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<document id="0070CA5D93B1321C5940DFD11AFFBDDB" ID-CLB-Dataset="42341" ID-DOI="10.11646/zootaxa.3911.3.1" ID-GBIF-Dataset="fc486911-8478-4171-8681-7f87863df5e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="254427" ID-ZooBank="071C75F3-4865-4CCE-AACA-21C82739E626" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460942063819" checkinUser="plazi" docAuthor="Anker, Arthur, Pratama, Idham Sumarto, Firdaus, Muhammad &amp; Rahayu, Dwi Listyo" docDate="2015" docId="AD437D0CF169AD5F94CE126CF2691B94" docLanguage="en" docName="zt03911p342.pdf" docOrigin="Zootaxa 3911 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laomedia astacina De Haan 1841" docType="treatment" docVersion="17" lastPageNumber="333" masterDocId="517A0574F174AD7F9459140AF653185C" masterDocTitle="On some interesting marine decapod crustaceans (Alpheidae, Laomediidae, Strahlaxiidae) from Lombok, Indonesia" masterLastPageNumber="342" masterPageNumber="301" pageNumber="330" updateTime="1698584489804" updateUser="ExternalLinkService">
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<mods:title id="69B491E8CA05B6CCF736E42E38EDA7D6">On some interesting marine decapod crustaceans (Alpheidae, Laomediidae, Strahlaxiidae) from Lombok, Indonesia</mods:title>
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<treatment id="AD437D0CF169AD5F94CE126CF2691B94" ID-DOI="http://doi.org/10.5281/zenodo.5073392" ID-GBIF-Taxon="119591579" ID-Zenodo-Dep="5073392" LSID="urn:lsid:plazi:treatment:AD437D0CF169AD5F94CE126CF2691B94" httpUri="http://treatment.plazi.org/id/AD437D0CF169AD5F94CE126CF2691B94" lastPageId="32" lastPageNumber="333" pageId="29" pageNumber="330">
<subSubSection id="6DF09F91F169AD6294CE126CF4A51EBB" pageId="29" pageNumber="330" type="nomenclature">
<paragraph id="2555CC1AF169AD6294CE126CF4691EDC" blockId="29.[151,570,1638,1698]" box="[151,570,1638,1664]" pageId="29" pageNumber="330">
<heading id="7E1D7B76F169AD6294CE126CF4691EDC" bold="true" box="[151,570,1638,1664]" fontSize="11" level="1" pageId="29" pageNumber="330" reason="1">
<taxonomicName id="E2EAB799F169AD6294CE126CF4691EDC" ID-CoL="c840ed71-2bff-40bf-95f2-f86c030b829f" authority="De Haan, 1841" authorityName="De Haan" authorityYear="1841" box="[151,570,1638,1664]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F169AD6294CE126CF4691EDC" bold="true" box="[151,570,1638,1664]" pageId="29" pageNumber="330">
<emphasis id="179E1008F169AD6294CE126CF7281EDC" bold="true" box="[151,379,1638,1664]" italics="true" pageId="29" pageNumber="330">Laomedia astacina</emphasis>
<collectingCountry id="5DFD8C8AF169AD6295DB126CF7F71EDC" box="[386,420,1638,1664]" name="South Georgia and the South Sandwich Islands" pageId="29" pageNumber="330">De</collectingCountry>
Haan, 1841
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="2555CC1AF169AD6294CE1283F6AB1EFE" blockId="29.[151,570,1638,1698]" box="[151,248,1673,1698]" pageId="29" pageNumber="330">
(
<figureCitation id="BDD1D09FF169AD6294C61283F6A21EFE" box="[159,241,1673,1698]" captionStart="FIGURE 23" captionStartId="31.[151,250,1612,1634]" captionTargetBox="[154,1436,151,1596]" captionTargetId="figure@31.[154,1439,151,1597]" captionTargetPageId="31" captionText="FIGURE 23. Laomedia astacina De Haan, 1841: female (cl 8.9 mm) from Lembar, Lombok, Indonesia (RMNH. CRUS. D. 56754), A—dorsal view; B—lateral view; close-up of frontal region, dorsal view." httpUri="https://zenodo.org/record/254448/files/figure.png" pageId="29" pageNumber="330">Fig. 23</figureCitation>
)
</paragraph>
<paragraph id="2555CC1AF169AD6294CE12DBF4A51EBB" blockId="29.[151,1436,1745,1920]" box="[151,758,1745,1767]" pageId="29" pageNumber="330">
Selected synonymy (for full synonymy see
<bibRefCitation id="417BB1EBF169AD62961612D8F4B91EBB" author="Ngoc-Ho" box="[591,746,1745,1767]" pageId="29" pageNumber="330" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae). Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho 1997</bibRefCitation>
):
</paragraph>
</subSubSection>
<subSubSection id="6DF09F91F169AD6294CE12FAF7271FDC" pageId="29" pageNumber="330" type="reference_group">
<paragraph id="2555CC1AF169AD6294CE12FAF7661F1F" blockId="29.[151,1436,1745,1920]" pageId="29" pageNumber="330">
<treatmentCitationGroup id="05FAEB34F169AD6294CE12FAF7661F1F" pageId="29" pageNumber="330">
<taxonomicName id="E2EAB799F169AD6294CE12FAF47D1F5A" ID-CoL="c840ed71-2bff-40bf-95f2-f86c030b829f" authority="De Haan 1841: 165" authorityName="De Haan" authorityPageNumber="165" authorityYear="1841" box="[151,558,1776,1798]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F169AD6294CE12FAF7091F5A" box="[151,346,1776,1798]" italics="true" pageId="29" pageNumber="330">Laomedia astacina</emphasis>
<collectingCountry id="5DFD8C8AF169AD62953B12FBF72C1F5A" box="[354,383,1777,1798]" name="South Georgia and the South Sandwich Islands" pageId="29" pageNumber="330">De</collectingCountry>
Haan 1841: 165
</taxonomicName>
, pl. 35, fig. 8, pl. N;
<treatmentCitation id="A44BEA0BF169AD62975212FAF5FA1F5A" author="Sakai" box="[779,937,1776,1798]" page="27" pageId="29" pageNumber="330" year="1962">
<bibRefCitation id="417BB1EBF169AD62975212FAF5D21F5A" author="Sakai" box="[779,897,1776,1798]" pageId="29" pageNumber="330" refString="Sakai K. (1962) Systematic studies on Thalassinidea. I. Laomedia astacina De Haan. Publications of the Seto Marine Biological Laboratory, 10, 27 - 34." type="journal article" year="1962">Sakai 1962</bibRefCitation>
: 27
</treatmentCitation>
, pls. 57, figs. 125;
<treatmentCitation id="A44BEA0BF169AD6290D312FAF3C41F5A" author="Sakai" box="[1162,1431,1776,1798]" page="86" pageId="29" pageNumber="330" year="1964">
<bibRefCitation id="417BB1EBF169AD6290D312FAF3221F5A" author="Sakai" box="[1162,1393,1776,1798]" pageId="29" pageNumber="330" refString="Sakai, K. &amp; Miyake, S. (1964) Description of the first zoea of Laomedia astacina de Haan (Decapoda Crustacea). Science Bulletin of the Faculty of Agriculture, Kyushu University, 21, 83 - 87." type="journal article" year="1964">Sakai &amp; Miyake 1964</bibRefCitation>
: 86
</treatmentCitation>
, figs. 13;
<treatmentCitation id="A44BEA0BF169AD6295761304F4021F78" author="Yaldwyn" box="[303,593,1806,1828]" page="137" pageId="29" pageNumber="330" year="1972">
<bibRefCitation id="417BB1EBF169AD6295761304F44D1F78" author="Yaldwyn" box="[303,542,1806,1828]" pageId="29" pageNumber="330" refString="Yaldwyn, J. C. &amp; Wear, R. G. (1972) The eastern Australian, burrowing mud-shrimp Laomedia healyi (Crustacea, Macrura Reptantia, Laomediidae) with notes on larvae of the genus Laomedia. Australian Zoologist, 17, 126 - 141." type="journal article" year="1972">Yaldwyn &amp; Wear 1972</bibRefCitation>
: 137
</treatmentCitation>
, figs. 1320;
<treatmentCitation id="A44BEA0BF169AD6296871304F52F1F78" author="Kim" box="[734,892,1806,1828]" page="589" pageId="29" pageNumber="330" year="1973">
<bibRefCitation id="417BB1EBF169AD6296871304F51B1F78" author="Kim" box="[734,840,1806,1828]" pageId="29" pageNumber="330" refString="Kim, H. S. (1973) Illustrated encyclopedia of fauna and flora of Korea. Vol. 14. Anomura and Brachyura. Samhwa Publishing Co. Ltd., Seoul, 694 pp." type="book" year="1973">Kim 1973</bibRefCitation>
: 589
</treatmentCitation>
, fig. 14;
<treatmentCitation id="A44BEA0BF169AD6297801304F2DB1F78" author="Fukuda" box="[985,1160,1806,1828]" page="19" pageId="29" pageNumber="330" year="1982">
<bibRefCitation id="417BB1EBF169AD6297801304F2311F78" author="Fukuda" box="[985,1122,1806,1828]" pageId="29" pageNumber="330" refString="Fukuda, Y. (1982) Zoeal stages of the burrowing mud shrimp Laomedia astacina De Haan (Decapoda: Thalassinidea: Laomediidae) reared in the laboratory. Proceedings of the Japanese Society of Systematic Zoology, 24, 19 - 31." type="journal article" year="1982">Fukuda 1982</bibRefCitation>
: 19
</treatmentCitation>
, figs. 17;
<treatmentCitation id="A44BEA0BF169AD6290A51305F6A31F1F" author="Ngoc-Ho" page="732" pageId="29" pageNumber="330" year="1997">
<bibRefCitation id="417BB1EBF169AD6290A51305F3CB1F78" author="Ngoc-Ho" box="[1276,1432,1806,1828]" pageId="29" pageNumber="330" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae). Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho 1997</bibRefCitation>
: 732
</treatmentCitation>
, fig. 1.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2555CC1AF169AD6294CE1347F7271FDC" blockId="29.[151,1436,1745,1920]" pageId="29" pageNumber="330">
<treatmentCitationGroup id="05FAEB34F169AD6294CE1347F7271FDC" pageId="29" pageNumber="330">
Not
<emphasis id="179E1008F169AD62949A1346F7C81F3E" box="[195,411,1868,1890]" italics="true" pageId="29" pageNumber="330">
<taxonomicName id="E2EAB799F169AD62949A1346F77B1F3E" ID-CoL="40689a9c-f8b4-44ce-9306-1a442d52759a" box="[195,296,1868,1890]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="330" phylum="Arthropoda" rank="genus">Laomedia</taxonomicName>
astacina—
</emphasis>
<treatmentCitation id="A44BEA0BF169AD6295C51346F40C1F3E" author="Sankolli" box="[412,607,1868,1890]" page="235" pageId="29" pageNumber="330" year="1971">
<bibRefCitation id="417BB1EBF169AD6295C51346F47F1F3E" author="Sankolli" box="[412,556,1868,1890]" pageId="29" pageNumber="330" refString="Sankolli, K. N. (1971) The Thalassinoidea (Crustacea, Anomura) of Maharashtra. Journal of the Bombay Natural History Society, 68, 94 - 106." type="journal article" year="1971">Sankolli 1971</bibRefCitation>
: 235
</treatmentCitation>
, figs. 1, 2 (=
<taxonomicName id="E2EAB799F169AD6296BF1346F5181F3E" ID-CoL="40689a9c-f8b4-44ce-9306-1a442d52759a" box="[742,843,1868,1890]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="330" phylum="Arthropoda" rank="genus">
<emphasis id="179E1008F169AD6296BF1346F5181F3E" box="[742,843,1868,1890]" italics="true" pageId="29" pageNumber="330">Laomedia</emphasis>
</taxonomicName>
sp., see below);
<treatmentCitation id="A44BEA0BF169AD6297AE1346F2E41F3E" author="Johnson" box="[1015,1207,1868,1890]" page="406" pageId="29" pageNumber="330" year="1972">
<bibRefCitation id="417BB1EBF169AD6297AE1346F2D51F3E" author="Johnson" box="[1015,1158,1868,1890]" pageId="29" pageNumber="330" refString="Johnson, D. S. (1972) The occurrence of the rare mud-shrimp, Laomedia astacina, in Malaysian waters. Journal of the marine Biological Association of India, 14, 406 - 408." type="journal article" year="1972">Johnson 1972</bibRefCitation>
: 406
</treatmentCitation>
(=
<taxonomicName id="E2EAB799F169AD6290821347F3CF1F3E" ID-CoL="4d2d954f-00af-4b80-b836-0f7258746ac5" box="[1243,1436,1868,1890]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="paucispinosa">
<emphasis id="179E1008F169AD6290821347F2A31F3E" box="[1243,1264,1869,1890]" italics="true" pageId="29" pageNumber="330">L.</emphasis>
cf.
<emphasis id="179E1008F169AD62914F1347F3CF1F3E" box="[1302,1436,1869,1890]" italics="true" pageId="29" pageNumber="330">paucispinosa</emphasis>
</taxonomicName>
<bibRefCitation id="417BB1EBF169AD62949E1361F73A1FDC" author="Ngoc-Ho" box="[199,361,1898,1920]" pageId="29" pageNumber="330" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae). Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho, 1997</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="71959C92F16AAD6194CE1358F4F61FDB" httpUri="https://zenodo.org/record/254447/files/figure.png" pageId="30" pageNumber="331" targetBox="[151,1436,195,1851]" targetPageId="30">
<paragraph id="2555CC1AF16AAD6194CE1358F4F61FDB" blockId="30.[151,1435,1874,1927]" pageId="30" pageNumber="331">
<emphasis id="179E1008F16AAD6194CE1358F74A1F3B" bold="true" box="[151,281,1874,1896]" pageId="30" pageNumber="331">FIGURE 22</emphasis>
.
<taxonomicName id="E2EAB799F16AAD61957F1358F4C81F34" authority="Banner, 1953" authorityName="Banner" authorityYear="1953" box="[294,667,1874,1896]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="tricristatus">
<emphasis id="179E1008F16AAD61957F1358F7C71F34" box="[294,404,1874,1896]" italics="true" pageId="30" pageNumber="331">Salmoneus</emphasis>
<emphasis id="179E1008F16AAD6195C31359F45A1F34" box="[410,521,1875,1896]" italics="true" pageId="30" pageNumber="331">tricristatus</emphasis>
Banner, 1953
</taxonomicName>
: ovigerous specimen (cl 5.6 mm) from Teluk Sekotong, Lombok, Indonesia (MZB Cru 4057), A—dorsal view; B—lateral view.
</paragraph>
</caption>
<caption id="71959C92F16BAD6094CE1246F2DE1EDD" httpUri="https://zenodo.org/record/254448/files/figure.png" pageId="31" pageNumber="332" targetBox="[154,1436,151,1596]" targetPageId="31">
<paragraph id="2555CC1AF16BAD6094CE1246F2DE1EDD" blockId="31.[151,1435,1612,1665]" pageId="31" pageNumber="332">
<emphasis id="179E1008F16BAD6094CE1246F77A1E3D" bold="true" box="[151,297,1612,1634]" pageId="31" pageNumber="332">FIGURE 23</emphasis>
.
<taxonomicName id="E2EAB799F16BAD60951E1246F4BE1E3E" authority="De Haan, 1841" authorityName="De Haan" authorityYear="1841" box="[327,749,1612,1634]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="332" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F16BAD60951E1246F44A1E3E" box="[327,537,1612,1634]" italics="true" pageId="31" pageNumber="332">Laomedia astacina</emphasis>
De Haan, 1841
</taxonomicName>
: female (cl 8.9 mm) from Lembar, Lombok, Indonesia (RMNH.CRUS.D.56754), A—dorsal view; B—lateral view; close-up of frontal region, dorsal view.
</paragraph>
</caption>
<subSubSection id="6DF09F91F16BAD6094CE12A7F7341F52" pageId="31" pageNumber="332" type="materials_examined">
<paragraph id="2555CC1AF16BAD6094CE12A7F7341F52" blockId="31.[151,1437,1709,2022]" pageId="31" pageNumber="332">
<emphasis id="179E1008F16BAD6094CE12A7F72A1E9A" bold="true" box="[151,377,1709,1734]" pageId="31" pageNumber="332">Material examined</emphasis>
.
<collectingCountry id="5DFD8C8AF16BAD6095DF12A7F7A41E9A" box="[390,503,1709,1734]" name="Indonesia" pageId="31" pageNumber="332">Indonesia</collectingCountry>
.
<specimenCount id="33EC0793F16BAD60965812A4F4311E9A" box="[513,610,1709,1734]" pageId="31" pageNumber="332" type="female">1 female</specimenCount>
(
<collectionCode id="43FB54DFF16BAD60962812A7F49F1E9A" LSID="urn:lsid:biocol.org:col:34992" box="[625,716,1709,1734]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="31" pageNumber="332">RMNH</collectionCode>
.
<collectionCode id="43FB54DFF16BAD60969612A7F54F1E9A" box="[719,796,1709,1734]" pageId="31" pageNumber="332">CRUS</collectionCode>
.D.56754), southern Lombok, Lembar, prawn ponds, mud flat, in burrows in banks of brackish stream, suction pump,
<quantity id="E21261FFF16BAD60971812D8F5CD1EB5" box="[833,926,1746,1770]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="31" pageNumber="332" unit="m" value="0.1" valueMax="0.2" valueMin="0.0">00.2 m</quantity>
at low tide, leg. D.L. Rahayu
<emphasis id="179E1008F16BAD6090A212D8F3661EB6" box="[1275,1333,1746,1770]" italics="true" pageId="31" pageNumber="332">et al.</emphasis>
,
<date id="5154EADAF16BAD60911D12D8F69C1F52" pageId="31" pageNumber="332" value="2014-05-14">14 May 2014</date>
[LB-St5-10].
</paragraph>
</subSubSection>
<subSubSection id="6DF09F91F16BAD60949E1313F7781FC2" pageId="31" pageNumber="332" type="description">
<paragraph id="2555CC1AF16BAD60949E1313F35A1F6E" blockId="31.[151,1437,1709,2022]" box="[199,1289,1817,1842]" pageId="31" pageNumber="332">
<emphasis id="179E1008F16BAD60949E1313F7021F6E" bold="true" box="[199,337,1817,1842]" pageId="31" pageNumber="332">Description</emphasis>
. See
<bibRefCitation id="417BB1EBF16BAD6095D61313F4BF1F6E" author="Sakai" box="[399,748,1817,1842]" pageId="31" pageNumber="332" refString="Sakai K. (1962) Systematic studies on Thalassinidea. I. Laomedia astacina De Haan. Publications of the Seto Marine Biological Laboratory, 10, 27 - 34." type="journal article" year="1962">Sakai (1962, with some errors)</bibRefCitation>
,
<bibRefCitation id="417BB1EBF16BAD6096AE1313F25E1F6E" author="Yaldwyn" box="[759,1037,1817,1842]" pageId="31" pageNumber="332" refString="Yaldwyn, J. C. &amp; Wear, R. G. (1972) The eastern Australian, burrowing mud-shrimp Laomedia healyi (Crustacea, Macrura Reptantia, Laomediidae) with notes on larvae of the genus Laomedia. Australian Zoologist, 17, 126 - 141." type="journal article" year="1972">Yaldwyn &amp; Wear (1972)</bibRefCitation>
, and
<bibRefCitation id="417BB1EBF16BAD6090111310F3551F6E" author="Ngoc-Ho" box="[1096,1286,1818,1842]" pageId="31" pageNumber="332" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae). Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho (1997)</bibRefCitation>
.
</paragraph>
<paragraph id="2555CC1AF16BAD60949E1337F7781FC2" blockId="31.[151,1437,1709,2022]" pageId="31" pageNumber="332">
<emphasis id="179E1008F16BAD60949E1337F7701F0A" bold="true" box="[199,291,1853,1878]" pageId="31" pageNumber="332">Habitat</emphasis>
. Mud flats and muddy sand flats, usually near estuaries or mangroves; in burrows or under rocks, mangrove wood, debris etc., also in fine mud in mangrove swamps, brackish prawn ponds. Depth range: intertidal to about
<specimenCount id="33EC0793F16BAD6094A0138FF7781FC2" box="[249,299,1925,1950]" pageId="31" pageNumber="332" type="generic">
<quantity id="E21261FFF16BAD6094A0138FF7741FC2" box="[249,295,1925,1950]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="332" unit="m" value="5.0">5 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="6DF09F91F16BAD60949E13A3F7D01FBA" pageId="31" pageNumber="332" type="distribution">
<paragraph id="2555CC1AF16BAD60949E13A3F7D01FBA" blockId="31.[151,1437,1709,2022]" pageId="31" pageNumber="332">
<emphasis id="179E1008F16BAD60949E13A3F70A1F9E" bold="true" box="[199,345,1961,1986]" pageId="31" pageNumber="332">Distribution</emphasis>
. Indo-West Pacific:
<collectingCountry id="5DFD8C8AF16BAD60961013A0F4DF1F9D" box="[585,652,1962,1985]" name="Japan" pageId="31" pageNumber="332">Japan</collectingCountry>
,
<collectingCountry id="5DFD8C8AF16BAD6096C313A0F4B21F9D" box="[666,737,1962,1985]" name="South Korea" pageId="31" pageNumber="332">Korea</collectingCountry>
,
<collectingCountry id="5DFD8C8AF16BAD6096B613A3F5661F9E" box="[751,821,1961,1986]" name="China" pageId="31" pageNumber="332">China</collectingCountry>
,
<collectingCountry id="5DFD8C8AF16BAD60971D13A3F5CA1F9E" box="[836,921,1961,1986]" name="Taiwan" pageId="31" pageNumber="332">Taiwan</collectingCountry>
,
<collectingCountry id="5DFD8C8AF16BAD6097FE13A3F25A1F9E" box="[935,1033,1961,1986]" name="Vietnam" pageId="31" pageNumber="332">Vietnam</collectingCountry>
,
<collectingCountry id="5DFD8C8AF16BAD60904E13A3F2D71F9E" box="[1047,1156,1961,1986]" name="Indonesia" pageId="31" pageNumber="332">Indonesia</collectingCountry>
(first record) (
<bibRefCitation id="417BB1EBF16BAD60916B13A0F7251FBA" author="Ngoc-Ho" pageId="31" pageNumber="332" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae). Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho 1997; present study</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6DF09F91F154AD5F949E149DF2691B94" pageId="32" pageNumber="333" type="discussion">
<paragraph id="2555CC1AF154AD5F949E149DF31419DB" blockId="32.[151,1437,151,968]" pageId="32" pageNumber="333">
<emphasis id="179E1008F154AD5F949E149DF76718EC" bold="true" box="[199,308,151,176]" pageId="32" pageNumber="333">Remarks</emphasis>
. Following the most recent assessment of
<taxonomicName id="E2EAB799F154AD5F97421492F5D818EC" box="[795,907,152,176]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="genus">
<emphasis id="179E1008F154AD5F97421492F5D818EC" box="[795,907,152,176]" italics="true" pageId="32" pageNumber="333">Laomedia</emphasis>
</taxonomicName>
in
<bibRefCitation id="417BB1EBF154AD5F97EB1492F23C18EC" author="Ngoc-Ho" box="[946,1135,152,176]" pageId="32" pageNumber="333" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae). Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho (1997)</bibRefCitation>
, the single specimen from Lombok is morphologically closest to the
<typeStatus id="FA5172B8F154AD5F962E14B7F4F41889" box="[631,679,189,213]" pageId="32" pageNumber="333">type</typeStatus>
species,
<taxonomicName id="E2EAB799F154AD5F975614B7F5DF1888" box="[783,908,189,212]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F154AD5F975614B7F5DF1888" box="[783,908,189,212]" italics="true" pageId="32" pageNumber="333">L. astacina</emphasis>
</taxonomicName>
. Based on the material examined by
<bibRefCitation id="417BB1EBF154AD5F916D14B7F6B018A4" author="Ngoc-Ho" pageId="32" pageNumber="333" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae). Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho (1997)</bibRefCitation>
,
<taxonomicName id="E2EAB799F154AD5F94A814EBF72318AB" box="[241,368,224,248]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F154AD5F94A814EBF72318AB" box="[241,368,224,248]" italics="true" pageId="32" pageNumber="333">L. astacina</emphasis>
</taxonomicName>
appeared to range from
<collectingCountry id="5DFD8C8AF154AD5F96D514EAF49F18AB" box="[652,716,224,247]" name="Japan" pageId="32" pageNumber="333">Japan</collectingCountry>
and
<collectingCountry id="5DFD8C8AF154AD5F975F14EAF51918AB" box="[774,842,224,247]" name="South Korea" pageId="32" pageNumber="333">Korea</collectingCountry>
south to
<collectingCountry id="5DFD8C8AF154AD5F97E114D5F24918A4" box="[952,1050,223,248]" name="Vietnam" pageId="32" pageNumber="333">Vietnam</collectingCountry>
; Johnsons (1972) specimen of
<taxonomicName id="E2EAB799F154AD5F91DC14EBF6A51940" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F154AD5F91DC14EBF6A51940" italics="true" pageId="32" pageNumber="333">L. astacina</emphasis>
</taxonomicName>
from
<collectingCountry id="5DFD8C8AF154AD5F951D150EF7F91941" box="[324,426,260,285]" name="Malaysia" pageId="32" pageNumber="333">Malaysia</collectingCountry>
was re-identified as
<taxonomicName id="E2EAB799F154AD5F96FF150FF5D61940" box="[678,901,260,285]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="paucispinosa">
<emphasis id="179E1008F154AD5F96FF150FF4EF1940" box="[678,700,261,284]" italics="true" pageId="32" pageNumber="333">L.</emphasis>
cf.
<emphasis id="179E1008F154AD5F96A8150FF5D61940" box="[753,901,261,284]" italics="true" pageId="32" pageNumber="333">paucispinosa</emphasis>
</taxonomicName>
<bibRefCitation id="417BB1EBF154AD5F97C9150FF2151940" author="Ngoc-Ho" box="[912,1094,260,285]" pageId="32" pageNumber="333" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae). Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho, 1997</bibRefCitation>
. Thus, prior to this study,
<taxonomicName id="E2EAB799F154AD5F91DC150FF6A51963" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F154AD5F91DC150FF6A51963" italics="true" pageId="32" pageNumber="333">L. astacina</emphasis>
</taxonomicName>
was believed to occur in temperate and subtropical waters of
<collectingCountry id="5DFD8C8AF154AD5F90591522F227191C" box="[1024,1140,295,320]" name="China" pageId="32" pageNumber="333">East Asia</collectingCountry>
, being “replaced” by
<taxonomicName id="E2EAB799F154AD5F91DC1523F7781938" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="paucispinosa">
<emphasis id="179E1008F154AD5F91DC1523F7781938" italics="true" pageId="32" pageNumber="333">L. paucispinosa</emphasis>
</taxonomicName>
in the more tropical parts of South-East Asia (
<bibRefCitation id="417BB1EBF154AD5F97381546F2401938" author="Ngoc-Ho" box="[865,1043,332,356]" pageId="32" pageNumber="333" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae). Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho 1997</bibRefCitation>
). However, our first record of
<taxonomicName id="E2EAB799F154AD5F91DC1547F6A519DB" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F154AD5F91DC1547F6A519DB" italics="true" pageId="32" pageNumber="333">L. astacina</emphasis>
</taxonomicName>
from central
<collectingCountry id="5DFD8C8AF154AD5F95D41565F7A919D4" box="[397,506,367,392]" name="Indonesia" pageId="32" pageNumber="333">Indonesia</collectingCountry>
suggests that this species extends well into the tropics south of the equator.
</paragraph>
<paragraph id="2555CC1AF154AD5F949E159EF4471AFB" blockId="32.[151,1437,151,968]" pageId="32" pageNumber="333">
The female from Lembar, southern Lombok, shows all the typical features of
<taxonomicName id="E2EAB799F154AD5F900A159FF28119F0" box="[1107,1234,405,428]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F154AD5F900A159FF28119F0" box="[1107,1234,405,428]" italics="true" pageId="32" pageNumber="333">L. astacina</emphasis>
</taxonomicName>
, as illustrated by
<bibRefCitation id="417BB1EBF154AD5F94CE15B2F700198C" author="Ngoc-Ho" box="[151,339,440,464]" pageId="32" pageNumber="333" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae). Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho (1997)</bibRefCitation>
, although a few differences were noted. For instance, in the female from Lombok, the postocular spines are blunter and less projecting compared to those of males from
<collectingCountry id="5DFD8C8AF154AD5F979A15D6F24619A9" box="[963,1045,476,501]" name="Taiwan" pageId="32" pageNumber="333">Taiwan</collectingCountry>
and
<collectingCountry id="5DFD8C8AF154AD5F901415D6F2FF19A9" box="[1101,1196,476,501]" name="Vietnam" pageId="32" pageNumber="333">Vietnam</collectingCountry>
(
<bibRefCitation id="417BB1EBF154AD5F90E515D6F68C1A44" author="Ngoc-Ho" pageId="32" pageNumber="333" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae). Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho 1997: fig. 1A, C</bibRefCitation>
). The chelipeds of the Lombok female are extremely hairy, covered by large and very dense tufts of setae (
<figureCitation id="BDD1D09FF154AD5F94C6162EF6A51A60" box="[159,246,548,573]" captionStart="FIGURE 23" captionStartId="31.[151,250,1612,1634]" captionTargetBox="[154,1436,151,1596]" captionTargetId="figure@31.[154,1439,151,1597]" captionTargetPageId="31" captionText="FIGURE 23. Laomedia astacina De Haan, 1841: female (cl 8.9 mm) from Lembar, Lombok, Indonesia (RMNH. CRUS. D. 56754), A—dorsal view; B—lateral view; close-up of frontal region, dorsal view." httpUri="https://zenodo.org/record/254448/files/figure.png" pageId="32" pageNumber="333">Fig. 23</figureCitation>
), completely concealing small spines on the dorsal surface of the palm and merus; Ngoc-Hos (1997) illustrations show chelipeds mostly denuded of setae. More material from
<collectingCountry id="5DFD8C8AF154AD5F9070164DF2C51A3C" box="[1065,1174,583,608]" name="Indonesia" pageId="32" pageNumber="333">Indonesia</collectingCountry>
and adjacent regions (
<collectingCountry id="5DFD8C8AF154AD5F94C71666F75A1AD9" box="[158,265,620,645]" name="Malaysia" pageId="32" pageNumber="333">Malaysia</collectingCountry>
,
<collectingCountry id="5DFD8C8AF154AD5F954C1666F7C51AD9" box="[277,406,620,645]" name="Philippines" pageId="32" pageNumber="333">Philippines</collectingCountry>
,
<collectingCountry id="5DFD8C8AF154AD5F95F81667F4281AD9" box="[417,635,620,645]" name="Papua New Guinea" pageId="32" pageNumber="333">Papua New Guinea</collectingCountry>
) is necessary to properly assess the status of what is currently seen as a tropical population of
<taxonomicName id="E2EAB799F154AD5F95C9169BF45F1AFB" box="[400,524,656,680]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F154AD5F95C9169BF45F1AFB" box="[400,524,656,680]" italics="true" pageId="32" pageNumber="333">L. astacina</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="2555CC1AF154AD5F949E16BEF2691B94" blockId="32.[151,1437,151,968]" pageId="32" pageNumber="333">
The record of
<taxonomicName id="E2EAB799F154AD5F953E16BFF7B71A90" box="[359,484,693,716]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F154AD5F953E16BFF7B71A90" box="[359,484,693,716]" italics="true" pageId="32" pageNumber="333">L. astacina</emphasis>
</taxonomicName>
from Maharashtra,
<collectingCountry id="5DFD8C8AF154AD5F969A16BEF4AC1A91" box="[707,767,692,717]" name="India" pageId="32" pageNumber="333">India</collectingCountry>
, by
<bibRefCitation id="417BB1EBF154AD5F977716BEF5B31A90" author="Sankolli" box="[814,992,692,717]" pageId="32" pageNumber="333" refString="Sankolli, K. N. (1971) The Thalassinoidea (Crustacea, Anomura) of Maharashtra. Journal of the Bombay Natural History Society, 68, 94 - 106." type="journal article" year="1971">Sankolli (1971)</bibRefCitation>
, although listed under the synonymy of
<taxonomicName id="E2EAB799F154AD5F94CE16D3F7441AB3" box="[151,279,728,752]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F154AD5F94CE16D3F7441AB3" box="[151,279,728,752]" italics="true" pageId="32" pageNumber="333">L. astacina</emphasis>
</taxonomicName>
by
<bibRefCitation id="417BB1EBF154AD5F951016D2F45A1AAC" author="Ngoc-Ho" box="[329,521,728,752]" pageId="32" pageNumber="333" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae). Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho (1997)</bibRefCitation>
, most likely refers to a different species, closely related to
<taxonomicName id="E2EAB799F154AD5F909E16D3F3141AB3" box="[1223,1351,728,752]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F154AD5F909E16D3F3141AB3" box="[1223,1351,728,752]" italics="true" pageId="32" pageNumber="333">L. astacina</emphasis>
</taxonomicName>
and
<taxonomicName id="E2EAB799F154AD5F91DC16D3F7781B48" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="paucispinosa">
<emphasis id="179E1008F154AD5F91DC16D3F7781B48" italics="true" pageId="32" pageNumber="333">L. paucispinosa</emphasis>
</taxonomicName>
. Sankollis single specimen differs from the Japanese specimens of
<taxonomicName id="E2EAB799F154AD5F900D16F7F2861B48" box="[1108,1237,765,788]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="astacina">
<emphasis id="179E1008F154AD5F900D16F7F2861B48" box="[1108,1237,765,788]" italics="true" pageId="32" pageNumber="333">L. astacina</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="417BB1EBF154AD5F914B16F6F3C41B49" author="Sakai" box="[1298,1431,764,789]" pageId="32" pageNumber="333" refString="Sakai K. (1962) Systematic studies on Thalassinidea. I. Laomedia astacina De Haan. Publications of the Seto Marine Biological Laboratory, 10, 27 - 34." type="journal article" year="1962">Sakai 1962</bibRefCitation>
;
<bibRefCitation id="417BB1EBF154AD5F94CE172AF7121B64" author="Ngoc-Ho" box="[151,321,800,824]" pageId="32" pageNumber="333" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae). Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho 1997</bibRefCitation>
) in the shape of the rostrum, which is narrowing distally, with two or three small, closely spaced teeth on the distolateral margins.
<bibRefCitation id="417BB1EBF154AD5F9646174EF4821B00" author="Sankolli" box="[543,721,836,861]" pageId="32" pageNumber="333" refString="Sankolli, K. N. (1971) The Thalassinoidea (Crustacea, Anomura) of Maharashtra. Journal of the Bombay Natural History Society, 68, 94 - 106." type="journal article" year="1971">Sankolli (1971)</bibRefCitation>
noted other differences between the Indian and the Japanese specimens, such as the absence of tubercles or teeth on the chelipeds, and the absence of a lateral notch on the uropod rami and telson. More material of
<taxonomicName id="E2EAB799F154AD5F96351786F48F1BF8" box="[620,732,908,932]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="333" phylum="Arthropoda" rank="genus">
<emphasis id="179E1008F154AD5F96351786F48F1BF8" box="[620,732,908,932]" italics="true" pageId="32" pageNumber="333">Laomedia</emphasis>
</taxonomicName>
from
<collectingCountry id="5DFD8C8AF154AD5F97781786F5091BF9" box="[801,858,908,933]" name="India" pageId="32" pageNumber="333">India</collectingCountry>
and other localities in the northern Indian Ocean is needed to clarify the taxonomic identity of the species reported by
<bibRefCitation id="417BB1EBF154AD5F97DD17A5F2651B94" author="Sankolli" box="[900,1078,943,968]" pageId="32" pageNumber="333" refString="Sankolli, K. N. (1971) The Thalassinoidea (Crustacea, Anomura) of Maharashtra. Journal of the Bombay Natural History Society, 68, 94 - 106." type="journal article" year="1971">Sankolli (1971)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>