426 lines
54 KiB
XML
426 lines
54 KiB
XML
<document id="2E598ECCBC282E11129A38F9F696C0EA" ID-DOI="10.1206/4015.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="12171426" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718827081284" checkinUser="felipe" docAuthor="Grimaldi, David A." docDate="2024" docId="0387351BFFE5FFA4F63A5DCDFC9AFCBC" docLanguage="en" docName="AmMusNovit.2024.4015.1-43.pdf" docOrigin="American Museum Novitates 2024 (4015)" docSource="https://bioone.org/journals/american-museum-novitates/volume-2024/issue-4015/4015.1/The-Drosophila-Sophophora-obscura-species-group-in-the-Americas-Diptera/10.1206/4015.1.full" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Drosophila (Sophophora) narragansett Sturtevant and Dobzhansky 1936" docType="treatment" docVersion="2" lastPageNumber="30" masterDocId="FFBE4D63FFFDFFB9F7345809FFBBFF97" masterDocTitle="The Drosophila (Sophophora) obscura species group in the Americas (Diptera: Drosophilidae): review, revisions, and three new species" masterLastPageNumber="44" masterPageNumber="1" pageNumber="25" updateTime="1718827702117" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="656C1F3E83AF1D377C731EA49FE4D550" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="7876E231630A89DD2FE0BE839D426ABE">
|
||
<mods:title id="3D987C21B513B79AB8C0C8A5F117DB82">The Drosophila (Sophophora) obscura species group in the Americas (Diptera: Drosophilidae): review, revisions, and three new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="C6434B80E3FBBD65E30ADD34C513FF92" type="personal">
|
||
<mods:role id="892E3E4928867CA1C457B9583ABF7D25">
|
||
<mods:roleTerm id="C53C20B2B11420796ABA8F678833D178">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="AC36AE2845C47EAD7D101A12485AA296">Grimaldi, David A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="109E04BA251DFC23C9028567B05F8800">text</mods:typeOfResource>
|
||
<mods:relatedItem id="C5EE1AA50319C8DC335EFBE22E297D7E" type="host">
|
||
<mods:titleInfo id="9F167CA9CA776022BF8E80F2CE02A094">
|
||
<mods:title id="FB11EB44FEB73B0F6B40C59FA7D9AE4E">American Museum Novitates</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="1BD9641463218CBB6B98CF2C7EABF461">
|
||
<mods:date id="1819C3F7D9B2B0C1F1719AFDAF0D0DD9">2024</mods:date>
|
||
<mods:detail id="1459759F42E4ADD9FA94660683E9D0D5" type="pubDate">
|
||
<mods:number id="94948FB38599CFB8DDDCE18C0ACF47FC">2024-05-16</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="236C5CC7CCA5E48630AD6A168B8E6FB7" type="volume">
|
||
<mods:number id="406986AA75BF1B825C5148AED8E0B45D">2024</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="82FD05DE23D8DBB5B9FA21EFCA56DBCD" type="issue">
|
||
<mods:number id="24338541E51A2AC9548BF97E51EF1A3B">4015</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="AD4821336316655F7F7E9BDAC9EA8F16" unit="page">
|
||
<mods:start id="5D9648BAA1D41B7F4A65D3AF49BA4C57">1</mods:start>
|
||
<mods:end id="F68C0E7CE8DEDCFA78ED8B140B52579D">44</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="0099668E0F73A2DA0F67DC5F2C06C9EA">
|
||
<mods:url id="484DA5678B714C3CB43121785EC64FDA">https://bioone.org/journals/american-museum-novitates/volume-2024/issue-4015/4015.1/The-Drosophila-Sophophora-obscura-species-group-in-the-Americas-Diptera/10.1206/4015.1.full</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="470A138050AF56251E8B43D97784D1B5">journal article</mods:classification>
|
||
<mods:identifier id="67149067C4D68F388BB3EA9C11E1F7AC" type="DOI">10.1206/4015.1</mods:identifier>
|
||
<mods:identifier id="96DB0F5CC9E60961BB4BA74FF79A9423" type="ISSN">0003-0082</mods:identifier>
|
||
<mods:identifier id="6BB72EB02804F4CB446CED67553BBAE2" type="Zenodo-Dep">12171426</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="0387351BFFE5FFA4F63A5DCDFC9AFCBC" LSID="urn:lsid:plazi:treatment:0387351BFFE5FFA4F63A5DCDFC9AFCBC" httpUri="http://treatment.plazi.org/id/0387351BFFE5FFA4F63A5DCDFC9AFCBC" lastPageId="29" lastPageNumber="30" pageId="24" pageNumber="25">
|
||
<subSubSection id="C334D786FFE5FFA1F63A5DCDFBA1FA49" box="[270,1050,1476,1502]" pageId="24" pageNumber="25" type="nomenclature">
|
||
<paragraph id="8B91840DFFE5FFA1F63A5DCDFBA1FA49" blockId="24.[270,1050,1476,1502]" box="[270,1050,1476,1502]" pageId="24" pageNumber="25">
|
||
<heading id="D0D93361FFE5FFA1F63A5DCDFBA1FA49" box="[270,1050,1476,1502]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="5">
|
||
<taxonomicName id="4C2EFF8EFFE5FFA1F63A5DCDFBA1FA49" ID-CoL="8BXDH" authority="Sturtevant and Dobzhansky" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[270,1050,1476,1502]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="narragansett" subGenus="Sophophora">
|
||
<emphasis id="B95A581FFFE5FFA1F63A5DCDFE37FA49" box="[270,396,1476,1502]" italics="true" pageId="24" pageNumber="25">Drosophila</emphasis>
|
||
(
|
||
<emphasis id="B95A581FFFE5FFA1F6AA5DCDFD9DFA49" box="[414,550,1476,1502]" italics="true" pageId="24" pageNumber="25">Sophophora</emphasis>
|
||
)
|
||
<emphasis id="B95A581FFFE5FFA1F50D5DCCFD70FA49" box="[569,715,1477,1502]" italics="true" pageId="24" pageNumber="25">narragansett</emphasis>
|
||
Sturtevant and Dobzhansky
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334D786FFE5FFA1F6A45DF5FC22F980" box="[400,921,1532,1559]" pageId="24" pageNumber="25" type="description">
|
||
<paragraph id="8B91840DFFE5FFA1F6A45DF5FC22F980" blockId="24.[400,921,1532,1559]" box="[400,921,1532,1559]" pageId="24" pageNumber="25">
|
||
<heading id="D0D93361FFE5FFA1F6A45DF5FC22F980" box="[400,921,1532,1559]" centered="true" fontSize="36" level="3" pageId="24" pageNumber="25" reason="7">
|
||
<figureCitation id="13159888FFE5FFA1F6A45DF5FD8EF980" box="[400,565,1532,1559]" captionStart="FIG" captionStartId="5.[109,149,1138,1159]" captionTargetBox="[108,1205,224,1111]" captionTargetId="figure-185@5.[108,1212,224,1111]" captionTargetPageId="5" captionText="FIG. 2. Heads and thoraces (lateral view) of representative New World species of the obscura group (A–D), and a male midleg of D. cuauhtemoci (E). A. D. chibcha holotype, n. sp. (ASG 05); B. D. maya holotype. C. D. narragansett paralectotype male. D. D. seminole holotype. E. D. cuauhtemoci, male midleg (lateral view), paratype." figureDoi="http://doi.org/10.5281/zenodo.12171434" httpUri="https://zenodo.org/record/12171434/files/figure.png" pageId="24" pageNumber="25">Figures 2C, D</figureCitation>
|
||
;
|
||
<figureCitation id="13159888FFE5FFA1F5775DF5FD33F980" box="[579,648,1532,1559]" captionStart="FIG" captionStartId="9.[108,150,1381,1402]" captionTargetBox="[108,1212,230,1353]" captionTargetId="figure-89@9.[108,1212,224,1354]" captionTargetPageId="9" captionText="FIG. 5. Frontal view of heads of Drosophila narragansett (including seminole, new synonym) and the related species D. olmeca, n. sp. A. D. narragansett, female paralectotype (ASG 11); B. D. narragansett, male paralectotype (ASG 10); C. D. seminole, male holotype; D. D. olmeca, n. sp., male holotype (ASG 25)." figureDoi="http://doi.org/10.5281/zenodo.12171459" httpUri="https://zenodo.org/record/12171459/files/figure.png" pageId="24" pageNumber="25">5A–C</figureCitation>
|
||
,
|
||
<figureCitation id="13159888FFE5FFA1F5A15DF4FD09F980" box="[661,690,1533,1559]" captionStart="FIG" captionStartId="15.[110,152,1343,1364]" captionTargetBox="[183,1175,246,1308]" captionTargetId="figure-174@15.[108,1212,224,1317]" captionTargetPageId="15" captionText="FIG. 7. Oviscapts and spermathecae (lateral views) of species in the affinis subgroup, with apical details for three species. A. D. affinis (ASG 28); B. D. athabasca (ASG 30); C. D. azteca (ASG 15); D. D. chibcha, n. sp. (ASG 17); E. D. dobzhanskii (ASG 33); F. D. narragansett (ASG 11); G. D. tolteca (ASG 24)." figureDoi="http://doi.org/10.5281/zenodo.12171461" httpUri="https://zenodo.org/record/12171461/files/figure.png" pageId="24" pageNumber="25">7F</figureCitation>
|
||
,
|
||
<figureCitation id="13159888FFE5FFA1F58A5DF5FD63F981" box="[702,728,1532,1558]" captionStart="FIG" captionStartId="17.[108,149,1608,1629]" captionTargetBox="[173,1210,226,1573]" captionTargetId="figure-6@17.[108,1212,224,1581]" captionTargetPageId="17" captionText="FIG. 9. Protarsal male sex combs (tarsomeres ta1 and ta2, and ta3 for D. chibcha from Peru) of species in the affinis subgroup. A. D. affinis (ASG 29); B. D. algonquin (ASG 32); C. D. athabasca (ASG 31); D. D. azteca (ASG 13: San Franciso, CA); E. D. azteca (ASG 06: Leticia, Colombia); F. D. chibcha, n. sp. (ASG 04: Zurqui, Costa Rica); G. D. chibcha, n. sp. (ASG 05: Las Alturas, Costa Rica); H. D. chibcha (ASG 07: Rancho Grande, Venezuela); I. D. chibcha (ASG 08: Wayqecha, Peru); J. D. narragansett (ASG 10); K. D. olmeca (ASG 25); L. D. tolteca (ASG 23). All to the same scale." figureDoi="http://doi.org/10.5281/zenodo.12171465" httpUri="https://zenodo.org/record/12171465/files/figure.png" pageId="24" pageNumber="25">9J</figureCitation>
|
||
;
|
||
<figureCitation id="13159888FFE5FFA1F5D15DF5FC8CF980" box="[741,823,1532,1559]" captionStart="FIG" captionStartId="20.[108,149,1681,1702]" captionTargetBox="[130,1151,230,1652]" captionTargetId="figure-6@20.[108,1173,224,1652]" captionTargetPageId="20" captionText="FIG. 12. Epandria and associated structures in representative species of the affinis subgroup. A. D. narragansett (ASG 10); B. Details of A; C. D. olmeca, n. sp. (ASG 25); D. Detail of C; E. D. tolteca (ASG 23)." figureDoi="http://doi.org/10.5281/zenodo.12171471" httpUri="https://zenodo.org/record/12171471/files/figure.png" pageId="24" pageNumber="25">12A, B</figureCitation>
|
||
;
|
||
<figureCitation id="13159888FFE5FFA1F4705DF5FC22F980" box="[836,921,1532,1559]" captionStart="FIG" captionStartId="26.[108,149,1461,1482]" captionTargetBox="[191,1202,241,1419]" captionTargetId="figure-64@26.[108,1212,226,1434]" captionTargetPageId="26" captionText="FIG. 15. Hypandrium, aedeagus and other appendages of representative species of the affinis subgroup (ventral views). A. D. narragansett (ASG 10: paralectotype, Massachusetts); B. D. olmeca (ASG 25: holotype); C. D. tolteca (ASG 23); D. D. narragansett (ASG: Virginia); E. D. olmeca (ASG 25: holotype, detail of B). A–C to same scale; D and E to same scale" figureDoi="http://doi.org/10.5281/zenodo.12171477" httpUri="https://zenodo.org/record/12171477/files/figure.png" pageId="24" pageNumber="25">15A, D</figureCitation>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334D786FFE5FFA1F7585E3FFBDBF9E7" pageId="24" pageNumber="25" type="reference_group">
|
||
<paragraph id="8B91840DFFE5FFA1F7585E3FFC11F9D9" blockId="24.[108,1120,1590,1648]" box="[108,938,1590,1614]" pageId="24" pageNumber="25">
|
||
<taxonomicName id="4C2EFF8EFFE5FFA1F7585E3FFC1DF9D9" ID-CoL="8BXDH" authority="Sturtevant and Dobzhansky, 1936: 577" authorityName="Sturtevant and Dobzhansky" authorityPageNumber="577" authorityYear="1936" box="[108,934,1590,1614]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="narragansett" subGenus="Sophophora">
|
||
<emphasis id="B95A581FFFE5FFA1F7585E3FFF64F9D9" box="[108,223,1590,1614]" italics="true" pageId="24" pageNumber="25">Drosophila</emphasis>
|
||
(
|
||
<emphasis id="B95A581FFFE5FFA1F7DB5E3FFED0F9D9" box="[239,363,1590,1614]" italics="true" pageId="24" pageNumber="25">Sophophora</emphasis>
|
||
)
|
||
<emphasis id="B95A581FFFE5FFA1F6485E3EFDBBF9D9" box="[380,512,1591,1614]" italics="true" pageId="24" pageNumber="25">narragansett</emphasis>
|
||
Sturtevant and Dobzhansky, 1936: 577
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B91840DFFE5FFA1F7585E51FBDBF9E7" blockId="24.[108,1120,1590,1648]" box="[108,1120,1624,1648]" pageId="24" pageNumber="25">
|
||
<taxonomicName id="4C2EFF8EFFE5FFA1F7585E51FCC7F9E7" ID-CoL="8BXHQ" authority="Sturtevant and Dobzhansky, 1936: 577" authorityName="Sturtevant and Dobzhansky" authorityPageNumber="577" authorityYear="1936" box="[108,892,1624,1648]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="seminole" subGenus="Sophophora">
|
||
<emphasis id="B95A581FFFE5FFA1F7585E51FF64F9E7" box="[108,223,1624,1648]" italics="true" pageId="24" pageNumber="25">Drosophila</emphasis>
|
||
(
|
||
<emphasis id="B95A581FFFE5FFA1F7DB5E51FED0F9E7" box="[239,363,1624,1648]" italics="true" pageId="24" pageNumber="25">Sophophora</emphasis>
|
||
)
|
||
<emphasis id="B95A581FFFE5FFA1F6485E51FE6CF9E7" box="[380,471,1624,1648]" italics="true" pageId="24" pageNumber="25">seminole</emphasis>
|
||
Sturtevant and Dobzhansky, 1936: 577
|
||
</taxonomicName>
|
||
. NEW SYNONYMY.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334D786FFE5FFA2F7A85E87FD5CFE06" lastPageId="27" lastPageNumber="28" pageId="24" pageNumber="25" type="diagnosis">
|
||
<paragraph id="8B91840DFFE5FFA2F7A85E87FD5CFE06" blockId="24.[108,1213,1678,1742]" lastBlockId="27.[108,1214,225,1745]" lastPageId="27" lastPageNumber="28" pageId="24" pageNumber="25">
|
||
DIAGNOSIS: Males immediately recognized among North American species of the group by the silvery frons, best seen in frontal view in dried/fresh specimens. Distinguished from
|
||
<taxonomicName id="4C2EFF8EFFE5FFA2F3965EBDFF07FF6C" authority="Grimaldi, 2024" authorityName="Grimaldi" authorityYear="2024" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" lastPageId="27" lastPageNumber="28" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="olmeca" status="n. sp.">
|
||
<emphasis id="B95A581FFFE5FFA2F3965EBDFF07FF6C" italics="true" lastPageId="27" lastPageNumber="28" pageId="24" pageNumber="25">D. olmeca</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A269E564FFE6FFA2F7FE58EBFEB3FF6B" box="[202,264,226,252]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
<collectingRegion id="49EA4AEFFFE6FFA2F62358E8FEC3FF6C" box="[279,376,225,251]" country="Mexico" name="Chiapas" pageId="27" pageNumber="28">Chiapas</collectingRegion>
|
||
<collectingRegion id="49EA4AEFFFE6FFA2F64B58E8FE60FF6C" box="[383,475,225,251]" country="Mexico" name="Mexico" pageId="27" pageNumber="28">Mexico</collectingRegion>
|
||
), also with a silvery male frons, as discussed under that species below.
|
||
<taxonomicName id="4C2EFF8EFFE6FFA2F78A590EFECFFEB7" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[190,372,263,288]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="narragansett">
|
||
<emphasis id="B95A581FFFE6FFA2F78A590EFECFFEB7" box="[190,372,263,288]" italics="true" pageId="27" pageNumber="28">D. narragansett</emphasis>
|
||
</taxonomicName>
|
||
further distinguished by the following: male ta1 with 4–5 teeth and ta2 with 1; ta1 1.3× length of ta2; base of inner ventral epandrial lobe with furrows, no microtrichia; surstylus with 9–10 prensisetae. Spermatheca distinctive, roughly cup-shaped, apical indentation short; sleeve large, conical, with fine annulations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF51D485FFE4FFA0F7585E55FD00F970" ID-DOI="http://doi.org/10.5281/zenodo.12171475" ID-Zenodo-Dep="12171475" httpUri="https://zenodo.org/record/12171475/files/figure.png" pageId="25" pageNumber="26" startId="25.[108,149,1628,1649]" targetBox="[238,1179,236,1592]" targetPageId="25" targetType="figure">
|
||
<paragraph id="8B91840DFFE4FFA0F7585E55FD00F970" blockId="25.[108,1212,1626,1767]" pageId="25" pageNumber="26">
|
||
FIG. 14. Hypandrium, aedeagus and other appendages of representative species of the
|
||
<taxonomicName id="4C2EFF8EFFE4FFA0F4F35E53FBD3F9E5" authority="subgroup" authorityName="SUBGROUP" box="[967,1128,1626,1650]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="12" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B95A581FFFE4FFA0F4F35E53FBB8F9E5" box="[967,1027,1626,1650]" italics="true" pageId="25" pageNumber="26">affinis</emphasis>
|
||
subgroup
|
||
</taxonomicName>
|
||
(ventral views). The valves of the aedeagal sheath and the pair of hypandrial lobes are highlighted in color; the postgonites are not.
|
||
<emphasis id="B95A581FFFE4FFA0F6245E9DFE92F93B" bold="true" box="[272,297,1684,1708]" pageId="25" pageNumber="26">A.</emphasis>
|
||
<taxonomicName id="4C2EFF8EFFE4FFA0F61A5E9CFE32F93A" box="[302,393,1685,1709]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B95A581FFFE4FFA0F61A5E9CFE32F93A" box="[302,393,1685,1709]" italics="true" pageId="25" pageNumber="26">D. affinis</emphasis>
|
||
</taxonomicName>
|
||
(ASG 29) (hypandrial lobes hidden);
|
||
<emphasis id="B95A581FFFE4FFA0F4395E9CFC98F93A" bold="true" box="[781,803,1685,1709]" pageId="25" pageNumber="26">B.</emphasis>
|
||
<taxonomicName id="4C2EFF8EFFE4FFA0F41D5E9CFC11F93B" box="[809,938,1684,1708]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="athabasca">
|
||
<emphasis id="B95A581FFFE4FFA0F41D5E9CFC11F93B" box="[809,938,1684,1708]" italics="true" pageId="25" pageNumber="26">D. athabasca</emphasis>
|
||
</taxonomicName>
|
||
(ASG 31);
|
||
<emphasis id="B95A581FFFE4FFA0F32A5E9DFB8DF93B" bold="true" box="[1054,1078,1684,1708]" pageId="25" pageNumber="26">C.</emphasis>
|
||
<taxonomicName id="4C2EFF8EFFE4FFA0F30F5E9CFB07F93B" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[1083,1212,1684,1708]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="algonquin">
|
||
<emphasis id="B95A581FFFE4FFA0F30F5E9CFB07F93B" box="[1083,1212,1684,1708]" italics="true" pageId="25" pageNumber="26">D. algonquin</emphasis>
|
||
</taxonomicName>
|
||
(ASG 32);
|
||
<emphasis id="B95A581FFFE4FFA0F7EA5EBBFF4CF95D" bold="true" box="[222,247,1714,1738]" pageId="25" pageNumber="26">D.</emphasis>
|
||
<taxonomicName id="4C2EFF8EFFE4FFA0F6345EBAFED7F95D" authority="Grimaldi, 2024" authorityName="Grimaldi" authorityYear="2024" box="[256,364,1714,1738]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="chibcha" status="n. sp.">
|
||
<emphasis id="B95A581FFFE4FFA0F6345EBAFED7F95D" box="[256,364,1714,1738]" italics="true" pageId="25" pageNumber="26">D. chibcha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A269E564FFE4FFA0F64E5EBBFE08F95E" box="[378,435,1714,1737]" pageId="25" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
|
||
(ASG 05), with detail of aedeagal valves of ASG 05 (above), and ASG 08 (below). All to the same scale except details of
|
||
<taxonomicName id="4C2EFF8EFFE4FFA0F5795ED9FD0EF970" authorityName="Grimaldi" authorityYear="2024" box="[589,693,1743,1767]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="chibcha">
|
||
<emphasis id="B95A581FFFE4FFA0F5795ED9FD0EF970" box="[589,693,1743,1767]" italics="true" pageId="25" pageNumber="26">D. chibcha</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF51D485FFE7FFA3F7585DBCFE7CF9B4" ID-DOI="http://doi.org/10.5281/zenodo.12171477" ID-Zenodo-Dep="12171477" httpUri="https://zenodo.org/record/12171477/files/figure.png" pageId="26" pageNumber="27" startId="26.[108,149,1461,1482]" targetBox="[191,1202,241,1419]" targetPageId="26" targetType="figure">
|
||
<paragraph id="8B91840DFFE7FFA3F7585DBCFE7CF9B4" blockId="26.[108,1213,1459,1571]" pageId="26" pageNumber="27">
|
||
FIG. 15. Hypandrium, aedeagus and other appendages of representative species of the
|
||
<taxonomicName id="4C2EFF8EFFE7FFA3F4F35DBDFBD3FA5C" authority="subgroup" authorityName="SUBGROUP" box="[967,1128,1459,1484]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="12" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B95A581FFFE7FFA3F4F35DBDFBB8FA5B" box="[967,1027,1460,1484]" italics="true" pageId="26" pageNumber="27">affinis</emphasis>
|
||
subgroup
|
||
</taxonomicName>
|
||
(ventral views).
|
||
<emphasis id="B95A581FFFE7FFA3F78F5DD8FF68FA7E" bold="true" box="[187,211,1489,1513]" pageId="26" pageNumber="27">A.</emphasis>
|
||
<taxonomicName id="4C2EFF8EFFE7FFA3F7EF5DDBFEC3FA7E" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[219,376,1490,1513]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="narragansett">
|
||
<emphasis id="B95A581FFFE7FFA3F7EF5DDBFEC3FA7E" box="[219,376,1490,1513]" italics="true" pageId="26" pageNumber="27">D. narragansett</emphasis>
|
||
</taxonomicName>
|
||
(ASG 10: paralectotype, Massachusetts);
|
||
<emphasis id="B95A581FFFE7FFA3F4105DD8FC80FA7E" bold="true" box="[804,827,1489,1513]" pageId="26" pageNumber="27">B.</emphasis>
|
||
<taxonomicName id="4C2EFF8EFFE7FFA3F4775DDBFC1CFA7E" authorityName="Grimaldi" authorityYear="2024" box="[835,935,1489,1513]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="olmeca">
|
||
<emphasis id="B95A581FFFE7FFA3F4775DDBFC1CFA7E" box="[835,935,1489,1513]" italics="true" pageId="26" pageNumber="27">D. olmeca</emphasis>
|
||
</taxonomicName>
|
||
(ASG 25: holotype);
|
||
<emphasis id="B95A581FFFE7FFA3F3B15DD8FB26FA7E" bold="true" box="[1157,1181,1489,1513]" pageId="26" pageNumber="27">C.</emphasis>
|
||
<taxonomicName id="4C2EFF8EFFE7FFA3F3915DDBFF16F991" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="tolteca">
|
||
<emphasis id="B95A581FFFE7FFA3F3915DDBFF16F991" italics="true" pageId="26" pageNumber="27">D. tolteca</emphasis>
|
||
</taxonomicName>
|
||
(ASG 23);
|
||
<emphasis id="B95A581FFFE7FFA3F6105DE7FE87F991" bold="true" box="[292,316,1518,1542]" pageId="26" pageNumber="27">D.</emphasis>
|
||
<taxonomicName id="4C2EFF8EFFE7FFA3F6705DE6FE59F991" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[324,482,1519,1542]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="narragansett">
|
||
<emphasis id="B95A581FFFE7FFA3F6705DE6FE59F991" box="[324,482,1519,1542]" italics="true" pageId="26" pageNumber="27">D. narragansett</emphasis>
|
||
</taxonomicName>
|
||
(ASG: Virginia);
|
||
<emphasis id="B95A581FFFE7FFA3F5AD5DE7FD15F991" bold="true" box="[665,686,1518,1542]" pageId="26" pageNumber="27">E.</emphasis>
|
||
<taxonomicName id="4C2EFF8EFFE7FFA3F5825DE6FCA1F991" authorityName="Grimaldi" authorityYear="2024" box="[694,794,1518,1542]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="olmeca">
|
||
<emphasis id="B95A581FFFE7FFA3F5825DE6FCA1F991" box="[694,794,1518,1542]" italics="true" pageId="26" pageNumber="27">D. olmeca</emphasis>
|
||
</taxonomicName>
|
||
(ASG 25: holotype, detail of B). A–C to same scale; D and E to same scale
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C334D786FFE6FFA5F7A85995FD18FDE6" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="8B91840DFFE6FFA2F7A85995FDEDFD76" blockId="27.[108,1214,225,1745]" pageId="27" pageNumber="28">
|
||
REDESCRIPTION: Coloration: Frons in
|
||
<specimenCount id="9D284F84FFE6FFA2F5535992FDC4FE21" box="[615,639,411,438]" count="1" pageId="27" pageNumber="28" type="male">♂</specimenCount>
|
||
silvery in frontal view, especially frontal vittae (fronto-orbital plates and ocellar triangle slightly less so); in dorsal view of
|
||
<specimenCount id="9D284F84FFE6FFA2F33B59C8FB9CFE4B" box="[1039,1063,449,476]" count="1" pageId="27" pageNumber="28" type="male">♂</specimenCount>
|
||
this silvery shagreenation diminished;
|
||
<specimenCount id="9D284F84FFE6FFA2F69F59EFFE78FD96" box="[427,451,486,513]" count="1" pageId="27" pageNumber="28" type="female">♀</specimenCount>
|
||
frons pollinose, slightly olive but not silvery, ocellar triangle and pair of spots at vertex lateral to ocelli without pollinosity; portion of frons near ptilinal suture dull, dark yellow. Antenna with pedicel light brown, flagellomere 1 darker brown; face, cheeks, and palps dull, tan; clypeus light brown. Scutum, scutellum, postpronotal lobe very light brown, dull, with dusting of pruinescence; notopleural area, anepisternum and anepimeron slightly darker brown, katepisternum lighter. Legs and halter knob dark yellowish tan; abdomen uniformly brown in both sexes, darker in
|
||
<specimenCount id="9D284F84FFE6FFA2F5015ACFFDF6FD76" box="[565,589,710,737]" count="1" pageId="27" pageNumber="28" type="male">♂</specimenCount>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B91840DFFE6FFA2F7A85AE5FDEDFB51" blockId="27.[108,1214,225,1745]" pageId="27" pageNumber="28">
|
||
Head: Arista with 4 dorsal, 2 ventral branches, plus terminal fork; pedicel with 4 larger setae. HD/HW 0.74 (mean of
|
||
<specimenCount id="9D284F84FFE6FFA2F6E45B18FE4DFCBB" box="[464,502,785,812]" count="4" pageId="27" pageNumber="28" type="male">4♂</specimenCount>
|
||
). Anterior reclinate orbital seta slightly posterolateral to the proclinate orbital; posterior reclinate slightly closer to proclinate than to inner vertical seta; proclinate 1.65× length of anterior reclinate, posterior reclinate 1.14× length of anterior reclinate. Ipsilateral vertical setae close together, sockets separated by distance ~2× their diameter; inner vertical in line with proclinate and posterior reclinate; vertical setae long, IV/OV 0.96. Ocellar seta socket on tangent between median and posterolateral ocelli; postocellars of medium length, parallel to convergent, pointing backward, shorter than ocellars (OC/POC 1.30); ~6 small setulae in ocellar triangle. Frons with 2–8 minute setulae near anterior margin. FL/LFW 0.94, UFW/LFW 1.47. Frontal W-index 2.85; carina very small, thin, short (half the length of face); vibrissa long, 1st genal seta small, GS1/VL 0.56, gena with 5–6 setae, decreasing in length posteriad. Cheek fairly narrow, ED/CD 7.5. Palp with 1 long preapical seta. Eye broadly oval in lateral view, EW/ED 0.81.
|
||
</paragraph>
|
||
<paragraph id="8B91840DFFE6FFA2F7A95CD8FD22F946" blockId="27.[108,1214,225,1745]" pageId="27" pageNumber="28">
|
||
Thorax: Length
|
||
<quantity id="4CD629E8FFE6FFA2F6515CD8FE6AFB7B" box="[357,465,1233,1260]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="27" pageNumber="28" unit="mm" value="0.88">0.88 mm</quantity>
|
||
(mean of
|
||
<specimenCount id="9D284F84FFE6FFA2F5655CD8FDCDFB7B" box="[593,630,1233,1260]" count="4" pageId="27" pageNumber="28" type="male">4♂</specimenCount>
|
||
). Acrostichals in 6 even rows between anterior dorsocentrals, lengths slightly increasing posteriad; acrostichals in front of scutellum or anterior dorsocentrals not enlarged. Anterior dorsocentrals 0.72× length of posterior ones; distance between ipsilateral dorsocentrals less than that between contralateral ones. Postpronotum with 2 strong setae, h-index 0.77; 2 strong notopleural setae near notopleural suture, plus 1 dorsally, another, short postsutural; katepisternum with 2 large setae, posterior one about 2× the size of other (S-index 0.57), sclerite with 5–7 small setulae. Anterior scutellar setae convergent, posterior ones crossing for about 0.2× their length, both pairs approximately equal in length (scut-index 0.96). Legs: Profemur with ventral row of 3–4 long setae on distal half (lengths approximately equal to femur width); mid and hind tibiae each with stout, ventroapical seta, thinner dorsal-preapical seta;
|
||
<specimenCount id="9D284F84FFE6FFA2F4995E4FFC7EF9F6" box="[941,965,1606,1633]" count="1" pageId="27" pageNumber="28" type="male">♂</specimenCount>
|
||
protarsus ta1 1.3× length of ta2; ta1 with 4–5 teeth, ta2 with 1; teeth stout, touching, length 1.3× tarsomere width. Wing of moderate length and width, ThL/WL 0.44, WL/WW 2.28, C-index 2.65, hb-index 2.49, 4V-index 2.23, 5X-index 2.15.
|
||
</paragraph>
|
||
<paragraph id="8B91840DFFE1FFA5F7A858E8FD18FDE6" blockId="28.[108,1214,225,1745]" pageId="28" pageNumber="29">Abdomen: Male terminalia: epandrium height slightly greater than width; cercus relatively narrow, with narrowed ventral lobe; outer lobes of ventral epandrial lobe relatively short (tips reaching to level of about midway to surstylus); margin of inner lobe of ventral epandrial lobe not well defined from surstylus, base of inner lobe with furrows; surstylus with row of 9–10 prensisetae; aedeagus and valves slightly shorter than (0.9× the length of) postgonites; gonites with faint elbow near middle of lateral margin. Aedeagal valves without ornamentation; membrane of aedeagus with very fine microtrichia. Hypandrium length 1.3× the width. Female terminalia: spermatheca distinct, width 1.5× the height; sleeve very large, funnel shaped, with fine annulations, extended into capsule 0.9× the capsule height, with small apical indentation protruding into end of sleeve. Oviscapt of moderate depth in lateral view, apex slightly narrowed; with ~13 ovisensilla pegs along margins.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334D786FFE1FFA5F7A95A74FB14FCC6" pageId="28" pageNumber="29" type="materials_examined">
|
||
<paragraph id="8B91840DFFE1FFA5F7A95A74FF4FFC91" blockId="28.[108,1214,225,1745]" pageId="28" pageNumber="29">
|
||
<materialsCitation id="3B468E50FFE1FFA5F7A95A74FF4FFC91" collectionCode="USNM" collectorName="A. H. Sturtevant Collection" pageId="28" pageNumber="29" specimenCount="26" specimenCount-female="15" specimenCount-male="11" typeStatus="lectotype">
|
||
TYPE:
|
||
<typeStatus id="54953AAFFFE1FFA5F7D15A74FEE0FD00" box="[229,347,637,663]" pageId="28" pageNumber="29" type="lectotype">Lectotype</typeStatus>
|
||
,
|
||
<specimenCount id="9D284F84FFE1FFA5F6535A72FEC4FD01" box="[359,383,635,662]" count="1" pageId="28" pageNumber="29" type="male">♂</specimenCount>
|
||
, selected by myself from a series of
|
||
<specimenCount id="9D284F84FFE1FFA5F4135A75FCF6FD01" box="[807,845,635,662]" count="9" pageId="28" pageNumber="29" type="male">9♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D284F84FFE1FFA5F4685A75FC2BFD01" box="[860,912,635,662]" count="14" pageId="28" pageNumber="29" type="female">14♀</specimenCount>
|
||
specimens, all labeled as: “WoodsHole, Mass [printed]/stock 25 [written]/
|
||
<collectorName id="26DBE1DBFFE1FFA5F5835AA8FC40FD2C" box="[695,1019,673,699]" pageId="28" pageNumber="29">A.H. Sturtevant Collection</collectorName>
|
||
, 1970 [printed.” Steyskal did not apply a note to any specimen as the type.
|
||
<specimenCount id="9D284F84FFE1FFA5F42C5ACEFCFDFD76" box="[792,838,710,737]" count="1" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D284F84FFE1FFA5F4625ACEFCC7FD76" box="[854,892,710,737]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
|
||
<typeStatus id="54953AAFFFE1FFA5F4B75ACEFB99FD76" box="[899,1058,711,737]" pageId="28" pageNumber="29" type="paralectotype">paralectotype</typeStatus>
|
||
dissected. In the
|
||
<collectionCode id="ED3F1CC8FFE1FFA5F7AE5AE5FF4BFC91" box="[154,240,748,774]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29" type="Museum">USNM</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B91840DFFE1FFA5F7A95B18FB14FCC6" blockId="28.[108,1214,225,1745]" pageId="28" pageNumber="29">
|
||
SPECIMENS EXAMINED: Besides the
|
||
<typeStatus id="54953AAFFFE1FFA5F5665B18FD05FCBC" box="[594,702,785,811]" pageId="28" pageNumber="29" type="lectotype">lectotype</typeStatus>
|
||
and the
|
||
<typeStatus id="54953AAFFFE1FFA5F4055B18FC69FCBC" box="[817,978,785,811]" pageId="28" pageNumber="29" type="paralectotype">paralectotype</typeStatus>
|
||
series (above), the
|
||
<typeStatus id="54953AAFFFE1FFA5F7585B3EFF68FCC6" box="[108,211,823,849]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F7C85B3EFEDBFCC6" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[252,352,823,849]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="seminole">
|
||
<emphasis id="B95A581FFFE1FFA5F7C85B3EFEDBFCC6" box="[252,352,823,849]" italics="true" pageId="28" pageNumber="29">seminole</emphasis>
|
||
</taxonomicName>
|
||
and several specimens identified as
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F43A5B3EFCC9FCC6" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[782,882,823,849]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="seminole">
|
||
<emphasis id="B95A581FFFE1FFA5F43A5B3EFCC9FCC6" box="[782,882,823,849]" italics="true" pageId="28" pageNumber="29">seminole</emphasis>
|
||
</taxonomicName>
|
||
(see in Comments below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334D786FFE1FFA5F7A85B55FE62FA31" pageId="28" pageNumber="29" type="distribution">
|
||
<paragraph id="8B91840DFFE1FFA5F7A85B55FC18FB36" blockId="28.[108,1214,225,1745]" pageId="28" pageNumber="29">
|
||
DISTRIBUTION: Currently a very rare species that historically extends from southern
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F3615B55FB07FCE1" box="[1109,1212,860,886]" country="United States of America" name="Alabama" pageId="28" pageNumber="29">Alabama</collectingRegion>
|
||
(Hartsell, Kushla),
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F6745B88FE05FC0C" box="[320,446,897,923]" country="United States of America" name="Mississippi" pageId="28" pageNumber="29">Mississippi</collectingRegion>
|
||
(Corinth), and mid-Florida (St. Petersburg) north to
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F32D5B88FB07FC0C" box="[1049,1212,897,923]" country="United States of America" name="Massachusetts" pageId="28" pageNumber="29">Massachusetts</collectingRegion>
|
||
(Amherst, Woods Hole), northern
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F6CB5BAEFDD6FC56" box="[511,621,935,961]" country="United States of America" name="Michigan" pageId="28" pageNumber="29">Michigan</collectingRegion>
|
||
(University of
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F42E5BAEFC33FC56" box="[794,904,935,961]" country="United States of America" name="Michigan" pageId="28" pageNumber="29">Michigan</collectingRegion>
|
||
Biological Station), and in the west to
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F7C45BC5FEE7FC71" box="[240,348,972,998]" country="United States of America" name="Nebraska" pageId="28" pageNumber="29">Nebraska</collectingRegion>
|
||
(Lincoln, Chadron St. Forest),
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F58B5BC5FC9EFC71" box="[703,805,972,998]" country="United States of America" name="Missouri" pageId="28" pageNumber="29">Missouri</collectingRegion>
|
||
(St. Louis) (
|
||
<bibRefCitation id="EFBFF9FCFFE1FFA5F4855BC5FBFBFC70" author="Miller, D. D." box="[945,1088,972,999]" pageId="28" pageNumber="29" pagination="52 - 70" refId="ref19748" refString="Miller, D. D. 1958. Geographical distribution of the American Drosophila affinis subgroup species. American Midland Naturalist 60: 52 - 70." type="journal article" year="1958">Miller, 1958</bibRefCitation>
|
||
), as well as
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F7585BF8FF7EFB9C" box="[108,197,1009,1035]" country="United States of America" name="Indiana" pageId="28" pageNumber="29">Indiana</collectingRegion>
|
||
(locality not specified) and
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F5325BFBFDFDFB9B" box="[518,582,1010,1036]" country="United States of America" name="Texas" pageId="28" pageNumber="29">Texas</collectingRegion>
|
||
(Bastrop St. Park) (
|
||
<bibRefCitation id="EFBFF9FCFFE1FFA5F41C5BF8FC41FB9C" author="Barrio, E. & A. Latorre & A. Moya & F. J. Ayala" box="[808,1018,1009,1036]" pageId="28" pageNumber="29" pagination="621 - 635" refId="ref18308" refString="Barrio, E., A. Latorre, A. Moya, and F. J. Ayala. 1992. Phylogenetic reconstruction of the Drosophila obscura group, on the basis of mitochondrial DNA. Molecular Biology and Evolution 9: 621 - 635." type="journal article" year="1992">Barrio et al., 1992</bibRefCitation>
|
||
). The record and figures in M. Miller et al. (2017) of
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F6CB5C1EFD15FBA7" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[511,686,1047,1072]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="narragansett">
|
||
<emphasis id="B95A581FFFE1FFA5F6CB5C1EFD15FBA7" box="[511,686,1047,1072]" italics="true" pageId="28" pageNumber="29">D. narragansett</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F5C05C1EFC86FBA6" box="[756,829,1047,1073]" country="United States of America" name="Maine" pageId="28" pageNumber="29">Maine</collectingRegion>
|
||
are incorrect. These are based on a series of specimens in the AMNH: “SE Guarette
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F5F45C35FCB0FBC1" box="[704,779,1084,1110]" country="United States of America" name="Maine" pageId="28" pageNumber="29">Maine</collectingRegion>
|
||
,
|
||
<date id="FF90A2CDFFE1FFA5F42D5C35FC3EFBC1" box="[793,901,1084,1110]" pageId="28" pageNumber="29" value="1948-07">July 1948</date>
|
||
/ M.R. Wheeler collector,” which were misidentified by Marshall Wheeler as
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F5895C6BFCF7FBEC" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[701,844,1122,1147]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="narragansett">
|
||
<emphasis id="B95A581FFFE1FFA5F5895C6BFCF7FBEC" box="[701,844,1122,1147]" italics="true" pageId="28" pageNumber="29">narragansett</emphasis>
|
||
</taxonomicName>
|
||
. The specimens are actually
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F3955C6BFF67FB36" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="athabasca">
|
||
<emphasis id="B95A581FFFE1FFA5F3955C6BFF67FB36" italics="true" pageId="28" pageNumber="29">D. athabasca</emphasis>
|
||
</taxonomicName>
|
||
, with a spermathecal capsule that is especially flat and broad.
|
||
</paragraph>
|
||
<paragraph id="8B91840DFFE1FFA5F7A95CA4FE62FA31" blockId="28.[108,1214,225,1745]" pageId="28" pageNumber="29">
|
||
In a study of abundances of
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F6CF5CA4FDDCFB51" box="[507,615,1196,1222]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B95A581FFFE1FFA5F6CF5CA4FDDCFB51" box="[507,615,1196,1222]" italics="true" pageId="28" pageNumber="29">D. affinis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F54C5CA4FCA9FB51" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[632,786,1196,1222]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="algonquin">
|
||
<emphasis id="B95A581FFFE1FFA5F54C5CA4FCA9FB51" box="[632,786,1196,1222]" italics="true" pageId="28" pageNumber="29">D. algonquin</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F4165CA4FC07FB51" box="[802,956,1196,1222]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="athabasca">
|
||
<emphasis id="B95A581FFFE1FFA5F4165CA4FC07FB51" box="[802,956,1196,1222]" italics="true" pageId="28" pageNumber="29">D. athabasca</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F3375CA4FEABFB7B" authority="(Miller, 1958)" baseAuthorityName="Miller" baseAuthorityYear="1958" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="narragansett">
|
||
<emphasis id="B95A581FFFE1FFA5F3375CA4FB06FB51" box="[1027,1213,1197,1222]" italics="true" pageId="28" pageNumber="29">D. narragansett</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBFF9FCFFE1FFA5F7405CD8FEB3FB7B" author="Miller, D. D." box="[116,264,1233,1260]" pageId="28" pageNumber="29" pagination="52 - 70" refId="ref19748" refString="Miller, D. D. 1958. Geographical distribution of the American Drosophila affinis subgroup species. American Midland Naturalist 60: 52 - 70." type="journal article" year="1958">Miller, 1958</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the last one was not found at most of the localities, and when present was less than 1% of the abundance of these four species, except at two sites: Corinth,
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F4CF5CFEFBC6FA86" box="[1019,1149,1271,1297]" country="United States of America" name="Mississippi" pageId="28" pageNumber="29">Mississippi</collectingRegion>
|
||
(3%) and Hartshell,
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F62E5D15FE38FAA1" box="[282,387,1308,1334]" country="United States of America" name="Alabama" pageId="28" pageNumber="29">Alabama</collectingRegion>
|
||
(7%). Astonishingly,
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F5495D14FCB4FAA1" baseAuthorityName="Miller" baseAuthorityYear="1958" box="[637,783,1309,1334]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="narragansett">
|
||
<emphasis id="B95A581FFFE1FFA5F5495D14FCB4FAA1" box="[637,783,1309,1334]" italics="true" pageId="28" pageNumber="29">narragansett</emphasis>
|
||
</taxonomicName>
|
||
has to my knowledge been collected only once within the past 60 years, despite its broad range: a male swept from over compost in
|
||
<date id="FF90A2CDFFE1FFA5F7585D6EFF5BFA16" box="[108,224,1383,1409]" pageId="28" pageNumber="29" value="2017-05">
|
||
May
|
||
<quantity id="4CD629E8FFE1FFA5F79C5D6EFF5BFA16" box="[168,224,1383,1409]" metricMagnitude="1" metricUnit="m" metricValue="5.12318" pageId="28" pageNumber="29" unit="in" value="2017.0">2017</quantity>
|
||
</date>
|
||
in Rochester
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F6B15D6EFE40FA16" box="[389,507,1383,1409]" country="United States of America" name="New York" pageId="28" pageNumber="29">New York</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFBFF9FCFFE1FFA5F53E5D6EFD47FA16" author="Werner, T. & T. Steenwinkel & J. Jaenike" box="[522,764,1383,1409]" pageId="28" pageNumber="29" refId="ref20464" refString="Werner, T., T. Steenwinkel, and J. Jaenike. 2020 a. The encyclopedia of North American Drosophilids, vol. 1. Drosophilids of the Midwest and Northeast. Houghton, MI: Pelt and Opie Library, Michigan Technical University." type="book" year="2020">Werner et al., 2020a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFBFF9FCFFE1FFA5F43D5D6EFCEDFA16" author="Werner, T. & T. Steenwinkel & J. Jaenike" box="[777,854,1383,1409]" pageId="28" pageNumber="29" refId="ref20512" refString="Werner, T., T. Steenwinkel, and J. Jaenike. 2020 b. The encyclopedia of North American Drosophilids, vol. 2. Drosophilids of the Southeast. Houghton, MI: Pelt and Opie Library, Michigan Technical University." type="book" year="2020">2020b</bibRefCitation>
|
||
). I have never collected it, nor have many other drosophilists.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C334D786FFE1FFA4F7A95DB8FC9AFCBC" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="discussion">
|
||
<paragraph id="8B91840DFFE1FFA4F7A95DB8FB8AFEFC" blockId="28.[108,1214,225,1745]" lastBlockId="29.[108,1213,225,812]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
|
||
COMMENTS: The
|
||
<typeStatus id="54953AAFFFE1FFA5F6565DB8FE7CFA5C" box="[354,455,1457,1483]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F6D95DB8FDF5FA5C" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[493,590,1457,1483]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="seminole">
|
||
<emphasis id="B95A581FFFE1FFA5F6D95DB8FDF5FA5C" box="[493,590,1457,1483]" italics="true" pageId="28" pageNumber="29">seminole</emphasis>
|
||
</taxonomicName>
|
||
, a male, is labeled: “Kushla, Ala[bama], My.14.22, oak trunk [in Sturtevant’s writing] / A.H. Sturtevant collection, 1970 [printed].” In USNM. This is the sole specimen from the type locality (Kushla), it has a penciled note by Steyskal (in his writing)
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F7595E28FF64F9AC" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[109,223,1569,1595]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="seminole">“seminole</taxonomicName>
|
||
type”; it was labeled
|
||
<date id="FF90A2CDFFE1FFA5F6EA5E28FDC2F9AC" box="[478,633,1569,1596]" pageId="28" pageNumber="29" value="2023-06-12">12 June 2023</date>
|
||
by myself as the
|
||
<typeStatus id="54953AAFFFE1FFA5F47E5E28FC0AF9AC" box="[842,945,1569,1595]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
|
||
; glue on the tip of the abdomen prevented it from being dissected for the genitalia. The glue also embedded the foretarsi, but fortunately it is clear and four long teeth are visible on protarsomere-1. The type of
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F3975E64FF75F93C" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="seminole">
|
||
<emphasis id="B95A581FFFE1FFA5F3975E64FF75F93C" italics="true" pageId="28" pageNumber="29">D. seminole</emphasis>
|
||
</taxonomicName>
|
||
is darker than specimens in the type series of
|
||
<taxonomicName id="4C2EFF8EFFE1FFA5F5C55E9BFC1DF93C" baseAuthorityName="Miller" baseAuthorityYear="1958" box="[753,934,1682,1708]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="31" phylum="Arthropoda" rank="species" species="narragansett">
|
||
D.
|
||
<emphasis id="B95A581FFFE1FFA5F4235E9BFC1DF93C" box="[791,934,1682,1707]" italics="true" pageId="28" pageNumber="29">narragansett</emphasis>
|
||
</taxonomicName>
|
||
and the specimen from
|
||
<collectingRegion id="49EA4AEFFFE1FFA5F75A5EBEFF74F946" box="[110,207,1719,1745]" country="United States of America" name="Virginia" pageId="28" pageNumber="29">Virginia</collectingRegion>
|
||
, but in all other respects it is identical to those others, including measured proportions of body structures and setae. The putative differences between
|
||
<taxonomicName id="4C2EFF8EFFE0FFA4F47858EBFC40FF6C" baseAuthorityName="Miller" baseAuthorityYear="1958" box="[844,1019,226,251]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="narragansett">
|
||
<emphasis id="B95A581FFFE0FFA4F47858EBFC40FF6C" box="[844,1019,226,251]" italics="true" pageId="29" pageNumber="30">D. narragansett</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2EFF8EFFE0FFA4F30D58EBFB07FF6C" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[1081,1212,225,251]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="seminole">
|
||
<emphasis id="B95A581FFFE0FFA4F30D58EBFB07FF6C" box="[1081,1212,225,251]" italics="true" pageId="29" pageNumber="30">D. seminole</emphasis>
|
||
</taxonomicName>
|
||
in the silvery frons, reported by Sturtevant and Dobzhansky (1936) and cited by
|
||
<bibRefCitation id="EFBFF9FCFFE0FFA4F319590EFEBDFED1" author="Sulerud, R. L. & D. D. Miller" pageId="29" pageNumber="30" pagination="446 - 474" refId="ref20220" refString="Sulerud, R. L., and D. D. Miller. 1966. A study of key characteristics for distinguishing several Drosophila affinis subgroup species, with a description of a new related species. American Midland Naturalist 75: 446 - 474." type="journal article" year="1966">Sulerud and Miller (1966)</bibRefCitation>
|
||
, are incorrect and one basis for confusion that
|
||
<taxonomicName id="4C2EFF8EFFE0FFA4F4155924FC1FFED1" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[801,932,300,326]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="seminole">
|
||
<emphasis id="B95A581FFFE0FFA4F4155924FC1FFED1" box="[801,932,300,326]" italics="true" pageId="29" pageNumber="30">D. seminole</emphasis>
|
||
</taxonomicName>
|
||
could be a separate species. The silvery male frons may not be apparent in specimens preserved in alcohol.
|
||
</paragraph>
|
||
<paragraph id="8B91840DFFE0FFA4F7A8597EFC9AFCBC" blockId="29.[108,1213,225,812]" pageId="29" pageNumber="30">
|
||
Two other specimens in the USNM collection identified as
|
||
<taxonomicName id="4C2EFF8EFFE0FFA4F454597EFC5CFE06" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[864,999,375,401]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="seminole">
|
||
<emphasis id="B95A581FFFE0FFA4F454597EFC5CFE06" box="[864,999,375,401]" italics="true" pageId="29" pageNumber="30">D. seminole</emphasis>
|
||
</taxonomicName>
|
||
are the following: one mentioned in the original publication (Sturtevant and Dobzhansky, 1936) labeled as: “Whistler Ala[bama] Oc[t.].20.24 [written in Sturtevant’s hand] / A.H. Sturtevant Collection 1970 [printed]/
|
||
<taxonomicName id="4C2EFF8EFFE0FFA4F62A59EEFDBEFD96" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[286,517,487,513]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="seminole">
|
||
<emphasis id="B95A581FFFE0FFA4F62A59EEFDBEFD96" box="[286,517,487,513]" italics="true" pageId="29" pageNumber="30">Drosophila seminole</emphasis>
|
||
</taxonomicName>
|
||
[written].” This specimen, a male with the genitalia everted, is not
|
||
<taxonomicName id="4C2EFF8EFFE0FFA4F7825A04FE80FDB1" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[182,315,524,550]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="seminole">
|
||
<emphasis id="B95A581FFFE0FFA4F7825A04FE80FDB1" box="[182,315,524,550]" italics="true" pageId="29" pageNumber="30">D. seminole</emphasis>
|
||
</taxonomicName>
|
||
(i.e.,
|
||
<taxonomicName id="4C2EFF8EFFE0FFA4F6485A04FD8BFDB1" baseAuthorityName="Miller" baseAuthorityYear="1958" box="[380,560,525,550]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="narragansett">
|
||
<emphasis id="B95A581FFFE0FFA4F6485A04FD8BFDB1" box="[380,560,525,550]" italics="true" pageId="29" pageNumber="30">D. narragansett</emphasis>
|
||
</taxonomicName>
|
||
); it lacks the distinctive silvery frons. A third specimen is a male with a silvery frons, labeled: “Mountain L[ake], Va., 22-VII-’
|
||
<specimenCount id="9D284F84FFE0FFA4F49E5A38FC4FFDDB" box="[938,1012,561,588]" count="40" pageId="29" pageNumber="30" type="generic">40 W.</specimenCount>
|
||
H. #2, L.J. & M.J. Milne /
|
||
<taxonomicName id="4C2EFF8EFFE0FFA4F7F85A5EFE0DFDE6" authorityName="Sturtevant and Dobzhansky" authorityYear="1936" box="[204,438,599,625]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="seminole">
|
||
<emphasis id="B95A581FFFE0FFA4F7F85A5EFE0DFDE6" box="[204,438,599,625]" italics="true" pageId="29" pageNumber="30">Drosophila seminole</emphasis>
|
||
</taxonomicName>
|
||
det. Steyskal ‘[19]44.” This specimen is
|
||
<taxonomicName id="4C2EFF8EFFE0FFA4F4A55A5EFBFEFDE7" baseAuthorityName="Miller" baseAuthorityYear="1958" box="[913,1093,599,624]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="narragansett">
|
||
<emphasis id="B95A581FFFE0FFA4F4A55A5EFBFEFDE7" box="[913,1093,599,624]" italics="true" pageId="29" pageNumber="30">D. narragansett</emphasis>
|
||
</taxonomicName>
|
||
, although it has some interesting distinctions from other measured specimens of
|
||
<taxonomicName id="4C2EFF8EFFE0FFA4F4815A74FBD3FD01" baseAuthorityName="Miller" baseAuthorityYear="1958" box="[949,1128,637,662]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="narragansett">
|
||
<emphasis id="B95A581FFFE0FFA4F4815A74FBD3FD01" box="[949,1128,637,662]" italics="true" pageId="29" pageNumber="30">D. narragansett</emphasis>
|
||
</taxonomicName>
|
||
: upper seta of postpronotal lobe smaller than lower one (h-index 0.62, vs. 0.80–0.87 for others), smaller anterior scutellars (Scut-index 0.88, vs. 0.97–1.02 for others), C-index smaller (2.48, vs. 2.65–2.71), and hb-index smaller (2.18, vs. 2.42–2.75). The male genitalia of the
|
||
<typeStatus id="54953AAFFFE0FFA4F3745AE4FBC8FC90" box="[1088,1139,749,775]" pageId="29" pageNumber="30">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C2EFF8EFFE0FFA4F7BF5B1BFEFBFCBC" baseAuthorityName="Miller" baseAuthorityYear="1958" box="[139,320,786,811]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="narragansett">
|
||
<emphasis id="B95A581FFFE0FFA4F7BF5B1BFEFBFCBC" box="[139,320,786,811]" italics="true" pageId="29" pageNumber="30">D. narragansett</emphasis>
|
||
</taxonomicName>
|
||
and the
|
||
<collectingRegion id="49EA4AEFFFE0FFA4F69D5B18FDB2FCBC" box="[425,521,785,811]" country="United States of America" name="Virginia" pageId="29" pageNumber="30">Virginia</collectingRegion>
|
||
specimen are identical.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |