[196,365,1127,1153]" pageId="2" pageNumber="3">
|
||
<typeStatus id="54B0BB80FFAD4B1FFF0C2718FE9AB1C0" box="[196,257,1127,1153]" pageId="2" pageNumber="3">Type</typeStatus>
|
||
locality:
|
||
</emphasis>
|
||
<collectingCountry id="F31C45B2FFAD4B1FFEBC2717FE54B1C3" box="[372,463,1128,1154]" name="Indonesia" pageId="2" pageNumber="3">Ambon</collectingCountry>
|
||
,
|
||
<collectingCountry id="F31C45B2FFAD4B1FFE132717FDCBB1C3" box="[475,592,1128,1154]" name="Indonesia" pageId="2" pageNumber="3">Moluccas</collectingCountry>
|
||
,
|
||
<collectingCountry id="F31C45B2FFAD4B1FFD942717FD49B1C3" box="[604,722,1128,1154]" name="Indonesia" pageId="2" pageNumber="3">Indonesia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB40522FFAD4B1FFF0C27F2FCA4B1E6" blockId="2.[151,1437,1015,2012]" box="[196,831,1165,1191]" pageId="2" pageNumber="3">
|
||
<emphasis id="B97FD930FFAD4B1FFF0C27F2FE1FB1E6" bold="true" box="[196,388,1165,1191]" pageId="2" pageNumber="3">Current status:</emphasis>
|
||
<taxonomicName id="4C0B7EA1FFAD4B1FFE4427F2FCA0B1E6" authority="Ludwig, 1882" authorityName="Ludwig" authorityYear="1882" box="[396,827,1165,1191]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="magnum">
|
||
<emphasis id="B97FD930FFAD4B1FFE4427F2FDE3B1E7" box="[396,632,1165,1190]" italics="true" pageId="2" pageNumber="3">Massinium magnum</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9A78D3FFAD4B1FFD4027F2FCA9B1E6" author="Ludwig" box="[648,818,1165,1191]" pageId="2" pageNumber="18" refString="Ludwig, H. (1882) List of the holothurians in the collection of the Leyden Museum. Notes Leyden Museum, 4 (10), 127 - 137." type="journal article" year="1882">Ludwig, 1882</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB40522FFAD4B1FFF0C27CDFC60B3CD" blockId="2.[151,1437,1015,2012]" pageId="2" pageNumber="3">
|
||
<emphasis id="B97FD930FFAD4B1FFF0C27CDFE54B18D" bold="true" box="[196,463,1202,1228]" pageId="2" pageNumber="3">Material examined:</emphasis>
|
||
NMNH-Leiden 1989 (
|
||
<typeStatus id="54B0BB80FFAD4B1FFD3B27CDFCF1B18D" box="[755,874,1202,1228]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
),
|
||
<collectingCountry id="F31C45B2FFAD4B1FFC4927CDFC45B18D" box="[897,990,1202,1228]" name="Indonesia" pageId="2" pageNumber="3">Ambon</collectingCountry>
|
||
(Molluccas,
|
||
<collectingCountry id="F31C45B2FFAD4B1FFB4F27CDFA93B18D" box="[1159,1288,1202,1228]" name="Indonesia" pageId="2" pageNumber="3">Indonesia</collectingCountry>
|
||
), depth and collection data unknown, coll. E.W.A. Ludeking;
|
||
<collectionCode id="ED1A9DE7FFAD4B1FFD2027A7FCA4B1B3" box="[744,831,1240,1266]" httpUri="http://grbio.org/cool/c09s-jk3y" name="Zoological Museum, University of Athens" pageId="2" pageNumber="3">ZMUA</collectionCode>
|
||
V.
|
||
<collectionCode id="ED1A9DE7FFAD4B1FFCA927A7FC3AB1B3" LSID="urn:lsid:biocol.org:col:12987" box="[865,929,1240,1266]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12987" name="Elmira College" pageId="2" pageNumber="3">ECH</collectionCode>
|
||
.H.3431 (1 complete specimen), Siboga St. 213, Saleyer Reef (
|
||
<collectingCountry id="F31C45B2FFAD4B1FFE432782FD96B056" box="[395,525,1277,1303]" name="Indonesia" pageId="2" pageNumber="3">Indonesia</collectingCountry>
|
||
), up to
|
||
<quantity id="4CF3A8C7FFAD4B1FFDA52782FD35B056" box="[621,686,1277,1303]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="3" unit="m" value="36.0">36 m</quantity>
|
||
deep,
|
||
<date id="FFB523E2FFAD4B1FFCCB2782FC16B056" box="[771,909,1277,1303]" pageId="2" pageNumber="3" value="1899-09-26">26.ix.1899</date>
|
||
, coll. Siboga Expedition;
|
||
<collectionCode id="ED1A9DE7FFAD4B1FFB102782FAA9B056" box="[1240,1330,1277,1303]" httpUri="http://grbio.org/cool/c09s-jk3y" name="Zoological Museum, University of Athens" pageId="2" pageNumber="3">ZMUA</collectionCode>
|
||
3430 (
|
||
<specimenCount id="9D0DCEABFFAD4B1FFA452782FE8FB07D" pageId="2" pageNumber="3" type="generic">3 specimens</specimenCount>
|
||
, one complete), Siboga St. 213, Saleyer Reef (
|
||
<collectingCountry id="F31C45B2FFAD4B1FFCF9265DFC32B07D" box="[817,937,1314,1340]" name="Indonesia" pageId="2" pageNumber="3">Indonesia</collectingCountry>
|
||
), up to
|
||
<quantity id="4CF3A8C7FFAD4B1FFC35265DFBA2B07D" box="[1021,1081,1314,1340]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="3" unit="m" value="36.0">36 m</quantity>
|
||
deep,
|
||
<date id="FFB523E2FFAD4B1FFB4D265DFA9EB07D" box="[1157,1285,1314,1340]" pageId="2" pageNumber="3" value="1899-09-26">26.ix.1899</date>
|
||
, coll. Siboga Expedition; IG 28251/169 (calcareous ring and associated structures only), Kudingaieng Keke (SW Sulawesi,
|
||
<collectingCountry id="F31C45B2FFAD4B1FFF5F2612FE88B0C6" box="[151,275,1389,1415]" name="Indonesia" pageId="2" pageNumber="3">Indonesia</collectingCountry>
|
||
),
|
||
<quantity id="4CF3A8C7FFAD4B1FFEE12612FEFCB0C6" box="[297,359,1389,1415]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="3" unit="m" value="14.0">14 m</quantity>
|
||
depth,
|
||
<date id="FFB523E2FFAD4B1FFE0C2612FDD1B0C6" box="[452,586,1389,1415]" pageId="2" pageNumber="3" value="1994-09-26">26.ix.1994</date>
|
||
, coll. C. Massin;
|
||
<collectionCode id="ED1A9DE7FFAD4B1FFCED2612FC1AB0C6" box="[805,897,1389,1415]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="3">NMNH</collectionCode>
|
||
<collectionCode id="ED1A9DE7FFAD4B1FFC412612FC51B0C6" LSID="urn:lsid:biocol.org:col:12987" box="[905,970,1389,1415]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12987" name="Elmira College" pageId="2" pageNumber="3">ECH</collectionCode>
|
||
. 6090 (calcareous ring and associated structures only), Barang Lompo (SW Sulawesi,
|
||
<collectingCountry id="F31C45B2FFAD4B1FFD0226EDFCDBB0ED" box="[714,832,1426,1452]" name="Indonesia" pageId="2" pageNumber="3">Indonesia</collectingCountry>
|
||
), depth
|
||
<quantity id="4CF3A8C7FFAD4B1FFC5426EDFC57B0ED" box="[924,972,1426,1452]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="3" unit="m" value="3.0">3 m</quantity>
|
||
,
|
||
<date id="FFB523E2FFAD4B1FFC1F26EDFBCCB0ED" box="[983,1111,1426,1452]" pageId="2" pageNumber="3" value="1994-09-23">23.ix.1994</date>
|
||
, coll. C. Massin; IG 28455/ 51 (calcareous ring and associated structures only), Laing Island (
|
||
<collectingCountry id="F31C45B2FFAD4B1FFC5426C7FB1DB093" box="[924,1158,1464,1490]" name="Papua New Guinea" pageId="2" pageNumber="3">Papua New Guinea</collectingCountry>
|
||
), depth
|
||
<quantity id="4CF3A8C7FFAD4B1FFB2B26C7FA88B093" box="[1251,1299,1464,1490]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="3" unit="m" value="5.0">5 m</quantity>
|
||
,
|
||
<date id="FFB523E2FFAD4B1FFAD726C7FA0CB093" box="[1311,1431,1464,1490]" pageId="2" pageNumber="3" value="1996-10-16">16.x.1996</date>
|
||
, coll. C. Massin; IG 28455/54 (calcareous ring and associated structures only), Laing Island (
|
||
<collectingCountry id="F31C45B2FFAD4B1FFAC626A2FF6BB35D" name="Papua New Guinea" pageId="2" pageNumber="3">Papua New Guinea</collectingCountry>
|
||
), depth
|
||
<quantity id="4CF3A8C7FFAD4B1FFE84257DFEE0B35D" box="[332,379,1538,1564]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="3" unit="m" value="6.0">6 m</quantity>
|
||
,
|
||
<date id="FFB523E2FFAD4B1FFE4F257DFE64B35D" box="[391,511,1538,1564]" pageId="2" pageNumber="3" value="1996-10-17">17.x.1996</date>
|
||
, coll. C. Massin; AM
|
||
<accessionNumber id="945898C1FFAD4B1FFCC9257DFCC6B35D" box="[769,861,1538,1564]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J13573" pageId="2" pageNumber="3" type="EnaNcbi">J 13573</accessionNumber>
|
||
(one complete specimen and one calcareous ring and associated structures) (identified as
|
||
<taxonomicName id="4C0B7EA1FFAD4B1FFDA72557FD60B300" box="[623,763,1576,1601]" class="Holothuroidea" family="Phyllophoridae" genus="Neothyonidium" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="species" species="perissum">
|
||
<emphasis id="B97FD930FFAD4B1FFDA72557FD60B300" box="[623,763,1576,1601]" italics="true" pageId="2" pageNumber="3">N. perissum</emphasis>
|
||
</taxonomicName>
|
||
by F.W.E. Rowe), Heron Island (Queensland,
|
||
<collectingCountry id="F31C45B2FFAD4B1FFAD72557FA14B303" box="[1311,1423,1576,1602]" name="Australia" pageId="2" pageNumber="3">Australia</collectingCountry>
|
||
), depth ±
|
||
<quantity id="4CF3A8C7FFAD4B1FFF392532FEB4B326" box="[241,303,1613,1639]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="3" unit="m" value="12.0">12 m</quantity>
|
||
,
|
||
<date id="FFB523E2FFAD4B1FFEF42532FE5FB326" box="[316,452,1613,1639]" pageId="2" pageNumber="3" value="1979-12-12">12.xii.1979</date>
|
||
, coll. H. Silver; AM J 9490 (calcareous ring and associated structures only), Heron Island (Queensland,
|
||
<collectingCountry id="F31C45B2FFAD4B1FFE40250DFE63B3CD" box="[392,504,1650,1676]" name="Australia" pageId="2" pageNumber="3">Australia</collectingCountry>
|
||
), depth
|
||
<quantity id="4CF3A8C7FFAD4B1FFD9C250DFD08B3CD" box="[596,659,1650,1676]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="m" value="10.0">10 m</quantity>
|
||
,
|
||
<date id="FFB523E2FFAD4B1FFD56250DFC85B3CD" box="[670,798,1650,1676]" pageId="2" pageNumber="3" value="1974-11-21">21.xi.1974</date>
|
||
, coll. N. Coleman.
|
||
</paragraph>
|
||
<paragraph id="8BB40522FFAD4B1EFF0C25E8FE7CB406" blockId="2.[151,1437,1015,2012]" lastBlockId="3.[151,1437,152,327]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
<emphasis id="B97FD930FFAD4B1FFF0C25E8FBBAB3F0" bold="true" box="[196,1057,1687,1713]" pageId="2" pageNumber="3">
|
||
<typeStatus id="54B0BB80FFAD4B1FFF0C25E8FEA6B3F0" box="[196,317,1687,1713]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
|
||
description (after
|
||
<bibRefCitation id="EF9A78D3FFAD4B1FFDFF25E8FD73B3F0" author="Ludwig" box="[567,744,1687,1713]" pageId="2" pageNumber="18" refString="Ludwig, H. (1882) List of the holothurians in the collection of the Leyden Museum. Notes Leyden Museum, 4 (10), 127 - 137." type="journal article" year="1882">Ludwig 1882</bibRefCitation>
|
||
: 132, amended herein):
|
||
</emphasis>
|
||
<typeStatus id="54B0BB80FFAD4B1FFBE225E7FB05B3F3" box="[1066,1182,1688,1714]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
|
||
consists of only the calcareous ring with its associated structures; length of complete fragment (excluding tentacles and Polian vesicles)
|
||
<quantity id="4CF3A8C7FFAD4B1FFEC0259DFEFBB3BD" box="[264,352,1762,1788]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="3" unit="mm" value="75.0">75 mm</quantity>
|
||
. Colour yellowish-brown, tube feet and tentacles darker brown. Mouth encircled by welldeveloped brownish-white peristomial membrane; tentacles 20, in two distinct circles, outer circle with five pairs of large (
|
||
<quantity id="4CF3A8C7FFAD4B1FFE8E2452FE51B206" box="[326,458,1837,1863]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="2" pageNumber="3" unit="mm" value="35.0" valueMax="45.0" valueMin="25.0">25–45 mm</quantity>
|
||
) tentacles, alternating with inner circle of five pairs of small (
|
||
<quantity id="4CF3A8C7FFAD4B1FFB7D2452FA80B206" box="[1205,1307,1837,1863]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="3" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
|
||
) tentacles; shaft large tentacles pale-brown with one dark banding proximally; tips large tentacles darker brown; shaft small tentacles uniform pale brown; tips small tentacles darker. Tube feet of introvert distributed in two radial rows over most of introvert length, decreasing in size proximally, where they also scatter over interradial areas. Calcareous ring (see
|
||
<bibRefCitation id="EF9A78D3FFAD4B1FFE2324BDFC87B29D" author="Heding" box="[491,796,1986,2012]" pageId="2" pageNumber="18" refString="Heding, S. G. & Panning, A. (1954) Phyllophoridae. Eine Bearbeitung der Polytentaculaten Dendrochiroten Holothurien des Zoologische Museums in Kopenhagen. Spolia Zoologica Musei Hauniensis, 13, 1 - 209, figs 1 - 102." type="journal article" year="1954">Heding & Panning 1954</bibRefCitation>
|
||
: 197, fig. 98a)
|
||
<quantity id="4CF3A8C7FFAD4B1FFC1124BDFBB4B29D" box="[985,1071,1986,2012]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="3" unit="mm" value="70.0">70 mm</quantity>
|
||
long, tubular, spirally coiled, broken into a mosaic of small pieces camouflaged by a thick enveloping membrane; radial and interradial plates fused over about two thirds of length of calacreous ring; radial plates with deep anterior notch, posteriorly prolonged, bifurcating, processes fused with processes of neigbouring radial plates; interradial plates anteriorly pointed. Polian vesicles four,
|
||
<quantity id="4CF3A8C7FFAC4B1EFD712277FCCEB463" box="[697,853,264,290]" metricMagnitude="-1" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="3" pageNumber="4" unit="mm" value="165.0" valueMax="180.0" valueMin="150.0">150–180 mm</quantity>
|
||
long, tubular. Stone canal
|
||
<quantity id="4CF3A8C7FFAC4B1EFB582277FB78B463" box="[1168,1251,264,290]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="4" unit="mm" value="20.0">20 mm</quantity>
|
||
long, ending in small truncated madreporite.
|
||
</paragraph>
|
||
<caption id="DF7455AAFFAC4B1EFF5F26D9FAA0B09F" httpUri="https://zenodo.org/record/194048/files/figure.png" pageId="3" pageNumber="4" targetBox="[152,1436,400,1408]" targetPageId="3">
|
||
<paragraph id="8BB40522FFAC4B1EFF5F26D9FAA0B09F" blockId="3.[151,1437,1446,1502]" pageId="3" pageNumber="4">
|
||
<emphasis id="B97FD930FFAC4B1EFF5F26D9FE80B0FF" bold="true" box="[151,283,1446,1470]" pageId="3" pageNumber="4">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C0B7EA1FFAC4B1EFEEB26D8FD28B0FF" authority="Ludwig, 1882" authorityName="Ludwig" authorityYear="1882" box="[291,691,1447,1470]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="magnum">
|
||
<emphasis id="B97FD930FFAC4B1EFEEB26D8FE65B0FF" box="[291,510,1447,1470]" italics="true" pageId="3" pageNumber="4">Massinium magnum</emphasis>
|
||
(Ludwig, 1882)
|
||
</taxonomicName>
|
||
, Holotype. A. Tables and scarce rosettes of introvert; B. Rosettes of introvert tube feet; C. Rods of introvert tube feet; D. Rosettes of peristome; E. Lattices and rosette of gonoduct.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB40522FFAC4B1EFF0C254DFCFAB2A7" blockId="3.[151,1437,1586,2025]" pageId="3" pageNumber="4">
|
||
<emphasis id="B97FD930FFAC4B1EFF0C254DFE37B30A" box="[196,428,1586,1611]" italics="true" pageId="3" pageNumber="4">Ossicle assemblage</emphasis>
|
||
. Introvert with tables and scarce rosettes (figs 1A, 2A). Tables with circular to ovoid to irregular disc with smooth rims and perforated by four large central holes and usually a single circle of alternating smaller holes, occasionally two. Table discs 60–80 Μm long and 55–60 Μm wide; spire low to moderate, of two pillars united by one to two cross-bars, ending in 4–6 tooth-like projections, sometimes subdivided. Rosettes 55–70 Μm long. Tube feet of introvert with numerous rosettes (figs 1B, 2B) and perforated irregular rods, sometimes forming plates (figs 1C, 2C). Rosettes 35–45 Μm long and 20–40 Μm wide, complex with distal endings swollen, not anastamosing. Rods 90–180 Μm long, with some large perforations and more smaller ones, ending with irregular branched projections. Peristome with numerous rosettes only (figs 1D, 2D), 30–65 Μm long, complex, roundish to elongate, endings distinctly swollen, never anastomosing. Shaft and tips of small and large tentacles with rosettes only, similar in size and shape to those of peristome (figs 2E–H). Longitudinal muscles of introvert devoid of ossicles. Gonoduct with rosettes, 25–30 Μm long, and lattice-like deposits, 80–100 Μm long (fig. 1E).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31156A9FFAB4B19FF0C23E8FC3DB4F6" pageId="4" pageNumber="5" type="discussion">
|
||
<paragraph id="8BB40522FFAB4B19FF0C23E8FC11B42D" blockId="4.[151,1437,151,439]" pageId="4" pageNumber="5">
|
||
<emphasis id="B97FD930FFAB4B19FF0C23E8FEDBB5F0" bold="true" box="[196,320,151,177]" pageId="4" pageNumber="5">Remarks.</emphasis>
|
||
Regrettably, the
|
||
<typeStatus id="54B0BB80FFAB4B19FDDA23E7FDE7B5F3" box="[530,636,152,178]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
lacks the main body of the animal and although its description is accurate, it is not accompanied by illustrations. Amongst the subsequent workers who examined the
|
||
<typeStatus id="54B0BB80FFAB4B19FAFE23C2FA07B596" box="[1334,1436,189,215]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
only
|
||
<bibRefCitation id="EF9A78D3FFAB4B19FF1D239DFD8FB5BD" author="Heding" box="[213,532,226,252]" pageId="4" pageNumber="18" refString="Heding, S. G. & Panning, A. (1954) Phyllophoridae. Eine Bearbeitung der Polytentaculaten Dendrochiroten Holothurien des Zoologische Museums in Kopenhagen. Spolia Zoologica Musei Hauniensis, 13, 1 - 209, figs 1 - 102." type="journal article" year="1954">Heding & Panning (1954)</bibRefCitation>
|
||
provided a figure of its calcareous ring, but they too did not illustrate the ossicles, apart from two table deposits copied from
|
||
<bibRefCitation id="EF9A78D3FFAB4B19FCCB2277FC41B463" author="Domantay" box="[771,986,264,290]" pageId="4" pageNumber="18" refString="Domantay, J. S. (1933) Littoral Holothurioidea of Port Galera Bay and Adjacent Waters. Natural Applied Science Bulletin, University Philippines. 3, 41 - 101, pls 1 - 4." type="journal article" year="1933">Domantay (1933)</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="EF9A78D3FFAB4B19FC2E2277FB15B463" author="Sluiter" box="[998,1166,264,290]" pageId="4" pageNumber="19" refString="Sluiter, C. P. (1901) Die Holothurien der Siboga Expedition. Siboga Expedition, 44, 1 - 142, 10 pls." type="journal article" year="1901">Sluiter (1901)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF9A78D3FFAB4B19FB012277FA07B463" author="Domantay" box="[1225,1436,264,290]" pageId="4" pageNumber="18" refString="Domantay, J. S. (1933) Littoral Holothurioidea of Port Galera Bay and Adjacent Waters. Natural Applied Science Bulletin, University Philippines. 3, 41 - 101, pls 1 - 4." type="journal article" year="1933">Domantay (1933)</bibRefCitation>
|
||
accurately described complete specimens, but failed to illustrate the ossicle assemblage in detail, thus adding to the difficulty in correctly assigning specimens to
|
||
<taxonomicName id="4C0B7EA1FFAB4B19FD3F222DFC18B42A" box="[759,899,338,363]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="species" species="magnum">
|
||
<emphasis id="B97FD930FFAB4B19FD3F222DFC18B42A" box="[759,899,338,363]" italics="true" pageId="4" pageNumber="5">M. magnum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB40522FFAB4B19FF0C2207FC3DB4F6" blockId="4.[151,1437,151,439]" pageId="4" pageNumber="5">
|
||
Therefore, to avoid confusion over the identity of
|
||
<taxonomicName id="4C0B7EA1FFAB4B19FCF72207FC4FB4D0" box="[831,980,376,401]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="species" species="magnum">
|
||
<emphasis id="B97FD930FFAB4B19FCF72207FC4FB4D0" box="[831,980,376,401]" italics="true" pageId="4" pageNumber="5">M. magnum</emphasis>
|
||
</taxonomicName>
|
||
we have here re-described the
|
||
<typeStatus id="54B0BB80FFAB4B19FA962207FA03B4D3" box="[1374,1432,376,402]" pageId="4" pageNumber="5">type</typeStatus>
|
||
, supplementing it with a description of a complete specimen below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7455AAFFAB4B19FF5F24F7FC6BB29E" httpUri="https://zenodo.org/record/194049/files/figure.png" pageId="4" pageNumber="5" targetBox="[283,1303,493,1907]" targetPageId="4">
|
||
<paragraph id="8BB40522FFAB4B19FF5F24F7FC6BB29E" blockId="4.[151,1436,1928,2015]" pageId="4" pageNumber="5">
|
||
<emphasis id="B97FD930FFAB4B19FF5F24F7FE82B2E1" bold="true" box="[151,281,1928,1952]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C0B7EA1FFAB4B19FEE824F7FD32B2DE" authority="Ludwig, 1882" authorityName="Ludwig" authorityYear="1882" box="[288,681,1928,1951]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="species" species="magnum">
|
||
<emphasis id="B97FD930FFAB4B19FEE824F7FE63B2DE" box="[288,504,1928,1951]" italics="true" pageId="4" pageNumber="5">Massinium magnum</emphasis>
|
||
(Ludwig, 1882)
|
||
</taxonomicName>
|
||
, Holotype. A. Tables of introvert; B. Rosettes of introvert tube feet; C. Rods of introvert tube feet; D. Rosettes of peristome; E. Rosettes of shaft of large tentacle; F. Rosette of tip of large tentacle; G. Rosette of shaft of small tentacle; H. Rosette of tip of small tentacle.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7455AAFFAA4B18FF5F2451FBD0B2A7" httpUri="https://zenodo.org/record/194050/files/figure.png" pageId="5" pageNumber="6" targetBox="[231,1358,202,1837]" targetPageId="5">
|
||
<paragraph id="8BB40522FFAA4B18FF5F2451FBD0B2A7" blockId="5.[151,1436,1838,2022]" pageId="5" pageNumber="6">
|
||
<emphasis id="B97FD930FFAA4B18FF5F2451FE89B207" bold="true" box="[151,274,1838,1862]" pageId="5" pageNumber="6">FIGURE 3</emphasis>
|
||
<taxonomicName id="4C0B7EA1FFAA4B18FED22450FD3FB207" authority="Ludwig, 1882" authorityName="Ludwig" authorityYear="1882" box="[282,676,1839,1862]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="5" pageNumber="6" phylum="Echinodermata" rank="species" species="magnum">
|
||
<emphasis id="B97FD930FFAA4B18FED22450FE69B207" box="[282,498,1839,1862]" italics="true" pageId="5" pageNumber="6">Massinium magnum</emphasis>
|
||
(Ludwig, 1882)
|
||
</taxonomicName>
|
||
, non-type material (complete specimen). A. Tables, pseudobuttons and rosettes of ventral body wall; B. Tables, pseudobuttons and rosettes of dorsal body wall; C. Perforated plate of ventral tube feet; D. Pseudobuttons of ventral tube feet; E. Perforated plate of dorsal tube feet; F. Pseudobuttons of dorsal tube feet; G. Tables and rare rosette of introvert; H. Perforated plates of introvert tube feet; J. Rosettes of introvert tube feet; K. Rosettes of peristome; L. Pseudobuttons and irregular rods of longitudinal muscles of body wall; M. Pseudobuttons and irregular rods of cloacal retractor muscles; N. Lattice-like plate and rosettes of gonoduct.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7455AAFFA94B1BFF5F2787FC02B06E" httpUri="https://zenodo.org/record/194051/files/figure.png" pageId="6" pageNumber="7" targetBox="[291,1296,183,1260]" targetPageId="6">
|
||
<paragraph id="8BB40522FFA94B1BFF5F2787FC02B06E" blockId="6.[151,1435,1272,1327]" pageId="6" pageNumber="7">
|
||
<emphasis id="B97FD930FFA94B1BFF5F2787FE83B051" bold="true" box="[151,280,1272,1296]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C0B7EA1FFA94B1BFED72787FD3DB04E" authority="Ludwig, 1882" authorityName="Ludwig" authorityYear="1882" box="[287,678,1272,1295]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="7" phylum="Echinodermata" rank="species" species="magnum">
|
||
<emphasis id="B97FD930FFA94B1BFED72787FE6DB04E" box="[287,502,1272,1295]" italics="true" pageId="6" pageNumber="7">Massinium magnum</emphasis>
|
||
(Ludwig, 1882)
|
||
</taxonomicName>
|
||
, non type material (complete specimen). A. Pseudobuttons and rosettes of ventral body wall; B. Pseudobuttons and rosettes of dorsal body wall.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31156A9FFA94B1AFF0C26FDFC7DB646" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
|
||
<paragraph id="8BB40522FFA94B1BFF0C26FDFCBFB2B3" blockId="6.[151,1436,1410,2034]" pageId="6" pageNumber="7">
|
||
<emphasis id="B97FD930FFA94B1BFF0C26FDFC27B0DD" bold="true" box="[196,956,1410,1436]" pageId="6" pageNumber="7">Description of complete specimen [ZMUA V.ECH.H.3431]:</emphasis>
|
||
Specimen entire, well-preserved. Body wall firm, but not thick (
|
||
<quantity id="4CF3A8C7FFA94B1BFE0E26D7FDABB083" box="[454,560,1448,1474]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="6" pageNumber="7" unit="mm" value="2.5" valueMax="4.0" valueMin="1.0">1–4 mm</quantity>
|
||
), slightly rough to the touch. Body form cylindrical, U-shaped, slightly contracted. Introvert well-extended, attached to main body by narrow, proximal end. Body length along ventral surface
|
||
<quantity id="4CF3A8C7FFA94B1BFE85268DFE2BB34D" box="[333,432,1522,1548]" metricMagnitude="-1" metricUnit="m" metricValue="1.18" pageId="6" pageNumber="7" unit="mm" value="118.0">118 mm</quantity>
|
||
, along dorsal surface
|
||
<quantity id="4CF3A8C7FFA94B1BFD7A268DFC93B34D" box="[690,776,1522,1548]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="7" unit="mm" value="90.0">90 mm</quantity>
|
||
, height of mid-body
|
||
<quantity id="4CF3A8C7FFA94B1BFBC8268DFBCCB34D" box="[1024,1111,1522,1548]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="7" unit="mm" value="60.0">60 mm</quantity>
|
||
, length of introvert
|
||
<quantity id="4CF3A8C7FFA94B1BFA89268DFA03B34D" box="[1345,1432,1522,1548]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="7" unit="mm" value="85.0">85 mm</quantity>
|
||
, length of constricted proximal part
|
||
<quantity id="4CF3A8C7FFA94B1BFDFF2567FD15B373" box="[567,654,1560,1586]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="7" unit="mm" value="13.0">13 mm</quantity>
|
||
. Colouration of body and introvert uniform yellowish-white. Tube feet of body wall yellowish, numerous, retracted, scattered more or less evenly over dorsal and ventral radial and interradial areas, suckers small. Tube feet of introvert brownish, aligned in radial areas in two welldefined rows, except proximally where they also occur in interradial areas, absent in distal
|
||
<quantity id="4CF3A8C7FFA94B1BFB0325F7FAB9B3E3" box="[1227,1314,1672,1698]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="7" unit="mm" value="30.0">30 mm</quantity>
|
||
. Tentacles 19, nine large, 10 small, arranged in two circles; large tentacles in outer circle, ±
|
||
<quantity id="4CF3A8C7FFA94B1BFB8C25D2FB0DB386" box="[1092,1174,1709,1735]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="7" unit="mm" value="60.0">60 mm</quantity>
|
||
long, in pairs in radial areas; small tentacles, in inner circle, ±
|
||
<quantity id="4CF3A8C7FFA94B1BFDB225ACFD54B3AC" box="[634,719,1747,1773]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="mm" value="10.0">10 mm</quantity>
|
||
long, in pairs in interradial areas. Tentacles well branched, shaft yellowish-brown, some tentacles slightly banded; branches uniform dark brown. Anus retracted into body, presence or absence of anal teeth not determined. Calcareous ring
|
||
<quantity id="4CF3A8C7FFA94B1BFC2D2462FBA3B276" box="[997,1080,1821,1847]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="7" unit="mm" value="80.0">80 mm</quantity>
|
||
long, tubular, fragmented into a mosaic of small pieces obscured by thick membrane; radial and interradial plates fused over about seven eights of length of calcareous ring; radial plates posteriorly prolonged, bifurcating into processes which fuse with those of neighbouring radial plates. Polian vesicles four, white,
|
||
<quantity id="4CF3A8C7FFA94B1BFC1224F2FBF1B2E6" box="[986,1130,1933,1959]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="6" pageNumber="7" unit="mm" value="105.0" valueMax="120.0" valueMin="90.0">90–120 mm</quantity>
|
||
long. Stone canal single, slightly contorted, free,
|
||
<quantity id="4CF3A8C7FFA94B1BFE7F24CCFD90B28C" box="[439,523,1971,1997]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="7" unit="mm" value="26.0">26 mm</quantity>
|
||
long, terminating in small globular madreporite. Introvert retractor muscles thick and short, originating at anterior third of the body.
|
||
</paragraph>
|
||
<paragraph id="8BB40522FFA84B1AFF0C23E7FCC4B7D6" blockId="7.[151,1437,152,1484]" pageId="7" pageNumber="8">
|
||
<emphasis id="B97FD930FFA84B1AFF0C23E7FE36B5F0" box="[196,429,152,177]" italics="true" pageId="7" pageNumber="8">Ossicle assemblage</emphasis>
|
||
. Body wall with rosettes, pseudobuttons and few tables, mostly reduced (figs 3A,B; 4A,B). Rosettes simple, slightly elongated, 25–40 Μm long; pseudobuttons of various shapes, with 2–10 small, randomly placed perforations; tables scarce, reduced to oval, smooth disc, generally with four large central holes and few smaller peripheral ones, occasionally with reduced spire. Tube feet of ventral and dorsal body wall with pseudobuttons, rosettes, elongated perforated plates surrounding endplate (fig 3C–F). Introvert with tables and occasionally rosettes (fig. 3G). Tables with oval, smooth disc, 45–80 Μm wide and 75–95 Μm long, perforated by four large central holes and usually one, occasionally two, circles of smaller peripheral holes, the larger of these alternating with the central holes, the other arranged more irregularly; spire consisting of two pillars, united by 1–2 cross-bars, terminating in some toothed projections. Tube feet of introvert with rosettes and irregular plates with branched ends (fig. 3H,J). Peristome with only complex rosettes, 50–75 Μm long (fig. 3K). Shaft and tips of both large and small tentacles sometimes with few rosettes. Longitudinal muscles of body wall with irregular perforated rods and pseudobuttons (fig. 3L). Longitudinal muscles of introvert devoid of ossicles. Cloacal retractor muscles with irregular deposits (fig 3M). Gonoduct with rosettes and lattice-like plates (fig. 3N).
|
||
</paragraph>
|
||
<paragraph id="8BB40522FFA84B1AFF0C21DDFC7DB646" blockId="7.[151,1437,152,1484]" pageId="7" pageNumber="8">
|
||
<emphasis id="B97FD930FFA84B1AFF0C21DDFEA9B7FD" bold="true" box="[196,306,674,700]" pageId="7" pageNumber="8">Ecology:</emphasis>
|
||
<bibRefCitation id="EF9A78D3FFA84B1AFEF321DDFD8DB7FD" author="Massin" box="[315,534,674,700]" pageId="7" pageNumber="19" refString="Massin, C. (1999) Reef-dwelling Holothuroidea (Echinodermata) of the Spermonde Archipelago (South-West Sulawesi, Indonesia). Zoologische Verhandelingen, 329, 1 - 144." type="journal article" year="1999">Massin (1999: 90)</bibRefCitation>
|
||
noted ‘…they live deeply buried in sand or coral rubble’ for material from Sulawesi and in 2005 (p. 69) ‘body always buried in sand, only tentacles and introvert visisble; tentacles coming out at sunset and at night’ for material from
|
||
<collectingCountry id="F31C45B2FFA84B1AFD342192FC79B646" box="[764,994,749,775]" name="Papua New Guinea" pageId="7" pageNumber="8">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31156A9FFA84B1AFF0C206DFBBAB08D" pageId="7" pageNumber="8" type="discussion">
|
||
<paragraph id="8BB40522FFA84B1AFF0C206DFD57B13D" blockId="7.[151,1437,152,1484]" pageId="7" pageNumber="8">
|
||
<emphasis id="B97FD930FFA84B1AFF0C206DFEDAB66D" bold="true" box="[196,321,786,812]" pageId="7" pageNumber="8">Remarks:</emphasis>
|
||
The ossicle assemblage of the complete specimen here described (V.Ech.H. 3431) differs in some aspects from the
|
||
<typeStatus id="54B0BB80FFA84B1AFE652047FD81B613" box="[429,538,824,850]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
. For instance the tables of the introvert of the
|
||
<typeStatus id="54B0BB80FFA84B1AFB832047FB2FB613" box="[1099,1204,824,850]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
have discs that are slightly smaller and less perforated than those of the complete specimen. However, comparison of the
|
||
<typeStatus id="54B0BB80FFA84B1AFF5F20FDFF65B6DD" box="[151,254,898,924]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
and the other specimens we examined showed that this variation is to be considered intra- rather than interspecific. Nevertheless, we want to note that the second of Sluiter’s (1901) complete specimens (V.Ech.H. 3430, partim) differs slightly from his best preserved specimen described above (V.Ech.H. 3431) in presenting five minute calcareous teeth. However, as the ossicle assemblage of the introvert and tentacles of all of Sluiter’s specimens largely corresponds with that of the
|
||
<typeStatus id="54B0BB80FFA84B1AFC832767FC18B173" box="[843,899,1048,1074]" pageId="7" pageNumber="8">type</typeStatus>
|
||
, except for the abundance of ossicles in the tentacles and the presence or absence of rosettes in the longitudinal muscles of the introvert, we conclude that all these specimens are referable to
|
||
<taxonomicName id="4C0B7EA1FFA84B1AFDF1271CFD5EB13D" box="[569,709,1123,1148]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="magnum">
|
||
<emphasis id="B97FD930FFA84B1AFDF1271CFD5EB13D" box="[569,709,1123,1148]" italics="true" pageId="7" pageNumber="8">M. magnum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB40522FFA84B1AFF0C27F7FBBAB08D" blockId="7.[151,1437,152,1484]" pageId="7" pageNumber="8">
|
||
Finally, an important difference between all of Sluiter’s voucher material and the
|
||
<typeStatus id="54B0BB80FFA84B1AFB4127F7FB68B1E3" box="[1161,1267,1160,1186]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
, together with Massin’s voucher materials from Sulawesi (1999) and
|
||
<collectingCountry id="F31C45B2FFA84B1AFCED27D2FB97B186" box="[805,1036,1197,1223]" name="Papua New Guinea" pageId="7" pageNumber="8">Papua New Guinea</collectingCountry>
|
||
(2005), rests with the banding of the tentacles. Whereas the former shows a more uniform brownish colouration (see also
|
||
<bibRefCitation id="EF9A78D3FFA84B1AFB6B27ADFAF8B1AD" author="Domantay" box="[1187,1379,1234,1260]" pageId="7" pageNumber="18" refString="Domantay, J. S. (1933) Littoral Holothurioidea of Port Galera Bay and Adjacent Waters. Natural Applied Science Bulletin, University Philippines. 3, 41 - 101, pls 1 - 4." type="journal article" year="1933">Domantay 1933</bibRefCitation>
|
||
), the latter show clear banding at the base of the tentacles. There are also some differences in the ossicle assemblage of the muscles of the introvert between the
|
||
<typeStatus id="54B0BB80FFA84B1AFCF02662FCF7B076" box="[824,876,1309,1335]" pageId="7" pageNumber="8">type</typeStatus>
|
||
and Sluiter’s and Massin’s voucher materials: while the
|
||
<typeStatus id="54B0BB80FFA84B1AFEC3263DFEDBB01D" box="[267,320,1346,1372]" pageId="7" pageNumber="8">type</typeStatus>
|
||
, Massin’s Sulawesi material and two of Sluiter’s specimens did not present these ossicles, two of Sluiter’s
|
||
<collectingCountry id="F31C45B2FFA84B1AFEFF2617FE23B0C3" box="[311,440,1384,1410]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
|
||
and both of Massin’s
|
||
<collectingCountry id="F31C45B2FFA84B1AFD2C2617FC79B0C3" box="[740,994,1384,1410]" name="Papua New Guinea" pageId="7" pageNumber="8">Papua New Guinea</collectingCountry>
|
||
specimens did show these. More representative material from the whole distribution range of the species may eventually allow the separation of these slightly different forms which, for now, we retain under
|
||
<emphasis id="B97FD930FFA84B1AFC4526CCFBBAB08D" box="[909,1057,1459,1484]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName id="4C0B7EA1FFA84B1AFC4526CCFB86B08D" box="[909,1053,1459,1484]" class="Holothuroidea" family="Phyllophoridae" genus="Massinium" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="magnum">M. magnum</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |