213 lines
22 KiB
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213 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo" docDate="2022" docId="A36AB90A04FB54DCA7842D21AA4A8089" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Metasarcus bolivianus Roewer 1913" docType="treatment" docVersion="1" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1660711556436" updateUser="pensoft">
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<mods:title>Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Benedetti, Alipio Rezende</mods:namePart>
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<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil & Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pinto-da-Rocha, Ricardo</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
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<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>80</mods:number>
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<mods:extent unit="page">
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<mods:start>309</mods:start>
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<mods:end>388</mods:end>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e73829</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e73829</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-80-309</mods:identifier>
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<mods:identifier type="Pensoft-UUID">11907D1B6F665D96B5FD2F9751431366</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:A36AB90A04FB54DCA7842D21AA4A8089" httpUri="http://treatment.plazi.org/id/A36AB90A04FB54DCA7842D21AA4A8089" lastPageNumber="309" pageId="0" pageNumber="309">
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<subSubSection pageId="0" pageNumber="309" type="nomenclature">
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<paragraph pageId="0" pageNumber="309">
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3.42.
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<taxonomicName LSID="A36AB90A-04FB-54DC-A784-2D21AA4A8089" authority="Roewer, 1913" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus bolivianus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bolivianus">Metasarcus bolivianus Roewer, 1913</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="description">
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<paragraph pageId="0" pageNumber="309">
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">Figs 5C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="reference_group">
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<paragraph pageId="0" pageNumber="309">
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<taxonomicName authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus bolivianus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bolivianus">Metasarcus bolivianus</taxonomicName>
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Roewer, 1913: 305 (desc.), fig.123 (female dorsal habitus, pedipalpus);
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<bibRefCitation author="Roewer, CF" journalOrPublisher="Gustav Fischer, Jena, 1" pageId="0" pageNumber="309" refId="B98" refString="Roewer, CF, 1923. Die Weberknechte der Erde. Systematische Bear-beitung der bisher bekannten Opiliones. Gustav Fischer, Jena, 1" title="Die Weberknechte der Erde. Systematische Bear-beitung der bisher bekannten Opiliones." year="1923">Roewer 1923</bibRefCitation>
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: 517 (rdesc.), fig. 647 (female dorsal habitus and pedipalpus);
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<bibRefCitation author="Soares, BAM" journalOrPublisher="Arquivos de Zoologia do Estado de Sao Paulo" pageId="0" pageNumber="309" pagination="149 - 240" refId="B115" refString="Soares, BAM, Soares, HEM, 1949. Monografia dos generos de opilioes neotropicos II. Arquivos de Zoologia do Estado de Sao Paulo 7 (2): 149 - 240" title="Monografia dos generos de opilioes neotropicos II." volume="7" year="1949">Soares and Soares 1949</bibRefCitation>
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: 234 (cat.);
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<bibRefCitation DOI="https://doi.org/10.1080/03946975.1994.10539264" author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" pagination="343 - 353" refId="B50" refString="Kury, AB, 1994. Early lineages fo Gonyleptidae (Arachinida, Opiliones, Laniatores). Tropical Zoology 7 (2): 343 - 353, DOI: https://doi.org/10.1080/03946975.1994.10539264" title="Early lineages fo Gonyleptidae (Arachinida, Opiliones, Laniatores)." url="https://doi.org/10.1080/03946975.1994.10539264" volume="7" year="1994">Kury 1994</bibRefCitation>
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: 349 (cit.);
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<bibRefCitation author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
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: 145 (cat.).
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</paragraph>
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<paragraph pageId="0" pageNumber="309">
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<taxonomicName class="Arachnida" family="Metasarcidae" genus="Chaconatus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chaconatus armatipalpus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="armatipalpus">Chaconatus armatipalpus</taxonomicName>
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Roewer, 1929: 275 (key), 276 (desc.), fig. 43 (female lateral habitus, pedipalpus), fig. 43a (ocularium);
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<bibRefCitation author="Soares, HEM" journalOrPublisher="Revista Brasileira de Entomologia" pageId="0" pageNumber="309" pagination="1 - 14" refId="B116" refString="Soares, HEM, Soares, BAM, Jim, RLS, 1992. Monografia dos generos de opilioes neotropicos IV. (Opiliones, Gonyleptidae, Prostygninae). Revista Brasileira de Entomologia 36 (1): 1 - 14" title="Monografia dos generos de opilioes neotropicos IV. (Opiliones, Gonyleptidae, Prostygninae)." volume="36" year="1992">Soares et al. 1992</bibRefCitation>
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: 5 (cat.);
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<bibRefCitation author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
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: 145 (cat.),
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<emphasis bold="true" pageId="0" pageNumber="309">syn. n.</emphasis>
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</paragraph>
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<paragraph pageId="0" pageNumber="309">
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<taxonomicName baseAuthorityName="Roewer" baseAuthorityYear="1929" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus armatipalpus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="armatipalpus">Metasarcus armatipalpus</taxonomicName>
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:
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<bibRefCitation DOI="https://doi.org/10.1080/03946975.1994.10539264" author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" pagination="343 - 353" refId="B50" refString="Kury, AB, 1994. Early lineages fo Gonyleptidae (Arachinida, Opiliones, Laniatores). Tropical Zoology 7 (2): 343 - 353, DOI: https://doi.org/10.1080/03946975.1994.10539264" title="Early lineages fo Gonyleptidae (Arachinida, Opiliones, Laniatores)." url="https://doi.org/10.1080/03946975.1994.10539264" volume="7" year="1994">Kury 1994</bibRefCitation>
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: 349 (sist.);
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<bibRefCitation author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
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: 145 (cat.).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="redescription">
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<paragraph pageId="0" pageNumber="309">Redescription.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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<emphasis bold="true" pageId="0" pageNumber="309">
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FEMALE:
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<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
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</emphasis>
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(
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<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
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=2) DSW: 5.4-5.6 (5.4); DSL: 5.2 -5.5 (5.5); CL: 2.2 (2.2). FIVL: 11.2-12.7 (12.7). ChL: 2.0-2.3 (2.0).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
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(according to Roewer, 1913): Body, chelicerae and pedipalpus yellow-brown, in dark tone. Black legs. Granules of areas I-II yellow.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
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. (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5C</figureCitation>
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) Gamma-type DSS, slightly concave with posterior margin of DS. Anterior margin of carapace with median elevation, covered with a few sparse granules. Ocularium medially depressed, unarmed. Carapace with few sparse granules. Areas I-IV with granules (in lower density on areas III-IV). Area I divided by a longitudinal groove. Areas I, II and III unarmed. Area III with a median pair of large spines, directed posteriorly located at granular elevations of integument. Lateral margins of dorsal scutum granular. Posterior margin of DS and free tergites I-II with a row of granules. Free tergite III with a median spiniform apophysis, longer than the length of this tergite, directed posteriorly, with sparse granules on its posterior margin.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
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: (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5C</figureCitation>
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) Segment I covered with small granules in the bulla. Segment II, finger with three teeth. Segment III with four teeth.
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<taxonomicName genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
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</taxonomicName>
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: With granules irregularly distributed over the entire length, except tibia and tarsus. Trochanter with a ventroapical spine. Femur with a small ventroapical setiferous tubercle; a ventral row of four spines throughout its length; a proventral spine. Patella with a proventral spine. Tibia: prolateral iIIi, retrolateral iIii. Tarsus: prolateral IiI, retrolateral II.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
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: Coxae I-IV with sparse granules. Coxa I with a median row of 3-4 small setiferous tubercles. Genital area with few granules sparsely distributed. Free sternites with a row of small tubercles. Anal operculum smooth.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
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. (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5C</figureCitation>
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) Coxae I-II each one with a prolateral and a retrolateral apophysis. Coxa III unarmed. Coxa IV with few granules. Trochanters I-III smooth. Trochanter IV with one-two retroapical tubercles with wide base, rhombus. Femora I-III straight and unarmed, with small granules. Femur IV slightly curved and unarmed, with small granules. Tarsal segmentation: (
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<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
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=2) 7, 14, 10, 11-12 (11). -
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<emphasis bold="true" pageId="0" pageNumber="309">MALE</emphasis>
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: unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="diagnosis">
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<paragraph pageId="0" pageNumber="309">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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Very similar to females of
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<taxonomicName authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus fellinii" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="fellinii">
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<emphasis italics="true" pageId="0" pageNumber="309">Metasarcus fellinii</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
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because a long spiniform apophysis in tergite III (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5C</figureCitation>
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). Differs from
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<taxonomicName class="Arachnida" genus="M." kingdom="Animalia" lsidName="M. fellinii" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="fellinii">
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<emphasis italics="true" pageId="0" pageNumber="309">M. fellinii</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
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because longer DS (DSL greater than 5); a sparser granulation in dorsal scutum (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5C</figureCitation>
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); longer legs (FIVL greater than 11) and coxa III unarmed (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5C</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="remarks">
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<paragraph pageId="0" pageNumber="309">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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It is intriguing how Roewer described the same species from the Bolivian Chaco not only in different species and genera, but in different subfamilies (
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<taxonomicName class="Arachnida" genus="M." kingdom="Animalia" lsidName="M. bolivianus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bolivianus">
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<emphasis italics="true" pageId="0" pageNumber="309">M. bolivianus</emphasis>
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</taxonomicName>
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was placed in
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<taxonomicName authorityName="Simon" authorityYear="1879" family="Metasarcidae" kingdom="Animalia" lsidName="" order="Opiliones" pageId="0" pageNumber="309" rank="subFamily" subFamily="Mitobatinae">Mitobatinae</taxonomicName>
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and
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<taxonomicName class="Arachnida" family="Metasarcidae" genus="Chaconatus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chaconatus armatipalpus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="armatipalpus">
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<emphasis italics="true" pageId="0" pageNumber="309">Chaconatus armatipalpus</emphasis>
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</taxonomicName>
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in
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<taxonomicName authorityName="Roewer" authorityYear="1913" family="Metasarcidae" kingdom="Animalia" lsidName="" order="Opiliones" pageId="0" pageNumber="309" rank="subFamily" subFamily="Prostygninae">Prostygninae</taxonomicName>
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). As
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<bibRefCitation DOI="https://doi.org/10.1080/03946975.1994.10539264" author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" pagination="343 - 353" refId="B50" refString="Kury, AB, 1994. Early lineages fo Gonyleptidae (Arachinida, Opiliones, Laniatores). Tropical Zoology 7 (2): 343 - 353, DOI: https://doi.org/10.1080/03946975.1994.10539264" title="Early lineages fo Gonyleptidae (Arachinida, Opiliones, Laniatores)." url="https://doi.org/10.1080/03946975.1994.10539264" volume="7" year="1994">Kury (1994</bibRefCitation>
|
||
: p. 349) noted "There is no positive evidence for either of the assignments, furthermore both species are very closely related, if not the same." Possibly, this fact can be interpreted because of the Roewerian taxonomic system (see Introduction). Upon closer examination of the type material of both species and the observation of a great morphological similarity, it was possible to conclude that
|
||
<normalizedToken originalValue="Kury’s">Kury's</normalizedToken>
|
||
suspicion was correct, and hence the synonymy made here.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="309" type="distribution">
|
||
<paragraph pageId="0" pageNumber="309">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="309">BOLIVIA. Chaco.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="309" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="309">Material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="309">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Type material</emphasis>
|
||
:
|
||
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="BOLIVIA" location="Chaco" specimenCount="♀" typeStatus="Holotype">
|
||
Of
|
||
<taxonomicName class="Arachnida" genus="M." kingdom="Animalia" lsidName="M. bolivianus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bolivianus">
|
||
<emphasis italics="true" pageId="0" pageNumber="309">M. bolivianus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis bold="true" pageId="0" pageNumber="309">
|
||
<typeStatus>Holotype</typeStatus>
|
||
</emphasis>
|
||
<specimenCount>♀</specimenCount>
|
||
, '
|
||
<collectingCountry name="Bolivia">BOLIVIA</collectingCountry>
|
||
, Gran Chaco, whitout date and leg. (SMF 873). Of
|
||
<taxonomicName class="Arachnida" genus="M." kingdom="Animalia" lsidName="M. armatipalpus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="armatipalpus">
|
||
<emphasis italics="true" pageId="0" pageNumber="309">M. armatipalpus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis bold="true" pageId="0" pageNumber="309">
|
||
<typeStatus>Holotype</typeStatus>
|
||
</emphasis>
|
||
<specimenCount>♀</specimenCount>
|
||
, '
|
||
<collectingCountry name="Bolivia">BOLIVIA</collectingCountry>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:A36AB90A04FB54DCA7842D21AA4A8089:7141322E4F6CD0C14D7EB7C4D7ECCE6F" country="BOLIVIA" name="Chaco">Chaco</location>
|
||
| whitout date and leg. (SMF 1003/2)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |