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<document id="306A66E1006A470CE476790D6281C866" ID-DOI="10.1649/0010-065X-70.2.217" ID-Zenodo-Dep="5337631" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630178801869" checkinUser="diego" docAuthor="Chandler, Donald S. &amp; Geromini, Sarah D." docDate="1962" docId="B34D1D494F21AE706B0D78912811FD71" docLanguage="en" docName="0010-065X-70.2.217.pdf" docOrigin="The Coleopterists Bulletin 1962 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-70.2.217" docStyle="DocumentStyle:A0A70CFE30726088A421CCB226FF86C7.4:TheColeopBull.2010-2016.journal_article.1cover" docStyleId="A0A70CFE30726088A421CCB226FF86C7" docStyleName="TheColeopBull.2010-2016.journal_article.1cover" docStyleVersion="4" docTitle="Custotychus spinosus Chandler and Geromini 1962, new species" docType="treatment" docVersion="3" lastPageNumber="242" masterDocId="4F7465314F2EAE60680778532A04FFA1" masterDocTitle="A Revision of the Eastern Nearctic GenusCustotychusPark and Wagner, 1962 (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="242" masterPageNumber="217" pageNumber="217" updateTime="1699176462293" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="2698536F26495C0EC743C943ABDA6C29" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="279CF8E1080EE8C57284153B93C09D31">
<mods:title id="28271BB8FE0E21208E90A5A662B7BA0D">A Revision of the Eastern Nearctic GenusCustotychusPark and Wagner, 1962 (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
</mods:titleInfo>
<mods:name id="82FF06C247AFFDE01CC22F248E681B09" type="personal">
<mods:role id="61310A392976D965EBC19CA8EFB2D84A">
<mods:roleTerm id="917933AFC1C1BFE1EE4EDA8CA9A22FD2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="89A5896FDC4632BDFF5C2E3EFF050DD8">Chandler, Donald S.</mods:namePart>
</mods:name>
<mods:name id="D8E99DBDA4EEF5FA9010842335D4B858" type="personal">
<mods:role id="7764AABCE1C8D497F588FD8FA2D418F8">
<mods:roleTerm id="EC4F0BCCCB25272D2D7EB0CB1B27A2A6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3988D5B4B02F4E5CFA20FD8D54A25A95">Geromini, Sarah D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D88182F4557240AC26BC107331231E75">text</mods:typeOfResource>
<mods:relatedItem id="D9BF25F16235C204A01B30488C48DC0B" type="host">
<mods:titleInfo id="657156FE409BD8DFCC953834E635108A">
<mods:title id="70C420628BE89BCF3953321731529051">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="790B7ED78763801043405CB31C2A49EB">
<mods:date id="7ADCA92D1FABE822DB91096F6BD87180">1962</mods:date>
<mods:detail id="28A5360EA786CFB9E8FCA5C15EA54401" type="pubDate">
<mods:number id="99786306D86D18820FF4F49499CB5BCC">2016-06-30</mods:number>
</mods:detail>
<mods:detail id="460134519E6C2974DD86F446D8816299" type="volume">
<mods:number id="2EDEE113DE1448BF69B22DFC6DB4C6D5">1962</mods:number>
</mods:detail>
<mods:detail id="74E303F5B28E569287593371AD2BFCCF" type="issue">
<mods:number id="80EA5453991B5537811EE0837BE5BE5D">2</mods:number>
</mods:detail>
<mods:extent id="164340789E88795533C98E219AF2ADD2" unit="page">
<mods:start id="4FE99D85EA6253451012C458BB22BB6C">217</mods:start>
<mods:end id="9608C611F2EF00D12EEE21A10B42AE9D">242</mods:end>
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</mods:relatedItem>
<mods:location id="0E7DEC973C3B3CCF2CA41DBABDBD3ECD">
<mods:url id="E85B1B3DC3C79068F067069DF5E1397F">http://dx.doi.org/10.1649/0010-065x-70.2.217</mods:url>
</mods:location>
<mods:classification id="279E5696219ADFFCDF2C166CA8244DF2">journal article</mods:classification>
<mods:identifier id="43939D54F6C10BF539EEC95D3C13389D" type="DOI">10.1649/0010-065X-70.2.217</mods:identifier>
<mods:identifier id="FFAB0B26788E640890ABAAFA29E4333F" type="Zenodo-Dep">5337631</mods:identifier>
</mods:mods>
<treatment id="B34D1D494F21AE706B0D78912811FD71" LSID="urn:lsid:plazi:treatment:B34D1D494F21AE706B0D78912811FD71" httpUri="http://treatment.plazi.org/id/B34D1D494F21AE706B0D78912811FD71" lastPageId="16" pageId="15">
<subSubSection id="73FEFFD44F21AE6F6B0D78912E44FF53" pageId="15" type="nomenclature">
<paragraph id="3B5BAC5F4F21AE6F6B0D78912E44FF53" blockId="15.[778,1121,194,269]" pageId="15">
<heading id="60131B334F21AE6F6B0D78912E44FF53" centered="true" fontSize="10" level="2" pageId="15" reason="2">
<emphasis id="0990704D4F21AE6F6B0D78912E44FF53" bold="true" pageId="15">
6.
<taxonomicName id="FCE4D7DC4F21AE6F6B22789129B9FF53" ID-CoL="32MKP" authority="Chandler and Geromini" authorityName="Chandler and Geromini" authorityYear="1962" class="Insecta" family="Staphylinidae" genus="Custotychus" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="spinosus" status="sp. nov.">
<emphasis id="0990704D4F21AE6F6B22789129FDFF76" bold="true" box="[805,1017,194,215]" italics="true" pageId="15">Custotychus spinosus</emphasis>
Chandler and Geromini
</taxonomicName>
,
<taxonomicNameLabel id="12A3CD364F21AE6F6BCD788E2E44FF53" box="[970,1088,221,242]" pageId="15" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="73FEFFD44F21AE706B4578A42846FEE0" lastPageId="16" pageId="15" type="description">
<paragraph id="3B5BAC5F4F21AE6F6B4578A42E2EFEAD" blockId="15.[778,1121,194,269]" box="[834,1066,247,269]" pageId="15">
(
<figureCitation id="A3DFB0DA4F21AE6F6B4E78A42994FEAD" box="[841,912,247,268]" captionStart="Figs" captionStartId="4.[175,215,1525,1544]" captionTargetBox="[182,1152,200,1496]" captionTargetId="figure-63@4.[151,1171,183,1501]" captionTargetPageId="4" captionText="Figs. 1–7. Custotychus species. 1) C. natchez, dorsal habitus; 2–7) Aedeagus, dorsal and left lateral views:" figureDoi="http://doi.org/10.5281/zenodo.5337633" httpUri="https://zenodo.org/record/5337633/files/figure.png" pageId="15">Figs. 7</figureCitation>
,
<figureCitation id="A3DFB0DA4F21AE6F6B9B78A429DCFEAD" box="[924,984,247,269]" captionStart="Figs" captionStartId="7.[197,239,1493,1512]" captionTargetBox="[173,1193,194,1466]" captionTargetId="figure-1@7.[172,1194,194,1470]" captionTargetPageId="7" captionText="Figs. 16–27. Custotychus species, scanning electron micrographs right lateral (a) and posterior (b) views of metasternal tubercle. 16) C. chickasaw; 17) C. daggyi; 18) C. lobatus; 19) C. turnbowi; 20) C. rothorum; 21) C. spinosus; 22) C. verticalis; 23) C. watrousi; 24) C. carloritaorum; 25) C. leei; 26) C. minor; 27) C. natchez. Grid for measurements of 20 or 50m at bottom of micrographs." figureDoi="http://doi.org/10.5281/zenodo.5337639" httpUri="https://zenodo.org/record/5337639/files/figure.png" pageId="15">21a–b</figureCitation>
,
<figureCitation id="A3DFB0DA4F21AE6F6BE378A429FAFEAD" box="[996,1022,247,268]" captionStart="Figs" captionStartId="8.[175,215,740,759]" captionTargetBox="[151,1171,194,717]" captionTargetId="figure-61@8.[151,1173,194,717]" captionTargetPageId="8" captionText="Figs. 28–32. Custotychus species. 28) C. pocahontas, scanning electron micrographs of right lateral (a) and posterior (b) views of metasternal tubercle. Grid for measurements of 20 or 50m at bottom of micrographs; 29) C. carloritaorum, mesotibial spurs; 30) C. spinosus, mesotibial spurs; 31) C. watrousi, abdominal ventrites, ventrolateral view; 32) C. daggyi, right maxillary palpus, ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5337643" httpUri="https://zenodo.org/record/5337643/files/figure.png" pageId="15">30</figureCitation>
,
<figureCitation id="A3DFB0DA4F21AE6F6C0D78A42E27FEAD" box="[1034,1059,247,268]" captionStart="Figs" captionStartId="9.[197,237,1662,1681]" captionTargetBox="[289,1077,194,1639]" captionTargetId="figure-1@9.[289,1078,194,1639]" captionTargetPageId="9" captionText="Figs. 34–36. Distribution of Custotychus species. 34) Distribution of species in the spinosus and verticalis species-groups: C. rothorum and C. spinosus, and C. cavei, C. verticalis, and C. watrousi; 35) Distribution of C. minor; 36) Distribution of other species in the minor species-group: C. carloritaorum, C. leei, C. natchez, and C. pocahontas." figureDoi="http://doi.org/10.5281/zenodo.5337649" httpUri="https://zenodo.org/record/5337649/files/figure.png" pageId="15">34</figureCitation>
)
</paragraph>
<paragraph id="3B5BAC5F4F21AE6F6ADC7971294FFC01" blockId="15.[707,1194,290,1729]" pageId="15">
<emphasis id="0990704D4F21AE6F6ADC797129C2FE96" bold="true" box="[731,966,290,311]" pageId="15">Specimens Examined.</emphasis>
<emphasis id="0990704D4F21AE6F6BD3797129E4FE96" box="[980,992,290,311]" italics="true" pageId="15">n</emphasis>
=
<materialsCitation id="8B8CA6024F21AE6F6C0E79712981FE1C" collectingDate="2003-05-28" collectionCode="FMNH" collectorName="T. King Berlese" elevation="285" latitude="33.88253" location="Blount Co. Nr. Highland Lake" longLatPrecision="1" longitude="-86.42185" pageId="15" specimenCode="LSAM 0293949" specimenCount="285" typeStatus="holotype">
2 1.
<emphasis id="0990704D4F21AE6F6C3D797128FEFEF0" bold="true" pageId="15">H o l o t y p e male:</emphasis>
// USA.AL.
<location id="3E3BFA844F21AE6F6B7A796F2EACFEF0" LSID="urn:lsid:plazi:treatment:B34D1D494F21AE706B0D78912811FD71:3E3BFA844F21AE6F6B7A796F2EACFEF0" box="[893,1192,316,337]" latitude="33.88253" longLatPrecision="1" longitude="-86.42185" name="Blount Co. Nr. Highland Lake" pageId="15">Blount Co. Nr. Highland Lake</location>
<geoCoordinate id="5ED0CA984F21AE6F6AC479042938FECD" box="[707,828,343,364]" degrees="33.88253" direction="north" orientation="latitude" pageId="15" precision="1" value="33.88253">33.88253°N</geoCoordinate>
<geoCoordinate id="5ED0CA984F21AE6F6B41790429C1FECD" box="[838,965,343,364]" degrees="86.42185" direction="west" orientation="longitude" pageId="15" precision="1" value="-86.42185">86.42185°W</geoCoordinate>
<specimenCount id="2DE267D64F21AE6F6BC879042E0BFECD" box="[975,1039,343,364]" pageId="15" type="generic">
<quantity id="FC1C01BA4F21AE6F6BC879042E0FFECD" box="[975,1035,343,364]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="15" unit="m" value="285.0">
<elevation id="B0C94B6C4F21AE6F6BC879042E0FFECD" box="[975,1035,343,364]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="15" unit="m" value="285.0">285m</elevation>
</quantity>
.
</specimenCount>
<date id="4F5A8A9F4F21AE6F6C1079042EA2FECD" box="[1047,1190,343,364]" pageId="15" value="2003-05-28">
<collectingDate id="5F1E73774F21AE6F6C1079042EA2FECD" box="[1047,1190,343,364]" pageId="15" value="2003-05-28">28 May 2003</collectingDate>
</date>
.
<collectorName id="9611C9894F21AE6F6AC479212968FE26" box="[707,876,370,391]" pageId="15">T. King Berlese</collectorName>
/
<specimenCode id="6B4204244F21AE6F6B7979212E2CFE26" box="[894,1064,370,391]" collectionCode="LSAM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" lsid="urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="15">LSAM 0293949</specimenCode>
/ [red label]
<typeStatus id="E45F12FD4F21AE6F6AC479DE2940FE03" box="[707,836,397,418]" pageId="15" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="FCE4D7DC4F21AE6F6B4979DE291FFE1C" authority="Chandler &amp; Geromini" authorityName="Chandler &amp; Geromini" authorityYear="1962" class="Insecta" family="Staphylinidae" genus="Custotychus" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="spinosus">Custotychus spinosus Chandler &amp; Geromini</taxonomicName>
// (
<collectionCode id="5DF5349A4F21AE6F6B3279FB297AFE1C" box="[821,894,424,445]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="15" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8B8CA6024F21AE6F6B8979FB2E1AFE53" collectingDate="2003-05-28" collectionCode="DSCC, LSAM" collectorName="T. King" county="Blount Co." elevation="285" location="Nr. Highland Lake" pageId="15" specimenCount="1" stateProvince="Alabama" typeStatus="paratype">
<emphasis id="0990704D4F21AE6F6B8979FB2E60FE1C" bold="true" box="[910,1124,424,445]" pageId="15">
<typeStatus id="E45F12FD4F21AE6F6B8979FB29F1FE1C" box="[910,1013,424,445]" pageId="15" type="paratype">Paratypes</typeStatus>
:
<collectingRegion id="F92062BD4F21AE6F6C0779FB2E64FE1C" box="[1024,1120,424,445]" country="United States of America" name="Alabama" pageId="15">Alabama</collectingRegion>
:
</emphasis>
<emphasis id="0990704D4F21AE6F6C6C79FB28E1FE79" italics="true" pageId="15">
<collectingCounty id="D23AD4D34F21AE6F6C6C79FB28E1FE79" pageId="15">Blount Co.</collectingCounty>
</emphasis>
:
<location id="3E3BFA844F21AE6F6AF7799029A5FE79" LSID="urn:lsid:plazi:treatment:B34D1D494F21AE706B0D78912811FD71:3E3BFA844F21AE6F6AF7799029A5FE79" box="[752,929,451,472]" county="Blount Co." name="Nr. Highland Lake" pageId="15" stateProvince="Alabama">Nr. Highland Lake</location>
,
<quantity id="FC1C01BA4F21AE6F6BAD799029E7FE79" box="[938,995,451,472]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="15" unit="m" value="285.0">
<elevation id="B0C94B6C4F21AE6F6BAD799029E7FE79" box="[938,995,451,472]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="15" unit="m" value="285.0">285m</elevation>
</quantity>
,
<date id="4F5A8A9F4F21AE6F6BEB79902E56FE79" box="[1004,1106,451,472]" pageId="15" value="2003-05-28">
<collectingDate id="5F1E73774F21AE6F6BEB79902E56FE79" box="[1004,1106,451,472]" pageId="15" value="2003-05-28">V-28-2003</collectingDate>
</date>
,
<collectorName id="9611C9894F21AE6F6C5B79902EA1FE79" box="[1116,1189,451,472]" pageId="15">T. King</collectorName>
, berlese (1M,
<collectionCode id="5DF5349A4F21AE6F6B4B798E298BFE53" box="[844,911,477,498]" pageId="15">DSCC</collectionCode>
; 8M,
<collectionCode id="5DF5349A4F21AE6F6BC9798E2E12FE53" box="[974,1046,477,498]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" lsid="urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="15">LSAM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8B8CA6024F21AE6F6C2D798D2EA2FDE2" collectingDate="2014-03-24" collectionCode="MEMC" collectorName="J. A. MacGown &amp; J. G. Lewis" county="Colbert Co." location="Cane Creek" pageId="15" specimenCount="1" stateProvince="Alabama" typeStatus="paratype">
<emphasis id="0990704D4F21AE6F6C2D798D2EA7FE52" box="[1066,1187,478,499]" italics="true" pageId="15">
<collectingCounty id="D23AD4D34F21AE6F6C2D798D2EA7FE52" box="[1066,1187,478,499]" pageId="15">Colbert Co.</collectingCounty>
</emphasis>
:
<location id="3E3BFA844F21AE6F6AC479AB293CFDAC" LSID="urn:lsid:plazi:treatment:B34D1D494F21AE706B0D78912811FD71:3E3BFA844F21AE6F6AC479AB293CFDAC" box="[707,824,504,525]" county="Colbert Co." name="Cane Creek" pageId="15" stateProvince="Alabama">Cane Creek</location>
Canyon N[ature] P[reserve],
<date id="4F5A8A9F4F21AE6F6C6F79AB28F1FD89" pageId="15" value="2014-03-24">
<collectingDate id="5F1E73774F21AE6F6C6F79AB28F1FD89" pageId="15" value="2014-03-24">III-24- 2014</collectingDate>
</date>
,
<collectorName id="9611C9894F21AE6F6B067A402991FD89" box="[769,917,531,552]" pageId="15">J.A. MacGown</collectorName>
&amp;
<collectorName id="9611C9894F21AE6F6BBE7A402E21FD89" box="[953,1061,531,552]" pageId="15">J.G. Lewis</collectorName>
, berlese litter from upland hardwood forest (1M 2F,
<collectionCode id="5DF5349A4F21AE6F6C577A7D2E9BFDE2" box="[1104,1183,558,579]" pageId="15">MEMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8B8CA6024F21AE6F6AC47A1A29D0FDD8" collectingDate="1972-05-16" collectionCode="CNCI" collectorName="S. &amp; J. Peck" county="Jackson Co." location="10 mi. N Scottsboro" pageId="15" specimenCount="1" stateProvince="Alabama" typeStatus="paratype">
<emphasis id="0990704D4F21AE6F6AC47A1A2945FDFF" box="[707,833,585,606]" italics="true" pageId="15">
<collectingCounty id="D23AD4D34F21AE6F6AC47A1A2945FDFF" box="[707,833,585,606]" pageId="15">Jackson Co.</collectingCounty>
</emphasis>
:
<location id="3E3BFA844F21AE6F6B557A1A2E2DFDFF" LSID="urn:lsid:plazi:treatment:B34D1D494F21AE706B0D78912811FD71:3E3BFA844F21AE6F6B557A1A2E2DFDFF" box="[850,1065,585,606]" county="Jackson Co." name="10 mi. N Scottsboro" pageId="15" stateProvince="Alabama">
<quantity id="FC1C01BA4F21AE6F6B557A1A2992FDFF" box="[850,918,585,606]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="15" unit="mi" value="10.0">10 mi.</quantity>
N Scottsboro
</location>
,
<date id="4F5A8A9F4F21AE6F6C307A1A2EA1FDFF" box="[1079,1189,585,606]" pageId="15" value="1972-05-16">
<collectingDate id="5F1E73774F21AE6F6C307A1A2EA1FDFF" box="[1079,1189,585,606]" pageId="15" value="1972-05-16">V-16-1972</collectingDate>
</date>
,
<collectorName id="9611C9894F21AE6F6AC47A372943FDD8" box="[707,839,612,633]" pageId="15">S. &amp; J. Peck</collectorName>
(1M,
<collectionCode id="5DF5349A4F21AE6F6B8A7A3729C9FDD8" box="[909,973,612,633]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="15">CNCI</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8B8CA6024F21AE6F6BE67A37299DFD0F" collectingDate="1984-06-04" collectionCode="FMNH" collectorName="T. King" county="Walker Co." location="Devil'" municipality="Jasper" pageId="15" specimenCount="1" stateProvince="Alabama" typeStatus="paratype">
<emphasis id="0990704D4F21AE6F6BE67A372E57FDD8" box="[993,1107,612,633]" italics="true" pageId="15">
<collectingCounty id="D23AD4D34F21AE6F6BE67A372E57FDD8" box="[993,1107,612,633]" pageId="15">Walker Co.</collectingCounty>
</emphasis>
:
<collectingMunicipality id="DB3F36254F21AE6F6C637A372EA1FDD8" box="[1124,1189,612,633]" pageId="15">Jasper</collectingMunicipality>
,
<location id="3E3BFA844F21AE6F6AC47A2C28FBFD35" LSID="urn:lsid:plazi:treatment:B34D1D494F21AE706B0D78912811FD71:3E3BFA844F21AE6F6AC47A2C28FBFD35" box="[707,767,638,660]" county="Walker Co." municipality="Jasper" name="Devil'" pageId="15" stateProvince="Alabama">Devil’</location>
s
<location id="3E3BFA844F21AE6F6B167A2C2953FD35" LSID="urn:lsid:plazi:treatment:B34D1D494F21AE706B0D78912811FD71:3E3BFA844F21AE6F6B167A2C2953FD35" box="[785,855,639,660]" county="Walker Co." municipality="Jasper" name="Ladder" pageId="15" stateProvince="Alabama">Ladder</location>
,
<date id="4F5A8A9F4F21AE6F6B637A2C29C8FD35" box="[868,972,639,660]" pageId="15" value="1984-06-04">
<collectingDate id="5F1E73774F21AE6F6B637A2C29C8FD35" box="[868,972,639,660]" pageId="15" value="1984-06-04">VI-4-1984</collectingDate>
</date>
,
<collectorName id="9611C9894F21AE6F6BDE7A2C2E23FD35" box="[985,1063,639,660]" pageId="15">T. King</collectorName>
, florest floor debris (1M,
<collectionCode id="5DF5349A4F21AE6F6B427ACA2995FD0F" box="[837,913,665,686]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="15" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8B8CA6024F21AE6F6BA17ACA2996FD45" collectingDate="2011-11-26" collectionCode="BBC" collectorName="B. Baldwin" county="Garland Co." location="Jessieville" pageId="15" specimenCount="1" stateProvince="Arkansas" typeStatus="paratype">
<emphasis id="0990704D4F21AE6F6BA17ACA2E12FD0F" bold="true" box="[934,1046,665,686]" pageId="15">
<collectingRegion id="F92062BD4F21AE6F6BA17ACA2E15FD0F" box="[934,1041,665,686]" country="United States of America" name="Arkansas" pageId="15">Arkansas</collectingRegion>
:
</emphasis>
<emphasis id="0990704D4F21AE6F6C187AC92EA6FD0E" box="[1055,1186,666,687]" italics="true" pageId="15">
<collectingCounty id="D23AD4D34F21AE6F6C187AC92EA6FD0E" box="[1055,1186,666,687]" pageId="15">Garland Co.</collectingCounty>
</emphasis>
:
<location id="3E3BFA844F21AE6F6AC47AE72920FD68" LSID="urn:lsid:plazi:treatment:B34D1D494F21AE706B0D78912811FD71:3E3BFA844F21AE6F6AC47AE72920FD68" box="[707,804,692,713]" county="Garland Co." name="Jessieville" pageId="15" stateProvince="Arkansas">Jessieville</location>
,
<date id="4F5A8A9F4F21AE6F6B297AE7299AFD68" box="[814,926,692,713]" pageId="15" value="2011-11-26">
<collectingDate id="5F1E73774F21AE6F6B297AE7299AFD68" box="[814,926,692,713]" pageId="15" value="2011-11-26">XI-26-2011</collectingDate>
</date>
,
<collectorName id="9611C9894F21AE6F6BAF7AE72E12FD68" box="[936,1046,692,713]" pageId="15">B. Baldwin</collectorName>
,
<collectingMethod id="E2A5D4484F21AE6F6C267AE72EADFD68" box="[1057,1193,692,713]" pageId="15">Berlese funnel</collectingMethod>
leaf litter (1M,
<collectionCode id="5DF5349A4F21AE6F6B5F7A9C298FFD45" box="[856,907,719,740]" pageId="15">BBC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8B8CA6024F21AE6F6B9A7A9C2E39FD5E" collectionCode="DSCC" county="Clarke Co." location="5 mi. W Athens" pageId="15" specimenCount="1" stateProvince="Georgia" typeStatus="paratype">
<emphasis id="0990704D4F21AE6F6B9A7A9C29F1FD45" bold="true" box="[925,1013,719,740]" pageId="15">
<collectingRegion id="F92062BD4F21AE6F6B9A7A9C29F5FD45" box="[925,1009,719,740]" country="United States of America" name="Georgia" pageId="15">Georgia</collectingRegion>
:
</emphasis>
<emphasis id="0990704D4F21AE6F6BF97A9C2E6CFD45" box="[1022,1128,719,740]" italics="true" pageId="15">
<collectingCounty id="D23AD4D34F21AE6F6BF97A9C2E6CFD45" box="[1022,1128,719,740]" pageId="15">Clarke Co.</collectingCounty>
</emphasis>
:
<location id="3E3BFA844F21AE6F6C717A9C2923FD5E" LSID="urn:lsid:plazi:treatment:B34D1D494F21AE706B0D78912811FD71:3E3BFA844F21AE6F6C717A9C2923FD5E" county="Clarke Co." name="5 mi. W Athens" pageId="15" stateProvince="Georgia">
<quantity id="FC1C01BA4F21AE6F6C717A9C2EAEFD45" box="[1142,1194,719,740]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="15" unit="mi" value="5.0">5 mi.</quantity>
W Athens
</location>
, II-8/21-1978 (1M,
<collectionCode id="5DF5349A4F21AE6F6BF47AB92E32FD5E" box="[1011,1078,746,767]" pageId="15">DSCC</collectionCode>
)
</materialsCitation>
, III-15/22- 1978 (1M, UGAC), XII-1/7-1977 (1M, UGAC), XII-8/16-1977 (1M, UGAC),
<materialsCitation id="8B8CA6024F21AE6F6BF77B732942FC01" collectionCode="FMNH" collectorName="C. L. Smith &amp; Bruce Hollow &amp; S. Peck &amp; A. Fiske" county="Edmonson Co." location="Mammoth Cave National Park" pageId="15" specimenCount="1" stateProvince="Kentucky" typeStatus="paratype">
<collectorName id="9611C9894F21AE6F6BF77B732E66FC94" box="[1008,1122,800,821]" pageId="15">C.L. Smith</collectorName>
,
<collectingMethod id="E2A5D4484F21AE6F6C687B7328F5FCEE" pageId="15">pitfall trap</collectingMethod>
.
<emphasis id="0990704D4F21AE6F6B077B69297DFCEE" bold="true" box="[768,889,826,847]" pageId="15">
<collectingRegion id="F92062BD4F21AE6F6B077B692970FCEE" box="[768,884,826,847]" country="United States of America" name="Kentucky" pageId="15">Kentucky</collectingRegion>
:
</emphasis>
<emphasis id="0990704D4F21AE6F6B827B682E2CFCF1" box="[901,1064,827,848]" italics="true" pageId="15">
<collectingCounty id="D23AD4D34F21AE6F6B827B682E2CFCF1" box="[901,1064,827,848]" pageId="15">Edmonson Co.</collectingCounty>
</emphasis>
:
<location id="3E3BFA844F21AE6F6C3C7B692988FCCB" LSID="urn:lsid:plazi:treatment:B34D1D494F21AE706B0D78912811FD71:3E3BFA844F21AE6F6C3C7B692988FCCB" county="Edmonson Co." name="Mammoth Cave National Park" pageId="15" stateProvince="Kentucky">Mammoth Cave National Park</location>
,
<collectorName id="9611C9894F21AE6F6B9E7B062E2CFCCB" box="[921,1064,853,874]" pageId="15">Bruce Hollow</collectorName>
, VIII-24/27- 1967,
<collectorName id="9611C9894F21AE6F6AF87B23294CFC24" box="[767,840,880,901]" pageId="15">S. Peck</collectorName>
&amp;
<collectorName id="9611C9894F21AE6F6B6D7B2329B9FC24" box="[874,957,880,901]" pageId="15">A. Fiske</collectorName>
, berlese, log stump litter (1M,
<collectionCode id="5DF5349A4F21AE6F6AFF7BD82944FC01" box="[760,832,907,928]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="15" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="3B5BAC5F4F21AE6F6ADC7BF62E32F92D" blockId="15.[707,1194,290,1729]" pageId="15">
<emphasis id="0990704D4F21AE6F6ADC7BF6299EFC1B" bold="true" box="[731,922,933,954]" pageId="15">Description. Body:</emphasis>
Brown;
<quantity id="FC1C01BA4F21AE6F6BEB7BF62E77FC1B" box="[1004,1139,933,955]" metricMagnitude="-3" metricUnit="m" metricValue="1.59" metricValueMax="1.62" metricValueMin="1.56" pageId="15" unit="mm" value="1.59" valueMax="1.62" valueMin="1.56">1.56–1.62 mm</quantity>
long.
<emphasis id="0990704D4F21AE6F6AC47B9328F8FC74" bold="true" box="[707,764,960,981]" pageId="15">Male:</emphasis>
Eyes with about 35 facets; metasternal tubercle (
<figureCitation id="A3DFB0DA4F21AE6F6AEE7B882951FC51" box="[745,853,987,1008]" captionStart="Figs" captionStartId="7.[197,239,1493,1512]" captionTargetBox="[173,1193,194,1466]" captionTargetId="figure-1@7.[172,1194,194,1470]" captionTargetPageId="7" captionText="Figs. 16–27. Custotychus species, scanning electron micrographs right lateral (a) and posterior (b) views of metasternal tubercle. 16) C. chickasaw; 17) C. daggyi; 18) C. lobatus; 19) C. turnbowi; 20) C. rothorum; 21) C. spinosus; 22) C. verticalis; 23) C. watrousi; 24) C. carloritaorum; 25) C. leei; 26) C. minor; 27) C. natchez. Grid for measurements of 20 or 50m at bottom of micrographs." figureDoi="http://doi.org/10.5281/zenodo.5337639" httpUri="https://zenodo.org/record/5337639/files/figure.png" pageId="15">Figs. 21a, b</figureCitation>
) at posterior margin of metasternum; in lateral view, curving anteriorly, apex narrowly rounded, posterior face flat; in posterior view, lateral margins to apex slightly broadening, apex rounded; tubercle height
<quantity id="FC1C01BA4F21AE6F6BCC7C152E33FBFA" box="[971,1079,1094,1115]" metricMagnitude="-5" metricUnit="m" metricValue="6.9" pageId="15" unit="mm" value="0.069">0.069 mm</quantity>
, maximum apical width
<quantity id="FC1C01BA4F21AE6F6B497C3229BFFBD7" box="[846,955,1121,1142]" metricMagnitude="-5" metricUnit="m" metricValue="3.2" pageId="15" unit="mm" value="0.032">0.032 mm</quantity>
, minimum stem width
<quantity id="FC1C01BA4F21AE6F6AC47C282921FB31" box="[707,805,1147,1168]" metricMagnitude="-5" metricUnit="m" metricValue="2.6" pageId="15" unit="mm" value="0.026">0.026 mm</quantity>
[tubercle measurements based on specimen from near Highland Lake, AL; LSAM]. Mesotibiae with large, slightly curved spine at middle on posteromesal margin (
<figureCitation id="A3DFB0DA4F21AE6F6B9D7C9F29E7FB40" box="[922,995,1228,1249]" captionStart="Figs" captionStartId="8.[175,215,740,759]" captionTargetBox="[151,1171,194,717]" captionTargetId="figure-61@8.[151,1173,194,717]" captionTargetPageId="8" captionText="Figs. 28–32. Custotychus species. 28) C. pocahontas, scanning electron micrographs of right lateral (a) and posterior (b) views of metasternal tubercle. Grid for measurements of 20 or 50m at bottom of micrographs; 29) C. carloritaorum, mesotibial spurs; 30) C. spinosus, mesotibial spurs; 31) C. watrousi, abdominal ventrites, ventrolateral view; 32) C. daggyi, right maxillary palpus, ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5337643" httpUri="https://zenodo.org/record/5337643/files/figure.png" pageId="15">Fig. 30</figureCitation>
), probably modified preapical spur, otherwise preapical spur lacking, apical spur short, bluntly pointed. Ventrite 2 broadly rounded at middle, ventrite 6 with small, faint, circular impression at middle. Aedeagus (
<figureCitation id="A3DFB0DA4F21AE6F6ACD7D02290DFAC7" box="[714,777,1361,1382]" captionStart="Figs" captionStartId="4.[175,215,1525,1544]" captionTargetBox="[182,1152,200,1496]" captionTargetId="figure-63@4.[151,1171,183,1501]" captionTargetPageId="4" captionText="Figs. 1–7. Custotychus species. 1) C. natchez, dorsal habitus; 2–7) Aedeagus, dorsal and left lateral views:" figureDoi="http://doi.org/10.5281/zenodo.5337633" httpUri="https://zenodo.org/record/5337633/files/figure.png" pageId="15">Fig. 7</figureCitation>
) 0.32 long mm; in dorsal view, median lobe elongate, narrow, lateral margins slightly concave throughout length, lateral arms of dorsal plate elongate but short, extending to point just beyond middle of median lobe, appearing shortly bifurcate at apex; in lateral view, median lobe pointed at apex, slightly swollen in apical half; lateral arms of dorsal plate with spur curved strongly ventrally, second spur curved weakly dorsally; parameres swollen in apical half, extending to point even with apex of median lobe, membranous areas small.
<emphasis id="0990704D4F21AE6F6B727E2429C3F92D" bold="true" box="[885,967,1655,1676]" pageId="15">Female:</emphasis>
Unknown.
</paragraph>
<paragraph id="3B5BAC5F4F21AE706ADC7EC12846FEE0" blockId="15.[707,1194,290,1729]" lastBlockId="16.[151,638,194,720]" lastPageId="16" pageId="15">
<emphasis id="0990704D4F21AE6F6ADC7EC129DCF906" bold="true" box="[731,984,1682,1703]" pageId="15">Collection Data Synopsis.</emphasis>
Individuals were collected from February to June, and in August and December. They were collected by processing forest floor debris/litter and stump litter with Berlese funnels, and several were taken by pitfall traps. Specimens are present in the following collections: BBC, CNCI, DSCC, FMNH, MEMC, and UGAC.
</paragraph>
</subSubSection>
<subSubSection id="73FEFFD44F3EAE7068A879142BBCFE0D" pageId="16" type="distribution">
<paragraph id="3B5BAC5F4F3EAE7068A879142BBCFE0D" blockId="16.[151,638,194,720]" pageId="16">
<emphasis id="0990704D4F3EAE7068A879142B36FEFD" bold="true" box="[175,306,327,348]" pageId="16">Distribution.</emphasis>
This species occurs in the southeastern states from
<collectingCountry id="43F3ECCF4F3EAE70695379312BA4FED6" box="[340,416,354,375]" name="Georgia" pageId="16">Georgia</collectingCountry>
to Arkansas (
<figureCitation id="A3DFB0DA4F3EAE706A2D79312877FED6" box="[554,627,354,375]" captionStart="Figs" captionStartId="9.[197,237,1662,1681]" captionTargetBox="[289,1077,194,1639]" captionTargetId="figure-1@9.[289,1078,194,1639]" captionTargetPageId="9" captionText="Figs. 34–36. Distribution of Custotychus species. 34) Distribution of species in the spinosus and verticalis species-groups: C. rothorum and C. spinosus, and C. cavei, C. verticalis, and C. watrousi; 35) Distribution of C. minor; 36) Distribution of other species in the minor species-group: C. carloritaorum, C. leei, C. natchez, and C. pocahontas." figureDoi="http://doi.org/10.5281/zenodo.5337649" httpUri="https://zenodo.org/record/5337649/files/figure.png" pageId="16">Fig. 34</figureCitation>
). Specimens were examined from the following states: AL, AR, GA, and KY.
</paragraph>
</subSubSection>
<subSubSection id="73FEFFD44F3EAE7068A879E2286DFE40" pageId="16" type="etymology">
<paragraph id="3B5BAC5F4F3EAE7068A879E2286DFE40" blockId="16.[151,638,194,720]" pageId="16">
<emphasis id="0990704D4F3EAE7068A879E22B26FE67" bold="true" box="[175,290,433,454]" pageId="16">Etymology.</emphasis>
The name for this species was suggested by the prominent spine of the mesotibiae.
</paragraph>
</subSubSection>
<subSubSection id="73FEFFD44F3EAE7068A879B52811FD71" pageId="16" type="discussion">
<paragraph id="3B5BAC5F4F3EAE7068A879B52811FD71" blockId="16.[151,638,194,720]" pageId="16">
<emphasis id="0990704D4F3EAE7068A879B52B20FE5A" bold="true" box="[175,292,486,507]" pageId="16">Comments.</emphasis>
<taxonomicName id="FCE4D7DC4F3EAE70693679B52807FE5A" authorityName="Chandler &amp; Geromini" authorityYear="1962" box="[305,515,486,507]" class="Insecta" family="Staphylinidae" genus="Custotychus" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="0990704D4F3EAE70693679B52807FE5A" box="[305,515,486,507]" italics="true" pageId="16">Custotychus spinosus</emphasis>
</taxonomicName>
is generally similar to
<taxonomicName id="FCE4D7DC4F3EAE7068FA7A522B70FDB7" authorityName="Chandler &amp; Geromini" authorityYear="1962" box="[253,372,513,534]" class="Insecta" family="Staphylinidae" genus="Custotychus" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="rothorum">
<emphasis id="0990704D4F3EAE7068FA7A522B70FDB7" box="[253,372,513,534]" italics="true" pageId="16">C. rothorum</emphasis>
</taxonomicName>
by form and placement of the metasternal tubercle, and by the general form of the genitalia. The presence of a median spine on the mesotibiae leads to rapid recognition of this species, and the median lobe has the lateral margins gently concave in dorsal view, while
<taxonomicName id="FCE4D7DC4F3EAE706A007AD5287AFD3A" authorityName="Chandler &amp; Geromini" authorityYear="1962" box="[519,638,646,667]" class="Insecta" family="Staphylinidae" genus="Custotychus" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="rothorum">
<emphasis id="0990704D4F3EAE706A007AD5287AFD3A" box="[519,638,646,667]" italics="true" pageId="16">C. rothorum</emphasis>
</taxonomicName>
lacks the median mesotibial spine and the lateral margins of the median lobe are straight.
</paragraph>
</subSubSection>
</treatment>
</document>
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