890 lines
125 KiB
XML
890 lines
125 KiB
XML
<document ID-DOI="10.11646/zootaxa.5256.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7745281" ID-ZooBank="CE0AC7CD-E55D-484C-9346-86F16C985DBE" approvalRequired="2" approvalRequired_for_taxonomicNames="2" checkinTime="1679054310250" checkinUser="plazi" docAuthor="Serrano, Artur R. M." docDate="2023" docId="AE0487C8F52FFFC60986A38B64B7FBF0" docLanguage="en" docName="zootaxa.5256.1.1.pdf" docOrigin="Zootaxa 5256 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrocheilus Hope 1838" docType="key" docVersion="3" lastPageNumber="31" masterDocId="523DFFB0F533FFD80911A31C612BFFA8" masterDocTitle="Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes" masterLastPageNumber="35" masterPageNumber="1" pageNumber="29" updateTime="1679078510330" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes</mods:title>
|
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Serrano, Artur R. M.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0003-4282-6073</mods:nameIdentifier>
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<mods:affiliation>cE 3 c-Center for Ecology, Evolution and Environmental Changes & CHANGE - Global Change and Sustainability Institute, Departamento de Biologia Animal, Faculdade de Ciências da Universidade de Lisboa, R. Ernesto de Vasconcelos, Ed. C 2, Campo Grande, 1749 - 016 Lisboa, PORTUGAL. aserrano @ fc. ul. pt; http: // orcid. org / 0000 - 0003 - 4282 - 6073</mods:affiliation>
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<mods:nameIdentifier type="email">aserrano@fc.ul.pt</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-16</mods:number>
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||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>5256</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>35</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5256.1.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5256.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7745281</mods:identifier>
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<mods:identifier type="ZooBank">CE0AC7CD-E55D-484C-9346-86F16C985DBE</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:AE0487C8F52FFFC60986A38B64B7FBF0" httpUri="http://treatment.plazi.org/id/AE0487C8F52FFFC60986A38B64B7FBF0" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
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<subSubSection box="[151,1017,151,178]" pageId="28" pageNumber="29" type="nomenclature">
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<paragraph blockId="28.[151,1017,151,178]" box="[151,1017,151,178]" pageId="28" pageNumber="29">
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<heading bold="true" box="[151,1017,151,178]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
|
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<emphasis bold="true" box="[151,1017,151,178]" pageId="28" pageNumber="29">
|
||
Key to the Afrotropical (excluding
|
||
<collectingCountry box="[586,743,151,177]" name="Madagascar" pageId="28" pageNumber="29">Madagascar</collectingCountry>
|
||
)
|
||
<taxonomicName ID-CoL="8KWL5" ID-ENA="177436" authority="Hope, 1838" authorityName="Hope" authorityYear="1838" box="[758,924,151,178]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[758,924,151,178]" italics="true" pageId="28" pageNumber="29">Macrocheilus</emphasis>
|
||
</taxonomicName>
|
||
species
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="28" pageNumber="29" type="description">
|
||
<paragraph blockId="28.[151,1437,223,466]" pageId="28" pageNumber="29">
|
||
An identification key for the Afrotropical species/subspecies is provided, including the new species. The
|
||
<taxonomicName pageId="28" pageNumber="29">
|
||
<emphasis italics="true" pageId="28" pageNumber="29">Macrocheilus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
/subspecies from
|
||
<collectingCountry box="[514,649,259,285]" name="Madagascar" pageId="28" pageNumber="29">Madagascar</collectingCountry>
|
||
can be identified through
|
||
<bibRefCitation author="Jeannel, R." box="[944,1110,259,285]" pageId="28" pageNumber="29" pagination="767 - 1146" refId="ref25594" refString="Jeannel, R. (1949) s. n. In: Faune de L`Empire Francais. XI. Coleopteres Carabiques de la Region Malgache. T roiseme Partie. Librairie Larose, Paris, pp. 767 - 1146." type="book chapter" year="1949">Jeannel (1949)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Basilewsky, P." box="[1164,1379,259,286]" pageId="28" pageNumber="29" pagination="53 - 65" refId="ref25094" refString="Basilewsky, P. (1953) Contributions a l`etude des coleopteres carabiques de la region malgache. I. Memoires de l'Institut Scientifique de Madagascar (E), 3, 53 - 65." type="journal article" year="1953">Basilewsky (1953)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,223,466]" pageId="28" pageNumber="29">
|
||
Since the diagnostic external morphological characters are similar for males and females, the key can be used for both sexes.
|
||
<taxonomicName box="[281,569,331,357]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="basilewskyi">
|
||
<emphasis box="[281,569,331,357]" italics="true" pageId="28" pageNumber="29">Macrocheilus basilewskyi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[621,748,331,357]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="dorsalis">
|
||
<emphasis box="[621,748,331,357]" italics="true" pageId="28" pageNumber="29">M. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
are identified in any alternative dichotomic step 5 [larger species (length ≥
|
||
<quantity box="[305,385,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="mm" value="10.0">10 mm</quantity>
|
||
)
|
||
<emphasis box="[400,423,368,393]" italics="true" pageId="28" pageNumber="29">vs</emphasis>
|
||
. smaller species (length<
|
||
<quantity box="[712,796,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="mm" value="10.0">10 mm</quantity>
|
||
)], since both can present specimens within these options. After reaching the identification of a species, other characters can be added, reinforcing its diagnosis (e.g. labrum and/or pronotum shapes/features, elytra patches, male and/or female genitalia).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="key">
|
||
<keyStep pageId="28" pageNumber="29">
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">
|
||
1. Last maxillary palpomere barrel twisted shaped; labrum asymmetric, with a pair of marginal or submarginal lateral setae in the middle (
|
||
<figureCitation box="[305,379,537,559]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="28" pageNumber="29">
|
||
Figs
|
||
<quantity box="[354,379,538,559]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="28" pageNumber="29" unit="g" value="1.0">1g</quantity>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation box="[390,415,538,559]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="28" pageNumber="29">2e</figureCitation>
|
||
,
|
||
<figureCitation box="[425,444,537,559]" captionStart="FIGURE 5" captionStartId="17.[152,255,1916,1940]" captionTargetBox="[178,1402,165,1877]" captionTargetId="figure-83@17.[166,1418,151,1890]" captionTargetPageId="17" captionText="FIGURE 5. Habitus of Macrocheilus (dorsal view). a, M. biguttatus, SEN (ACC); b, M. bimaculatus, GBI (ASC); c, M. clasispilus (holotype, plus labels); d, M. diplospilus (holotype, plus labels); e, M. dorsalis, ZAM (TAC); f, M. dorsiger, BFO (PSC); g, M. elegantulus (holotype, plus labels); h, M. hybridus, ZIM (TAC); i, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745293" httpUri="https://zenodo.org/record/7745293/files/figure.png" pageId="28" pageNumber="29">5i</figureCitation>
|
||
,
|
||
<figureCitation box="[455,478,537,559]" captionStart="FIGURE 6" captionStartId="18.[152,255,1916,1940]" captionTargetBox="[171,1427,204,1867]" captionTargetId="figure-60@18.[151,1436,194,1885]" captionTargetPageId="18" captionText="FIGURE 6. Labrum of Macrocheilus (dorsal view). a, M. diplospilus, GBI (ASC); b, M. dorsalis, ZAM (TAC); c, M. dorsiger, BFO (PSC); d, M. hybridus, ZIM (TAC); e, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745295" httpUri="https://zenodo.org/record/7745295/files/figure.png" pageId="28" pageNumber="29">6e</figureCitation>
|
||
)............................................................................. 2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">
|
||
- Last maxillary palpomere more or less securiform shaped; labrum symmetric, with more than one pair of setae (e.g.
|
||
<figureCitation box="[1358,1432,567,589]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="28" pageNumber="29">Figs 1b</figureCitation>
|
||
,
|
||
<figureCitation box="[226,249,596,617]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="28" pageNumber="29">2a</figureCitation>
|
||
)................................................................................................ 3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="28" pageNumber="29">
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">
|
||
2. Elytra black, intervals smooth, one pair of a more or less ovoid orange-yellowish spots in the end of the basal half throughout the 3
|
||
<superScript attach="left" box="[275,287,654,666]" fontSize="5" pageId="28" pageNumber="29">rd</superScript>
|
||
to 6
|
||
<superScript attach="left" box="[332,343,654,666]" fontSize="5" pageId="28" pageNumber="29">th</superScript>
|
||
or 7
|
||
<superScript attach="left" box="[390,401,654,666]" fontSize="5" pageId="28" pageNumber="29">th</superScript>
|
||
intervals; head and pronotum more densely punctured; posterior angles of pronotum slightly dentiform at apex, with a slight short expansion just before the angles; male and female genitalia as in figures 7c and 8d, respectively..................................................................................
|
||
<taxonomicName authority="(Dejean, 1831)" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[1172,1436,713,735]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="labrosus">
|
||
<emphasis box="[1172,1286,713,735]" italics="true" pageId="28" pageNumber="29">M. labrosus</emphasis>
|
||
(Dejean, 1831)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">
|
||
- Elytra totally black, intervals with a very thin polygonal microreticulation; head and pronotum less densely punctured; posterior angles of pronotum rounded, without any expansion before the angles; male genitalia as in figure 3d..................................................................................................
|
||
<taxonomicName authority="Burgeon, 1937" authorityName="Burgeon" authorityYear="1937" box="[1176,1436,801,823]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="alluaudi">
|
||
<emphasis box="[1176,1288,801,823]" italics="true" pageId="28" pageNumber="29">M. alluaudi</emphasis>
|
||
<bibRefCitation author="Burgeon, L." box="[1294,1436,802,823]" pageId="28" pageNumber="29" pagination="319 - 405" refId="ref25317" refString="Burgeon, L. (1937) Catalogues raisonnes de la faune entomologique du Congo Belge. Coleopteres Carabides. Troisieme Partie. Annales du Musee du Congo Belge, Zoology, Serie 3 (2), 2 (5), 319 - 405." type="journal article" year="1937">Burgeon, 1937</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="28" pageNumber="29">
|
||
<paragraph blockId="28.[151,1437,508,2026]" box="[151,1436,831,853]" pageId="28" pageNumber="29">
|
||
<keyLead box="[151,1436,831,853]" pageId="28" pageNumber="29">3. Elytra microreticulate.................................................................................. 4</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,508,2026]" box="[151,1436,860,882]" pageId="28" pageNumber="29">
|
||
<keyLead box="[151,1436,860,882]" pageId="28" pageNumber="29">- Elytra smooth, without microreticulation................................................................... 7</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="28" pageNumber="29">
|
||
<paragraph blockId="28.[151,1437,508,2026]" box="[151,1436,889,911]" pageId="28" pageNumber="29">
|
||
<keyLead box="[151,1436,889,911]" pageId="28" pageNumber="29">4. Posterior angles of pronotum slightly obtuse; elytra moderately microreticulate.................................... 5</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,508,2026]" box="[151,1436,919,941]" pageId="28" pageNumber="29">
|
||
<keyLead box="[151,1436,919,941]" pageId="28" pageNumber="29">- Posterior angles of pronotum strongly obtuse; elytra strongly microreticulate..................................... 6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="28" pageNumber="29">
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">
|
||
5. Head, pronotum and elytra black; posterior angles of pronotum slightly dentiform at apex; one pair of orange spots jagged in the median discal zone (
|
||
<figureCitation box="[458,540,977,999]" captionStart="FIGURE 15" captionStartId="31.[152,255,1943,1967]" captionTargetBox="[245,1343,189,1903]" captionTargetId="figure-16@31.[221,1366,181,1920]" captionTargetPageId="31" captionText="FIGURE 15. Habitus of Macrocheilus (dorsal view). a, M. angustatus (holotype, plus labels); b, M. biplagiatus (MRAC, plus labels); c, M. burgeoni (holotype, plus labels); d, M. crampeli (paratype, plus labels); e, M. cruciatus (MRAC, plus labels); f, M. moraisi (holotype, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745315" httpUri="https://zenodo.org/record/7745315/files/figure.png" pageId="28" pageNumber="29">Fig. 15d</figureCitation>
|
||
)....................................................
|
||
<taxonomicName authority="Alluaud, 1916" authorityName="Alluaud" authorityYear="1916" box="[1177,1436,977,999]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="crampeli">
|
||
<emphasis box="[1177,1294,977,999]" italics="true" pageId="28" pageNumber="29">M. crampeli</emphasis>
|
||
Alluaud, 1916
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">
|
||
- Head and pronotum dark brown, almost black; elytra dark brown-reddish; posterior angles of pronotum slightly rounded at apex; one pair of orange spots round shaped in the median discal zone, with a black hole (
|
||
<figureCitation box="[1089,1169,1036,1058]" captionStart="FIGURE 16" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[242,1350,192,1911]" captionTargetId="figure-19@32.[226,1362,181,1920]" captionTargetPageId="32" captionText="FIGURE 16. Habitus of Macrocheilus (dorsal view). a, M. ocellatus (holotype, plus labels); b, M. overlaeti (holotype, plus labels); c, M. varians varians (holotype); d, M. varians eugrammus (holotype); e, M. vinctus (paratype)." figureDoi="http://doi.org/10.5281/zenodo.7745317" httpUri="https://zenodo.org/record/7745317/files/figure.png" pageId="28" pageNumber="29">Fig. 16a</figureCitation>
|
||
)................................................................................................
|
||
<taxonomicName authority="Basilewsky, 1953" authorityName="Basilewsky" authorityYear="1953" box="[1142,1436,1065,1087]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ocellatus">
|
||
<emphasis box="[1142,1261,1065,1087]" italics="true" pageId="28" pageNumber="29">M. ocellatus</emphasis>
|
||
<bibRefCitation author="Basilewsky, P." box="[1267,1436,1065,1087]" pageId="28" pageNumber="29" pagination="53 - 65" refId="ref25094" refString="Basilewsky, P. (1953) Contributions a l`etude des coleopteres carabiques de la region malgache. I. Memoires de l'Institut Scientifique de Madagascar (E), 3, 53 - 65." type="journal article" year="1953">Basilewsky, 1953</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="28" pageNumber="29">
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">
|
||
6. Legs brown, or at least coxae, trochanters and femora brown; pronotum strongly sinuate just anterior to posterior angles (
|
||
<figureCitation captionStart="FIGURE 11" captionStartId="25.[152,255,1923,1947]" captionTargetBox="[359,1259,530,1904]" captionTargetId="figure-221@25.[333,1270,517,1916]" captionTargetPageId="25" captionText="FIGURE 11. Habitus of Macrocheilus (dorsal view). a, M. scapularis (MRAC, plus labels); b, M. spectandus (holotype); c, M. spectandus, TAN (TAC); d, M. taedatus (paratype); e, M. taedatus, TAN (TAC); f, M. viduatus katanganus (holotype, plus labels); g, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745305" httpUri="https://zenodo.org/record/7745305/files/figure.png" pageId="28" pageNumber="29">Figs 11f, g</figureCitation>
|
||
); anterior half of labrum strongly ogive shaped, three marginal/submarginal pairs of setae (
|
||
<figureCitation box="[1172,1257,1124,1146]" captionStart="FIGURE 12" captionStartId="26.[152,255,1275,1299]" captionTargetBox="[176,1405,161,1252]" captionTargetId="figure-327@26.[166,1413,151,1264]" captionTargetPageId="26" captionText="FIGURE 12. Labrum of Macrocheilus (dorsal view). a, M. scapularis (MRAC); b, M. spectandus, TAN (TAC); c, M. taedatus, TAN (TAC); d, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745307" httpUri="https://zenodo.org/record/7745307/files/figure.png" pageId="28" pageNumber="29">Fig. 12d</figureCitation>
|
||
); male and female genitalia as in figures 13d and 14d, respectively (in the nominotypical subspecies the legs are totally black)................................................................................
|
||
<taxonomicName authority="Basilewsky, 1960" authorityName="Basilewsky" authorityYear="1960" box="[1031,1436,1183,1205]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="viduatus" subSpecies="katanganus">
|
||
<emphasis box="[1031,1261,1183,1204]" italics="true" pageId="28" pageNumber="29">M. viduatus katanganus</emphasis>
|
||
<bibRefCitation author="Basilewsky, P." box="[1267,1436,1183,1205]" pageId="28" pageNumber="29" pagination="39 - 51" refId="ref25130" refString="Basilewsky, P. (1960) Sur le genre Macrocheilus Hope (Col. Carabidae Helluoninae). Bulletin et Annales de la Societe Royale d`Entomologie de Belgique, 96 (3 - 4), 39 - 51." type="journal article" year="1960">Basilewsky, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">
|
||
- Legs black; pronotum less sinuate just anterior to posterior angles (
|
||
<figureCitation box="[883,956,1212,1234]" captionStart="FIGURE 5" captionStartId="17.[152,255,1916,1940]" captionTargetBox="[178,1402,165,1877]" captionTargetId="figure-83@17.[166,1418,151,1890]" captionTargetPageId="17" captionText="FIGURE 5. Habitus of Macrocheilus (dorsal view). a, M. biguttatus, SEN (ACC); b, M. bimaculatus, GBI (ASC); c, M. clasispilus (holotype, plus labels); d, M. diplospilus (holotype, plus labels); e, M. dorsalis, ZAM (TAC); f, M. dorsiger, BFO (PSC); g, M. elegantulus (holotype, plus labels); h, M. hybridus, ZIM (TAC); i, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745293" httpUri="https://zenodo.org/record/7745293/files/figure.png" pageId="28" pageNumber="29">Fig. 5b</figureCitation>
|
||
); anterior half of labrum less ogive shaped, three marginal/submarginal pairs of setae (
|
||
<figureCitation box="[581,653,1241,1263]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="28" pageNumber="29">Fig. 2h</figureCitation>
|
||
); male genitalia as in
|
||
<bibRefCitation author="Serrano, A. R. M." box="[863,1002,1242,1263]" pageId="28" pageNumber="29" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano (2000</bibRefCitation>
|
||
,
|
||
<figureCitation box="[1012,1103,1241,1263]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="7.[152,255,1916,1940]" captionStartId-1="8.[152,255,1463,1487]" captionTargetBox-0="[165,1321,160,1875]" captionTargetBox-1="[162,1421,162,1440]" captionTargetId-0="figure-80@7.[151,1340,151,1890]" captionTargetId-1="figure-282@8.[151,1436,151,1453]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." captionText-1="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi-0="http://doi.org/10.5281/zenodo.7745283" figureDoi-1="http://doi.org/10.5281/zenodo.7745285" httpUri-0="https://zenodo.org/record/7745283/files/figure.png" httpUri-1="https://zenodo.org/record/7745285/files/figure.png" pageId="28" pageNumber="29">Figs. 1–2</figureCitation>
|
||
); female genitalia as in figure 8a............................................................................
|
||
<taxonomicName authority="(Dejean, 1831)" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[1137,1436,1271,1293]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bimaculatus">
|
||
<emphasis box="[1137,1286,1271,1292]" italics="true" pageId="28" pageNumber="29">M. bimaculatus</emphasis>
|
||
(Dejean, 1831)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="28" pageNumber="29">
|
||
<paragraph blockId="28.[151,1437,508,2026]" box="[151,1436,1300,1322]" pageId="28" pageNumber="29">
|
||
<keyLead box="[151,1436,1300,1322]" pageId="28" pageNumber="29">
|
||
7. Larger species (length ≥
|
||
<quantity box="[458,527,1300,1321]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="mm" value="10.0">10 mm</quantity>
|
||
)......................................................................... 8
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,508,2026]" box="[151,1436,1329,1351]" pageId="28" pageNumber="29">
|
||
<keyLead box="[151,1436,1329,1351]" pageId="28" pageNumber="29">
|
||
- Smaller species (length <
|
||
<quantity box="[470,538,1330,1351]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="mm" value="10.0">10 mm</quantity>
|
||
)....................................................................... 29
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="28" pageNumber="29">
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">8. Head black and pronotum ferruginous or head and pronotum black or dark brown; elytra black or dark brown with orange or yellowish spots...................................................................................... 9</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,508,2026]" box="[151,1436,1417,1439]" pageId="28" pageNumber="29">
|
||
<keyLead box="[151,1436,1417,1439]" pageId="28" pageNumber="29">- Head and pronotum ferruginous, orange or brownish; elytra with dark patches and orange or yellowish spots........... 17</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="28" pageNumber="29">
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">
|
||
9. Head black and pronotum ferruginous; pronotum slightly transverse; elytra black with a pair of ovoid orange-yellowish spots in the end of first half (
|
||
<figureCitation box="[448,529,1476,1498]" captionStart="FIGURE 16" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[242,1350,192,1911]" captionTargetId="figure-19@32.[226,1362,181,1920]" captionTargetPageId="32" captionText="FIGURE 16. Habitus of Macrocheilus (dorsal view). a, M. ocellatus (holotype, plus labels); b, M. overlaeti (holotype, plus labels); c, M. varians varians (holotype); d, M. varians eugrammus (holotype); e, M. vinctus (paratype)." figureDoi="http://doi.org/10.5281/zenodo.7745317" httpUri="https://zenodo.org/record/7745317/files/figure.png" pageId="28" pageNumber="29">Fig. 16b</figureCitation>
|
||
)....................................................
|
||
<taxonomicName authority="Burgeon, 1937" authorityName="Burgeon" authorityYear="1937" box="[1171,1436,1476,1498]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="overlaeti">
|
||
<emphasis box="[1171,1288,1476,1498]" italics="true" pageId="28" pageNumber="29">M. overlaeti</emphasis>
|
||
<bibRefCitation author="Burgeon, L." box="[1294,1436,1476,1498]" pageId="28" pageNumber="29" pagination="319 - 405" refId="ref25317" refString="Burgeon, L. (1937) Catalogues raisonnes de la faune entomologique du Congo Belge. Coleopteres Carabides. Troisieme Partie. Annales du Musee du Congo Belge, Zoology, Serie 3 (2), 2 (5), 319 - 405." type="journal article" year="1937">Burgeon, 1937</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,508,2026]" box="[151,1436,1505,1527]" pageId="28" pageNumber="29">
|
||
<keyLead box="[151,1436,1505,1527]" pageId="28" pageNumber="29">- Head and pronotum black or dark brown; elytra black or dark brown with orange or yellowish spots.................. 10</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="28" pageNumber="29">
|
||
<paragraph blockId="28.[151,1437,508,2026]" box="[151,1436,1535,1557]" pageId="28" pageNumber="29">
|
||
<keyLead box="[151,1436,1535,1557]" pageId="28" pageNumber="29">10. Pronotum slightly transverse (1.1–1.2 times wider than long); elytra slender...................................... 11</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,508,2026]" box="[151,1436,1564,1586]" pageId="28" pageNumber="29">
|
||
<keyLead box="[151,1436,1564,1586]" pageId="28" pageNumber="29">- Pronotum more transverse (1.45–1.75 times wider than long); elytra broader..................................... 12</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="28" pageNumber="29">
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">
|
||
11. Posterior angles of pronotum strongly obtuse, sharpish, without expansion before the angles; elytra 1.9 times longer than wide, one pair of orange spots in the median discal zone (
|
||
<figureCitation box="[719,791,1623,1645]" captionStart="FIGURE 5" captionStartId="17.[152,255,1916,1940]" captionTargetBox="[178,1402,165,1877]" captionTargetId="figure-83@17.[166,1418,151,1890]" captionTargetPageId="17" captionText="FIGURE 5. Habitus of Macrocheilus (dorsal view). a, M. biguttatus, SEN (ACC); b, M. bimaculatus, GBI (ASC); c, M. clasispilus (holotype, plus labels); d, M. diplospilus (holotype, plus labels); e, M. dorsalis, ZAM (TAC); f, M. dorsiger, BFO (PSC); g, M. elegantulus (holotype, plus labels); h, M. hybridus, ZIM (TAC); i, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745293" httpUri="https://zenodo.org/record/7745293/files/figure.png" pageId="28" pageNumber="29">Fig. 5h</figureCitation>
|
||
); anterior half of labrum ogive shaped, two pairs of marginal setae, one pair of setae in the middle of the disc, margins of posterior half subparallel (
|
||
<figureCitation box="[996,1068,1652,1674]" captionStart="FIGURE 6" captionStartId="18.[152,255,1916,1940]" captionTargetBox="[171,1427,204,1867]" captionTargetId="figure-60@18.[151,1436,194,1885]" captionTargetPageId="18" captionText="FIGURE 6. Labrum of Macrocheilus (dorsal view). a, M. diplospilus, GBI (ASC); b, M. dorsalis, ZAM (TAC); c, M. dorsiger, BFO (PSC); d, M. hybridus, ZIM (TAC); e, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745295" httpUri="https://zenodo.org/record/7745295/files/figure.png" pageId="28" pageNumber="29">Fig. 6d</figureCitation>
|
||
); female genitalia as in figure 8c................................................................................
|
||
<taxonomicName authority="Peringuey, 1896" authorityName="Peringuey" authorityYear="1896" box="[1158,1436,1681,1703]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hybridus">
|
||
<emphasis box="[1158,1274,1681,1703]" italics="true" pageId="28" pageNumber="29">M. hybridus</emphasis>
|
||
<bibRefCitation author="Peringuey, L." box="[1280,1436,1681,1703]" pageId="28" pageNumber="29" pagination="125 - 623" refId="ref25799" refString="Peringuey, L. (1896) A descriptive catalogue of the Coleoptera of South Africa. Part II. Carabidae. Transactions of the South African Philosophical Society, 7, 125 - 623, pls. III - X. https: // doi. org / 10.5962 / bhl. title. 9230" type="journal article" year="1896">Péringuey, 1896</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">
|
||
- Posterior angles of pronotum obtuse, rounded at apex, a slight short expansion just before the angles; elytra 2.1 times longer than wide, one pair of orange spots in the median discal zone (
|
||
<figureCitation box="[826,906,1740,1762]" captionStart="FIGURE 15" captionStartId="31.[152,255,1943,1967]" captionTargetBox="[245,1343,189,1903]" captionTargetId="figure-16@31.[221,1366,181,1920]" captionTargetPageId="31" captionText="FIGURE 15. Habitus of Macrocheilus (dorsal view). a, M. angustatus (holotype, plus labels); b, M. biplagiatus (MRAC, plus labels); c, M. burgeoni (holotype, plus labels); d, M. crampeli (paratype, plus labels); e, M. cruciatus (MRAC, plus labels); f, M. moraisi (holotype, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745315" httpUri="https://zenodo.org/record/7745315/files/figure.png" pageId="28" pageNumber="29">Fig. 15a</figureCitation>
|
||
).................
|
||
<taxonomicName authority="Basilewsky, 1949" authorityName="Basilewsky" authorityYear="1949" box="[1125,1436,1740,1762]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="angustatus">
|
||
<emphasis box="[1125,1261,1741,1762]" italics="true" pageId="28" pageNumber="29">M. angustatus</emphasis>
|
||
Basilewsky, 1949
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="28" pageNumber="29">
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">
|
||
12. Legs with trochanters and femora brown, tibia and tarsi blackish; anterior half of labrum ogive shaped, two marginal pairs of setae, one submarginal pair of setae (
|
||
<figureCitation box="[577,648,1799,1821]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="28" pageNumber="29">
|
||
Fig.
|
||
<quantity box="[621,648,1799,1820]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="28" pageNumber="29" unit="g" value="2.0">2g</quantity>
|
||
</figureCitation>
|
||
); posterior angles of pronotum strongly obtuse, dentiform, a slight short expansion just before the angles; male and female genitalia as in figures 3e and 4e,f, respectively.........
|
||
<taxonomicName authority="(Gory, 1832)" baseAuthorityName="Gory" baseAuthorityYear="1832" box="[1178,1436,1828,1850]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="biguttatus">
|
||
<emphasis box="[1178,1306,1828,1850]" italics="true" pageId="28" pageNumber="29">M. biguttatus</emphasis>
|
||
(Gory, 1832)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,508,2026]" box="[151,1436,1857,1879]" pageId="28" pageNumber="29">
|
||
<keyLead box="[151,1436,1857,1879]" pageId="28" pageNumber="29">- Legs totally black; body slightly or densely punctured....................................................... 13</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="28" pageNumber="29">
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">
|
||
13. Pronotum strongly sinuate just anterior to posterior angles, which are strongly obtuse, not dentiform at apex; elytra with one pair of large orange or yellowish spots (3
|
||
<superScript attach="left" box="[620,632,1915,1927]" fontSize="5" pageId="28" pageNumber="29">rd</superScript>
|
||
to 7
|
||
<superScript attach="left" box="[674,685,1915,1927]" fontSize="5" pageId="28" pageNumber="29">th</superScript>
|
||
intervals) (
|
||
<figureCitation box="[794,866,1916,1938]" captionStart="FIGURE 5" captionStartId="17.[152,255,1916,1940]" captionTargetBox="[178,1402,165,1877]" captionTargetId="figure-83@17.[166,1418,151,1890]" captionTargetPageId="17" captionText="FIGURE 5. Habitus of Macrocheilus (dorsal view). a, M. biguttatus, SEN (ACC); b, M. bimaculatus, GBI (ASC); c, M. clasispilus (holotype, plus labels); d, M. diplospilus (holotype, plus labels); e, M. dorsalis, ZAM (TAC); f, M. dorsiger, BFO (PSC); g, M. elegantulus (holotype, plus labels); h, M. hybridus, ZIM (TAC); i, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745293" httpUri="https://zenodo.org/record/7745293/files/figure.png" pageId="28" pageNumber="29">
|
||
Fig.
|
||
<quantity box="[839,866,1916,1938]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="28" pageNumber="29" unit="g" value="5.0">5g</quantity>
|
||
</figureCitation>
|
||
); anterior half of labrum ogive shaped, three marginal pairs of setae; male genitalia as in figure 7b.............................................
|
||
<taxonomicName authority="Burgeon, 1937" authorityName="Burgeon" authorityYear="1937" box="[1145,1436,1945,1967]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="elegantulus">
|
||
<emphasis box="[1145,1288,1945,1967]" italics="true" pageId="28" pageNumber="29">M. elegantulus</emphasis>
|
||
<bibRefCitation author="Burgeon, L." box="[1294,1436,1946,1967]" pageId="28" pageNumber="29" pagination="319 - 405" refId="ref25317" refString="Burgeon, L. (1937) Catalogues raisonnes de la faune entomologique du Congo Belge. Coleopteres Carabides. Troisieme Partie. Annales du Musee du Congo Belge, Zoology, Serie 3 (2), 2 (5), 319 - 405." type="journal article" year="1937">Burgeon, 1937</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="28.[151,1437,508,2026]" pageId="28" pageNumber="29">
|
||
<keyLead pageId="28" pageNumber="29">- Pronotum not strongly sinuate just anterior to posterior angles which are strongly obtuse, dentiform or rounded; elytra with one pair of smaller orange or yellowish spots................................................................. 14</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
14. Anterior half of labrum round shaped, two marginal pairs of setae and one discal pair of setae (
|
||
<figureCitation box="[1149,1220,151,173]" captionStart="FIGURE 6" captionStartId="18.[152,255,1916,1940]" captionTargetBox="[171,1427,204,1867]" captionTargetId="figure-60@18.[151,1436,194,1885]" captionTargetPageId="18" captionText="FIGURE 6. Labrum of Macrocheilus (dorsal view). a, M. diplospilus, GBI (ASC); b, M. dorsalis, ZAM (TAC); c, M. dorsiger, BFO (PSC); d, M. hybridus, ZIM (TAC); e, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745295" httpUri="https://zenodo.org/record/7745295/files/figure.png" pageId="29" pageNumber="30">Fig. 6a</figureCitation>
|
||
); pronotum transverse (1.4 times wider than long), posterior angles of pronotum rounded; elytral intervals almost flat, second interval with two rows of punctures; male genitalia as in figure
|
||
<specimenCount box="[613,649,210,232]" pageId="29" pageNumber="30" type="generic">3f.</specimenCount>
|
||
.......................................
|
||
<taxonomicName authority="Basilewsky, 1967" authorityName="Basilewsky" authorityYear="1967" box="[1124,1436,210,232]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="diplospilus">
|
||
<emphasis box="[1124,1261,210,232]" italics="true" pageId="29" pageNumber="30">M. diplospilus</emphasis>
|
||
<bibRefCitation author="Basilewsky, P." box="[1267,1436,210,232]" pageId="29" pageNumber="30" pagination="319 - 330" refId="ref25218" refString="Basilewsky, P. (1967) Coleopteres Carabidae africains nouveaux. XVI. Revue de Zoologie et Botanique Africaines, 75 (3 - 4), 319 - 330." type="journal article" year="1967">Basilewsky, 1967</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" box="[151,1436,239,261]" pageId="29" pageNumber="30">
|
||
<keyLead box="[151,1436,239,261]" pageId="29" pageNumber="30">
|
||
- Anterior half of labrum ogive shaped, three marginal pairs of setae (e.g.
|
||
<figureCitation box="[921,984,239,261]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="29" pageNumber="30">Fig. 2i</figureCitation>
|
||
).................................. 15
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
15. Head and pronotum not so densely punctured; posterior angles of pronotum slightly obtuse, dentiform at apex, a slight expansion before the posterior angles; elytra with one pair of small orange spots (3
|
||
<superScript attach="left" box="[970,982,297,309]" fontSize="5" pageId="29" pageNumber="30">rd</superScript>
|
||
to 6
|
||
<superScript attach="left" box="[1026,1037,297,309]" fontSize="5" pageId="29" pageNumber="30">th</superScript>
|
||
intervals) (
|
||
<figureCitation box="[1147,1230,298,320]" captionStart="FIGURE 15" captionStartId="31.[152,255,1943,1967]" captionTargetBox="[245,1343,189,1903]" captionTargetId="figure-16@31.[221,1366,181,1920]" captionTargetPageId="31" captionText="FIGURE 15. Habitus of Macrocheilus (dorsal view). a, M. angustatus (holotype, plus labels); b, M. biplagiatus (MRAC, plus labels); c, M. burgeoni (holotype, plus labels); d, M. crampeli (paratype, plus labels); e, M. cruciatus (MRAC, plus labels); f, M. moraisi (holotype, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745315" httpUri="https://zenodo.org/record/7745315/files/figure.png" pageId="29" pageNumber="30">Fig. 15c</figureCitation>
|
||
); male genitalia as in
|
||
<bibRefCitation author="Basilewsky, P." box="[226,401,327,349]" pageId="29" pageNumber="30" pagination="319 - 330" refId="ref25218" refString="Basilewsky, P. (1967) Coleopteres Carabidae africains nouveaux. XVI. Revue de Zoologie et Botanique Africaines, 75 (3 - 4), 319 - 330." type="journal article" year="1967">Basilewsky (1967</bibRefCitation>
|
||
,
|
||
<figureCitation box="[411,488,327,349]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="29" pageNumber="30">Figs. 2c</figureCitation>
|
||
,
|
||
<figureCitation box="[499,522,328,349]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="29" pageNumber="30">3c</figureCitation>
|
||
)..................................................
|
||
<taxonomicName authority="Basilewsky, 1967" authorityName="Basilewsky" authorityYear="1967" box="[1143,1436,327,349]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="burgeoni">
|
||
<emphasis box="[1143,1261,327,349]" italics="true" pageId="29" pageNumber="30">M. burgeoni</emphasis>
|
||
<bibRefCitation author="Basilewsky, P." box="[1267,1436,327,349]" pageId="29" pageNumber="30" pagination="319 - 330" refId="ref25218" refString="Basilewsky, P. (1967) Coleopteres Carabidae africains nouveaux. XVI. Revue de Zoologie et Botanique Africaines, 75 (3 - 4), 319 - 330." type="journal article" year="1967">Basilewsky, 1967</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" box="[151,1436,357,379]" pageId="29" pageNumber="30">
|
||
<keyLead box="[151,1436,357,379]" pageId="29" pageNumber="30">- Head and pronotum strongly and densely punctured; male genitalia with a different shape.......................... 16</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
16. Elytra with one pair of small orange spots (3
|
||
<superScript attach="left" box="[658,670,385,397]" fontSize="5" pageId="29" pageNumber="30">rd</superScript>
|
||
to 7
|
||
<superScript attach="left" box="[714,725,385,397]" fontSize="5" pageId="29" pageNumber="30">th</superScript>
|
||
intervals) (
|
||
<figureCitation box="[835,905,386,408]" captionStart="FIGURE 5" captionStartId="17.[152,255,1916,1940]" captionTargetBox="[178,1402,165,1877]" captionTargetId="figure-83@17.[166,1418,151,1890]" captionTargetPageId="17" captionText="FIGURE 5. Habitus of Macrocheilus (dorsal view). a, M. biguttatus, SEN (ACC); b, M. bimaculatus, GBI (ASC); c, M. clasispilus (holotype, plus labels); d, M. diplospilus (holotype, plus labels); e, M. dorsalis, ZAM (TAC); f, M. dorsiger, BFO (PSC); g, M. elegantulus (holotype, plus labels); h, M. hybridus, ZIM (TAC); i, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745293" httpUri="https://zenodo.org/record/7745293/files/figure.png" pageId="29" pageNumber="30">Fig. 5c</figureCitation>
|
||
); Male genitalia as in
|
||
<bibRefCitation author="Basilewsky, P." box="[1121,1297,386,408]" pageId="29" pageNumber="30" pagination="319 - 330" refId="ref25218" refString="Basilewsky, P. (1967) Coleopteres Carabidae africains nouveaux. XVI. Revue de Zoologie et Botanique Africaines, 75 (3 - 4), 319 - 330." type="journal article" year="1967">Basilewsky (1967</bibRefCitation>
|
||
,
|
||
<figureCitation box="[1307,1387,386,408]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="29" pageNumber="30">Figs. 2b</figureCitation>
|
||
,
|
||
<figureCitation box="[1398,1424,386,408]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="29" pageNumber="30">3b</figureCitation>
|
||
); female genitalia as in figure 8b.................................................
|
||
<taxonomicName authority="Basilewsky, 1967" authorityName="Basilewsky" authorityYear="1967" box="[1128,1436,415,437]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="clasispilus">
|
||
<emphasis box="[1128,1261,415,437]" italics="true" pageId="29" pageNumber="30">M. clasispilus</emphasis>
|
||
<bibRefCitation author="Basilewsky, P." box="[1267,1436,415,437]" pageId="29" pageNumber="30" pagination="319 - 330" refId="ref25218" refString="Basilewsky, P. (1967) Coleopteres Carabidae africains nouveaux. XVI. Revue de Zoologie et Botanique Africaines, 75 (3 - 4), 319 - 330." type="journal article" year="1967">Basilewsky, 1967</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
- Elytra with one pair of small orange spots (3
|
||
<superScript attach="left" box="[654,666,444,456]" fontSize="5" pageId="29" pageNumber="30">rd</superScript>
|
||
to 6
|
||
<superScript attach="left" box="[708,719,444,456]" fontSize="5" pageId="29" pageNumber="30">th</superScript>
|
||
intervals) (
|
||
<figureCitation box="[829,911,445,467]" captionStart="FIGURE 16" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[242,1350,192,1911]" captionTargetId="figure-19@32.[226,1362,181,1920]" captionTargetPageId="32" captionText="FIGURE 16. Habitus of Macrocheilus (dorsal view). a, M. ocellatus (holotype, plus labels); b, M. overlaeti (holotype, plus labels); c, M. varians varians (holotype); d, M. varians eugrammus (holotype); e, M. vinctus (paratype)." figureDoi="http://doi.org/10.5281/zenodo.7745317" httpUri="https://zenodo.org/record/7745317/files/figure.png" pageId="29" pageNumber="30">Fig. 16e</figureCitation>
|
||
); Male genitalia as in
|
||
<bibRefCitation author="Basilewsky, P." box="[1124,1298,445,467]" pageId="29" pageNumber="30" pagination="319 - 330" refId="ref25218" refString="Basilewsky, P. (1967) Coleopteres Carabidae africains nouveaux. XVI. Revue de Zoologie et Botanique Africaines, 75 (3 - 4), 319 - 330." type="journal article" year="1967">Basilewsky (1967</bibRefCitation>
|
||
,
|
||
<figureCitation box="[1308,1386,445,467]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="29" pageNumber="30">Figs. 2a</figureCitation>
|
||
,
|
||
<figureCitation box="[1396,1420,445,466]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="29" pageNumber="30">3a</figureCitation>
|
||
).............................................................................
|
||
<taxonomicName authority="Basilewsky, 1960" authorityName="Basilewsky" authorityYear="1960" box="[1161,1436,474,496]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vinctus">
|
||
<emphasis box="[1161,1261,475,496]" italics="true" pageId="29" pageNumber="30">M. vinctus</emphasis>
|
||
<bibRefCitation author="Basilewsky, P." box="[1267,1436,474,496]" pageId="29" pageNumber="30" pagination="39 - 51" refId="ref25130" refString="Basilewsky, P. (1960) Sur le genre Macrocheilus Hope (Col. Carabidae Helluoninae). Bulletin et Annales de la Societe Royale d`Entomologie de Belgique, 96 (3 - 4), 39 - 51." type="journal article" year="1960">Basilewsky, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" box="[151,1436,503,525]" pageId="29" pageNumber="30">
|
||
<keyLead box="[151,1436,503,525]" pageId="29" pageNumber="30">
|
||
17. Elytra with medium to large orange or yellowish spots (e.g.
|
||
<figureCitation box="[803,872,503,525]" captionStart="FIGURE 9" captionStartId="22.[152,255,1916,1940]" captionTargetBox="[238,1445,163,1876]" captionTargetId="figure-82@22.[219,1454,151,1889]" captionTargetPageId="22" captionText="FIGURE 9. Habitus of Macrocheilus (dorsal view). a, M. longicollis (holotype); b, M. longicollis, MOZ (PSC); c, M. longicollis, ZAM (TAC); d, M. persimilis (holotype, plus labels); e, M. persimilis, ZAM (TAC); f, M. proximus (holotype); g, M. proximus, ETI (TAC); h, M. quadrinotatus (holotype, plus labels); i, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745301" httpUri="https://zenodo.org/record/7745301/files/figure.png" pageId="29" pageNumber="30">Fig. 9b</figureCitation>
|
||
)............................................ 18
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" box="[151,1436,533,555]" pageId="29" pageNumber="30">
|
||
<keyLead box="[151,1436,533,555]" pageId="29" pageNumber="30">
|
||
- Elytra with small orange or yellowish spots (e.g.
|
||
<figureCitation box="[698,777,533,555]" captionStart="FIGURE 11" captionStartId="25.[152,255,1923,1947]" captionTargetBox="[359,1259,530,1904]" captionTargetId="figure-221@25.[333,1270,517,1916]" captionTargetPageId="25" captionText="FIGURE 11. Habitus of Macrocheilus (dorsal view). a, M. scapularis (MRAC, plus labels); b, M. spectandus (holotype); c, M. spectandus, TAN (TAC); d, M. taedatus (paratype); e, M. taedatus, TAN (TAC); f, M. viduatus katanganus (holotype, plus labels); g, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745305" httpUri="https://zenodo.org/record/7745305/files/figure.png" pageId="29" pageNumber="30">Fig. 11b</figureCitation>
|
||
)................................................... 27
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" box="[151,1436,562,584]" pageId="29" pageNumber="30">
|
||
<keyLead box="[151,1436,562,584]" pageId="29" pageNumber="30">18. Elytra with one pair of spots........................................................................... 19</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" box="[151,1436,591,613]" pageId="29" pageNumber="30">
|
||
<keyLead box="[151,1436,591,613]" pageId="29" pageNumber="30">- Elytra with two pairs of spots, separated or coalescent....................................................... 20</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
19. Anterior half of labrum ogive shaped; pronotum as wide as long, not heart-shaped; elytra with orange spots as in figure 1e...............................................................................
|
||
<taxonomicName authority="Basilewsky, 1957" authorityName="Basilewsky" authorityYear="1957" box="[1165,1436,650,672]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="allardi">
|
||
<emphasis box="[1165,1261,650,672]" italics="true" pageId="29" pageNumber="30">M. allardi</emphasis>
|
||
Basilewsky, 1957
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
- Anterior half of labrum round shaped; pronotum transverse (about 1.30 wider than long), heart-shaped; elytra with orange spots as in figure 15b............................................................
|
||
<taxonomicName authority="(Boheman, 1848)" baseAuthorityName="Boheman" baseAuthorityYear="1848" box="[1122,1436,709,731]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="biplagiatus">
|
||
<emphasis box="[1122,1262,709,730]" italics="true" pageId="29" pageNumber="30">M. biplagiatus</emphasis>
|
||
(Boheman, 1848)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
20. Pronotum as wide as long, posterior angles strongly obtuse, slightly sharp at apex; anterior half of labrum ogive shaped, two marginal pairs of setae, one submarginal pair of setae (
|
||
<figureCitation box="[753,836,767,789]" captionStart="FIGURE 10" captionStartId="23.[152,255,1844,1868]" captionTargetBox="[166,1426,707,1780]" captionTargetId="figure-241@23.[152,1436,697,1791]" captionTargetPageId="23" captionText="FIGURE 10. Labrum of Macrocheilus (dorsal view). a, M. longicollis, MOZ (PSC); b, M. persimilis, ZAM (TAC); c, M. proximus, ETI (TAC); d, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745303" httpUri="https://zenodo.org/record/7745303/files/figure.png" pageId="29" pageNumber="30">Fig. 10a</figureCitation>
|
||
); elytra with orange spots and dark patches as in figures 9b, c; male genitalia as in figure 7d....................................................
|
||
<taxonomicName authority="Peringuey, 1904" authorityName="Peringuey" authorityYear="1904" box="[1141,1436,797,819]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="longicollis">
|
||
<emphasis box="[1141,1274,797,818]" italics="true" pageId="29" pageNumber="30">M. longicollis</emphasis>
|
||
<bibRefCitation author="Peringuey, L." box="[1280,1436,797,819]" pageId="29" pageNumber="30" pagination="167 - 206" refId="ref25855" refString="Peringuey, L. (1904). VII. Sixth contribution to the South Africa coleopterous fauna. Description of new species of Coleoptera in the collection of the South African Museum. Annals of the South African Museum, 3, 167 - 206." type="journal article" year="1904">Péringuey, 1904</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" box="[151,1436,826,848]" pageId="29" pageNumber="30">
|
||
<keyLead box="[151,1436,826,848]" pageId="29" pageNumber="30">- Pronotum transverse, wider than long (1.22–1.46 times)...................................................... 21</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" box="[151,1436,855,877]" pageId="29" pageNumber="30">
|
||
<keyLead box="[151,1436,855,877]" pageId="29" pageNumber="30">21. Anterior half of labrum round shaped.................................................................... 22</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
- Anterior half of labrum triangular or ogive shaped, two marginal or submarginal or discal pairs of setae (e.g.
|
||
<figureCitation box="[1272,1370,885,907]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="29" pageNumber="30">Figs. 2c, f</figureCitation>
|
||
,
|
||
<figureCitation box="[1380,1405,885,907]" captionStart="FIGURE 6" captionStartId="18.[152,255,1916,1940]" captionTargetBox="[171,1427,204,1867]" captionTargetId="figure-60@18.[151,1436,194,1885]" captionTargetPageId="18" captionText="FIGURE 6. Labrum of Macrocheilus (dorsal view). a, M. diplospilus, GBI (ASC); b, M. dorsalis, ZAM (TAC); c, M. dorsiger, BFO (PSC); d, M. hybridus, ZIM (TAC); e, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745295" httpUri="https://zenodo.org/record/7745295/files/figure.png" pageId="29" pageNumber="30">6b</figureCitation>
|
||
). .................................................................................................. 23
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
22. Anterior half of labrum with two marginal pairs of setae, one pair of setae beneath the anterior margin near apex (
|
||
<figureCitation box="[1340,1425,943,965]" captionStart="FIGURE 10" captionStartId="23.[152,255,1844,1868]" captionTargetBox="[166,1426,707,1780]" captionTargetId="figure-241@23.[152,1436,697,1791]" captionTargetPageId="23" captionText="FIGURE 10. Labrum of Macrocheilus (dorsal view). a, M. longicollis, MOZ (PSC); b, M. persimilis, ZAM (TAC); c, M. proximus, ETI (TAC); d, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745303" httpUri="https://zenodo.org/record/7745303/files/figure.png" pageId="29" pageNumber="30">Fig. 10b</figureCitation>
|
||
); elytra with orange spots and dark patches as in figures 9d, e; male genitalia as in figure 13a......................................................................................................
|
||
<taxonomicName authority="Basilewsky, 1970" authorityName="Basilewsky" authorityYear="1970" box="[1134,1436,1002,1024]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="persimilis">
|
||
<emphasis box="[1134,1261,1002,1024]" italics="true" pageId="29" pageNumber="30">M. persimilis</emphasis>
|
||
<bibRefCitation author="Basilewsky, P." box="[1267,1436,1002,1024]" pageId="29" pageNumber="30" pagination="337 - 348" refId="ref25250" refString="Basilewsky, P. (1970) Coleopteres Carabidae africains nouveaux. XVIII. Revue de Zoologie et Botanique Africaines, 81 (3 - 4), 337 - 348." type="journal article" year="1970">Basilewsky, 1970</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
- Anterior half of labrum with three marginal pairs of setae; elytra with yellowish spots and dark patches as in figure 15e....................................................................................
|
||
<taxonomicName authority="(Marc, 1840)" baseAuthorityName="Marc" baseAuthorityYear="1840" box="[1182,1436,1061,1082]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cruciatus">
|
||
<emphasis box="[1182,1303,1061,1082]" italics="true" pageId="29" pageNumber="30">M. cruciatus</emphasis>
|
||
(Marc, 1840)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" box="[151,1436,1090,1112]" pageId="29" pageNumber="30">
|
||
<keyLead box="[151,1436,1090,1112]" pageId="29" pageNumber="30">
|
||
23. Larger species (length>
|
||
<quantity box="[458,546,1090,1112]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="29" pageNumber="30" unit="mm" value="11.5">11.5 mm</quantity>
|
||
); anterior half of labrum almost triangular shaped............................... 24
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" box="[151,1436,1119,1141]" pageId="29" pageNumber="30">
|
||
<keyLead box="[151,1436,1119,1141]" pageId="29" pageNumber="30">
|
||
- Smaller species (
|
||
<quantity box="[389,508,1119,1141]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.15" metricValueMin="1.0" pageId="29" pageNumber="30" unit="mm" value="10.75" valueMax="11.5" valueMin="10.0">10–11.5mm</quantity>
|
||
); anterior half of labrum ogive shaped............................................ 25
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
24. Margins of posterior half of labrum subparallel (see
|
||
<bibRefCitation author="Felix, R. F. F. L. & Muilwijk, J." box="[726,951,1149,1171]" pageId="29" pageNumber="30" pagination="83 - 86" refId="ref25418" refString="Felix, R. F. F. L. & Muilwijk, J. (2007) A new Macrocheilus Hope, 1838 from Yemen (Coleotera, Carabidae). Boletin Sociedad Entomologica Aragonesa, 41, 83 - 86." type="journal article" year="2007">Felix & Muilwijk 2007</bibRefCitation>
|
||
,
|
||
<figureCitation box="[961,1019,1149,1171]" captionStart="FIGURE 4" captionStartId="13.[152,255,1917,1941]" captionTargetBox="[203,1362,163,1877]" captionTargetId="figure-76@13.[197,1378,151,1890]" captionTargetPageId="13" captionText="FIGURE 4. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. assmanni (holotype); b, M. philippeorum (paratype); c, M. schuelei (paratype); d, M. basilewskyi, ZIM (TAC); e, M. biguttatus, ETI (TAC); f, M. biguttatus, SEN (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745289" httpUri="https://zenodo.org/record/7745289/files/figure.png" pageId="29" pageNumber="30">Fig. 4</figureCitation>
|
||
); orange spots and dark patches of elytra as in
|
||
<bibRefCitation author="Felix, R. F. F. L. & Muilwijk, J." box="[251,484,1178,1200]" pageId="29" pageNumber="30" pagination="83 - 86" refId="ref25418" refString="Felix, R. F. F. L. & Muilwijk, J. (2007) A new Macrocheilus Hope, 1838 from Yemen (Coleotera, Carabidae). Boletin Sociedad Entomologica Aragonesa, 41, 83 - 86." type="journal article" year="2007">Felix & Muilwijk (2007</bibRefCitation>
|
||
,
|
||
<figureCitation box="[494,549,1178,1200]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="29" pageNumber="30">Fig. 1</figureCitation>
|
||
).........................................
|
||
<taxonomicName authority="Felix & Muilwijk, 2007" authorityName="Felix & Muilwijk" authorityYear="2007" box="[1065,1436,1178,1200]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanharteni">
|
||
<emphasis box="[1065,1201,1178,1200]" italics="true" pageId="29" pageNumber="30">M. vanharteni</emphasis>
|
||
<bibRefCitation author="Felix, R. F. F. L. & Muilwijk, J." box="[1207,1436,1178,1200]" pageId="29" pageNumber="30" pagination="83 - 86" refId="ref25418" refString="Felix, R. F. F. L. & Muilwijk, J. (2007) A new Macrocheilus Hope, 1838 from Yemen (Coleotera, Carabidae). Boletin Sociedad Entomologica Aragonesa, 41, 83 - 86." type="journal article" year="2007">Felix & Muilwijk, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
- Margins of posterior half of labrum parallel (
|
||
<figureCitation box="[651,720,1207,1229]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="29" pageNumber="30">Fig. 2c</figureCitation>
|
||
); elytra with orange spots and dark patches as in figure 1c; male and female genitalia as in figures 3b and 4b, respectively..............................................
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[1215,1376,1237,1258]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="philippeorum" status="n. sp.">
|
||
<emphasis box="[1215,1376,1237,1258]" italics="true" pageId="29" pageNumber="30">M. philippeorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1382,1437,1237,1258]" pageId="29" pageNumber="30">
|
||
<taxonomicNameLabel box="[1382,1437,1237,1258]" pageId="29" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
25. Posterior angles of pronotum slightly obtuse, rounded or very slightly sharp at apex, without expansion just before the angles; anterior half of labrum with one discal pair of setae (
|
||
<figureCitation box="[727,794,1295,1317]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="29" pageNumber="30">Fig. 2f</figureCitation>
|
||
); orange spots and dark patches as in figure 1h; male genitalia as in
|
||
<bibRefCitation author="Serrano, A. R. M." box="[226,365,1325,1346]" pageId="29" pageNumber="30" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano (2000</bibRefCitation>
|
||
,
|
||
<figureCitation box="[374,454,1325,1347]" captionStart="FIGURE 13" captionStartId="27.[152,255,872,896]" captionTargetBox="[370,1262,159,835]" captionTargetId="figure-117@27.[365,1272,151,843]" captionTargetPageId="27" captionText="FIGURE 13. Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. persimilis, ZAM (TAC); b, M. scapularis (MRAC); c, M. spectandus, ZAM (TAC); d, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745311" httpUri="https://zenodo.org/record/7745311/files/figure.png" pageId="29" pageNumber="30">Figs. 13</figureCitation>
|
||
,
|
||
<figureCitation box="[464,490,1325,1346]" captionStart="FIGURE 14" captionStartId="27.[152,255,1916,1940]" captionTargetBox="[421,1245,991,1878]" captionTargetId="figure-120@27.[413,1253,983,1890]" captionTargetPageId="27" captionText="FIGURE 14. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. proximus, ETI (TAC);; b, M. quadrinotatus, ANG (ASC); c, M. taedatus, TAN (TAC); d, M. viduatus katanganus, RSA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745313" httpUri="https://zenodo.org/record/7745313/files/figure.png" pageId="29" pageNumber="30">14</figureCitation>
|
||
), female genitalia as in figure 4d.............................
|
||
<taxonomicName authority="Serrano, 2000" authorityName="Serrano" authorityYear="2000" box="[1153,1436,1325,1346]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="basilewskyi">
|
||
<emphasis box="[1153,1296,1325,1346]" italics="true" pageId="29" pageNumber="30">M. basilewskyi</emphasis>
|
||
<bibRefCitation author="Serrano, A. R. M." box="[1302,1436,1325,1346]" pageId="29" pageNumber="30" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
- Posterior angles of pronotum strongly obtuse and dentiform at apex, a slight short expansion just before the angles; anterior half of labrum without a discal pair of setae (
|
||
<figureCitation box="[614,694,1383,1405]" captionStart="FIGURE 6" captionStartId="18.[152,255,1916,1940]" captionTargetBox="[171,1427,204,1867]" captionTargetId="figure-60@18.[151,1436,194,1885]" captionTargetPageId="18" captionText="FIGURE 6. Labrum of Macrocheilus (dorsal view). a, M. diplospilus, GBI (ASC); b, M. dorsalis, ZAM (TAC); c, M. dorsiger, BFO (PSC); d, M. hybridus, ZIM (TAC); e, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745295" httpUri="https://zenodo.org/record/7745295/files/figure.png" pageId="29" pageNumber="30">Figs. 6b</figureCitation>
|
||
,
|
||
<figureCitation box="[704,741,1383,1405]" captionStart="FIGURE 10" captionStartId="23.[152,255,1844,1868]" captionTargetBox="[166,1426,707,1780]" captionTargetId="figure-241@23.[152,1436,697,1791]" captionTargetPageId="23" captionText="FIGURE 10. Labrum of Macrocheilus (dorsal view). a, M. longicollis, MOZ (PSC); b, M. persimilis, ZAM (TAC); c, M. proximus, ETI (TAC); d, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745303" httpUri="https://zenodo.org/record/7745303/files/figure.png" pageId="29" pageNumber="30">10d</figureCitation>
|
||
)...................................................... 26
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
26. Sutural dark patch of elytra becoming a discoidal one, reaching the 4
|
||
<superScript attach="left" box="[897,908,1412,1424]" fontSize="5" pageId="29" pageNumber="30">th</superScript>
|
||
striae from the base to about one-fourth of the length, widening obliquely from there to near both the lateral and apical margins (
|
||
<figureCitation box="[934,1033,1442,1464]" captionStart="FIGURE 9" captionStartId="22.[152,255,1916,1940]" captionTargetBox="[238,1445,163,1876]" captionTargetId="figure-82@22.[219,1454,151,1889]" captionTargetPageId="22" captionText="FIGURE 9. Habitus of Macrocheilus (dorsal view). a, M. longicollis (holotype); b, M. longicollis, MOZ (PSC); c, M. longicollis, ZAM (TAC); d, M. persimilis (holotype, plus labels); e, M. persimilis, ZAM (TAC); f, M. proximus (holotype); g, M. proximus, ETI (TAC); h, M. quadrinotatus (holotype, plus labels); i, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745301" httpUri="https://zenodo.org/record/7745301/files/figure.png" pageId="29" pageNumber="30">Figs. 9f, g</figureCitation>
|
||
); male genitalia as in
|
||
<bibRefCitation author="Serrano, A. R. M." box="[1241,1380,1442,1463]" pageId="29" pageNumber="30" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano (2000</bibRefCitation>
|
||
,
|
||
<figureCitation captionStart="FIGURE 15" captionStartId="31.[152,255,1943,1967]" captionTargetBox="[245,1343,189,1903]" captionTargetId="figure-16@31.[221,1366,181,1920]" captionTargetPageId="31" captionText="FIGURE 15. Habitus of Macrocheilus (dorsal view). a, M. angustatus (holotype, plus labels); b, M. biplagiatus (MRAC, plus labels); c, M. burgeoni (holotype, plus labels); d, M. crampeli (paratype, plus labels); e, M. cruciatus (MRAC, plus labels); f, M. moraisi (holotype, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745315" httpUri="https://zenodo.org/record/7745315/files/figure.png" pageId="29" pageNumber="30">Figs. 15</figureCitation>
|
||
,
|
||
<figureCitation box="[262,288,1471,1493]" captionStart="FIGURE 16" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[242,1350,192,1911]" captionTargetId="figure-19@32.[226,1362,181,1920]" captionTargetPageId="32" captionText="FIGURE 16. Habitus of Macrocheilus (dorsal view). a, M. ocellatus (holotype, plus labels); b, M. overlaeti (holotype, plus labels); c, M. varians varians (holotype); d, M. varians eugrammus (holotype); e, M. vinctus (paratype)." figureDoi="http://doi.org/10.5281/zenodo.7745317" httpUri="https://zenodo.org/record/7745317/files/figure.png" pageId="29" pageNumber="30">16</figureCitation>
|
||
), female genitalia as in figure 14a.............................................
|
||
<taxonomicName authority="Peringuey, 1896" authorityName="Peringuey" authorityYear="1896" box="[1154,1436,1471,1493]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="proximus">
|
||
<emphasis box="[1154,1274,1472,1493]" italics="true" pageId="29" pageNumber="30">M. proximus</emphasis>
|
||
<bibRefCitation author="Peringuey, L." box="[1280,1436,1471,1493]" pageId="29" pageNumber="30" pagination="125 - 623" refId="ref25799" refString="Peringuey, L. (1896) A descriptive catalogue of the Coleoptera of South Africa. Part II. Carabidae. Transactions of the South African Philosophical Society, 7, 125 - 623, pls. III - X. https: // doi. org / 10.5962 / bhl. title. 9230" type="journal article" year="1896">Péringuey, 1896</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
- Dark drop-like patch of elytra along the suture reaching almost the lateral margins in its largest wide, getting too far from the apical margin (
|
||
<figureCitation box="[370,439,1530,1552]" captionStart="FIGURE 5" captionStartId="17.[152,255,1916,1940]" captionTargetBox="[178,1402,165,1877]" captionTargetId="figure-83@17.[166,1418,151,1890]" captionTargetPageId="17" captionText="FIGURE 5. Habitus of Macrocheilus (dorsal view). a, M. biguttatus, SEN (ACC); b, M. bimaculatus, GBI (ASC); c, M. clasispilus (holotype, plus labels); d, M. diplospilus (holotype, plus labels); e, M. dorsalis, ZAM (TAC); f, M. dorsiger, BFO (PSC); g, M. elegantulus (holotype, plus labels); h, M. hybridus, ZIM (TAC); i, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745293" httpUri="https://zenodo.org/record/7745293/files/figure.png" pageId="29" pageNumber="30">Fig. 5e</figureCitation>
|
||
); male genitalia as in figure 7a......................................
|
||
<taxonomicName authority="Klug, 1834" authorityName="Klug" authorityYear="1834" box="[1213,1436,1530,1552]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="dorsalis">
|
||
<emphasis box="[1213,1322,1530,1552]" italics="true" pageId="29" pageNumber="30">M. dorsalis</emphasis>
|
||
Klug, 1834
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
27. Anterior half of labrum strongly ogive or even triangular shaped, with three pairs of marginal setae (
|
||
<figureCitation box="[1226,1311,1559,1581]" captionStart="FIGURE 12" captionStartId="26.[152,255,1275,1299]" captionTargetBox="[176,1405,161,1252]" captionTargetId="figure-327@26.[166,1413,151,1264]" captionTargetPageId="26" captionText="FIGURE 12. Labrum of Macrocheilus (dorsal view). a, M. scapularis (MRAC); b, M. spectandus, TAN (TAC); c, M. taedatus, TAN (TAC); d, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745307" httpUri="https://zenodo.org/record/7745307/files/figure.png" pageId="29" pageNumber="30">Fig. 12c</figureCitation>
|
||
); elytra dark brown, about twice as long as wide, with 2 orange spots (
|
||
<figureCitation box="[794,912,1589,1611]" captionStart="FIGURE 11" captionStartId="25.[152,255,1923,1947]" captionTargetBox="[359,1259,530,1904]" captionTargetId="figure-221@25.[333,1270,517,1916]" captionTargetPageId="25" captionText="FIGURE 11. Habitus of Macrocheilus (dorsal view). a, M. scapularis (MRAC, plus labels); b, M. spectandus (holotype); c, M. spectandus, TAN (TAC); d, M. taedatus (paratype); e, M. taedatus, TAN (TAC); f, M. viduatus katanganus (holotype, plus labels); g, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745305" httpUri="https://zenodo.org/record/7745305/files/figure.png" pageId="29" pageNumber="30">Figs. 11d, e</figureCitation>
|
||
); male genitalia as in
|
||
<bibRefCitation author="Basilewsky, P." box="[1130,1307,1589,1611]" pageId="29" pageNumber="30" pagination="39 - 51" refId="ref25130" refString="Basilewsky, P. (1960) Sur le genre Macrocheilus Hope (Col. Carabidae Helluoninae). Bulletin et Annales de la Societe Royale d`Entomologie de Belgique, 96 (3 - 4), 39 - 51." type="journal article" year="1960">Basilewsky (1960</bibRefCitation>
|
||
,
|
||
<figureCitation box="[1318,1424,1589,1611]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="29" pageNumber="30">Figs. 2b, e</figureCitation>
|
||
); female genitalia as in figure 14c..................................................
|
||
<taxonomicName authority="Basilewsky, 1960" authorityName="Basilewsky" authorityYear="1960" box="[1148,1436,1618,1640]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="taedatus">
|
||
<emphasis box="[1148,1261,1618,1640]" italics="true" pageId="29" pageNumber="30">M. taedatus</emphasis>
|
||
<bibRefCitation author="Basilewsky, P." box="[1267,1436,1618,1640]" pageId="29" pageNumber="30" pagination="39 - 51" refId="ref25130" refString="Basilewsky, P. (1960) Sur le genre Macrocheilus Hope (Col. Carabidae Helluoninae). Bulletin et Annales de la Societe Royale d`Entomologie de Belgique, 96 (3 - 4), 39 - 51." type="journal article" year="1960">Basilewsky, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
- Anterior half of labrum ogive shaped, two pairs of marginal setae, one pair of discal setae (e.g.
|
||
<figureCitation box="[1156,1240,1647,1669]" captionStart="FIGURE 12" captionStartId="26.[152,255,1275,1299]" captionTargetBox="[176,1405,161,1252]" captionTargetId="figure-327@26.[166,1413,151,1264]" captionTargetPageId="26" captionText="FIGURE 12. Labrum of Macrocheilus (dorsal view). a, M. scapularis (MRAC); b, M. spectandus, TAN (TAC); c, M. taedatus, TAN (TAC); d, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745307" httpUri="https://zenodo.org/record/7745307/files/figure.png" pageId="29" pageNumber="30">Fig. 12b</figureCitation>
|
||
); elytra brownish or blackish, about 1.6–1.7 times longer than wide, with 4 orange spots (
|
||
<figureCitation box="[887,1000,1677,1699]" captionStart="FIGURE 11" captionStartId="25.[152,255,1923,1947]" captionTargetBox="[359,1259,530,1904]" captionTargetId="figure-221@25.[333,1270,517,1916]" captionTargetPageId="25" captionText="FIGURE 11. Habitus of Macrocheilus (dorsal view). a, M. scapularis (MRAC, plus labels); b, M. spectandus (holotype); c, M. spectandus, TAN (TAC); d, M. taedatus (paratype); e, M. taedatus, TAN (TAC); f, M. viduatus katanganus (holotype, plus labels); g, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745305" httpUri="https://zenodo.org/record/7745305/files/figure.png" pageId="29" pageNumber="30">Figs. 11b, c</figureCitation>
|
||
,
|
||
<figureCitation box="[1011,1044,1677,1699]" captionStart="FIGURE 15" captionStartId="31.[152,255,1943,1967]" captionTargetBox="[245,1343,189,1903]" captionTargetId="figure-16@31.[221,1366,181,1920]" captionTargetPageId="31" captionText="FIGURE 15. Habitus of Macrocheilus (dorsal view). a, M. angustatus (holotype, plus labels); b, M. biplagiatus (MRAC, plus labels); c, M. burgeoni (holotype, plus labels); d, M. crampeli (paratype, plus labels); e, M. cruciatus (MRAC, plus labels); f, M. moraisi (holotype, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745315" httpUri="https://zenodo.org/record/7745315/files/figure.png" pageId="29" pageNumber="30">15f</figureCitation>
|
||
)............................. 28
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
28. Larger species (
|
||
<quantity box="[381,491,1706,1727]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.9" metricValueMin="1.4" pageId="29" pageNumber="30" unit="mm" value="16.5" valueMax="19.0" valueMin="14.0">14–19 mm</quantity>
|
||
); Body more densely punctured; margins of posterior half of labrum subround (
|
||
<figureCitation box="[1278,1362,1706,1728]" captionStart="FIGURE 12" captionStartId="26.[152,255,1275,1299]" captionTargetBox="[176,1405,161,1252]" captionTargetId="figure-327@26.[166,1413,151,1264]" captionTargetPageId="26" captionText="FIGURE 12. Labrum of Macrocheilus (dorsal view). a, M. scapularis (MRAC); b, M. spectandus, TAN (TAC); c, M. taedatus, TAN (TAC); d, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745307" httpUri="https://zenodo.org/record/7745307/files/figure.png" pageId="29" pageNumber="30">Fig. 12c</figureCitation>
|
||
); elytra orange spots as in figures 11b, c; male genitalia as in figure 13c (see also
|
||
<bibRefCitation author="Serrano, A. R. M." box="[931,1062,1736,1757]" pageId="29" pageNumber="30" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano 2000</bibRefCitation>
|
||
,
|
||
<figureCitation box="[1072,1139,1735,1757]" captionStart="FIGURE 9" captionStartId="22.[152,255,1916,1940]" captionTargetBox="[238,1445,163,1876]" captionTargetId="figure-82@22.[219,1454,151,1889]" captionTargetPageId="22" captionText="FIGURE 9. Habitus of Macrocheilus (dorsal view). a, M. longicollis (holotype); b, M. longicollis, MOZ (PSC); c, M. longicollis, ZAM (TAC); d, M. persimilis (holotype, plus labels); e, M. persimilis, ZAM (TAC); f, M. proximus (holotype); g, M. proximus, ETI (TAC); h, M. quadrinotatus (holotype, plus labels); i, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745301" httpUri="https://zenodo.org/record/7745301/files/figure.png" pageId="29" pageNumber="30">Figs. 9</figureCitation>
|
||
,
|
||
<figureCitation box="[1149,1174,1736,1757]" captionStart="FIGURE 10" captionStartId="23.[152,255,1844,1868]" captionTargetBox="[166,1426,707,1780]" captionTargetId="figure-241@23.[152,1436,697,1791]" captionTargetPageId="23" captionText="FIGURE 10. Labrum of Macrocheilus (dorsal view). a, M. longicollis, MOZ (PSC); b, M. persimilis, ZAM (TAC); c, M. proximus, ETI (TAC); d, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745303" httpUri="https://zenodo.org/record/7745303/files/figure.png" pageId="29" pageNumber="30">10</figureCitation>
|
||
)...............................................................................................
|
||
<taxonomicName authority="Peringuey, 1904" authorityName="Peringuey" authorityYear="1904" box="[1136,1436,1765,1787]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="spectandus">
|
||
<emphasis box="[1136,1275,1765,1786]" italics="true" pageId="29" pageNumber="30">M. spectandus</emphasis>
|
||
<bibRefCitation author="Peringuey, L." box="[1280,1436,1765,1787]" pageId="29" pageNumber="30" pagination="167 - 206" refId="ref25855" refString="Peringuey, L. (1904). VII. Sixth contribution to the South Africa coleopterous fauna. Description of new species of Coleoptera in the collection of the South African Museum. Annals of the South African Museum, 3, 167 - 206." type="journal article" year="1904">Péringuey, 1904</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
- Smaller species (
|
||
<quantity box="[390,533,1794,1816]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" metricValueMax="1.18" metricValueMin="1.16" pageId="29" pageNumber="30" unit="mm" value="11.7" valueMax="11.8" valueMin="11.6">11.6–11.8 mm</quantity>
|
||
); Body less densely punctured; margins of posterior half of labrum subparallel; elytra orange spots as in figure 15f; male genitalia as in
|
||
<bibRefCitation author="Serrano, A. R. M." box="[632,771,1824,1845]" pageId="29" pageNumber="30" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano (2000</bibRefCitation>
|
||
,
|
||
<figureCitation box="[780,848,1823,1845]" captionStart="FIGURE 7" captionStartId="19.[152,255,1531,1555]" captionTargetBox="[163,1424,429,1485]" captionTargetId="figure-63@19.[151,1436,421,1492]" captionTargetPageId="19" captionText="FIGURE 7.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. dorsalis, ZAM (TAC); b, M. elegantulus, DOC? (MRAC); c, M. labrosus, RCA (PSC); d, M. longicollis, MOZ (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745297" httpUri="https://zenodo.org/record/7745297/files/figure.png" pageId="29" pageNumber="30">Figs. 7</figureCitation>
|
||
,
|
||
<figureCitation box="[858,870,1824,1845]" captionStart="FIGURE 8" captionStartId="20.[152,255,1122,1146]" captionTargetBox="[408,1189,193,1080]" captionTargetId="figure-346@20.[401,1197,181,1088]" captionTargetPageId="20" captionText="FIGURE 8. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. bimaculatus, GBI (ASC); b, M. clasispilus, RCI (ACC); c, M. hybridus, ZIM (TAC); d, M. labrosus, RCI (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745299" httpUri="https://zenodo.org/record/7745299/files/figure.png" pageId="29" pageNumber="30">8</figureCitation>
|
||
).........................
|
||
<taxonomicName authority="Serrano, 2000" authorityName="Serrano" authorityYear="2000" box="[1190,1436,1824,1845]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="moraisi">
|
||
<emphasis box="[1190,1295,1824,1845]" italics="true" pageId="29" pageNumber="30">M. moraisi</emphasis>
|
||
<bibRefCitation author="Serrano, A. R. M." box="[1302,1436,1824,1845]" pageId="29" pageNumber="30" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
|
||
<keyLead pageId="29" pageNumber="30">
|
||
29. Body totally black; anterior half of labrum strongly rounded, two pairs of marginal setae, one pair of setae in the middle of the disc, margins of posterior half rounded (
|
||
<figureCitation box="[619,691,1882,1904]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="29" pageNumber="30">Fig. 2a</figureCitation>
|
||
); pronotum slightly cordiform, posterior angles slightly obtuse and rounded, anterior submarginal area wrinkled; female genitalia as in figure 4a..............................
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[1254,1377,1912,1933]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="assmanni" status="n. sp.">
|
||
<emphasis box="[1254,1377,1912,1933]" italics="true" pageId="29" pageNumber="30">M. assmanni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1382,1437,1912,1933]" pageId="29" pageNumber="30">
|
||
<taxonomicNameLabel box="[1382,1437,1912,1933]" pageId="29" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,2021]" box="[151,1436,1941,1963]" pageId="29" pageNumber="30">
|
||
<keyLead box="[151,1436,1941,1963]" pageId="29" pageNumber="30">- Body colour not as above.............................................................................. 30</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||
<paragraph blockId="29.[151,1437,151,2021]" lastBlockId="30.[151,1437,151,1112]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||
<keyLead lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||
30. Head blackish; pronotum testaceous-redish; elytra blackish concolorous (
|
||
<figureCitation box="[933,1014,1970,1992]" captionStart="FIGURE 16" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[242,1350,192,1911]" captionTargetId="figure-19@32.[226,1362,181,1920]" captionTargetPageId="32" captionText="FIGURE 16. Habitus of Macrocheilus (dorsal view). a, M. ocellatus (holotype, plus labels); b, M. overlaeti (holotype, plus labels); c, M. varians varians (holotype); d, M. varians eugrammus (holotype); e, M. vinctus (paratype)." figureDoi="http://doi.org/10.5281/zenodo.7745317" httpUri="https://zenodo.org/record/7745317/files/figure.png" pageId="29" pageNumber="30">Fig. 16c</figureCitation>
|
||
) [elytra with one pair of a longitudinal redyellowish patch spread throughout the 4
|
||
<superScript attach="none" box="[611,622,1999,2011]" fontSize="5" pageId="29" pageNumber="30">th</superScript>
|
||
, 5
|
||
<superScript attach="left" box="[646,657,1999,2011]" fontSize="5" pageId="29" pageNumber="30">th</superScript>
|
||
, 6
|
||
<superScript attach="left" box="[687,698,1999,2011]" fontSize="5" pageId="29" pageNumber="30">th</superScript>
|
||
and part of 7
|
||
<superScript attach="left" box="[826,837,1999,2011]" fontSize="5" pageId="29" pageNumber="30">th</superScript>
|
||
intervals since the base to the median region of the disc in the subspecies
|
||
<taxonomicName authority="Basilewsky, 1960" authorityName="Basilewsky" authorityYear="1960" box="[334,732,151,173]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="varians" subSpecies="eugrammus">
|
||
<emphasis box="[334,556,152,173]" italics="true" pageId="30" pageNumber="31">M. varians eugrammus</emphasis>
|
||
<bibRefCitation author="Basilewsky, P." box="[562,732,151,173]" pageId="30" pageNumber="31" pagination="39 - 51" refId="ref25130" refString="Basilewsky, P. (1960) Sur le genre Macrocheilus Hope (Col. Carabidae Helluoninae). Bulletin et Annales de la Societe Royale d`Entomologie de Belgique, 96 (3 - 4), 39 - 51." type="journal article" year="1960">Basilewsky, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[745,829,151,173]" captionStart="FIGURE 16" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[242,1350,192,1911]" captionTargetId="figure-19@32.[226,1362,181,1920]" captionTargetPageId="32" captionText="FIGURE 16. Habitus of Macrocheilus (dorsal view). a, M. ocellatus (holotype, plus labels); b, M. overlaeti (holotype, plus labels); c, M. varians varians (holotype); d, M. varians eugrammus (holotype); e, M. vinctus (paratype)." figureDoi="http://doi.org/10.5281/zenodo.7745317" httpUri="https://zenodo.org/record/7745317/files/figure.png" pageId="30" pageNumber="31">Fig. 16d</figureCitation>
|
||
)]...................
|
||
<taxonomicName authority="Peringuey, 1904" authorityName="Peringuey" authorityYear="1904" box="[1092,1436,151,173]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="varians" subSpecies="varians">
|
||
<emphasis box="[1092,1274,152,173]" italics="true" pageId="30" pageNumber="31">M. varians varians</emphasis>
|
||
<bibRefCitation author="Peringuey, L." box="[1280,1436,151,173]" pageId="30" pageNumber="31" pagination="167 - 206" refId="ref25855" refString="Peringuey, L. (1904). VII. Sixth contribution to the South Africa coleopterous fauna. Description of new species of Coleoptera in the collection of the South African Museum. Annals of the South African Museum, 3, 167 - 206." type="journal article" year="1904">Péringuey, 1904</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="30.[151,1437,151,1112]" box="[151,1436,181,203]" pageId="30" pageNumber="31">
|
||
<keyLead box="[151,1436,181,203]" pageId="30" pageNumber="31">- Body colour not as above............................................................................. 31</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="30" pageNumber="31">
|
||
<paragraph blockId="30.[151,1437,151,1112]" box="[151,1436,210,232]" pageId="30" pageNumber="31">
|
||
<keyLead box="[151,1436,210,232]" pageId="30" pageNumber="31">31. Head and pronotum black or dark brown; elytra black or dark brown with orange-brownish or yellowish spots.......... 32</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="30.[151,1437,151,1112]" pageId="30" pageNumber="31">
|
||
<keyLead pageId="30" pageNumber="31">- Head and pronotum orange-yellowish or brownish; almost all elytra brownish or yellowish with drop-like dark markings... .................................................................................................. 35</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="30" pageNumber="31">
|
||
<paragraph blockId="30.[151,1437,151,1112]" pageId="30" pageNumber="31">
|
||
<keyLead pageId="30" pageNumber="31">
|
||
32. Elytra with two pairs of orange-brownish spots (
|
||
<figureCitation box="[691,790,298,320]" captionStart="FIGURE 9" captionStartId="22.[152,255,1916,1940]" captionTargetBox="[238,1445,163,1876]" captionTargetId="figure-82@22.[219,1454,151,1889]" captionTargetPageId="22" captionText="FIGURE 9. Habitus of Macrocheilus (dorsal view). a, M. longicollis (holotype); b, M. longicollis, MOZ (PSC); c, M. longicollis, ZAM (TAC); d, M. persimilis (holotype, plus labels); e, M. persimilis, ZAM (TAC); f, M. proximus (holotype); g, M. proximus, ETI (TAC); h, M. quadrinotatus (holotype, plus labels); i, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745301" httpUri="https://zenodo.org/record/7745301/files/figure.png" pageId="30" pageNumber="31">Figs. 9h, i</figureCitation>
|
||
), one row of punctures in the first interval; margins of anterior half of labrum strongly rounded, two pairs of marginal setae, one pair of setae in the middle of the disc, margins of posterior half rounded (
|
||
<figureCitation box="[318,402,357,379]" captionStart="FIGURE 10" captionStartId="23.[152,255,1844,1868]" captionTargetBox="[166,1426,707,1780]" captionTargetId="figure-241@23.[152,1436,697,1791]" captionTargetPageId="23" captionText="FIGURE 10. Labrum of Macrocheilus (dorsal view). a, M. longicollis, MOZ (PSC); b, M. persimilis, ZAM (TAC); c, M. proximus, ETI (TAC); d, M. quadrinotatus, ANG (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745303" httpUri="https://zenodo.org/record/7745303/files/figure.png" pageId="30" pageNumber="31">Fig. 10d</figureCitation>
|
||
); pronotum slightly cordiform, posterior angles right and rounded; female genitalia as in figure 14b................................................................................
|
||
<taxonomicName authority="Burgeon, 1937" authorityName="Burgeon" authorityYear="1937" box="[1121,1436,386,408]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="quadrinotatus">
|
||
<emphasis box="[1121,1288,386,408]" italics="true" pageId="30" pageNumber="31">M. quadrinotatus</emphasis>
|
||
<bibRefCitation author="Burgeon, L." box="[1294,1436,386,408]" pageId="30" pageNumber="31" pagination="319 - 405" refId="ref25317" refString="Burgeon, L. (1937) Catalogues raisonnes de la faune entomologique du Congo Belge. Coleopteres Carabides. Troisieme Partie. Annales du Musee du Congo Belge, Zoology, Serie 3 (2), 2 (5), 319 - 405." type="journal article" year="1937">Burgeon, 1937</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="30.[151,1437,151,1112]" box="[151,1436,415,437]" pageId="30" pageNumber="31">
|
||
<keyLead box="[151,1436,415,437]" pageId="30" pageNumber="31">- Elytra with one pair of orange-brownish spots............................................................. 33</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="30" pageNumber="31">
|
||
<paragraph blockId="30.[151,1437,151,1112]" pageId="30" pageNumber="31">
|
||
<keyLead pageId="30" pageNumber="31">
|
||
33 Elytra with one pair of ovoid orange-brownish spots in the middle region; body densely punctured; anterior half of labrum ogive shaped, two marginal pairs of setae, one submarginal pair of setae (
|
||
<figureCitation box="[917,988,474,496]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="30" pageNumber="31">Fig. 2b</figureCitation>
|
||
); posterior angles of pronotum strongly obtuse, dentiform at apex, a slight short expansion just before the angles; legs totally brown; male genitalia as in figure 3a..............................................................................................
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[1270,1377,533,554]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="kudrnai" status="n. sp.">
|
||
<emphasis box="[1270,1377,533,554]" italics="true" pageId="30" pageNumber="31">M. kudrnai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1382,1437,533,554]" pageId="30" pageNumber="31">
|
||
<taxonomicNameLabel box="[1382,1437,533,554]" pageId="30" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="30.[151,1437,151,1112]" box="[151,1436,562,584]" pageId="30" pageNumber="31">
|
||
<keyLead box="[151,1436,562,584]" pageId="30" pageNumber="31">- Elytra with one pair of longitudinal orange-brownish spots spread throughout the first half not reaching the base........ 34</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="30" pageNumber="31">
|
||
<paragraph blockId="30.[151,1437,151,1112]" pageId="30" pageNumber="31">
|
||
<keyLead pageId="30" pageNumber="31">
|
||
34. Pronotum strongly cordiform, posterior angles acute and rounded; anterior half of labrum slightly ogive shaped, three pairs of marginal setae (
|
||
<figureCitation box="[378,460,621,643]" captionStart="FIGURE 12" captionStartId="26.[152,255,1275,1299]" captionTargetBox="[176,1405,161,1252]" captionTargetId="figure-327@26.[166,1413,151,1264]" captionTargetPageId="26" captionText="FIGURE 12. Labrum of Macrocheilus (dorsal view). a, M. scapularis (MRAC); b, M. spectandus, TAN (TAC); c, M. taedatus, TAN (TAC); d, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745307" httpUri="https://zenodo.org/record/7745307/files/figure.png" pageId="30" pageNumber="31">Fig. 12a</figureCitation>
|
||
); knee, tibia and tarsi blackish, femora brownish (
|
||
<figureCitation box="[940,1021,621,643]" captionStart="FIGURE 11" captionStartId="25.[152,255,1923,1947]" captionTargetBox="[359,1259,530,1904]" captionTargetId="figure-221@25.[333,1270,517,1916]" captionTargetPageId="25" captionText="FIGURE 11. Habitus of Macrocheilus (dorsal view). a, M. scapularis (MRAC, plus labels); b, M. spectandus (holotype); c, M. spectandus, TAN (TAC); d, M. taedatus (paratype); e, M. taedatus, TAN (TAC); f, M. viduatus katanganus (holotype, plus labels); g, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745305" httpUri="https://zenodo.org/record/7745305/files/figure.png" pageId="30" pageNumber="31">Fig. 11a</figureCitation>
|
||
); male genitalia as in figure 13b.....................................................................................
|
||
<taxonomicName authority="(Reiche, 1843)" baseAuthorityName="Reiche" baseAuthorityYear="1843" box="[1156,1436,650,672]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="scapularis">
|
||
<emphasis box="[1156,1288,650,672]" italics="true" pageId="30" pageNumber="31">M. scapularis</emphasis>
|
||
(Reiche, 1843)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="30.[151,1437,151,1112]" pageId="30" pageNumber="31">
|
||
<keyLead pageId="30" pageNumber="31">
|
||
- Pronotum slightly cordiform, posterior angles right or even slightly obtuse and rounded at apex; anterior half of labrum ogive shaped, two pairs of marginal setae, one submarginal intermedium pair (
|
||
<figureCitation box="[937,1011,709,731]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="30" pageNumber="31">Fig. 2d</figureCitation>
|
||
); legs brownish (
|
||
<figureCitation box="[1181,1253,709,731]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="30" pageNumber="31">Fig. 1d</figureCitation>
|
||
); male and female genitalia as in figures 3c and 4c, respectively.................................................
|
||
<taxonomicName authority="Serrano, 2023" authorityName="Serrano" authorityYear="2023" box="[1266,1377,738,760]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="schuelei" status="n. sp.">
|
||
<emphasis box="[1266,1377,738,760]" italics="true" pageId="30" pageNumber="31">M. schuelei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1382,1437,738,759]" pageId="30" pageNumber="31">
|
||
<taxonomicNameLabel box="[1382,1437,738,759]" pageId="30" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="30" pageNumber="31">
|
||
<paragraph blockId="30.[151,1437,151,1112]" pageId="30" pageNumber="31">
|
||
<keyLead pageId="30" pageNumber="31">
|
||
35. Posterior angles of pronotum slightly obtuse, rounded or very slightly sharp at apex, without expansion just before the angles; anterior half of labrum with one discal pair of setae (
|
||
<figureCitation box="[727,794,797,819]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="30" pageNumber="31">Fig. 2f</figureCitation>
|
||
); orange spots and dark patches as in figure 1h; male genitalia as in
|
||
<bibRefCitation author="Serrano, A. R. M." box="[226,365,826,847]" pageId="30" pageNumber="31" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano (2000</bibRefCitation>
|
||
,
|
||
<figureCitation box="[374,454,826,848]" captionStart="FIGURE 13" captionStartId="27.[152,255,872,896]" captionTargetBox="[370,1262,159,835]" captionTargetId="figure-117@27.[365,1272,151,843]" captionTargetPageId="27" captionText="FIGURE 13. Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. persimilis, ZAM (TAC); b, M. scapularis (MRAC); c, M. spectandus, ZAM (TAC); d, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745311" httpUri="https://zenodo.org/record/7745311/files/figure.png" pageId="30" pageNumber="31">Figs. 13</figureCitation>
|
||
,
|
||
<figureCitation box="[464,490,826,847]" captionStart="FIGURE 14" captionStartId="27.[152,255,1916,1940]" captionTargetBox="[421,1245,991,1878]" captionTargetId="figure-120@27.[413,1253,983,1890]" captionTargetPageId="27" captionText="FIGURE 14. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. proximus, ETI (TAC);; b, M. quadrinotatus, ANG (ASC); c, M. taedatus, TAN (TAC); d, M. viduatus katanganus, RSA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745313" httpUri="https://zenodo.org/record/7745313/files/figure.png" pageId="30" pageNumber="31">14</figureCitation>
|
||
), female genitalia as in figure 4d.............................
|
||
<taxonomicName authority="Serrano, 2000" authorityName="Serrano" authorityYear="2000" box="[1153,1436,826,848]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="basilewskyi">
|
||
<emphasis box="[1153,1296,826,848]" italics="true" pageId="30" pageNumber="31">M. basilewskyi</emphasis>
|
||
<bibRefCitation author="Serrano, A. R. M." box="[1302,1436,826,847]" pageId="30" pageNumber="31" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="30.[151,1437,151,1112]" box="[151,1436,855,877]" pageId="30" pageNumber="31">
|
||
<keyLead box="[151,1436,855,877]" pageId="30" pageNumber="31">- Posterior angles of pronotum obtuse and dentiform at apex; orange spots and dark patches not as in figure 1h........... 36</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="30" pageNumber="31">
|
||
<paragraph blockId="30.[151,1437,151,1112]" pageId="30" pageNumber="31">
|
||
<keyLead pageId="30" pageNumber="31">
|
||
36. Larger species (
|
||
<quantity box="[380,476,885,906]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="30" pageNumber="31" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
|
||
); head and pronotum densely punctured (
|
||
<figureCitation box="[878,949,885,907]" captionStart="FIGURE 5" captionStartId="17.[152,255,1916,1940]" captionTargetBox="[178,1402,165,1877]" captionTargetId="figure-83@17.[166,1418,151,1890]" captionTargetPageId="17" captionText="FIGURE 5. Habitus of Macrocheilus (dorsal view). a, M. biguttatus, SEN (ACC); b, M. bimaculatus, GBI (ASC); c, M. clasispilus (holotype, plus labels); d, M. diplospilus (holotype, plus labels); e, M. dorsalis, ZAM (TAC); f, M. dorsiger, BFO (PSC); g, M. elegantulus (holotype, plus labels); h, M. hybridus, ZIM (TAC); i, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745293" httpUri="https://zenodo.org/record/7745293/files/figure.png" pageId="30" pageNumber="31">Fig. 5e</figureCitation>
|
||
); pronotum more transverse (1.3 times larger than long), a slight short expansion just before the posterior angles; anterior half of labrum ogive shaped, three pairs of marginal setae (
|
||
<figureCitation box="[288,361,943,965]" captionStart="FIGURE 6" captionStartId="18.[152,255,1916,1940]" captionTargetBox="[171,1427,204,1867]" captionTargetId="figure-60@18.[151,1436,194,1885]" captionTargetPageId="18" captionText="FIGURE 6. Labrum of Macrocheilus (dorsal view). a, M. diplospilus, GBI (ASC); b, M. dorsalis, ZAM (TAC); c, M. dorsiger, BFO (PSC); d, M. hybridus, ZIM (TAC); e, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745295" httpUri="https://zenodo.org/record/7745295/files/figure.png" pageId="30" pageNumber="31">Fig. 6b</figureCitation>
|
||
); dark drop-like patch of elytra along the suture reaching almost the margins in its largest wide (
|
||
<figureCitation box="[1299,1370,943,965]" captionStart="FIGURE 5" captionStartId="17.[152,255,1916,1940]" captionTargetBox="[178,1402,165,1877]" captionTargetId="figure-83@17.[166,1418,151,1890]" captionTargetPageId="17" captionText="FIGURE 5. Habitus of Macrocheilus (dorsal view). a, M. biguttatus, SEN (ACC); b, M. bimaculatus, GBI (ASC); c, M. clasispilus (holotype, plus labels); d, M. diplospilus (holotype, plus labels); e, M. dorsalis, ZAM (TAC); f, M. dorsiger, BFO (PSC); g, M. elegantulus (holotype, plus labels); h, M. hybridus, ZIM (TAC); i, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745293" httpUri="https://zenodo.org/record/7745293/files/figure.png" pageId="30" pageNumber="31">Fig. 5e</figureCitation>
|
||
); male genitalia as in figure 7a..............................................................
|
||
<taxonomicName authority="Klug, 1834" authorityName="Klug" authorityYear="1834" box="[1213,1436,973,995]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="dorsalis">
|
||
<emphasis box="[1213,1322,973,994]" italics="true" pageId="30" pageNumber="31">M. dorsalis</emphasis>
|
||
Klug, 1834
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="30.[151,1437,151,1112]" pageId="30" pageNumber="31">
|
||
<keyLead pageId="30" pageNumber="31">
|
||
- Smaller species (
|
||
<quantity box="[388,471,1002,1024]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="30" pageNumber="31" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
|
||
); head and pronotum less densely punctured (
|
||
<figureCitation box="[906,973,1002,1024]" captionStart="FIGURE 5" captionStartId="17.[152,255,1916,1940]" captionTargetBox="[178,1402,165,1877]" captionTargetId="figure-83@17.[166,1418,151,1890]" captionTargetPageId="17" captionText="FIGURE 5. Habitus of Macrocheilus (dorsal view). a, M. biguttatus, SEN (ACC); b, M. bimaculatus, GBI (ASC); c, M. clasispilus (holotype, plus labels); d, M. diplospilus (holotype, plus labels); e, M. dorsalis, ZAM (TAC); f, M. dorsiger, BFO (PSC); g, M. elegantulus (holotype, plus labels); h, M. hybridus, ZIM (TAC); i, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745293" httpUri="https://zenodo.org/record/7745293/files/figure.png" pageId="30" pageNumber="31">Fig. 5f</figureCitation>
|
||
); pronotum less transverse (1.2 times larger than long), without short expansion just before the posterior angles; anterior two thirds of labrum strongly triangular shaped, three pairs of marginal setae (
|
||
<figureCitation box="[464,536,1061,1083]" captionStart="FIGURE 6" captionStartId="18.[152,255,1916,1940]" captionTargetBox="[171,1427,204,1867]" captionTargetId="figure-60@18.[151,1436,194,1885]" captionTargetPageId="18" captionText="FIGURE 6. Labrum of Macrocheilus (dorsal view). a, M. diplospilus, GBI (ASC); b, M. dorsalis, ZAM (TAC); c, M. dorsiger, BFO (PSC); d, M. hybridus, ZIM (TAC); e, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745295" httpUri="https://zenodo.org/record/7745295/files/figure.png" pageId="30" pageNumber="31">Fig. 6c</figureCitation>
|
||
); dark drop-like patch of elytra along the suture narrower in its largest wide (
|
||
<figureCitation box="[1300,1370,1061,1083]" captionStart="FIGURE 5" captionStartId="17.[152,255,1916,1940]" captionTargetBox="[178,1402,165,1877]" captionTargetId="figure-83@17.[166,1418,151,1890]" captionTargetPageId="17" captionText="FIGURE 5. Habitus of Macrocheilus (dorsal view). a, M. biguttatus, SEN (ACC); b, M. bimaculatus, GBI (ASC); c, M. clasispilus (holotype, plus labels); d, M. diplospilus (holotype, plus labels); e, M. dorsalis, ZAM (TAC); f, M. dorsiger, BFO (PSC); g, M. elegantulus (holotype, plus labels); h, M. hybridus, ZIM (TAC); i, M. labrosus, RCA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745293" httpUri="https://zenodo.org/record/7745293/files/figure.png" pageId="30" pageNumber="31">Fig. 5f</figureCitation>
|
||
); male genitalia as in
|
||
<bibRefCitation author="Serrano, A. R. M." box="[365,505,1090,1111]" pageId="30" pageNumber="31" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano (2000</bibRefCitation>
|
||
,
|
||
<figureCitation box="[514,580,1090,1112]" captionStart="FIGURE 3" captionStartId="9.[152,255,1844,1868]" captionTargetBox="[159,1421,165,1805]" captionTargetId="figure-75@9.[151,1436,151,1815]" captionTargetPageId="9" captionText="FIGURE 3.Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. kudrnai n. sp. (holotype); b, M. philippeorum n. sp. (holotype); c, M. schuelei n. sp. (holotype); d, M. alluaudi, CON (PSC); e, M. biguttatus, GBI (ASC); f, M. diplospilus, GBI (ASC)." figureDoi="http://doi.org/10.5281/zenodo.7745287" httpUri="https://zenodo.org/record/7745287/files/figure.png" pageId="30" pageNumber="31">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation box="[591,604,1090,1111]" captionStart="FIGURE 4" captionStartId="13.[152,255,1917,1941]" captionTargetBox="[203,1362,163,1877]" captionTargetId="figure-76@13.[197,1378,151,1890]" captionTargetPageId="13" captionText="FIGURE 4. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. assmanni (holotype); b, M. philippeorum (paratype); c, M. schuelei (paratype); d, M. basilewskyi, ZIM (TAC); e, M. biguttatus, ETI (TAC); f, M. biguttatus, SEN (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745289" httpUri="https://zenodo.org/record/7745289/files/figure.png" pageId="30" pageNumber="31">4</figureCitation>
|
||
)............................................
|
||
<taxonomicName authority="(Chaudoir, 1876)" baseAuthorityName="Chaudoir" baseAuthorityYear="1876" box="[1153,1436,1090,1112]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="dorsiger">
|
||
<emphasis box="[1153,1266,1090,1112]" italics="true" pageId="30" pageNumber="31">M. dorsiger</emphasis>
|
||
(Chaudoir, 1876)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |