treatments-xml/data/C4/0A/B7/C40AB79FBC4459E2ADE9335BDF63398D.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1123.90141" ID-GBIF-Dataset="d6d98bd3-2433-44ff-b560-dffef5ebd40e" ID-Pensoft-Pub="1313-2970-1123-1" ID-Pensoft-UUID="5378DBF1DEED5922910377DB1774B3B4" ID-ZooBank="B80C686A001C4A729E8A7DED3FEE9515" ModsDocID="1313-2970-1123-1" checkinTime="1664474094714" checkinUser="pensoft" docAuthor="Schneider, Scott A. &amp; LaPolla, John S." docDate="2022" docId="C40AB79FBC4459E2ADE9335BDF63398D" docLanguage="en" docName="ZooKeys 1123: 1-30" docOrigin="ZooKeys 1123" docPubDate="2022-09-29" docSource="http://dx.doi.org/10.3897/zookeys.1123.90141" docTitle="Ripersiella illicians Schneider &amp; LaPolla 2022, sp. nov." docType="treatment" docUuid="7A49B97A-43A7-48D4-B8DF-8D9408E9A882" docUuidSource="ZooBank" docVersion="2" id="5378DBF1DEED5922910377DB1774B3B4" lastPageNumber="1" masterDocId="5378DBF1DEED5922910377DB1774B3B4" masterDocTitle="A Neotropical complex of Ripersiella species (Hemiptera, Coccomorpha, Rhizoecidae) collected from the nests of Acropyga ants (Hymenoptera, Formicidae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="1" updateTime="1664474713734" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A Neotropical complex of Ripersiella species (Hemiptera, Coccomorpha, Rhizoecidae) collected from the nests of Acropyga ants (Hymenoptera, Formicidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schneider, Scott A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9679-8437</mods:nameIdentifier>
<mods:affiliation>United States Department of Agriculture, Agricultural Research Service, Henry A. Wallace Beltsville Agricultural Research Center, Systematic Entomology Laboratory, 10300 Baltimore Avenue, Beltsville, MD, 20705, USA</mods:affiliation>
<mods:nameIdentifier type="email">scott.schneider@ars.usda.gov</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>LaPolla, John S.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7602-1527</mods:nameIdentifier>
<mods:affiliation>Department of Biological Sciences, Towson University, 8000 York Road, Towson, MD, 21252, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1123</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>30</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1123.90141</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1123.90141</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1123-1</mods:identifier>
<mods:identifier type="ZooBank">B80C686A001C4A729E8A7DED3FEE9515</mods:identifier>
<mods:identifier type="Pensoft-UUID">5378DBF1DEED5922910377DB1774B3B4</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:7A49B97A-43A7-48D4-B8DF-8D9408E9A882" httpUri="http://treatment.plazi.org/id/C40AB79FBC4459E2ADE9335BDF63398D" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/7A49B97A-43A7-48D4-B8DF-8D9408E9A882" authority="Schneider &amp; LaPolla" authorityName="Schneider &amp; LaPolla" authorityYear="2022" class="Insecta" family="Rhizoecidae" genus="Ripersiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ripersiella illicians" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="illicians" status="sp. nov.">Ripersiella illicians Schneider &amp; LaPolla</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Ripersiella illicians Schneider &amp; LaPolla sp. nov. Adult female, full body view, illustrated from holotype by T. Litwak (USDA ARS SEL), with edits by SAS." figureDoi="10.3897/zookeys.1123.90141.figure5" httpUri="https://binary.pensoft.net/fig/750176" pageId="0" pageNumber="1">Figures 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Ripersiella illicians Schneider &amp; LaPolla sp. nov. Scanning electron micrograph (SEM) by J. Mowery (USDA ARS SEL). Dorsal anterior view of adult female head and thorax, showing the absence of anterior ostioles and the presence of curled wax deposits and domed dermal micro-bumps in intersegmental areas." figureDoi="10.3897/zookeys.1123.90141.figure6" httpUri="https://binary.pensoft.net/fig/750177" pageId="0" pageNumber="1">, 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2019-06-10" collectorName="JS LaPolla, SA Schneider" country="Peru" latitude="-11.8833" location="Cocha Cashu Biological Station" longLatPrecision="7" longitude="-71.4" municipality="Manu National Park" specimenCount="1" specimenCount-adult="1" stateProvince="Madre de Dios" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry name="Peru">Peru</collectingCountry>
• 
<specimenCount type="adult">1 adult</specimenCount>
<specimenCount>♀</specimenCount>
; 
<collectingRegion country="Peru" name="Madre de Dios">Madre de Dios</collectingRegion>
, 
<collectingMunicipality>Manu National Park</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:C40AB79FBC4459E2ADE9335BDF63398D:18884711B08C833ECF09E1B2C95EC5F3" country="Peru" latitude="-11.8833" longLatPrecision="7" longitude="-71.4" municipality="Manu National Park" name="Cocha Cashu Biological Station" stateProvince="Madre de Dios">Cocha Cashu Biological Station</location>
, near trail marker 27:1150; 
<geoCoordinate degrees="11.8833" direction="south" orientation="latitude" precision="5" value="-11.8833">-11.8833</geoCoordinate>
, 
<geoCoordinate degrees="71.4000" direction="west" orientation="longitude" precision="5" value="-71.4">-71.4000</geoCoordinate>
; 
<collectingDate value="2019-06-10">10.vi.2019</collectingDate>
; 
<collectorName>JS LaPolla</collectorName>
, 
<collectorName>SA Schneider</collectorName>
leg.; upland forest, from large nest of 
<taxonomicName authorityName="Forel" authorityYear="1893" class="Insecta" family="Formicidae" genus="Acropyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acropyga goeldii" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="goeldii">
<emphasis italics="true" pageId="0" pageNumber="1">Acropyga goeldii</emphasis>
</taxonomicName>
(group) at base of tree; USNM (nest PER25-01: prep S0426E)
</materialsCitation>
. 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="JS LaPolla &amp; SA Schneider" country="Peru" latitude="-11.8833" location="Peru" longLatPrecision="7" longitude="-71.4" municipality="Manu National Park" specimenCount="3" specimenCount-adult="3" stateProvince="Madre de Dios" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry name="Peru">Peru</collectingCountry>
• 
<specimenCount type="adult">3 adult</specimenCount>
<specimenCount>♀♀</specimenCount>
; same data as holotype; USNM (nest PER25-01: preps S0426B,D,F)
</materialsCitation>
• 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="JS LaPolla &amp; SA Schneider" country="Peru" latitude="-11.8833" location="Cocha Cashu Biological Station" longLatPrecision="7" longitude="-71.4" municipality="Manu National Park" specimenCount="1" specimenCount-adult="1" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount type="adult">1 adult</specimenCount>
<specimenCount>♀</specimenCount>
; same data as holotype; UNMSM (nest PER25-01: prep S0426C)
</materialsCitation>
• 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="JS LaPolla &amp; SA Schneider" country="Peru" latitude="-11.8833" location="Cocha Cashu Biological Station" longLatPrecision="7" longitude="-71.4" municipality="Manu National Park" specimenCount="1" specimenCount-adult="1" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount type="adult">1 adult</specimenCount>
<specimenCount>♀</specimenCount>
; same data as holotype; UNAB (nest PER25-01: prep S0426G)
</materialsCitation>
• 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="JS LaPolla &amp; SA Schneider" country="Peru" latitude="-11.8833" location="Cocha Cashu Biological Station" longLatPrecision="7" longitude="-71.4" municipality="Manu National Park" specimenCount="1" specimenCount-adult="1" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount type="adult">1 adult</specimenCount>
<specimenCount>♀</specimenCount>
; same data as holotype; UKMNH (nest PER25-01: prep S0426A)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1123.90141.figure5" httpUri="https://binary.pensoft.net/fig/750176" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
<taxonomicName authorityName="Schneider &amp; LaPolla" authorityYear="2022" class="Insecta" family="Rhizoecidae" genus="Ripersiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ripersiella illicians" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="illicians">
<emphasis italics="true" pageId="0" pageNumber="1">Ripersiella illicians</emphasis>
</taxonomicName>
Schneider &amp; LaPolla sp. nov. Adult female, full body view, illustrated from 
<typeStatus>holotype</typeStatus>
by T. Litwak (USDA ARS SEL), with edits by SAS.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adult female (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 7). In life, body bright white to cream colored and visibly coated in powdery white wax.
</paragraph>
<caption doi="10.3897/zookeys.1123.90141.figure6" httpUri="https://binary.pensoft.net/fig/750177" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
<taxonomicName authorityName="Schneider &amp; LaPolla" authorityYear="2022" class="Insecta" family="Rhizoecidae" genus="Ripersiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ripersiella illicians" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="illicians">
<emphasis italics="true" pageId="0" pageNumber="1">Ripersiella illicians</emphasis>
</taxonomicName>
Schneider &amp; LaPolla sp. nov. Scanning electron micrograph (SEM) by J. Mowery (USDA ARS SEL). Dorsal anterior view of adult female head and thorax, showing the absence of anterior ostioles and the presence of curled wax deposits and domed dermal micro-bumps in intersegmental areas.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">General.</emphasis>
Mounted on microscope slide, body broadly oval and membranous, 0.70-0.83 (0.78) mm long, widest at metathorax and abdominal segments I-II, 0.46-0.61 (0.55) mm wide. Abdomen rounded and gently tapering toward posterior apex; abdominal segment VIII 148-196 (196) 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide at base. Anal lobes poorly developed with several flagellate setae on dorsum, 57-60 
<normalizedToken originalValue="μm">μm</normalizedToken>
long. Body setae flagellate, 15-27 
<normalizedToken originalValue="μm">μm</normalizedToken>
on head, 17-33 
<normalizedToken originalValue="μm">μm</normalizedToken>
on thorax, 15-40 
<normalizedToken originalValue="μm">μm</normalizedToken>
on abdominal segments. Trilocular pores abundant and distributed among body setae; multilocular disc pores absent; bitubular cerores absent; oral collar tubular ducts absent. Microtrichia apparently present only on dorsum of segments VI-VIII; rounded dermal micro-bumps present in intersegmental areas and surroundings of appendages. Internal genital organ sclerotized in some specimens of type series. Eyes absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Venter</emphasis>
.
</emphasis>
Cephalic plate present. Labium with 3 segments; 82 
<normalizedToken originalValue="μm">μm</normalizedToken>
long and 41 
<normalizedToken originalValue="μm">μm</normalizedToken>
at widest point. Antennae geniculate, 6-segmented, closely situated near midline on ventral submargin of head; overall length 130-151 (151) 
<normalizedToken originalValue="μm">μm</normalizedToken>
; length of segment I: 30-37 (37) 
<normalizedToken originalValue="μm">μm</normalizedToken>
; segment II: 14-18 (18) 
<normalizedToken originalValue="μm">μm</normalizedToken>
; segment III: 17-20 (20) 
<normalizedToken originalValue="μm">μm</normalizedToken>
; segment IV: 14-19 (19) 
<normalizedToken originalValue="μm">μm</normalizedToken>
; segment V: 13-17 (17) 
<normalizedToken originalValue="μm">μm</normalizedToken>
; segment VI: 39-40 (40) 
<normalizedToken originalValue="μm">μm</normalizedToken>
; apical segment with 4 falcate stout sensory setate; a few flagellate setae present on each antennal segment, 19-30 
<normalizedToken originalValue="μm">μm</normalizedToken>
long. Legs well developed; overall length of hind leg 274-290 (288) 
<normalizedToken originalValue="μm">μm</normalizedToken>
; length of hind coxa 43-44 (44) 
<normalizedToken originalValue="μm">μm</normalizedToken>
; length of hind trochanter + femur 102-108 (106) 
<normalizedToken originalValue="μm">μm</normalizedToken>
; length of hind tibia + tarsus 102-110 (110) 
<normalizedToken originalValue="μm">μm</normalizedToken>
; length of hind claw 26-31 (28) 
<normalizedToken originalValue="μm">μm</normalizedToken>
; each claw with stout setose digitule 1-2 
<normalizedToken originalValue="μm">μm</normalizedToken>
long; flagellate setae present on each segment approximately 23 
<normalizedToken originalValue="μm">μm</normalizedToken>
long. Single conical circulus present between abdominal segments III and IV, 22 
<normalizedToken originalValue="μm">μm</normalizedToken>
in diameter; inner margin of orifice crenulated or with rugose projections.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Dorsum</emphasis>
.
</emphasis>
Anal ring situated on dorsal surface separated from posterior body margin by approximately one-half diameter of ring, 59-61 
<normalizedToken originalValue="μm">μm</normalizedToken>
in diameter; with oval cells lacking spicules; bearing 3 pairs of setae about 50 
<normalizedToken originalValue="μm">μm</normalizedToken>
long. Posterior pair of ostioles present, diameter of orifice measured along longitudinal axis approximately 39 
<normalizedToken originalValue="μm">μm</normalizedToken>
; anterior pair of ostioles absent.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="informal synonyms">
<paragraph pageId="0" pageNumber="1">Informal synonyms.</paragraph>
<paragraph pageId="0" pageNumber="1">
Specimens of 
<taxonomicName lsidName="R. illicians" pageId="0" pageNumber="1" rank="species" species="illicians">
<emphasis italics="true" pageId="0" pageNumber="1">R. illicians</emphasis>
</taxonomicName>
have been previously referred to in the literature as &quot; 
<taxonomicName authorityName="Tinsley" authorityYear="1899" class="Insecta" family="Rhizoecidae" genus="Ripersiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ripersiella" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Ripersiella</emphasis>
</taxonomicName>
undescribed (i)&quot; (
<bibRefCitation DOI="https://doi.org/10.1007/s13744-022-00948-9" author="Schneider, SA" journalOrPublisher="Neotropical Entomology" pageId="0" pageNumber="1" pagination="641 - 647" refId="B25" refString="Schneider, SA, Sodano, J, LaPolla, JS, 2022. Distinguishing symbiotic partners of Acropyga ants from free-living soil inhabitants. Neotropical Entomology 51 (4): 641 - 647, DOI: https://doi.org/10.1007/s13744-022-00948-9" title="Distinguishing symbiotic partners of Acropyga ants from free-living soil inhabitants." url="https://doi.org/10.1007/s13744-022-00948-9" volume="51" year="2022">Schneider et al. 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The species epithet is an adjective formed from 
<emphasis italics="true" pageId="0" pageNumber="1">illicium</emphasis>
, meaning attraction or enticement, and its use alludes to the apparent tendency of 
<taxonomicName authorityName="Roger" authorityYear="1862" class="Insecta" family="Formicidae" genus="Acropyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acropyga" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Acropyga</emphasis>
</taxonomicName>
colonies to gain rhizoecid partners through horizontal acquisitions. Such colonies have been figuratively attracted away from their primary associates in 
<taxonomicName class="Insecta" family="Xenococcidae" higherTaxonomySource="GBIF" kingdom="Metazoa" lsidName="" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Xenococcidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Schneider &amp; LaPolla" authorityYear="2022" class="Insecta" family="Rhizoecidae" genus="Ripersiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ripersiella illicians" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="illicians">
<emphasis italics="true" pageId="0" pageNumber="1">Ripersiella illicians</emphasis>
</taxonomicName>
sp. nov. is similar to 
<taxonomicName lsidName="R. colombiensis" pageId="0" pageNumber="1" rank="species" species="colombiensis">
<emphasis italics="true" pageId="0" pageNumber="1">R. colombiensis</emphasis>
</taxonomicName>
as both species have a subcircular body shape, bearing one circulus, and both are lacking multilocular disc pores and an anterior pair of dorsal ostioles. However, 
<taxonomicName lsidName="R. illicians" pageId="0" pageNumber="1" rank="species" species="illicians">
<emphasis italics="true" pageId="0" pageNumber="1">R. illicians</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName lsidName="R. colombiensis" pageId="0" pageNumber="1" rank="species" species="colombiensis">
<emphasis italics="true" pageId="0" pageNumber="1">R. colombiensis</emphasis>
</taxonomicName>
as follows (character states for 
<taxonomicName lsidName="R. colombiensis" pageId="0" pageNumber="1" rank="species" species="colombiensis">
<emphasis italics="true" pageId="0" pageNumber="1">R. colombiensis</emphasis>
</taxonomicName>
are given in parentheses): having 6-segmented antennae (5-segmented antennae); having comparatively long body setae, ranging from 15-40 
<normalizedToken originalValue="μm">μm</normalizedToken>
(comparatively short, ranging from 15-25 
<normalizedToken originalValue="μm">μm</normalizedToken>
); and having anal lobe setae that are distinctly longer than body setae, 57-60 
<normalizedToken originalValue="μm">μm</normalizedToken>
(similar in length to body setae, 18-28 
<normalizedToken originalValue="μm">μm</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Schneider &amp; LaPolla" authorityYear="2022" class="Insecta" family="Rhizoecidae" genus="Ripersiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ripersiella illicians" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="illicians">
<emphasis italics="true" pageId="0" pageNumber="1">Ripersiella illicians</emphasis>
</taxonomicName>
was discovered from a large nest of 
<taxonomicName authorityName="Forel" authorityYear="1893" class="Insecta" family="Formicidae" genus="Acropyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acropyga goeldii" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="goeldii">
<emphasis italics="true" pageId="0" pageNumber="1">Acropyga goeldii</emphasis>
</taxonomicName>
(group). Root mealybugs were abundant in the nest, and wax could be seen on their body using a hand lens. Their direct association was confirmed through observation of the colony using a nest-box, as described by 
<bibRefCitation DOI="https://doi.org/10.1007/s13744-022-00948-9" author="Schneider, SA" journalOrPublisher="Neotropical Entomology" pageId="0" pageNumber="1" pagination="641 - 647" refId="B25" refString="Schneider, SA, Sodano, J, LaPolla, JS, 2022. Distinguishing symbiotic partners of Acropyga ants from free-living soil inhabitants. Neotropical Entomology 51 (4): 641 - 647, DOI: https://doi.org/10.1007/s13744-022-00948-9" title="Distinguishing symbiotic partners of Acropyga ants from free-living soil inhabitants." url="https://doi.org/10.1007/s13744-022-00948-9" volume="51" year="2022">Schneider et al. (2022)</bibRefCitation>
. After specimens were collected into a nest-box, worker ants gathered trophobionts into a protective cluster and were actively engaged in attending to them.
</paragraph>
</subSubSection>
</treatment>
</document>