treatments-xml/data/C2/46/C6/C246C6390F96596CA5F63331DCEABC5D.xml

<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e81961" ID-Pensoft-Pub="1864-8312-81-79" ID-Pensoft-UUID="CA16EA0C5A33560FBC955981DFC9D502" ID-ZooBank="5C1FDC828F9F4869ADDD83FA96E507ED" ModsDocID="1864-8312-81-79" checkinTime="1674309819629" checkinUser="pensoft" docAuthor="Domahovski, Alexandre Cruz &amp; Cavichioli, Rodney Ramiro" docDate="2023" docId="C246C6390F96596CA5F63331DCEABC5D" docLanguage="en" docName="ArthropodSystPhyl 81: 79-164" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/asp.81.e81961" docTitle="Coarctana rana Domahovski &amp; Cavichioli 2023, comb. nov." docType="treatment" docVersion="1" id="CA16EA0C5A33560FBC955981DFC9D502" lastPageNumber="79" masterDocId="CA16EA0C5A33560FBC955981DFC9D502" masterDocTitle="Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data" masterLastPageNumber="164" masterPageNumber="79" pageNumber="79" updateTime="1674309819629" updateUser="pensoft">
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<mods:title>Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Domahovski, Alexandre Cruz</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4588-4236</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
<mods:nameIdentifier type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cavichioli, Rodney Ramiro</mods:namePart>
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<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
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<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date>2023</mods:date>
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<mods:url>http://dx.doi.org/10.3897/asp.81.e81961</mods:url>
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<taxonomicName ID-CoL="78R62" LSID="C246C639-0F96-596C-A5F6-3331DCEABC5D" authority="(DeLong &amp; Freytag)" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" baseAuthorityName="Domahovski &amp; Cavichioli" baseAuthorityYear="2023" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Coarctana rana" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="rana" status="comb. nov.">Coarctana rana (DeLong &amp; Freytag)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="79">comb. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">Figs 17</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">, 32C, D</figureCitation>
</paragraph>
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<subSubSection pageId="0" pageNumber="79" type="reference_group">
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" baseAuthorityName="Domahovski &amp; Cavichioli" baseAuthorityYear="2023" class="Insecta" family="Cicadellidae" genus="Acuera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuera rana" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="rana" status="comb. nov.">Acuera rana</taxonomicName>
DeLong &amp; Freytag, 1982: 309.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="diagnosis">
<paragraph pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="79">
Pygofer, in dorsal view (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17g</figureCitation>
), with a small subapical process directed inwards. Subgenital plate (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17H</figureCitation>
) with long filiform setae. Style (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17K</figureCitation>
) with apex expanded forming rounded lobe ventrally and small dorsal process, curved anterad. Aedeagus (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17L, M</figureCitation>
) with apodemal process very narrow, strongly curved dorsally; shaft slender; apex slightly inflated, with two pairs of short processes: a subapical pair on posterior surface, directed dorsally and a apical pair directed ventrally, about twice as long as the subapical pair.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="79">Total length: males 7.6-8.1 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Description.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head, in frontal view (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17B</figureCitation>
), frons 1.1 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; frontogenal suture distant from eye margin by approximately the width of scape; clypeus 1.3 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; lateral margins straight and parallel; apex straight. Head, in lateral view (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17C</figureCitation>
), crown-face transition moderately thick, with 5-6 transverse carinae; clypeus not inflated. Profemur AV row with 5-6 setae; PV row with 3-4 setae. Protibia PD row with 4 setae and undifferentiated intercalary setae; PV row with 5 setae and undifferentiated intercalary setae. Metatibia PD, AD and AV rows with 22-24, 12 and 13-15 macrosetae respectively. Metatarsomere I 4.5 
<normalizedToken originalValue="×">x</normalizedToken>
longer than apical width; inner row of the ventral surface with 7-8 setae. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="coloration">
<paragraph pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head and thorax (Figs 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A-D</figureCitation>
, 
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">32C, D</figureCitation>
) yellowish-brown. Head (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
) with pair of small circular black spots, near posterior margin, behind ocelli; coronal suture black; ocellus surrounded by black maculae in more melanic specimens. Face (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17B</figureCitation>
) yellowish, without maculae. Pronotum (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
) with black punctures and irregular black spots near anterior margin; proepimeron (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17C</figureCitation>
), with black band below pronotal carina. Mesonotum (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
) with dark-brown lateral angles, pair of small median black spots near scutoscutellar suture. Forewing (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17D</figureCitation>
) strongly marked by dark-brown mottling; veins yellowish-brown, outlined by dark-brown. Metatibia (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="male terminalia">
<paragraph pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
Sternite VIII (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17E</figureCitation>
) 1.5 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; lateral margins parallel; posterior margin produced medially forming short, rounded lobe. Valve (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17F</figureCitation>
) narrow, 1.1 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; posterior margin tapered and rounded. Pygofer, in lateral view (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17G</figureCitation>
), 1.4 
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum height; basodorsal process present but reduced; anteroventral and posterodorsal margins straight; posteroventral margin broadly rounded; apex broad, slightly excavated; in dorsal view (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17g</figureCitation>
), with small subapical process directed inwards. Subgenital plate, in lateral view (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17G</figureCitation>
), surpassing pygofer apex; in ventral view (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17H</figureCitation>
), elongated, 3.8 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, maximum width medially; outer margin with long filiform setae; inner margin slightly rounded; outer margin straight; apex weakly tapered and rounded. Connective (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17I</figureCitation>
) wider than long; anterior margin excavated; dorsal keel and stem reduced. Style, in dorsal view (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17J</figureCitation>
), with outer lobe small and rounded; strongly sinuate between outer lobe and blade; in lateral view (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17K</figureCitation>
), with blade slightly wider near base and tapered toward apex; dorsal and ventral margins slightly sinuate; ventral margin with apical half serrated; apex expanded forming rounded lobe ventrally and small dorsal process, curved anterad. Aedeagus (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17L, M</figureCitation>
) preatrium weakly developed; dorsal apodeme with dorsal margin slightly excavated, lateral margins produced ventrolaterally; apodemal process very narrow, strongly curved dorsally, apex tapered and acute; shaft slender, cylindrical, curved dorsally near base; apex slightly inflated, with two pairs of short processes: a subapical pair on posterior surface, directed dorsally and a apical pair directed ventrally, about twice as long as the subapical pair. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="female">
<paragraph pageId="0" pageNumber="79">Female.</paragraph>
<paragraph pageId="0" pageNumber="79">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="materials_examined">
<paragraph pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="79">
<materialsCitation collectingDate="2002-06-10" collectingDateMax="2002-07-03" collectingDateMin="2002-06-10" country="Bolivia" latitude="-16.915277" location="Vila Tunari" longLatPrecision="21" longitude="-65.36833" municipality="Bolivia" specimenCount="3" specimenCount-male="3" stateProvince="Cochabamba">
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingCountry name="Bolivia">Bolivia</collectingCountry>
</emphasis>
: 
<specimenCount type="male">3♂</specimenCount>
, 
<normalizedToken originalValue="“Bolívia">
&quot; 
<collectingMunicipality>Bolivia</collectingMunicipality>
</normalizedToken>
, 
<collectingRegion country="Bolivia" name="Cochabamba">Cochabamba</collectingRegion>
,\ 
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:FB3CF4D60D8497A4C450613F7C623833" country="Bolivia" latitude="-16.915277" longLatPrecision="21" longitude="-65.36833" municipality="Bolivia" name="Vila Tunari" stateProvince="Cochabamba">Vila Tunari</location>
\ 
<geoCoordinate degrees="16" direction="south" minutes="54" orientation="latitude" precision="15" seconds="55" value="-16.915277">16°54′55″S</geoCoordinate>
<geoCoordinate degrees="65" direction="west" minutes="22" orientation="longitude" precision="15" seconds="06" value="-65.36833">65°22′06″W</geoCoordinate>
\ 
<collectingDate value="2002-06-10" valueMax="2002-07-03" valueMin="2002-06-10">10.VI-03.VII.2002</collectingDate>
\ malaise Helmut Heidas&quot; (DZRJ)
</materialsCitation>
; 
<materialsCitation collectingDate="2002-09-07" collectingDateMax="2002-09-20" collectingDateMin="2002-09-07" country="Bolivia" latitude="-16.915277" location="Vila Tunari" longLatPrecision="21" longitude="-65.36833" municipality="Bolivia" specimenCount="1" specimenCount-male="1" stateProvince="Cochabamba">
<specimenCount type="male">1♂</specimenCount>
, same data as preceding except 
<collectingDate value="2002-09-07" valueMax="2002-09-20" valueMin="2002-09-07">07-20.IX.2002</collectingDate>
(DZUP)
</materialsCitation>
. - 
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Brazil" location="Rondonia" specimenCount="1" stateProvince="Rondonia">
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingCountry name="Brazil">Brazil</collectingCountry>
: 
<normalizedToken originalValue="Rondônia">
<collectingRegion country="Brazil" name="Rondonia">Rondonia</collectingRegion>
</normalizedToken>
</emphasis>
</materialsCitation>
: 
<materialsCitation collectingDate="1987-10-31" collectorName="C. Elias" country="Peru" location="Ouro Preto" specimenCount="1" specimenCount-male="1">
<specimenCount type="male">1♂</specimenCount>
, &quot; 
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:0A8AC486653B468A2623A5D820078CF8" country="Peru" name="Ouro Preto">Ouro Preto</location>
\ d′Oeste, RO.\ 
<collectingDate value="1987-10-31">31.X.1987</collectingDate>
\ 
<collectorName>C. Elias</collectorName>
, leg&quot; (DZUP). - 
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingCountry name="Peru">Peru</collectingCountry>
</emphasis>
</materialsCitation>
: 
<materialsCitation collectingDate="2005-03-06" collectingDateMax="2005-03-15" collectingDateMin="2005-03-06" country="Peru" county="Peru" elevation="335" latitude="-7.081333" location="Mun. Zona Barreal" longLatPrecision="13" longitude="-79.314835" municipality="Cons." specimenCount="2" specimenCount-male="2" stateProvince="San Martin">
<specimenCount type="male">2♂</specimenCount>
, &quot; 
<collectingCounty>Peru</collectingCounty>
: 
<collectingRegion country="Peru" name="San Martin">San Martin</collectingRegion>
, 
<collectingMunicipality>Cons.</collectingMunicipality>
\ 
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:AAC657CE158BADB295BC5DF75C9F13C4" country="Peru" county="Peru" latitude="-7.081333" longLatPrecision="13" longitude="-79.314835" municipality="Cons." name="Mun. Zona Barreal" stateProvince="San Martin">Mun. Zona Barreal</location>
23Km S\ 
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:04E941E52304E5D8EA2661A92C923242" country="Peru" county="Peru" latitude="-7.081333" longLatPrecision="13" longitude="-79.314835" municipality="Cons." name="Picota" stateProvince="San Martin">Picota</location>
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.35" unit="m" value="335.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.35" unit="m" value="335.0">335m</elevation>
</quantity>
, 
<geoCoordinate degrees="7" direction="south" minutes="04.88" orientation="latitude" precision="9" value="-7.081333">7°04.88′S</geoCoordinate>
, 
<geoCoordinate degrees="79" direction="west" minutes="18.89" orientation="longitude" precision="9" value="-79.314835">79°18.89′W</geoCoordinate>
<collectingDate value="2005-03-06" valueMax="2005-03-15" valueMin="2005-03-06">6-15Mar2005</collectingDate>
,\ ME Irwin, JD Vasquez,\ Malaise, dry Florest PE 11-02&quot; (DZRJ)
</materialsCitation>
; 
<materialsCitation collectingDate="2009-01-23" collectingDateMax="2009-01-30" collectingDateMin="2009-01-23" country="Peru" county="Peru" elevation="1125" latitude="-5.689722" location="Distr. Aguas Verdes" longLatPrecision="21" longitude="-77.63694" municipality="Dept" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas">
<specimenCount type="male">1♂</specimenCount>
, &quot; 
<collectingCounty>Peru</collectingCounty>
: 
<collectingMunicipality>Dept</collectingMunicipality>
<collectingRegion country="Peru" name="Amazonas">Amazonas</collectingRegion>
,\ 
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:145EFE81248CB68D1BB83221C6EC6CB8" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Dept" name="Distr. Aguas Verdes" stateProvince="Amazonas">
Distr. 
<normalizedToken originalValue="Águas">Aguas</normalizedToken>
Verdes
</location>
,\ 
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:4580DFDF3751AC418563BAA9F00DA064" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Dept" name="Bagua Taparoto Rd" stateProvince="Amazonas">Bagua Taparoto Rd</location>
(5N)\ at 
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:254D4982CD4A163BD955FAB78A57133D" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Dept" name="Km" stateProvince="Amazonas">Km</location>
403; 
<collectingDate value="2009-01-23" valueMax="2009-01-30" valueMin="2009-01-23">23-30.I.2009</collectingDate>
\ ME Irwin, GA Amaya,\ 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.125" unit="m" value="1125.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.125" unit="m" value="1125.0">1125m</elevation>
</quantity>
; 
<geoCoordinate degrees="05" direction="south" minutes="41" orientation="latitude" precision="15" seconds="23" value="-5.689722">05°41′23″S</geoCoordinate>
<geoCoordinate degrees="77" direction="west" minutes="38" orientation="longitude" precision="15" seconds="13" value="-77.63694">77°38′13″W</geoCoordinate>
&quot; (DZRJ)
</materialsCitation>
; 
<materialsCitation collectingDate="2009-01-30" collectingDateMax="2009-02-06" collectingDateMin="2009-01-30" country="Peru" county="Peru" elevation="1125" latitude="-5.689722" location="Distr. Aguas Verdes" longLatPrecision="21" longitude="-77.63694" municipality="Dept" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas">
<specimenCount type="male">1♂</specimenCount>
, same data as preceding except 
<collectingDate value="2009-01-30" valueMax="2009-02-06" valueMin="2009-01-30">30.I-06.II.2009</collectingDate>
(DZRJ)
</materialsCitation>
; 
<materialsCitation collectingDate="2009-05-15" collectingDateMax="2009-05-22" collectingDateMin="2009-05-15" country="Peru" county="Peru" elevation="1125" latitude="-5.689722" location="Distr. Aguas Verdes" longLatPrecision="21" longitude="-77.63694" municipality="Dept" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas">
<specimenCount type="male">1♂</specimenCount>
, same data as preceding except 
<collectingDate value="2009-05-15" valueMax="2009-05-22" valueMin="2009-05-15">15-22.V.2009</collectingDate>
(DZUP)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="remarks">
<paragraph pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="79">
Our analysis recovered 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. rana" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="rana">
<emphasis italics="true" pageId="0" pageNumber="79">C. rana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
as sister to 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. serpenta" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="serpenta">
<emphasis italics="true" pageId="0" pageNumber="79">C. serpenta</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
with low branch support (SR &lt;50) (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phylogenetic relationships of Acuera found in the implied weighting analysis for the two equally most parsimonious trees, K = 3.700 and 4.284, length = 514, consistency index = 0.362, retention index = 0.771. Symmetric resampling values (&lt;50) are marked next to branches; red stars represent the type species of the three subgenera of Acuera. Synapomorphies are marked with black circles, while homoplasies are marked with empty circles." figureDoi="10.3897/asp.81.e81961.figure7" httpUri="https://binary.pensoft.net/fig/798160" pageId="0" pageNumber="79">7</figureCitation>
). These species are similar in the overall coloration (Fig. 
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">32C-F</figureCitation>
); in having a small digitiform processes at pygofer apex (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17g</figureCitation>
); the subgenital plates (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17H</figureCitation>
) with long filiform setae; and in the shape of the apodemal processes of aedeagus, slender and strongly curved (Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17L</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>