treatments-xml/data/94/E2/6D/94E26DE30DE754CE8A7BCBBCE029BE57.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.923.48420" ID-GBIF-Dataset="40ddbb19-1279-49e9-adb0-61ff8f27c186" ID-PMC="PMC7142171" ID-Pensoft-Pub="1313-2970-923-91" ID-Pensoft-UUID="D7C45185A29A573DBFA61CED9A510411" ID-PubMed="32292273" ID-ZooBank="F138D1ED8A51462888299617AC5D3029" ModsDocID="1313-2970-923-91" checkinTime="1585826875964" checkinUser="pensoft" docAuthor="Braganca, Pedro H. N., Zeeventer, Ryan M. van, Bills, Roger, Tweddle, Denis &amp; Chakona, Albert" docDate="2020" docId="94E26DE30DE754CE8A7BCBBCE029BE57" docLanguage="en" docName="ZooKeys 923: 91-113" docOrigin="ZooKeys 923" docSource="http://dx.doi.org/10.3897/zookeys.923.48420" docTitle="Lacustricola johnstoni" docType="treatment" docVersion="4" id="D7C45185A29A573DBFA61CED9A510411" lastPageNumber="91" masterDocId="D7C45185A29A573DBFA61CED9A510411" masterDocTitle="Diversity of the southern Africa Lacustricola Myers, 1924 and redescription of Lacustricola johnstoni (Guenther, 1894) and Lacustricola myaposae (Boulenger, 1908) (Cyprinodontiformes, Procatopodidae)" masterLastPageNumber="113" masterPageNumber="91" pageNumber="91" updateTime="1668168681547" updateUser="ExternalLinkService">
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<mods:title>Diversity of the southern Africa Lacustricola Myers, 1924 and redescription of Lacustricola johnstoni (Guenther, 1894) and Lacustricola myaposae (Boulenger, 1908) (Cyprinodontiformes, Procatopodidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Braganca, Pedro H. N.</mods:namePart>
<mods:affiliation>South African Institute for Aquatic Biodiversity, Private Bag 1015, Grahamstown, 6140, South Africa</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zeeventer, Ryan M. van</mods:namePart>
<mods:affiliation>South African Institute for Aquatic Biodiversity, Private Bag 1015, Grahamstown, 6140, South Africa</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bills, Roger</mods:namePart>
<mods:affiliation>South African Institute for Aquatic Biodiversity, Private Bag 1015, Grahamstown, 6140, South Africa</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tweddle, Denis</mods:namePart>
<mods:affiliation>South African Institute for Aquatic Biodiversity, Private Bag 1015, Grahamstown, 6140, South Africa</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chakona, Albert</mods:namePart>
<mods:affiliation>South African Institute for Aquatic Biodiversity, Private Bag 1015, Grahamstown, 6140, South Africa &amp; Department of Ichthyology and Fisheries Science, Rhodes University, Grahamstown, South Africa</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>923</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>91</mods:start>
<mods:end>113</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.923.48420</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.923.48420</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-923-91</mods:identifier>
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<mods:identifier type="Pensoft-UUID">D7C45185A29A573DBFA61CED9A510411</mods:identifier>
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<treatment ID-GBIF-Taxon="163421225" LSID="urn:lsid:plazi:treatment:94E26DE30DE754CE8A7BCBBCE029BE57" httpUri="http://treatment.plazi.org/id/94E26DE30DE754CE8A7BCBBCE029BE57" lastPageNumber="91" pageId="0" pageNumber="91">
<subSubSection pageId="0" pageNumber="91" type="nomenclature">
<paragraph pageId="0" pageNumber="91">
<taxonomicName LSID="94E26DE3-0DE7-54CE-8A7B-CBBCE029BE57" authority="(Guenther, 1894)" baseAuthorityName="Guenther" baseAuthorityYear="1894" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lacustricola johnstoni" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="species" species="johnstoni">
Lacustricola johnstoni (
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
, 1894)
</taxonomicName>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. &quot; Lacustricola &quot; johnstoni preserved colouration: A male SAIAB 118776, 34.7 mm SL B female SAIAB 118776, 32.0 mm SL; from Shire River at Liwonde Barrage, Malawi." figureDoi="10.3897/zookeys.923.48420.figure3" httpUri="https://binary.pensoft.net/fig/396600" pageId="0" pageNumber="91">Figures 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. &quot; Lacustricola &quot; johnstoni colouration in life: A male SAIAB 118776, 31.0 mm SL B female SAIAB 118776, 26 mm SL; from Shire River at Liwonde Barrage, Malawi." figureDoi="10.3897/zookeys.923.48420.figure4" httpUri="https://binary.pensoft.net/fig/396601" pageId="0" pageNumber="91">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="reference_group">
<paragraph pageId="0" pageNumber="91">
<taxonomicName class="Actinopterygii" family="Poeciliidae" genus="Haplochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplochilus johnstoni" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="species" species="johnstoni">Haplochilus johnstoni</taxonomicName>
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
, 1894: 627 [original description: Mangochi (former Fort Johnston), Malawi].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="materials_examined">
<paragraph pageId="0" pageNumber="91">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="91">
BMNH 1893.11.15.95, Lectotype; BMNH 1893.11.15.92-94,96-99, 7 Paralectotypes; Mangochi (former Fort Johnston), Malawi. Examined by photographs and x-rays - SAIAB 35820, 18 (5 C&amp;S), 24.7-35.4 mm SL; Upper Shire River, Mangochi, Malawi, 
<geoCoordinate degrees="14" direction="south" minutes="26" orientation="latitude" precision="15" seconds="60" value="-14.45">14°26'60&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="35" direction="east" minutes="15" orientation="longitude" precision="15" seconds="60" value="35.266666">35°15'60&quot;E</geoCoordinate>
; col: D. Tweddle &amp; N. G. Willoughby; 19 Sep. 1971. - SAIAB 8311, 3, 31.3-32.5 mm SL; Shire River, Liwonde, Malawi; col: D. Tweddle &amp; N. G. Willoughby; 20 Oct. 1975. - SAIAB 34384, 1, 29.1 mm SL; Shire River, Liwonde Barrage, Malawi, 
<geoCoordinate degrees="15" direction="south" minutes="04" orientation="latitude" precision="925" value="-15.066667">15°04'S</geoCoordinate>
, 
<geoCoordinate degrees="35" direction="east" minutes="13" orientation="longitude" precision="925" value="35.216667">35°13'E</geoCoordinate>
; col: D. Tweddle &amp; P. Skelton; 26 Oct. 1989. - SAIAB 34388, 1, 35.7 mm SL; Monkey Bay, Lake Malawi, Malawi, 
<geoCoordinate degrees="14" direction="south" minutes="04" orientation="latitude" precision="925" value="-14.066667">14°04'S</geoCoordinate>
, 
<geoCoordinate degrees="34" direction="east" minutes="55" orientation="longitude" precision="925" value="34.916668">34°55'E</geoCoordinate>
; 17 Oct. 1989. - SAIAB 40800, 15 (4 C&amp;S), 30.1-33.2 mm SL; Bridge over Dwambadzi River, Malawi, 
<geoCoordinate degrees="12" direction="south" minutes="14" orientation="latitude" precision="925" value="-12.233334">12°14'S</geoCoordinate>
, 
<geoCoordinate degrees="33" direction="east" minutes="59" orientation="longitude" precision="925" value="33.983334">33°59'E</geoCoordinate>
; col: D. Tweddle &amp; P. Skelton; 06 Sep. 1992. - SAIAB 11237, 7, 26.2-36.2 mm SL; Monkey Bay, Lake Malawi, Malawi, 
<geoCoordinate degrees="14" direction="south" minutes="3" orientation="latitude" precision="15" seconds="00" value="-14.05">14°3'00&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="34" direction="east" minutes="55" orientation="longitude" precision="15" seconds="00" value="34.916668">34°55'00&quot;E</geoCoordinate>
; col: D.H. Eccles; 31 Oct. 1974. - SAIAB 11876, 8, 27.2-34.7 mm SL; Shire River, Liwonde Barrage, Malawi, 
<geoCoordinate degrees="15" direction="south" minutes="3" orientation="latitude" precision="15" seconds="37" value="-15.060278">15°3'37&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="35" direction="east" minutes="13" orientation="longitude" precision="15" seconds="7" value="35.218613">35°13'7&quot;E</geoCoordinate>
; col: D. Tweddle &amp; T. Makinen; 27 May. 2011.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="diagnosis">
<paragraph pageId="0" pageNumber="91">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="91">
&quot; 
<taxonomicName authorityName="Myers" authorityYear="1924" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lacustricola" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="91">Lacustricola</emphasis>
</taxonomicName>
&quot; 
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1894" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="johnstoni" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="species" species="johnstoni">
<emphasis italics="true" pageId="0" pageNumber="91">johnstoni</emphasis>
</taxonomicName>
is distinguished from all congeners from the &quot; 
<taxonomicName genus="L." lsidName="L." pageId="0" pageNumber="91" rank="genus">L.</taxonomicName>
&quot; 
<taxonomicName lsidName="katangae" pageId="0" pageNumber="91" rank="species" species="katangae">katangae</taxonomicName>
clade by the absence of a zigzag black mark along the flank (vs. presence); and from congeners belonging to the &quot; 
<taxonomicName genus="L." lsidName="L." pageId="0" pageNumber="91" rank="genus">L.</taxonomicName>
&quot; 
<taxonomicName lsidName="hutereaui" pageId="0" pageNumber="91" rank="species" species="hutereaui">hutereaui</taxonomicName>
clade by the absence of a barred dorsal, anal and caudal-fins and also by the absence of a conspicuous reticulate pattern on scale margins. It is further distinguished from all congeners except &quot; 
<taxonomicName genus="L." lsidName="L." pageId="0" pageNumber="91" rank="genus">L.</taxonomicName>
&quot; 
<taxonomicName lsidName="myaposae" pageId="0" pageNumber="91" rank="species" species="myaposae">
<emphasis italics="true" pageId="0" pageNumber="91">myaposae</emphasis>
</taxonomicName>
and &quot; 
<taxonomicName genus="L." lsidName="L." pageId="0" pageNumber="91" rank="genus">L.</taxonomicName>
&quot; 
<emphasis italics="true" pageId="0" pageNumber="91">moeruensis</emphasis>
by the presence of orange dorsal, anal and caudal-fins in females (vs. hyaline); it is distinguished from &quot; 
<taxonomicName genus="L." lsidName="L." pageId="0" pageNumber="91" rank="genus">L.</taxonomicName>
&quot; 
<taxonomicName lsidName="myaposae" pageId="0" pageNumber="91" rank="species" species="myaposae">
<emphasis italics="true" pageId="0" pageNumber="91">myaposae</emphasis>
</taxonomicName>
by the presence of a bluish colouration in the posterior region of flank (vs. light purple colouration); a slender body profile, male body depth 20.6-24.4% of SL (vs. 26.0-30.9% of SL), female body depth 19.7-22.5% of SL (vs. 22.8-25.1% of SL); a shorter dorsal-fin base length in males 8.7-11.6% of SL (vs. 11.9-13.1% of SL) and in females 7.3-10.1% of SL (vs. 10.8-11.6% SL); a less deep head in males 61.9-67.0% of HL (vs. 70.7-79.8% of HL) and in females 59.4-63.5% of HL (vs. 66.6-69.6% of HL); and a hyaline pectoral-fin in males (vs. orange). Other morphometric characters presenting a slight overlap but useful in distinguishing &quot; 
<taxonomicName genus="L." lsidName="L." pageId="0" pageNumber="91" rank="genus">L.</taxonomicName>
&quot; 
<taxonomicName lsidName="johnstoni" pageId="0" pageNumber="91" rank="species" species="johnstoni">
<emphasis italics="true" pageId="0" pageNumber="91">johnstoni</emphasis>
</taxonomicName>
from &quot; 
<taxonomicName genus="L." lsidName="L." pageId="0" pageNumber="91" rank="genus">L.</taxonomicName>
&quot; 
<taxonomicName lsidName="myaposae" pageId="0" pageNumber="91" rank="species" species="myaposae">
<emphasis italics="true" pageId="0" pageNumber="91">myaposae</emphasis>
</taxonomicName>
are: a comparatively narrow caudal peduncle, depth of 12.6-14.5% of SL in males and 11.2-12.7% of SL in females (vs. 14.0-17.1% of SL in males and 12.9-13.7% of SL in females); a comparatively elongated caudal-fin, 30.1-33.8% of SL in males and 28.9-31.2% of SL in females (vs. 27.5-30.2% of SL in males and 25.3-28.5% of SL in females); and a comparatively deep, laterally compressed head, 57.1-63.2% of HL in males and 59.0-64.7% of HL in females (vs. 63.1-67.9% of HL in males and 65.3-68.1% of HL in females). &quot; 
<taxonomicName authorityName="Myers" authorityYear="1924" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lacustricola" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="91">Lacustricola</emphasis>
</taxonomicName>
&quot; 
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1894" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="johnstoni" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="species" species="johnstoni">
<emphasis italics="true" pageId="0" pageNumber="91">johnstoni</emphasis>
</taxonomicName>
is distinguished from &quot; 
<taxonomicName genus="L." lsidName="L." pageId="0" pageNumber="91" rank="genus">L.</taxonomicName>
&quot; 
<emphasis italics="true" pageId="0" pageNumber="91">moeruensis</emphasis>
by a comparatively slender body and a more backward positioned dorsal-fin, first proximal radial of dorsal-fin between neural spine of vertebrae 16 and 17 (vs. 13 and 14).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="description">
<paragraph pageId="0" pageNumber="91">Description.</paragraph>
<paragraph pageId="0" pageNumber="91">
Morphometric data are presented in Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data of &quot; Lacustricola &quot; johnstoni and &quot; L. &quot; myaposae." httpUri="http://table.plazi.org/id/E1C45C9DC3A89D5F3C3F7C9ED6876531" pageId="0" pageNumber="91" tableUuid="E1C45C9DC3A89D5F3C3F7C9ED6876531">1</tableCitation>
. Maximum recorded adult size 35.6 mm SL. Dorsal profile of body approximately straight to slightly convex from snout tip to dorsal-fin origin; convex along dorsal-fin base. Ventral profile convex from lower jaw to beginning of anal-fin base; slightly convex along the anal-fin base and nearly straight on caudal peduncle. Anterior portion of body laterally compressed, becoming more compressed behind anal-fin origin.
</paragraph>
<caption ID-Table-UUID="E1C45C9DC3A89D5F3C3F7C9ED6876531" httpUri="http://table.plazi.org/id/E1C45C9DC3A89D5F3C3F7C9ED6876531" pageId="0" pageNumber="91" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="91">
<emphasis bold="true" pageId="0" pageNumber="91">Table 1.</emphasis>
Morphometric data of &quot; 
<taxonomicName authorityName="Myers" authorityYear="1924" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lacustricola" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="91">Lacustricola</emphasis>
</taxonomicName>
&quot; 
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1894" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="johnstoni" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="species" species="johnstoni">
<emphasis italics="true" pageId="0" pageNumber="91">johnstoni</emphasis>
</taxonomicName>
and &quot; 
<taxonomicName genus="L." lsidName="L." pageId="0" pageNumber="91" rank="genus">L.</taxonomicName>
&quot; 
<taxonomicName lsidName="myaposae" pageId="0" pageNumber="91" rank="species" species="myaposae">
<emphasis italics="true" pageId="0" pageNumber="91">myaposae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="91">
<table pageId="0" pageNumber="91">
<tr pageId="0" pageNumber="91">
<th colspan="1" isEmpty="true" pageId="0" pageNumber="91" rowspan="2">-</th>
<th colspan="2" pageId="0" pageNumber="91" rowspan="1">
&quot; 
<taxonomicName authorityName="Myers" authorityYear="1924" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lacustricola" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="91">Lacustricola</emphasis>
</taxonomicName>
&quot; 
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1894" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="johnstoni" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="species" species="johnstoni">
<emphasis italics="true" pageId="0" pageNumber="91">johnstoni</emphasis>
</taxonomicName>
</th>
<th colspan="2" pageId="0" pageNumber="91" rowspan="1">
&quot; 
<taxonomicName authorityName="Myers" authorityYear="1924" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lacustricola" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="91">Lacustricola</emphasis>
</taxonomicName>
&quot; 
<taxonomicName baseAuthorityName="Boulenger" baseAuthorityYear="1908" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="myaposae" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="species" species="myaposae">
<emphasis italics="true" pageId="0" pageNumber="91">myaposae</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="91">
<th colspan="1" pageId="0" pageNumber="91" rowspan="1">
males (
<emphasis italics="true" pageId="0" pageNumber="91">N</emphasis>
= 12)
</th>
<th colspan="1" pageId="0" pageNumber="91" rowspan="1">
females (
<emphasis italics="true" pageId="0" pageNumber="91">N</emphasis>
= 10)
</th>
<th colspan="1" pageId="0" pageNumber="91" rowspan="1">
males (
<emphasis italics="true" pageId="0" pageNumber="91">N</emphasis>
= 7)
</th>
<th colspan="1" pageId="0" pageNumber="91" rowspan="1">
females (
<emphasis italics="true" pageId="0" pageNumber="91">N</emphasis>
= 4)
</th>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Standard length (mm)</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">29.0-34.6</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">27.7-35.6</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">31.3-38.1</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">30.2-39.0</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="5" pageId="0" pageNumber="91" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="91">Percent of standard length</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Body depth</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">20.6-24.4</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">19.7-22.5</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">26.0-30.9</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">22.8-25.1</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Caudal peduncle depth</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">12.6-14.5</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">11.2-12.7</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">14.0-17.1</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">12.9-13.7</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Pre-dorsal length</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">67.6-73.9</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">71.6-73.9</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">67.5-72.3</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">68.5-70.7</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Pre-pelvic length</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">43.0-48.6</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">44.9-49.7</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">45.1-49.5</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">46.4-47.8</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Length of dorsal-fin base</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">8.7-11.6</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">7.3-10.1</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">11.9-13.1</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">10.8-11.6</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Length of anal-fin base</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">16.8-21.1</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">13.2-16.6</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">16.0-20.0</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">15.0-16.2</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Caudal-fin length</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">30.1-33.8</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">28.9-31.2</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">27.5-30.2</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">25.3-28.5</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Pectoral-fin length</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">18.6-22.1</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">18.8-20.8</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">18.7-22.4</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">17.5-19.5</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Pelvic-fin length</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">15.5-22.8</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">13.2-14.6</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">14.0-18.0</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">11.9-12.9</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Head length (mm)</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">6.9-8.4</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">6.2-7.9</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">8.2-9.5</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">7.2-9.8</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="5" pageId="0" pageNumber="91" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="91">Percent of head length</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Head depth</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">61.9-67.9</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">59.4-63.5</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">70.7-79.8</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">66.6-69.6</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Head width</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">57.1-63.2</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">59.0-64.7</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">63.1-67.9</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">65.3-68.1</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Snout length</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">21.0-23.9</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">19.2-23.4</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">20.7-22.8</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">22.2-24.4</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Lower jaw length</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">9.5-12.3</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">8.7-11.0</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">9.8-14.3</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">8.9-11.2</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Eye diameter</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">35.7-39.5</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">36.2-41.1</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">35.8-38.5</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">37.2-39.2</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="91">
There is clear sexual dimorphism in fin shape and size (Figs 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. &quot; Lacustricola &quot; johnstoni preserved colouration: A male SAIAB 118776, 34.7 mm SL B female SAIAB 118776, 32.0 mm SL; from Shire River at Liwonde Barrage, Malawi." figureDoi="10.3897/zookeys.923.48420.figure3" httpUri="https://binary.pensoft.net/fig/396600" pageId="0" pageNumber="91">3</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. &quot; Lacustricola &quot; johnstoni colouration in life: A male SAIAB 118776, 31.0 mm SL B female SAIAB 118776, 26 mm SL; from Shire River at Liwonde Barrage, Malawi." figureDoi="10.3897/zookeys.923.48420.figure4" httpUri="https://binary.pensoft.net/fig/396601" pageId="0" pageNumber="91">4</figureCitation>
). In males, dorsal-fin is rounded and elongated, almost reaching the caudal-fin base; its origin in vertical between 7th and 9th anal-fin rays; anal-fin rounded and elongated, reaching middle of the caudal peduncle. Pelvic-fin length variable, reaching between urogenital papillae aperture and the base of third anal-fin ray. In females, dorsal and anal-fins are not elongated and do not extend posteriorly to caudal-fin base. Caudal-fin slender in both sexes. Pectoral-fin elliptical, in both males and females, its posterior margin reaching vertical just behind pelvic-fin base. In females, pelvic-fin shorter than in males, tip reaching region just before urogenital opening. In both males and females, dorsal-fin rays 7(17), 8(13) and 9(2); anal-fin rays 12(4), 13(12), 14(14) and 15(2); caudal-fin rays 19(2), 20(10), 21(17), 22(2) and 23(1); pectoral-fin rays 12(13), 13(18) and 14(1); pelvic-fin rays 6.
</paragraph>
<caption doi="10.3897/zookeys.923.48420.figure3" httpUri="https://binary.pensoft.net/fig/396600" pageId="0" pageNumber="91" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="91">
<emphasis bold="true" pageId="0" pageNumber="91">Figure 3.</emphasis>
&quot; 
<taxonomicName authorityName="Myers" authorityYear="1924" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lacustricola" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="91">Lacustricola</emphasis>
</taxonomicName>
&quot; 
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1894" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="johnstoni" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="species" species="johnstoni">
<emphasis italics="true" pageId="0" pageNumber="91">johnstoni</emphasis>
</taxonomicName>
preserved colouration: 
<emphasis bold="true" pageId="0" pageNumber="91">A</emphasis>
male SAIAB 118776, 34.7 mm SL 
<emphasis bold="true" pageId="0" pageNumber="91">B</emphasis>
female SAIAB 118776, 32.0 mm SL; from Shire River at Liwonde Barrage, Malawi.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.923.48420.figure4" httpUri="https://binary.pensoft.net/fig/396601" pageId="0" pageNumber="91" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="91">
<emphasis bold="true" pageId="0" pageNumber="91">Figure 4.</emphasis>
&quot; 
<taxonomicName authorityName="Myers" authorityYear="1924" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lacustricola" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="91">Lacustricola</emphasis>
</taxonomicName>
&quot; 
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1894" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="johnstoni" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="species" species="johnstoni">
<emphasis italics="true" pageId="0" pageNumber="91">johnstoni</emphasis>
</taxonomicName>
colouration in life: 
<emphasis bold="true" pageId="0" pageNumber="91">A</emphasis>
male SAIAB 118776, 31.0 mm SL 
<emphasis bold="true" pageId="0" pageNumber="91">B</emphasis>
female SAIAB 118776, 26 mm SL; from Shire River at Liwonde Barrage, Malawi.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="91">
Frontal squamation G-patterned (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cephalic pores and head squamation pattern: A, B head dorsal and lateral view of &quot; L. &quot; johnstoni C head lateral view of &quot; L. &quot; myaposae." figureDoi="10.3897/zookeys.923.48420.figure5" httpUri="https://binary.pensoft.net/fig/396602" pageId="0" pageNumber="91">5</figureCitation>
). Head neuromasts placed in shallow grooves. Cephalic lateral line system: anterior portion of supraorbital sensory canal open, with three neuromasts, anteriormost one anteriorly displaced from the other two; posterior portion open, with three exposed neuromasts; anterior infra-orbital canal partially closed, with two pores and one free neuromast, but in juveniles and subadults all anterior infraorbital canal can be open; median portion of infra-orbital region with series of nine to eleven minute neuromasts; posterior infra-orbital canal closed, with two pores; preopercular canal closed in both dorsal and ventral portions with seven or eight pores; mandibular canal represented by two neuromasts, one in vertical through corner of mouth and the other anteriorly positioned in the lower jaw ventral portion. Longitudinal series of scales 27(3), 28(18), 29(9); transverse series of scales 7; circumpeduncular scales 10; predorsal scales 17(1), 18(18), 19(11).
</paragraph>
<caption doi="10.3897/zookeys.923.48420.figure5" httpUri="https://binary.pensoft.net/fig/396602" pageId="0" pageNumber="91" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="91">
<emphasis bold="true" pageId="0" pageNumber="91">Figure 5.</emphasis>
Cephalic pores and head squamation pattern: 
<emphasis bold="true" pageId="0" pageNumber="91">A, B</emphasis>
head dorsal and lateral view of &quot; 
<taxonomicName genus="L." lsidName="L." pageId="0" pageNumber="91" rank="genus">L.</taxonomicName>
&quot; 
<taxonomicName lsidName="johnstoni" pageId="0" pageNumber="91" rank="species" species="johnstoni">
<emphasis italics="true" pageId="0" pageNumber="91">johnstoni</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="91">C</emphasis>
head lateral view of &quot; 
<taxonomicName genus="L." lsidName="L." pageId="0" pageNumber="91" rank="genus">L.</taxonomicName>
&quot; 
<taxonomicName lsidName="myaposae" pageId="0" pageNumber="91" rank="species" species="myaposae">
<emphasis italics="true" pageId="0" pageNumber="91">myaposae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="osteology">
<paragraph pageId="0" pageNumber="91">Osteology.</paragraph>
<paragraph pageId="0" pageNumber="91">
Osteological structures are presented in Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Osteological plate of &quot; L. &quot; johnstoni (SAIAB 35820) from the Upper Shire River, Mangochi, Malawi. A Lachrymal B neurocranium, ventral view C neurocranium dorsal view D ventral post-cleithrum and first pleural rib, lateral view E anal-fin radials and proximal radials, left lateral view F caudal-fin skeleton, left lateral view G dorsal-fin radials and proximal radials, left lateral view H left branchial arches ventral portion, ventral view I right dorsal portion of branchial arches, ventral view J urohyal, left lateral view K left basipterygium, dorsal view L left shoulder girdle, lateral view M right hyoid bar, lateral view N left jaws, jaws suspensorium and opercular apparatus, lateral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.923.48420.figure6" httpUri="https://binary.pensoft.net/fig/396603" pageId="0" pageNumber="91">6</figureCitation>
. Mesethmoid and vomer absent. Frontals, anterior margin, extending anteriorly between nasals. Parasphenoid medial process short, not contacting pterophenoid; anterior margin rounded. Lateral ethmoid overlapping with anterior portion of parasphenoid. Posterior process of supraoccipital long, reaching first vertebra. Lachrymal rectangular. Premaxillary and dentary teeth well developed. Retroarticular triangular or subtriangular. Dentary deep. Dorsal process of maxilla broad overlapping the ascending process of premaxilla; ventral process greatly reduced, consisting of a rounded ventromedial bulge. Ventral arm of maxilla broad, laterally expanded. Medial surface of premaxilla ascending process with a straight or slightly concave profile. Entopterygoid posterior portion and sympletic bone keel deep. Opercle triangular, anterodorsal process present. Anterior process of anterior ceratohyal does not extend ventrally to ventral hypohyal. Urohyal ventral margin concave. First and second basibranchials with expanded lateral bone flanges. Cartilaginous portion of basihyal shorter than osseous portion. Fourth ceratobranchial anterior third with teeth. Second pharyngobranchial plate with teeth. First epibranchial base, broad, more than three times anterior portion width. Supracleithrum rounded. Posttemporal rod-like, ventral arm absent. Cleithrum bony flange not covering scapula foramen. Ventral postcleithrum, slender, similar in width to adjacent first pelural rib. Basipterygium, posterior process, shorter or about the same size of medial process. Anal-fin proximal radials about the same length and parallel to each other. Hypurals completely fused. Parahypural proximal end overlapping the preural centrum. Total vertebrae 30(4) and 31(5), precaudal 13(2) and 14(7) and caudal 16(1), 17(4) and 18(4). First proximal radial of dorsal-fin between neural spine of vertebrae 16 and 17. First proximal radial of anal-fin between pleural rib of vertebrae 11 and 13. Gill rakers on first branchial arch 10(1) and 11(3). Branchiostegal rays 5.
</paragraph>
<caption doi="10.3897/zookeys.923.48420.figure6" httpUri="https://binary.pensoft.net/fig/396603" pageId="0" pageNumber="91" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="91">
<emphasis bold="true" pageId="0" pageNumber="91">Figure 6.</emphasis>
Osteological plate of &quot; 
<taxonomicName genus="L." lsidName="L." pageId="0" pageNumber="91" rank="genus">L.</taxonomicName>
&quot; 
<taxonomicName lsidName="johnstoni" pageId="0" pageNumber="91" rank="species" species="johnstoni">
<emphasis italics="true" pageId="0" pageNumber="91">johnstoni</emphasis>
</taxonomicName>
(SAIAB 35820) from the Upper Shire River, Mangochi, Malawi. 
<emphasis bold="true" pageId="0" pageNumber="91">A</emphasis>
Lachrymal 
<emphasis bold="true" pageId="0" pageNumber="91">B</emphasis>
neurocranium, ventral view 
<emphasis bold="true" pageId="0" pageNumber="91">C</emphasis>
neurocranium dorsal view 
<emphasis bold="true" pageId="0" pageNumber="91">D</emphasis>
ventral post-cleithrum and first pleural rib, lateral view 
<emphasis bold="true" pageId="0" pageNumber="91">E</emphasis>
anal-fin radials and proximal radials, left lateral view 
<emphasis bold="true" pageId="0" pageNumber="91">F</emphasis>
caudal-fin skeleton, left lateral view 
<emphasis bold="true" pageId="0" pageNumber="91">G</emphasis>
dorsal-fin radials and proximal radials, left lateral view 
<emphasis bold="true" pageId="0" pageNumber="91">H</emphasis>
left branchial arches ventral portion, ventral view 
<emphasis bold="true" pageId="0" pageNumber="91">I</emphasis>
right dorsal portion of branchial arches, ventral view 
<emphasis bold="true" pageId="0" pageNumber="91">J</emphasis>
urohyal, left lateral view 
<emphasis bold="true" pageId="0" pageNumber="91">K</emphasis>
left basipterygium, dorsal view 
<emphasis bold="true" pageId="0" pageNumber="91">L</emphasis>
left shoulder girdle, lateral view 
<emphasis bold="true" pageId="0" pageNumber="91">M</emphasis>
right hyoid bar, lateral view 
<emphasis bold="true" pageId="0" pageNumber="91">N</emphasis>
left jaws, jaws suspensorium and opercular apparatus, lateral view. Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="colouration in alcohol">
<paragraph pageId="0" pageNumber="91">Colouration in alcohol.</paragraph>
<paragraph pageId="0" pageNumber="91">
Overall colouration of body pale brown yellow with minute chromatophores sparsely distributed, and some organised chromatophores forming an inconspicuous reticulate pattern along flank scales margin (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. &quot; Lacustricola &quot; johnstoni preserved colouration: A male SAIAB 118776, 34.7 mm SL B female SAIAB 118776, 32.0 mm SL; from Shire River at Liwonde Barrage, Malawi." figureDoi="10.3897/zookeys.923.48420.figure3" httpUri="https://binary.pensoft.net/fig/396600" pageId="0" pageNumber="91">3</figureCitation>
). Ventral surface scarcely pigmented. Pale brown chromatophores along whole mid-body line of flank. Head overall colouration yellowish brown. Brown chromatophores on dorsum of head. Minute chromatophores concentrated in the lower jaw and pre-orbital region, forming a distinct darker region. Iris silver, darker close to pupil; dark pigment concentrated on dorsal margin of eye. All fins hyaline in females, with melanophores sparsely concentrated only on fins membranes and along fin rays; melanophores forming small inconspicuous spots on male dorsal-fins, rarely forming two distinct bands; melanophores forming two distinct parallel dark bands in the anal-fin medial portion; region close to anal-fin rays insertion hyaline; caudal-fin with conspicuous dark blotches in the middle rays that could be organised in distinct bands or not; pectoral and pelvic-fin with melanophores sparsely concentrated on fins membranes and along fin rays. Female urogenital opening pocket scales with dark brown chromatophores.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="colouration in life">
<paragraph pageId="0" pageNumber="91">Colouration in life.</paragraph>
<paragraph pageId="0" pageNumber="91">
<emphasis bold="true" pageId="0" pageNumber="91">Males</emphasis>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. &quot; Lacustricola &quot; johnstoni colouration in life: A male SAIAB 118776, 31.0 mm SL B female SAIAB 118776, 26 mm SL; from Shire River at Liwonde Barrage, Malawi." figureDoi="10.3897/zookeys.923.48420.figure4" httpUri="https://binary.pensoft.net/fig/396601" pageId="0" pageNumber="91">4A</figureCitation>
). Flanks bright blue, with small scattered bright green dots. Cupric iridescent blotch on region just posterior to pectoral-fin. Dorsum yellow-brown. Ventral surface white between head and region anterior to pelvic-fin origin; bright blue between pelvic-fin and the end of the anal-fin insertion; ventral region of caudal peduncle whitish yellow. Side of head predominantly bluish silver, dorsal portion yellow-brown, post-orbital region with a distinct green bright blotch. Iris dark grey, light yellow close to pupil. Eye bright silver on dorsal portion. Lower jaw and pre-orbital region dark brown-grey, forming a distinct horizontal band. Pectoral-fin hyaline; all other fins yellowish brown with brown dots on fins arranged in distinct rows. Dorsal-fin with two to four rows; anal-fin with two to three rows; and caudal-fin with four to five rows, but in some specimens the brown dots are scattered over the fin and not arranged in distinct rows. Some males may present a distinct black distal margin on anal, dorsal and caudal-fins.
</paragraph>
<paragraph pageId="0" pageNumber="91">
<emphasis bold="true" pageId="0" pageNumber="91">Females</emphasis>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. &quot; Lacustricola &quot; johnstoni colouration in life: A male SAIAB 118776, 31.0 mm SL B female SAIAB 118776, 26 mm SL; from Shire River at Liwonde Barrage, Malawi." figureDoi="10.3897/zookeys.923.48420.figure4" httpUri="https://binary.pensoft.net/fig/396601" pageId="0" pageNumber="91">4B</figureCitation>
). Flanks bright blue. Cupric iridescent blotch on region just posterior to pectoral-fin. Dorsum yellow-brown. Venter white between head and region just anterior to urogenital opening; bright blue between region just anterior to urogenital opening and anal-fin insertion; whitish yellow along the anal-fin insertion to caudal peduncle. Scales around urogenital opening covered with dark chromatophores. Side of head predominantly bluish silver, ventral portion white, dorsal portion yellow-brown, post-orbital region with a distinct green bright blotch. Iris dark grey, light yellow close to pupil. Eye bright silver on dorsal portion. Lower jaw and pre-orbital region dark brown-grey, forming a distinct horizontal band. Pectoral and pelvic-fins hyaline; dorsal-fin orange, distal margin hyaline; anal-fin base hyaline, distal portion orange; caudal-fin orange, distal region hyaline.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="distribution">
<paragraph pageId="0" pageNumber="91">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="91">
&quot; 
<taxonomicName authorityName="Myers" authorityYear="1924" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lacustricola" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="91">Lacustricola</emphasis>
</taxonomicName>
&quot; 
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1894" class="Actinopterygii" family="Poeciliidae" genus="Lacustricola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="johnstoni" order="Cyprinodontiformes" pageId="0" pageNumber="91" phylum="Chordata" rank="species" species="johnstoni">
<emphasis italics="true" pageId="0" pageNumber="91">johnstoni</emphasis>
</taxonomicName>
is a widespread species occurring in the Lower, Middle and Upper Zambezi River, including the Shire River and Lake Malawi, the Limpopo River, and is also present in the Okavango system (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Southern Africa &quot; Lacustricola &quot; distribution maps. Spots correspond to the exact localities for the haplotypes included in this study, and the shaded area refers to the inferred distribution for each species. Map A yellow - &quot; Lacustricola &quot; hutereaui; grey - &quot; L. &quot; johnstoni; brown - &quot; L. &quot; myaposae; and purple - &quot; L. &quot; jubbi. Map B dark green - &quot; L. &quot; macrurus; orange - &quot; L. &quot; mediolateralis; red - &quot; L. &quot; katangae; light blue - &quot; L. &quot; centralis; blue - &quot; L. &quot; aff. johnstoni ' Okavango'; light green - &quot; L. &quot; aff. johnstoni ' Congo'." figureDoi="10.3897/zookeys.923.48420.figure2" httpUri="https://binary.pensoft.net/fig/396599" pageId="0" pageNumber="91">2</figureCitation>
). The species is usually found associated with marginal vegetation along the banks of small and large rivers, or in swampy areas, as well as along the shores of Lake Malawi and Lake Kariba.
</paragraph>
</subSubSection>
</treatment>
</document>