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<document id="B6E54607DD1446419222AF2DEB3CF24B" ID-CLB-Dataset="26002" ID-DOI="10.11646/zootaxa.4657.1.2" ID-GBIF-Dataset="a7c3cd9d-d4be-4686-a198-3255c948beb8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3369801" ID-ZooBank="F27CB967-C001-436A-9F25-184D5484363C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565968506341" checkinUser="plazi" docAuthor="Perkins, Philip D. &amp; Bergsten, Johannes" docDate="2019" docId="AD076747FFD9FFCEFF6DF8F5B16B96AF" docLanguage="en" docName="zootaxa.4657.1.2.pdf" docOrigin="Zootaxa 4657 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Incoltorrida quintacostata Perkins &amp; Bergsten 2019, new species" docType="treatment" docVersion="12" lastPageNumber="70" masterDocId="513E1F3FFFD2FFC3FFFAFFB9B27A9138" masterDocTitle="New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae)" masterLastPageNumber="96" masterPageNumber="57" pageNumber="68" updateTime="1698751487288" updateUser="ExternalLinkService">
<mods:mods id="B183EDDEDA2FC2DBEF5CB7BFD5B6747B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DC54A6F2203A4BD38491A48FBC160434">
<mods:title id="0B575825E4781C604C32C2339709EEA1">New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae)</mods:title>
</mods:titleInfo>
<mods:name id="8E53303CAF6FC72911F1AD1999BF18D3" type="personal">
<mods:role id="61DEDFB933F70E6BBA3AA9A9583D7407">
<mods:roleTerm id="F1657E20ACDD6350676155C4A320346C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EB0FD3801EAE0E8EF11EE77EA2E48CA0">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:name id="279C474F5E2F66C27801F94C33FAF666" type="personal">
<mods:role id="118505E96788E3119BD2F81AA42ADF59">
<mods:roleTerm id="48160645653D4B0F012AA3F8B91CE69E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="743F79BAC15FBEEE5FB0B8545705B6D6">Bergsten, Johannes</mods:namePart>
</mods:name>
<mods:typeOfResource id="6647D7D7BEACD57047B6BCEB35C8EDD4">text</mods:typeOfResource>
<mods:relatedItem id="9CB928AC627697312B3434EAD21571D4" type="host">
<mods:titleInfo id="461B6811EB87B132FB5CDDF22F4C0A3E">
<mods:title id="A857DCC0AD6BD2BFD93B74EB0E36102E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="99E90650EAC5CD17ED2682DCF3D39144">
<mods:date id="62245563349BB57D75E70837BACE4D88">2019</mods:date>
<mods:detail id="E73D39A2E7C8FC8985F2795271658F57" type="pubDate">
<mods:number id="89E2314A9CF627E1B20862F6F4C2FA9F">2019-08-16</mods:number>
</mods:detail>
<mods:detail id="6F786E6CC2CD9DBD2A113A39121E16F5" type="volume">
<mods:number id="45745621254ABBDC52234F19F52B8638">4657</mods:number>
</mods:detail>
<mods:detail id="B93B9655D85345B7F6EF958A529A8FCC" type="issue">
<mods:number id="F997CEEED7081037A60CFCDD0CBAE6B6">1</mods:number>
</mods:detail>
<mods:extent id="97AFD36CAC71B32D87DC59C3078F5701" unit="page">
<mods:start id="06F55A44A571F346560C7B5380CF5556">57</mods:start>
<mods:end id="CC9E879D5F2A8954AEDADFC887A6419D">96</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="287E21320040E60A12A7A1EA1F448C0E">journal article</mods:classification>
<mods:identifier id="A25E61680EA593934E8DA4542E7AD21D" type="CLB-Dataset">26002</mods:identifier>
<mods:identifier id="702B4FFD9EC7E0ACF3EBB4029DF2770C" type="DOI">10.11646/zootaxa.4657.1.2</mods:identifier>
<mods:identifier id="1A963ED18C6A9310C23457EBEDD73225" type="GBIF-Dataset">a7c3cd9d-d4be-4686-a198-3255c948beb8</mods:identifier>
<mods:identifier id="1F00899B887A7969955B477D1AE18ADC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8ECCFFA80A8AD72880BF0E4EE228415A" type="Zenodo-Dep">3369801</mods:identifier>
<mods:identifier id="D56E6EED9FF5CED63B69375652323037" type="ZooBank">F27CB967-C001-436A-9F25-184D5484363C</mods:identifier>
</mods:mods>
<treatment id="AD076747FFD9FFCEFF6DF8F5B16B96AF" ID-DOI="http://doi.org/10.5281/zenodo.5934251" ID-GBIF-Taxon="159131432" ID-Zenodo-Dep="5934251" LSID="urn:lsid:plazi:treatment:AD076747FFD9FFCEFF6DF8F5B16B96AF" httpUri="http://treatment.plazi.org/id/AD076747FFD9FFCEFF6DF8F5B16B96AF" lastPageId="13" lastPageNumber="70" pageId="11" pageNumber="68">
<subSubSection id="6DB485DAFFD9FFC8FF6DF8F5B009965F" box="[151,627,1868,1895]" pageId="11" pageNumber="68" type="nomenclature">
<paragraph id="2511D651FFD9FFC8FF6DF8F5B009965F" blockId="11.[151,1000,1868,1932]" box="[151,627,1868,1895]" pageId="11" pageNumber="68">
<heading id="7E59613DFFD9FFC8FF6DF8F5B009965F" bold="true" box="[151,627,1868,1895]" fontSize="11" level="1" pageId="11" pageNumber="68" reason="1">
<emphasis id="17DA0A43FFD9FFC8FF6DF8F5B009965F" bold="true" box="[151,627,1868,1895]" pageId="11" pageNumber="68">
<taxonomicName id="E2AEADD2FFD9FFC8FF6DF8F5B3AE965F" authority="Perkins &amp; Bergsten, 2019" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[151,468,1868,1895]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="68" phylum="Arthropoda" rank="species" species="quintacostata" status="sp. nov.">
<emphasis id="17DA0A43FFD9FFC8FF6DF8F5B3AE965F" bold="true" box="[151,468,1868,1895]" italics="true" pageId="11" pageNumber="68">Incoltorrida quintacostata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="0CE9B738FFD9FFC8FE19F8F4B009965F" box="[483,627,1869,1895]" pageId="11" pageNumber="68" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6DB485DAFFD9FFC8FF6DF8C8B19296B4" box="[151,1000,1905,1932]" pageId="11" pageNumber="68" type="description">
<paragraph id="2511D651FFD9FFC8FF6DF8C8B19296B4" blockId="11.[151,1000,1868,1932]" box="[151,1000,1905,1932]" pageId="11" pageNumber="68">
<figureCitation id="BD95CAD4FFD9FFC8FF6DF8C8B29F96B3" box="[151,229,1905,1931]" captionStart="FIGURES 2–4" captionStartId="3.[151,264,2010,2035]" captionTargetBox="[271,1317,378,1988]" captionTargetId="figure@3.[271,1317,378,1988]" captionTargetPageId="3" captionText="FIGURES 2–4. Dorsal morphology. (2) Incoltorrida quintacostata; (3–4) I. magna." httpUri="https://zenodo.org/record/3369809/files/figure.png" pageId="11" pageNumber="68">Figs. 2</figureCitation>
,
<figureCitation id="BD95CAD4FFD9FFC8FF0BF8C8B28596B3" box="[241,255,1905,1931]" captionStart="FIGURE 7" captionStartId="9.[151,250,1697,1722]" captionTargetBox="[182,1398,192,1651]" captionTargetId="figure@9.[166,1421,181,1677]" captionTargetPageId="9" captionText="FIGURE 7. Incoltorrida quintacostata, habitus views of holotype." httpUri="https://zenodo.org/record/3369815/files/figure.png" pageId="11" pageNumber="68">7</figureCitation>
(habitus), 17, 18 (genitalia), 23 (larva), 34 (map), 42, 43 (habitat),
</paragraph>
</subSubSection>
<subSubSection id="6DB485DAFFD9FFCEFF6DF800B16B96AF" lastPageId="13" lastPageNumber="70" pageId="11" pageNumber="68" type="materials_examined">
<paragraph id="2511D651FFD9FFC8FF6DF800B17B96CF" blockId="11.[151,1437,1976,2039]" pageId="11" pageNumber="68">
<emphasis id="17DA0A43FFD9FFC8FF6DF800B33296EB" bold="true" box="[151,328,1976,2003]" pageId="11" pageNumber="68">Type Material.</emphasis>
<materialsCitation id="95C6DC0CFFD9FFC8FEAAF800B17B96CF" ID-GBIF-Occurrence="2332527092" collectingDate="2014-11-05" collectionCode="NHRS" collectorName="P. D. Perkins" elevation="1471" latitude="-20.5974" location="Ivato" longLatPrecision="8" longitude="47.213" pageId="11" pageNumber="68" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="FA1568F3FFD9FFC8FEAAF800B3C296EB" box="[336,440,1977,2003]" pageId="11" pageNumber="68" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="17DA0A43FFD9FFC8FDEDF801B0CC96EB" bold="true" box="[535,694,1976,2003]" pageId="11" pageNumber="68">
<collectingRegion id="E76A18B3FFD9FFC8FDEDF801B0CC96EB" box="[535,694,1976,2003]" country="Madagascar" name="Fianarantsoa" pageId="11" pageNumber="68">Fianarantsoa</collectingRegion>
</emphasis>
,
<quantity id="E2567BB4FFD9FFC8FD3EF801B17196EB" box="[708,779,1976,2003]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="68" unit="km" value="3.5">3.5km</quantity>
N
<location id="2071808AFFD9FFC8FCD4F800B11096EB" LSID="urn:lsid:plazi:treatment:AD076747FFD9FFCEFF6DF8F5B16B96AF:2071808AFFD9FFC8FCD4F800B11096EB" box="[814,874,1977,2003]" latitude="-20.5974" longLatPrecision="8" longitude="47.213" name="Ivato" pageId="11" pageNumber="68" stateProvince="Fianarantsoa">Ivato</location>
,
<geoCoordinate id="409AB096FFD9FFC8FC8FF800B67796EB" box="[885,1037,1976,2003]" degrees="20" direction="south" minutes="35.844" orientation="latitude" pageId="11" pageNumber="68" precision="1" value="-20.5974">20º 35.844’ S</geoCoordinate>
<geoCoordinate id="409AB096FFD9FFC8FBEEF800B6DA96EB" box="[1044,1184,1977,2003]" degrees="47" direction="east" minutes="12.78" orientation="longitude" pageId="11" pageNumber="68" precision="9" value="47.213">47º 12.78’ E</geoCoordinate>
, rock face seep beside hwy. 7, elev.
<quantity id="E2567BB4FFD9FFC8FED0F864B3FB96CF" box="[298,385,2013,2039]" metricMagnitude="3" metricUnit="m" metricValue="1.4709999999999999" pageId="11" pageNumber="68" unit="m" value="1471.0">
<elevation id="AE833162FFD9FFC8FED0F864B3FB96CF" box="[298,385,2013,2039]" metricMagnitude="3" metricUnit="m" metricValue="1.4709999999999999" pageId="11" pageNumber="68" unit="m" value="1471.0">1471 m</elevation>
</quantity>
,
<date id="5110F091FFD9FFC8FE77F865B38096CF" box="[397,506,2012,2039]" pageId="11" pageNumber="68" value="2014-11-05">
<collectingDate id="41540979FFD9FFC8FE77F865B38096CF" box="[397,506,2012,2039]" pageId="11" pageNumber="68" value="2014-11-05">5 xi 2014</collectingDate>
</date>
,
<collectorName id="885BB387FFD9FFC8FDFFF864B0EC96CF" box="[517,662,2013,2039]" pageId="11" pageNumber="68">P. D. Perkins</collectorName>
(
<collectionCode id="43BF4E94FFD9FFC8FD5FF864B08C96CF" box="[677,758,2013,2039]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="11" pageNumber="68">NHRS</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="71D186D9FFDEFFCFFF6DF8FEB17C9658" box="[151,774,1863,1888]" httpUri="https://zenodo.org/record/3369823/files/figure.png" pageId="12" pageNumber="69" startId="12.[151,250,1863,1888]" targetBox="[175,1403,208,1802]" targetPageId="12">
<paragraph id="2511D651FFDEFFCFFF6DF8FEB17C9658" blockId="12.[151,774,1863,1888]" box="[151,774,1863,1888]" pageId="12" pageNumber="69">
<emphasis id="17DA0A43FFDEFFCFFF6DF8FEB35A9658" bold="true" box="[151,288,1863,1888]" pageId="12" pageNumber="69">FIGURE 10.</emphasis>
<taxonomicName id="E2AEADD2FFDEFFCFFEDCF8F1B3979658" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[294,493,1864,1888]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="69" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="17DA0A43FFDEFFCFFEDCF8F1B3979658" box="[294,493,1864,1888]" italics="true" pageId="12" pageNumber="69">Incoltorrida magna</emphasis>
</taxonomicName>
, habitus views of holotype.
</paragraph>
</caption>
<paragraph id="2511D651FFDEFFCEFF3DF836B05C9323" blockId="12.[151,1437,1934,2033]" lastBlockId="13.[151,1437,153,1943]" lastPageId="13" lastPageNumber="70" pageId="12" pageNumber="69">
<materialsCitation id="95C6DC0CFFDEFFCFFF3DF836B1149691" ID-GBIF-Occurrence="2332527072" box="[199,878,1934,1961]" collectionCode="NHRS, MCZ" location="Same" pageId="12" pageNumber="69" specimenCount="2" typeStatus="paratype">
<emphasis id="17DA0A43FFDEFFCFFF3DF836B3459691" bold="true" box="[199,319,1935,1961]" pageId="12" pageNumber="69">
<typeStatus id="FA1568F3FFDEFFCFFF3DF836B3459691" box="[199,319,1935,1961]" pageId="12" pageNumber="69" type="paratype">Paratypes</typeStatus>
</emphasis>
(193):
<location id="2071808AFFDEFFCFFE68F836B3AA9691" LSID="urn:lsid:plazi:treatment:AD076747FFD9FFCEFF6DF8F5B16B96AF:2071808AFFDEFFCFFE68F836B3AA9691" box="[402,464,1935,1961]" name="Same" pageId="12" pageNumber="69">Same</location>
data as
<typeStatus id="FA1568F3FFDEFFCFFDD7F837B0F59690" box="[557,655,1934,1960]" pageId="12" pageNumber="69" type="holotype">holotype</typeStatus>
(19:
<collectionCode id="43BF4E94FFDEFFCFFD36F836B1639691" box="[716,793,1935,1961]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="12" pageNumber="69">NHRS</collectionCode>
,
<collectionCode id="43BF4E94FFDEFFCFFCDFF836B11D9691" box="[805,871,1935,1961]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="12" pageNumber="69" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="95C6DC0CFFDEFFCFFC80F837B73D96C9" ID-GBIF-Occurrence="2332527089" collectingDate="2004-11-23" collectionCode="BMNH" collectorName="Balke" county="Anjiabe Ambony" elevation="303" location="Antsabe" longitude="13.6093" municipality="Ambilobe" pageId="12" pageNumber="69" specimenCode="MD16, BMNH 670734, BMNH 670735" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<emphasis id="17DA0A43FFDEFFCFFC80F837B6759691" bold="true" box="[890,1039,1934,1961]" pageId="12" pageNumber="69">
<collectingRegion id="E76A18B3FFDEFFCFFC80F837B6759691" box="[890,1039,1934,1961]" country="Madagascar" name="Antsiranana" pageId="12" pageNumber="69">Antsiranana</collectingRegion>
</emphasis>
,
<collectingCounty id="CC70AEDDFFDEFFCFFBE7F837B69B9690" box="[1053,1249,1934,1960]" pageId="12" pageNumber="69">Anjiabe Ambony</collectingCounty>
:
<collectingMunicipality id="C5754C2BFFDEFFCFFB11F837B7259690" box="[1259,1375,1934,1960]" pageId="12" pageNumber="69">Ambilobe</collectingMunicipality>
:
<location id="2071808AFFDEFFCFFA93F836B2B396F4" LSID="urn:lsid:plazi:treatment:AD076747FFD9FFCEFF6DF8F5B16B96AF:2071808AFFDEFFCFFA93F836B2B396F4" county="Anjiabe Ambony" longitude="13.6093" municipality="Ambilobe" name="Antsabe" pageId="12" pageNumber="69" stateProvince="Antsiranana">Antsabe</location>
stairways-like cascade with vertical (!) steps, exposed, extremely hot day, N:
<geoCoordinate id="409AB096FFDEFFCFFBD0F80BB6CD96F5" box="[1066,1207,1970,1997]" pageId="12" pageNumber="69">-13.60930 E</geoCoordinate>
:
<geoCoordinate id="409AB096FFDEFFCFFB38F80AB75496F5" box="[1218,1326,1971,1997]" pageId="12" pageNumber="69">48.72120</geoCoordinate>
, elev.
<quantity id="E2567BB4FFDEFFCFFA88F80AB2D596C9" metricMagnitude="2" metricUnit="m" metricValue="3.03" pageId="12" pageNumber="69" unit="m" value="303.0">
<elevation id="AE833162FFDEFFCFFA88F80AB2D596C9" metricMagnitude="2" metricUnit="m" metricValue="3.03" pageId="12" pageNumber="69" unit="m" value="303.0">303 m</elevation>
</quantity>
,
<date id="5110F091FFDEFFCFFF40F86EB34996C9" box="[186,307,2006,2033]" pageId="12" pageNumber="69" value="2004-11-23">
<collectingDate id="41540979FFDEFFCFFF40F86EB34996C9" box="[186,307,2006,2033]" pageId="12" pageNumber="69" value="2004-11-23">23 xi 2004</collectingDate>
</date>
,
<collectorName id="885BB387FFDEFFCFFEC7F86FB30496C8" box="[317,382,2006,2032]" pageId="12" pageNumber="69">Balke</collectorName>
<emphasis id="17DA0A43FFDEFFCFFE7EF86EB3CE96C8" box="[388,436,2006,2032]" italics="true" pageId="12" pageNumber="69">et al</emphasis>
. (P25
<specimenCode id="75087E2AFFDEFFCFFE0CF86FB03B96C9" box="[502,577,2006,2033]" collectionCode="MD" country="Germany" name="Museum Donaueschingen" pageId="12" pageNumber="69">MD16</specimenCode>
) (8
<collectionCode id="43BF4E94FFDEFFCFFD96F86EB0BE96C9" box="[620,708,2007,2033]" pageId="12" pageNumber="69">BMNH</collectionCode>
; 2 DNA extractions, #’s
<specimenCode id="75087E2AFFDEFFCFFC2BF86EB6F896C8" box="[977,1154,2006,2033]" collectionCode="BMNH" pageId="12" pageNumber="69">BMNH 670734</specimenCode>
,
<specimenCode id="75087E2AFFDEFFCFFB76F86EB74596C9" box="[1164,1343,2006,2033]" collectionCode="BMNH" pageId="12" pageNumber="69">BMNH 670735</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="95C6DC0CFFDEFFCEFAA8F86FB0F191C3" ID-GBIF-Occurrence="2332527093" collectingDate="2012-11-28" collectionCode="NHRS" collectorName="Bergsten &amp; R. Bukontaite &amp; J. H. Randriamihaja &amp; T. Ranarilalatiana" county="Diana" elevation="296" lastPageId="13" lastPageNumber="70" location="Galoko mountains" longitude="13.60931" municipality="Antsaba" pageId="12" pageNumber="69" specimenCode="MAD12-31" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<collectingCounty id="CC70AEDDFFDEFFCFFAA8F86FB7E396C8" box="[1362,1433,2006,2032]" pageId="12" pageNumber="69">Diana</collectingCounty>
:
<collectingMunicipality id="C5754C2BFFDFFFCEFF6DFF20B28D918B" box="[151,247,153,179]" pageId="13" pageNumber="70">Antsaba</collectingMunicipality>
:
<location id="2071808AFFDFFFCEFEFBFF20B3A8918B" LSID="urn:lsid:plazi:treatment:AD076747FFD9FFCEFF6DF8F5B16B96AF:2071808AFFDFFFCEFEFBFF20B3A8918B" box="[257,466,153,179]" county="Diana" longitude="13.60931" municipality="Antsaba" name="Galoko mountains" pageId="13" pageNumber="70" stateProvince="Antsiranana">Galoko mountains</location>
,
<quantity id="E2567BB4FFDFFFCEFE26FF20B053918B" box="[476,553,153,179]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="13" pageNumber="70" unit="km" value="3.4">
<locationDeviation id="34A3CF73FFDFFFCEFE26FF20B053918B" box="[476,553,153,179]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="13" pageNumber="70" unit="km" value="3.4">3.4 km</locationDeviation>
</quantity>
NW from
<location id="2071808AFFDFFFCEFD65FF20B084918B" LSID="urn:lsid:plazi:treatment:AD076747FFD9FFCEFF6DF8F5B16B96AF:2071808AFFDFFFCEFD65FF20B084918B" box="[671,766,153,179]" county="Diana" longitude="13.60931" municipality="Antsaba" name="Anstaba" pageId="13" pageNumber="70" stateProvince="Antsiranana">Anstaba</location>
,
<geoCoordinate id="409AB096FFDFFFCEFCF2FF20B67E918B" box="[776,1028,153,179]" pageId="13" pageNumber="70">S13.60931 E48.72129</geoCoordinate>
, aspirator, forceps, sieves: hygropetric rocks and pools, elev.
<quantity id="E2567BB4FFDFFFCEFE55FF04B38D91EF" box="[431,503,189,215]" metricMagnitude="2" metricUnit="m" metricValue="2.96" pageId="13" pageNumber="70" unit="m" value="296.0">
<elevation id="AE833162FFDFFFCEFE55FF04B38D91EF" box="[431,503,189,215]" metricMagnitude="2" metricUnit="m" metricValue="2.96" pageId="13" pageNumber="70" unit="m" value="296.0">296 m</elevation>
</quantity>
,
<date id="5110F091FFDFFFCEFDF8FF04B00191EF" box="[514,635,189,215]" pageId="13" pageNumber="70" value="2012-11-28">
<collectingDate id="41540979FFDFFFCEFDF8FF04B00191EF" box="[514,635,189,215]" pageId="13" pageNumber="70" value="2012-11-28">28 xi 2012</collectingDate>
</date>
, elev.
<quantity id="E2567BB4FFDFFFCEFD45FF04B17D91EF" box="[703,775,189,215]" metricMagnitude="2" metricUnit="m" metricValue="2.96" pageId="13" pageNumber="70" unit="m" value="296.0">
<elevation id="AE833162FFDFFFCEFD45FF04B17D91EF" box="[703,775,189,215]" metricMagnitude="2" metricUnit="m" metricValue="2.96" pageId="13" pageNumber="70" unit="m" value="296.0">296 m</elevation>
</quantity>
,
<date id="5110F091FFDFFFCEFCE8FF04B1F191EF" box="[786,907,189,215]" pageId="13" pageNumber="70" value="2012-11-28">
<collectingDate id="41540979FFDFFFCEFCE8FF04B1F191EF" box="[786,907,189,215]" pageId="13" pageNumber="70" value="2012-11-28">28 xi 2012</collectingDate>
</date>
, J.
<collectorName id="885BB387FFDFFFCEFC56FF04B66891EF" box="[940,1042,189,215]" pageId="13" pageNumber="70">Bergsten</collectorName>
,
<collectorName id="885BB387FFDFFFCEFBE7FF04B6C091EF" box="[1053,1210,189,215]" pageId="13" pageNumber="70">R. Bukontaite</collectorName>
,
<collectorName id="885BB387FFDFFFCEFB3EFF04B7E691EF" box="[1220,1436,189,215]" pageId="13" pageNumber="70">J.H. Randriamihaja</collectorName>
&amp;
<collectorName id="885BB387FFDFFFCEFF48FF58B30D91C3" box="[178,375,225,251]" pageId="13" pageNumber="70">T. Ranarilalatiana</collectorName>
(
<specimenCode id="75087E2AFFDFFFCEFE7EFF58B07091C3" box="[388,522,225,251]" collectionCode="MAD" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="13" pageNumber="70">MAD12-31</specimenCode>
) (5
<collectionCode id="43BF4E94FFDFFFCEFDCEFF58B0FE91C3" box="[564,644,225,251]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="13" pageNumber="70">NHRS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="95C6DC0CFFDFFFCEFD6FFF58B1E2907B" ID-GBIF-Occurrence="2332527075" collectingDate="2012-11-25" collectionCode="NHRS, BMNH, PBZT, MBC" collectorName="Bergsten &amp; Bukontaite &amp; J. H. Randriamihaja &amp; T. Ranarilalatiana" county="Diana" elevation="263" location="Galoko mountains" longitude="13.60974" municipality="Antsaba" pageId="13" pageNumber="70" specimenCode="MAD12-26" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<collectingCounty id="CC70AEDDFFDFFFCEFD6FFF58B0A691C3" box="[661,732,225,251]" pageId="13" pageNumber="70">Diana</collectingCounty>
:
<collectingMunicipality id="C5754C2BFFDFFFCEFD19FF58B13991C3" box="[739,835,225,251]" pageId="13" pageNumber="70">Antsaba</collectingMunicipality>
:
<location id="2071808AFFDFFFCEFCB6FF58B66691C3" LSID="urn:lsid:plazi:treatment:AD076747FFD9FFCEFF6DF8F5B16B96AF:2071808AFFDFFFCEFCB6FF58B66691C3" box="[844,1052,225,251]" county="Diana" longitude="13.60974" municipality="Antsaba" name="Galoko mountains" pageId="13" pageNumber="70" stateProvince="Antsiranana">Galoko mountains</location>
,
<geoCoordinate id="409AB096FFDFFFCEFBDFFF58B75B91C3" box="[1061,1313,224,251]" pageId="13" pageNumber="70">S13.60974 E48.72175</geoCoordinate>
, sieves and aspirator: hygropetric rocks and water pools, elev.
<quantity id="E2567BB4FFDFFFCEFD3EFEBCB1719027" box="[708,779,261,287]" metricMagnitude="2" metricUnit="m" metricValue="2.63" pageId="13" pageNumber="70" unit="m" value="263.0">
<elevation id="AE833162FFDFFFCEFD3EFEBCB1719027" box="[708,779,261,287]" metricMagnitude="2" metricUnit="m" metricValue="2.63" pageId="13" pageNumber="70" unit="m" value="263.0">263 m</elevation>
</quantity>
,
<date id="5110F091FFDFFFCEFCECFEBDB1F79027" box="[790,909,260,287]" pageId="13" pageNumber="70" value="2012-11-25">
<collectingDate id="41540979FFDFFFCEFCECFEBDB1F79027" box="[790,909,260,287]" pageId="13" pageNumber="70" value="2012-11-25">25 xi 2012</collectingDate>
</date>
, J.
<collectorName id="885BB387FFDFFFCEFC54FEBCB66E9027" box="[942,1044,261,287]" pageId="13" pageNumber="70">Bergsten</collectorName>
, R.
<collectorName id="885BB387FFDFFFCEFBC7FEBCB6C19027" box="[1085,1211,261,287]" pageId="13" pageNumber="70">Bukontaite</collectorName>
,
<collectorName id="885BB387FFDFFFCEFB3FFEBCB7E69027" box="[1221,1436,261,287]" pageId="13" pageNumber="70">J.H. Randriamihaja</collectorName>
&amp;
<collectorName id="885BB387FFDFFFCEFF49FE90B303907B" box="[179,377,297,323]" pageId="13" pageNumber="70">T. Ranarilalatiana</collectorName>
(
<specimenCode id="75087E2AFFDFFFCEFE7DFE90B077907C" box="[391,525,297,324]" collectionCode="MAD" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="13" pageNumber="70">MAD12-26</specimenCode>
) (62
<collectionCode id="43BF4E94FFDFFFCEFDBCFE90B0E9907B" box="[582,659,297,323]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="13" pageNumber="70">NHRS</collectionCode>
,
<collectionCode id="43BF4E94FFDFFFCEFD65FE90B08C907B" box="[671,758,297,323]" pageId="13" pageNumber="70">BMNH</collectionCode>
,
<collectionCode id="43BF4E94FFDFFFCEFCFAFE90B13D907B" box="[768,839,297,323]" country="Madagascar" httpUri="http://grbio.org/cool/a6tz-5rcd" name="Parc Botanique et Zoologique de Tsimbazaza" pageId="13" pageNumber="70">PBZT</collectionCode>
/
<collectionCode id="43BF4E94FFDFFFCEFCAAFE90B1EA907B" box="[848,912,297,323]" country="USA" httpUri="http://grbio.org/cool/tqbp-wnpd" name="Montgomery Botanical Center" pageId="13" pageNumber="70">MBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="95C6DC0CFFDFFFCEFC58FE91B33790B3" ID-GBIF-Occurrence="2332527073" collectingDate="2014-11-05" collectionCode="NHRS, MCZ" collectorName="Perkins" elevation="1415" latitude="-20.233906" location="3.2 km S Ambohimanjaka" longLatPrecision="1" longitude="47.09319" pageId="13" pageNumber="70" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<emphasis id="17DA0A43FFDFFFCEFC58FE91B63B907B" bold="true" box="[930,1089,296,323]" pageId="13" pageNumber="70">
<collectingRegion id="E76A18B3FFDFFFCEFC58FE91B63B907B" box="[930,1089,296,323]" country="Madagascar" name="Fianarantsoa" pageId="13" pageNumber="70">Fianarantsoa</collectingRegion>
</emphasis>
,
<location id="2071808AFFDFFFCEFBB4FE90B714907B" LSID="urn:lsid:plazi:treatment:AD076747FFD9FFCEFF6DF8F5B16B96AF:2071808AFFDFFFCEFBB4FE90B714907B" box="[1102,1390,297,323]" latitude="-20.233906" longLatPrecision="1" longitude="47.09319" name="3.2 km S Ambohimanjaka" pageId="13" pageNumber="70" stateProvince="Fianarantsoa">
<quantity id="E2567BB4FFDFFFCEFBB4FE90B6EF907B" box="[1102,1173,297,323]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="70" unit="km" value="3.2">3.2km</quantity>
S Ambohimanjaka
</location>
,
<geoCoordinate id="409AB096FFDFFFCEFA82FE90B36A905F" degrees="20" direction="south" minutes="14.0343" orientation="latitude" pageId="13" pageNumber="70" precision="1" value="-20.233906">20º 14.0343’ S</geoCoordinate>
<geoCoordinate id="409AB096FFDFFFCEFEECFEF4B3C4905F" box="[278,446,332,359]" degrees="47" direction="east" minutes="5.59145" orientation="longitude" pageId="13" pageNumber="70" precision="1" value="47.09319">47º 5.59145’ E</geoCoordinate>
, waterfall and hygropetric habitat near hwy. 7, elev.
<quantity id="E2567BB4FFDFFFCEFBFDFEF5B624905F" box="[1031,1118,332,359]" metricMagnitude="3" metricUnit="m" metricValue="1.415" pageId="13" pageNumber="70" unit="m" value="1415.0">
<elevation id="AE833162FFDFFFCEFBFDFEF5B624905F" box="[1031,1118,332,359]" metricMagnitude="3" metricUnit="m" metricValue="1.415" pageId="13" pageNumber="70" unit="m" value="1415.0">1415 m</elevation>
</quantity>
,
<date id="5110F091FFDFFFCEFB93FEF5B6AF905F" box="[1129,1237,332,359]" pageId="13" pageNumber="70" value="2014-11-05">
<collectingDate id="41540979FFDFFFCEFB93FEF5B6AF905F" box="[1129,1237,332,359]" pageId="13" pageNumber="70" value="2014-11-05">5 xi 2014</collectingDate>
</date>
, P. D.
<collectorName id="885BB387FFDFFFCEFAE6FEF4B70A905F" box="[1308,1392,333,359]" pageId="13" pageNumber="70">Perkins</collectorName>
(95
<collectionCode id="43BF4E94FFDFFFCEFF6DFEC8B29890B3" box="[151,226,369,395]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="13" pageNumber="70">NHRS</collectionCode>
&amp;
<collectionCode id="43BF4E94FFDFFFCEFEFEFEC8B33C90B3" box="[260,326,369,395]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="13" pageNumber="70" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="95C6DC0CFFDFFFCEFEACFEC8B34290EB" ID-GBIF-Occurrence="2332527087" collectingDate="2013-12-14" collectionCode="NHRS" collectorName="R. S. Manombo &amp; Randriamihaja &amp; Ranarilalatiana" county="Atsimo Antsinanana" elevation="21" location="Rearatra" longitude="23.006184" municipality="Parcelle" pageId="13" pageNumber="70" specimenCode="MAD13- 73" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<collectingCounty id="CC70AEDDFFDFFFCEFEACFEC8B04790B3" box="[342,573,369,395]" pageId="13" pageNumber="70">Atsimo Antsinanana</collectingCounty>
:
<collectorName id="885BB387FFDFFFCEFDBCFEC8B08F90B3" box="[582,757,369,395]" pageId="13" pageNumber="70">R.S. Manombo</collectorName>
:
<collectingMunicipality id="C5754C2BFFDFFFCEFD07FEC8B12D90B3" box="[765,855,369,395]" pageId="13" pageNumber="70">Parcelle</collectingMunicipality>
I,
<location id="2071808AFFDFFFCEFC8EFEC8B1AC90B3" LSID="urn:lsid:plazi:treatment:AD076747FFD9FFCEFF6DF8F5B16B96AF:2071808AFFDFFFCEFC8EFEC8B1AC90B3" box="[884,982,369,395]" county="Atsimo Antsinanana" longitude="23.006184" municipality="Parcelle" name="Rearatra" pageId="13" pageNumber="70" stateProvince="Fianarantsoa">Rearatra</location>
,
<location id="2071808AFFDFFFCEFC1AFEC8B66C90B3" LSID="urn:lsid:plazi:treatment:AD076747FFD9FFCEFF6DF8F5B16B96AF:2071808AFFDFFFCEFC1AFEC8B66C90B3" box="[992,1046,369,395]" county="Atsimo Antsinanana" longitude="23.006184" municipality="Parcelle" name="Piste" pageId="13" pageNumber="70" stateProvince="Fianarantsoa">Piste</location>
56:
<geoCoordinate id="409AB096FFDFFFCEFBBCFEC8B71190B3" box="[1094,1387,369,395]" pageId="13" pageNumber="70">S23.006183 E47.7338833</geoCoordinate>
, GB nets and sieves, forest stream with pools, elev.
<quantity id="E2567BB4FFDFFFCEFD5AFE2CB0A19097" box="[672,731,405,431]" metricMagnitude="1" metricUnit="m" metricValue="2.1" pageId="13" pageNumber="70" unit="m" value="21.0">
<elevation id="AE833162FFDFFFCEFD5AFE2CB0A19097" box="[672,731,405,431]" metricMagnitude="1" metricUnit="m" metricValue="2.1" pageId="13" pageNumber="70" unit="m" value="21.0">21 m</elevation>
</quantity>
,
<date id="5110F091FFDFFFCEFD1CFE2CB11D9097" box="[742,871,405,431]" pageId="13" pageNumber="70" value="2013-12-14">
<collectingDate id="41540979FFDFFFCEFD1CFE2CB11D9097" box="[742,871,405,431]" pageId="13" pageNumber="70" value="2013-12-14">14 xii 2013</collectingDate>
</date>
, J.H.
<collectorName id="885BB387FFDFFFCEFC6DFE2CB63E9097" box="[919,1092,405,431]" pageId="13" pageNumber="70">Randriamihaja</collectorName>
&amp; T.
<collectorName id="885BB387FFDFFFCEFB80FE2CB75C9097" box="[1146,1318,405,431]" pageId="13" pageNumber="70">Ranarilalatiana</collectorName>
(
<specimenCode id="75087E2AFFDFFFCEFACEFE2CB2CE90EB" collectionCode="MAD" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="13" pageNumber="70">MAD13- 73</specimenCode>
) (3
<collectionCode id="43BF4E94FFDFFFCEFF1BFE00B34B90EB" box="[225,305,441,467]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="13" pageNumber="70">NHRS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="95C6DC0CFFDFFFCEFEB9FE00B3DF9323" ID-GBIF-Occurrence="2332527066" collectingDate="2014-11-01" collectionCode="MAD" collectorName="Amoron'i Mania &amp; J. Bergsten &amp; T. Ranarilalatiana &amp; Holmgren" elevation="1700" latitude="-20.7722" location="Ankazomivady forest" longLatPrecision="7" longitude="47.1809" municipality="Ambalamanakana" pageId="13" pageNumber="70" specimenCode="MAD14-02" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<geoCoordinate id="409AB096FFDFFFCEFEB9FE00B3D790EB" box="[323,429,441,467]" degrees="20.7722" direction="south" orientation="latitude" pageId="13" pageNumber="70" precision="5" value="-20.7722">20.7722S</geoCoordinate>
<geoCoordinate id="409AB096FFDFFFCEFE4EFE00B05990EB" box="[436,547,441,467]" degrees="47.1809" direction="east" orientation="longitude" pageId="13" pageNumber="70" precision="5" value="47.1809">47.1809E</geoCoordinate>
;
<collectorName id="885BB387FFDFFFCEFDD6FE00B09190EB" box="[556,747,441,467]" pageId="13" pageNumber="70">Amoron’i Mania</collectorName>
,
<quantity id="E2567BB4FFDFFFCEFD0CFE00B15290EB" box="[758,808,441,467]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="70" unit="km" value="3.0">3km</quantity>
south of
<collectingMunicipality id="C5754C2BFFDFFFCEFC77FE00B62D90EB" box="[909,1111,441,467]" pageId="13" pageNumber="70">Ambalamanakana</collectingMunicipality>
next to RN7,
<location id="2071808AFFDFFFCEFB0BFE00B2A090CF" LSID="urn:lsid:plazi:treatment:AD076747FFD9FFCEFF6DF8F5B16B96AF:2071808AFFDFFFCEFB0BFE00B2A090CF" latitude="-20.7722" longLatPrecision="7" longitude="47.1809" municipality="Ambalamanakana" name="Ankazomivady forest" pageId="13" pageNumber="70" stateProvince="Fianarantsoa">Ankazomivady forest</location>
, hygropetric rocks and marsh with vegetation, elev.
<quantity id="E2567BB4FFDFFFCEFCD1FE64B1FE90CF" box="[811,900,477,503]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="70" unit="m" value="1700.0">
<elevation id="AE833162FFDFFFCEFCD1FE64B1FE90CF" box="[811,900,477,503]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="70" unit="m" value="1700.0">1700 m</elevation>
</quantity>
,
<date id="5110F091FFDFFFCEFC6BFE64B67A90CF" box="[913,1024,477,503]" pageId="13" pageNumber="70" value="2014-11-01">
<collectingDate id="41540979FFDFFFCEFC6BFE64B67A90CF" box="[913,1024,477,503]" pageId="13" pageNumber="70" value="2014-11-01">1 xi 2014</collectingDate>
</date>
,
<collectorName id="885BB387FFDFFFCEFBF6FE64B6F690CF" box="[1036,1164,477,503]" pageId="13" pageNumber="70">J. Bergsten</collectorName>
,
<collectorName id="885BB387FFDFFFCEFB62FE64B71A90CF" box="[1176,1376,477,503]" pageId="13" pageNumber="70">T. Ranarilalatiana</collectorName>
&amp; S.
<collectorName id="885BB387FFDFFFCEFF6DFDB8B3739323" box="[151,265,513,539]" pageId="13" pageNumber="70">Holmgren</collectorName>
(
<specimenCode id="75087E2AFFDFFFCEFEE2FDB8B3E79323" box="[280,413,513,539]" collectionCode="MAD" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="13" pageNumber="70">
<collectionCode id="43BF4E94FFDFFFCEFEE2FDB8B3249323" box="[280,350,513,539]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="13" pageNumber="70">MAD</collectionCode>
14-02
</specimenCode>
)
</materialsCitation>
(1
<collectionCode id="43BF4E94FFDFFFCEFE30FDB8B0609323" box="[458,538,513,539]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="13" pageNumber="70">NHRS</collectionCode>
).
</paragraph>
<paragraph id="2511D651FFDFFFCEFF3DFD9DB67F929F" blockId="13.[151,1437,153,1943]" pageId="13" pageNumber="70">
<emphasis id="17DA0A43FFDFFFCEFF3DFD9DB3A89307" bold="true" box="[199,466,548,575]" pageId="13" pageNumber="70">Differential Diagnosis.</emphasis>
Differentiated from all other known
<taxonomicName id="E2AEADD2FFDFFFCEFC91FD9CB18E9307" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[875,1012,549,575]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="17DA0A43FFDFFFCEFC91FD9CB18E9307" box="[875,1012,549,575]" italics="true" pageId="13" pageNumber="70">Incoltorrida</emphasis>
</taxonomicName>
, except
<taxonomicName id="E2AEADD2FFDFFFCEFBAAFD9FB6C89307" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[1104,1202,549,575]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="70" phylum="Arthropoda" rank="species" species="galoko">
<emphasis id="17DA0A43FFDFFFCEFBAAFD9FB6C89307" box="[1104,1202,549,575]" italics="true" pageId="13" pageNumber="70">I. galoko</emphasis>
</taxonomicName>
, by the elytral sculpture. Each elytron has only five costae: #1, #2, #3, #5, and #8; costa #4, #6, and #7, which are present in other species, are absent in
<taxonomicName id="E2AEADD2FFDFFFCEFE63FDD7B03693BF" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[409,588,622,647]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="70" phylum="Arthropoda" rank="species" species="quintacostata">
<emphasis id="17DA0A43FFDFFFCEFE63FDD7B03693BF" box="[409,588,622,647]" italics="true" pageId="13" pageNumber="70">I. quintacostata</emphasis>
</taxonomicName>
(
<figureCitation id="BD95CAD4FFDFFFCEFD9AFDD4B0D393BF" box="[608,681,621,647]" captionStart="FIGURES 2–4" captionStartId="3.[151,264,2010,2035]" captionTargetBox="[271,1317,378,1988]" captionTargetId="figure@3.[271,1317,378,1988]" captionTargetPageId="3" captionText="FIGURES 2–4. Dorsal morphology. (2) Incoltorrida quintacostata; (3–4) I. magna." httpUri="https://zenodo.org/record/3369809/files/figure.png" pageId="13" pageNumber="70">Fig. 2</figureCitation>
). The serial punctures are minute and very indistinct (microslide preparation necessary to see the punctures clearly). In this species the elytral costae are well developed in height, but comparatively narrow; the areas between the costae are quite flat. The transverse ridge that connects elytral costae #5 and #8 is more or less distinct (depending on the population). The anterior depressions of the pronotum are well developed and the foveae forming the medial margins of the ridges are narrowly separated and the intervening surface is transversely rounded, but there is no separate median carina. The pronotal sculpture and adult size of
<taxonomicName id="E2AEADD2FFDFFFCEFA69FC9BB3559267" authorityName="Perkins &amp; Bergsten" authorityYear="2019" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="70" phylum="Arthropoda" rank="species" species="quintacostata">
<emphasis id="17DA0A43FFDFFFCEFA69FC9BB3559267" italics="true" pageId="13" pageNumber="70">I quintacostata</emphasis>
</taxonomicName>
differ markedly from those of
<taxonomicName id="E2AEADD2FFDFFFCEFD68FCFFB08D9267" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[658,759,837,863]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="70" phylum="Arthropoda" rank="species" species="galoko">
<emphasis id="17DA0A43FFDFFFCEFD68FCFFB08D9267" box="[658,759,837,863]" italics="true" pageId="13" pageNumber="70">I. galoko</emphasis>
</taxonomicName>
, which also has five costae of each elytron. The ridges and impressions of the pronotum are similar to those of
<taxonomicName id="E2AEADD2FFDFFFCEFD1CFCD3B1FF92BB" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[742,901,873,899]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="70" phylum="Arthropoda" rank="species" species="benesculpta">
<emphasis id="17DA0A43FFDFFFCEFD1CFCD3B1FF92BB" box="[742,901,873,899]" italics="true" pageId="13" pageNumber="70">I. benesculpta</emphasis>
</taxonomicName>
, a much smaller species that has very different elytral sculpture; the aedeagi of the two species markedly differ (
<figureCitation id="BD95CAD4FFDFFFCEFC95FC34B1B6929F" box="[879,972,909,935]" captionStart="FIGURES 14–18" captionStartId="16.[151,264,1745,1770]" captionTargetBox="[177,1384,429,1687]" captionTargetId="figure@16.[177,1411,345,1726]" captionTargetPageId="16" captionText="FIGURES 14–18. Incoltorrida spp.; (14) I. madagassica, male genitalia of holotype (after Steffan 1973, permanent slide mount). (15–17) male genitalia apex showing pores of I. madagassica and spines of I. benesculpta and I. quintacostata; (18) I. quintacostata, aedeagus of holotype." httpUri="https://zenodo.org/record/3369827/files/figure.png" pageId="13" pageNumber="70">Figs. 18</figureCitation>
,
<figureCitation id="BD95CAD4FFDFFFCEFC22FC34B182929F" box="[984,1016,909,935]" captionStart="FIGURES 19–20" captionStartId="17.[151,264,1761,1786]" captionTargetBox="[235,1353,486,1744]" captionTargetId="figure@17.[235,1353,486,1744]" captionTargetPageId="17" captionText="FIGURES 19–20. (19) Incoltorrida madagassica, male genitalia of non-type; (20) Incoltorrida benesculpta, male genitalia of holotype." httpUri="https://zenodo.org/record/3369829/files/figure.png" pageId="13" pageNumber="70">20</figureCitation>
).
</paragraph>
<paragraph id="2511D651FFDFFFCEFF3DFC09B00F952B" blockId="13.[151,1437,153,1943]" pageId="13" pageNumber="70">
<emphasis id="17DA0A43FFDFFFCEFF3DFC09B32B92F3" bold="true" box="[199,337,944,971]" pageId="13" pageNumber="70">Description</emphasis>
. Size:
<typeStatus id="FA1568F3FFDFFFCEFE5EFC08B07C92F3" box="[420,518,945,971]" pageId="13" pageNumber="70" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.33/1.32; head width 0.47; pronotum 0.51/0.93, elytra 1.50/1.32. Dorsum dark brown to black, venter brown to dark brown, legs brown except femoral-tibial articulations and tarsi black.
</paragraph>
<paragraph id="2511D651FFDFFFCEFF3DFBA4B1909563" blockId="13.[151,1437,153,1943]" pageId="13" pageNumber="70">Sides of frontoclypeal shield very slightly arcuate, almost parallel-sided, and only very slightly and gradually narrowed from frons vertex to the transverse anterior margin of the clypeus.</paragraph>
<paragraph id="2511D651FFDFFFCEFF3DFBDCB6AE959B" blockId="13.[151,1437,153,1943]" pageId="13" pageNumber="70">Short oblique carinae on anterior 1/3 of pronotum are well developed; no indication of a midlongitudinal carina. Basal 1/3 of pronotum with distinctive carina on each side, each slightly oblique toward midline.</paragraph>
<paragraph id="2511D651FFDFFFCEFF3DFB14B7109437" blockId="13.[151,1437,153,1943]" pageId="13" pageNumber="70">Each elytron with only five costae: #1, #2, #3, #5, and #8; costae #4, #6, and #7 absent. Areas between costae quite flat, and serial punctures minute, very indistinct, a unilinear row located adjacent to the lateral margin of each costa. Transverse ridge linking costae #5 and #8 present, but more distinctive in some populations than others.</paragraph>
<paragraph id="2511D651FFDFFFCEFF3DFAA0B617946F" blockId="13.[151,1437,153,1943]" pageId="13" pageNumber="70">Lateral margins of the metaventral tabella straight for most of their length, arcuate only very near base. Midlongitudinal groove in posterior 1/2 of the metaventral tabella is narrow and parallel-sided.</paragraph>
<paragraph id="2511D651FFDFFFCEFF3DFAD8B3F394A7" blockId="13.[151,1437,153,1943]" pageId="13" pageNumber="70">Midlongitudinal carina of the first abdominal ventrite is moderately strong but usually does not extend the full length of the ventrite.</paragraph>
<paragraph id="2511D651FFDFFFCEFF3DFA10B11894FB" blockId="13.[151,1437,153,1943]" box="[199,866,1449,1475]" pageId="13" pageNumber="70">
Male genitalia long and slender (
<figureCitation id="BD95CAD4FFDFFFCEFDC2FA10B0BA94FB" box="[568,704,1449,1475]" captionStart="FIGURES 14–18" captionStartId="16.[151,264,1745,1770]" captionTargetBox="[177,1384,429,1687]" captionTargetId="figure@16.[177,1411,345,1726]" captionTargetPageId="16" captionText="FIGURES 14–18. Incoltorrida spp.; (14) I. madagassica, male genitalia of holotype (after Steffan 1973, permanent slide mount). (15–17) male genitalia apex showing pores of I. madagassica and spines of I. benesculpta and I. quintacostata; (18) I. quintacostata, aedeagus of holotype." httpUri="https://zenodo.org/record/3369827/files/figure.png" pageId="13" pageNumber="70">Figs. 17, 18</figureCitation>
); see remarks.
</paragraph>
<paragraph id="2511D651FFDFFFCEFF3DFA74B1D594DF" blockId="13.[151,1437,153,1943]" box="[199,943,1485,1511]" pageId="13" pageNumber="70">
<emphasis id="17DA0A43FFDFFFCEFF3DFA74B33294DF" bold="true" box="[199,328,1485,1511]" pageId="13" pageNumber="70">Etymology</emphasis>
. Named in reference to the five costae of each elytron.
</paragraph>
<paragraph id="2511D651FFDFFFCEFF3DFA48B7F29717" blockId="13.[151,1437,153,1943]" pageId="13" pageNumber="70">
<emphasis id="17DA0A43FFDFFFCEFF3DFA48B3419733" bold="true" box="[199,315,1521,1547]" pageId="13" pageNumber="70">Remarks.</emphasis>
Some specimens, such as the
<typeStatus id="FA1568F3FFDFFFCEFD77FA48B0889733" box="[653,754,1521,1547]" pageId="13" pageNumber="70" type="holotype">holotype</typeStatus>
, have the posterior angles of the pronotum very weakly shallowly notched; other specimens do not have this weak indentation, and some specimens have it only on one side.
</paragraph>
<paragraph id="2511D651FFDFFFCEFF3DF980B65B97DB" blockId="13.[151,1437,153,1943]" pageId="13" pageNumber="70">
The male genitalia in ventral view is long and comparatively slender, with a quite spinose apical tip (
<figureCitation id="BD95CAD4FFDFFFCEFA9EF980B298974F" captionStart="FIGURES 14–18" captionStartId="16.[151,264,1745,1770]" captionTargetBox="[177,1384,429,1687]" captionTargetId="figure@16.[177,1411,345,1726]" captionTargetPageId="16" captionText="FIGURES 14–18. Incoltorrida spp.; (14) I. madagassica, male genitalia of holotype (after Steffan 1973, permanent slide mount). (15–17) male genitalia apex showing pores of I. madagassica and spines of I. benesculpta and I. quintacostata; (18) I. quintacostata, aedeagus of holotype." httpUri="https://zenodo.org/record/3369827/files/figure.png" pageId="13" pageNumber="70">Figs. 17, 18</figureCitation>
). Males vary considerably in body length, ca.
<quantity id="E2567BB4FFDFFFCEFD03F9E4B1DB974F" box="[761,929,1629,1655]" metricMagnitude="-3" metricUnit="m" metricValue="2.24" metricValueMax="2.39" metricValueMin="2.09" pageId="13" pageNumber="70" unit="mm" value="2.24" valueMax="2.39" valueMin="2.09">2.09–2.39 mm</quantity>
. The length of the aedeagus varies, roughly, in proportion to the body size; for example: (aedeagus length/body length): 0.73/2.18 (=0.33); 0.75/2.15 (=0.35); 0.77/2.09 (=0.37);0.80/2.24 (=0.36); 0.93/2.39 (=0.39, the
<typeStatus id="FA1568F3FFDFFFCEFCD0F91CB1EB97F8" box="[810,913,1701,1728]" pageId="13" pageNumber="70" type="holotype">holotype</typeStatus>
). The shape of the aedeagus does not vary significantly. This is the most commonly collected species of
<taxonomicName id="E2AEADD2FFDFFFCEFCD2F970B1CB97DB" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[808,945,1737,1763]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="17DA0A43FFDFFFCEFCD2F970B1CB97DB" box="[808,945,1737,1763]" italics="true" pageId="13" pageNumber="70">Incoltorrida</emphasis>
</taxonomicName>
(
<figureCitation id="BD95CAD4FFDFFFCEFC3AF970B66F97DB" box="[960,1045,1737,1763]" captionStart="FIGURES 34–37" captionStartId="26.[151,264,1901,1926]" captionTargetId="figure@26.[293,1294,201,1882]" captionTargetPageId="26" captionText="FIGURES 34–37. Incoltorrida distribution maps." httpUri="https://zenodo.org/record/3387378/files/figure.png" pageId="13" pageNumber="70">Fig. 34</figureCitation>
).
</paragraph>
<paragraph id="2511D651FFDFFFCEFF3DF954B2C09613" blockId="13.[151,1437,153,1943]" pageId="13" pageNumber="70">
Four larvae (
<collectionCode id="43BF4E94FFDFFFCEFEADF954B3DF963F" box="[343,421,1773,1799]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="13" pageNumber="70">NHRS</collectionCode>
) were collected at the
<typeStatus id="FA1568F3FFDFFFCEFD5EF954B0AE963F" box="[676,724,1773,1799]" pageId="13" pageNumber="70">type</typeStatus>
locality, and
<specimenCount id="33A81DD8FFDFFFCEFC97F955B1AE963F" box="[877,980,1772,1799]" pageId="13" pageNumber="70" type="larva">54 larvae</specimenCount>
(
<collectionCode id="43BF4E94FFDFFFCEFC1EF954B648963F" box="[996,1074,1773,1799]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="13" pageNumber="70">NHRS</collectionCode>
) were collected at site
<collectionCode id="43BF4E94FFDFFFCEFACFF954B700963F" box="[1333,1402,1773,1799]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="13" pageNumber="70">MAD</collectionCode>
12- 26.
</paragraph>
<paragraph id="2511D651FFDFFFCEFF3DF88CB16B96AF" blockId="13.[151,1437,153,1943]" pageId="13" pageNumber="70">
DNA sequences of partial CO1 (Acc. Nos.
<accessionNumber id="3AFD4BB2FFDFFFCEFD44F88CB1579677" box="[702,813,1845,1871]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819700" pageId="13" pageNumber="70">FJ819700</accessionNumber>
(
<collectionCode id="43BF4E94FFDFFFCEFCC4F88CB1EE9668" box="[830,916,1845,1872]" pageId="13" pageNumber="70">BMNH</collectionCode>
670734) and
<accessionNumber id="3AFD4BB2FFDFFFCEFBD4F88CB6E69677" box="[1070,1180,1845,1871]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819701" pageId="13" pageNumber="70">FJ819701</accessionNumber>
(
<collectionCode id="43BF4E94FFDFFFCEFB57F88DB7799677" box="[1197,1283,1844,1871]" pageId="13" pageNumber="70">BMNH</collectionCode>
670735)) and partial 28S (
<accessionNumber id="3AFD4BB2FFDFFFCEFED9F8E1B3E8964B" box="[291,402,1880,1907]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ818156" pageId="13" pageNumber="70">
<collectionCode id="43BF4E94FFDFFFCEFED9F8E1B344964B" box="[291,318,1880,1907]" pageId="13" pageNumber="70">FJ</collectionCode>
818156
</accessionNumber>
(
<collectionCode id="43BF4E94FFDFFFCEFE59F8E0B383964C" box="[419,505,1881,1908]" pageId="13" pageNumber="70">BMNH</collectionCode>
670734) and
<accessionNumber id="3AFD4BB2FFDFFFCEFD75F8E1B084964B" box="[655,766,1880,1907]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ818157" pageId="13" pageNumber="70">FJ818157</accessionNumber>
(
<collectionCode id="43BF4E94FFDFFFCEFCF4F8E1B11E964B" box="[782,868,1880,1907]" pageId="13" pageNumber="70">BMNH</collectionCode>
670735)) published by
<bibRefCitation id="413FABA0FFDFFFCEFB96F8E0B708964B" author="Monaghan, M. T. &amp; Wild, R. &amp; Elliot, M. &amp; Fujisawa, T. &amp; Balke, M. &amp; Inward, D. J. &amp; Lees, D. C. &amp; Ranaivosolo, R. &amp; Eggleton, P. &amp; Barraclough, T. G. &amp; Vogler, A. P." box="[1132,1394,1881,1907]" pageId="13" pageNumber="70" pagination="298 - 311" refId="ref13383" refString="Monaghan, M. T., Wild, R., Elliot, M., Fujisawa, T., Balke, M., Inward, D. J., Lees, D. C., Ranaivosolo, R., Eggleton, P., Barraclough, T. G., Vogler, A. P., (2009) Accelerated species inventory on Madagascar using coalescent-based models of species delineation. Systematic Biology, 58, 298 - 311." type="journal article" year="2009">
Monaghan
<emphasis id="17DA0A43FFDFFFCEFB16F8E3B765964B" box="[1260,1311,1881,1907]" italics="true" pageId="13" pageNumber="70">et al</emphasis>
(2009)
</bibRefCitation>
are based on adult specimens, and are available in Genbank.
</paragraph>
</subSubSection>
</treatment>
</document>
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