175 lines
17 KiB
XML
175 lines
17 KiB
XML
<document id="64D3A82892AD1C617134D148E00E937C" ID-DOI="10.11646/zootaxa.3999.1.2" ID-GBIF-Dataset="12d1122e-3201-4fb2-a139-cb58026f2146" ID-ISSN="1175-5326" ID-Zenodo-Dep="234596" ID-ZooBank="FBF20477-7DF3-4927-A415-D4BC98736B98" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461279299333" checkinUser="plazi" docAuthor="Heppner, John B. & Bae, Yang-Seop" docDate="2015" docId="8B2F87A12C16DF7FFF42D7D7FC9CFB82" docLanguage="en" docName="zt03999p040.pdf" docOrigin="Zootaxa 3999 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nexosa hexaphala Meyrick 1912" docType="treatment" docVersion="8" masterDocId="7716FFD92C15DF7CFFD5D740FFACFFF2" masterDocTitle="Review of Nexosa Diakonoff in Vietnam, with a new species and a new subspecies, and transfer to the tribe Archipini (Lepidoptera: Tortricidae: Tortricinae: Archipini)" updateTime="1698613642155" updateUser="plazi">
|
||
<mods:mods id="EBDE7C96A8C21987AABD3904946988D3" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="9BED60476C49F3B1032D76DD52CDE17A">
|
||
<mods:title id="4CF9EA9D3A2C3DE69B1FF479D0161FCD">Review of Nexosa Diakonoff in Vietnam, with a new species and a new subspecies, and transfer to the tribe Archipini (Lepidoptera: Tortricidae: Tortricinae: Archipini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="997E106CCBD19C17976BA6A6022BCFBE" type="personal">
|
||
<mods:role id="0E695D6723998CD1D829B998769B0A51">
|
||
<mods:roleTerm id="402F777EF01149F6AF76833585AEC9CB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F5F0FBDB99B27DE1AD63E3209B91F11A">Heppner, John B.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="0ED4575EF48106A7FDFAAE87AA5EE94B" type="personal">
|
||
<mods:role id="610FD6B6BBF6C56EE549C401993B8030">
|
||
<mods:roleTerm id="9530CC7E94F7840A9CD0E9F42DC84369">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="EBAA55FA4ECF4F4734D1A8457E8FEB86">Bae, Yang-Seop</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="EDB9F95D816B40A62348E457764C8AF4">text</mods:typeOfResource>
|
||
<mods:relatedItem id="461D12B67447966DD9AA249E51121CDF" type="host">
|
||
<mods:titleInfo id="6142C6E56ED425431198074245CAC048">
|
||
<mods:title id="FFCBFE3B3048A2D1802BA597BF8663F4">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="99A7BAAD5213050A468B546BA398E7F4">
|
||
<mods:date id="4B1AF7AEC577361BC5F56F6D2275C1E5">2015</mods:date>
|
||
<mods:detail id="0862242271F7A92D745EA4307AD30130" type="volume">
|
||
<mods:number id="FFE73102379DF06FEE517F87E78528E7">3999</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="661921933D4F5113C9D0D644AA02B507" type="issue">
|
||
<mods:number id="616E045880A9763B9D360093C4A64937">1</mods:number>
|
||
</mods:detail>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="70223E1D121BCFFF446EC57D810B7754">journal volume</mods:classification>
|
||
<mods:identifier id="6CA71ABB4544E002AE0C424A5014C457" type="DOI">10.11646/zootaxa.3999.1.2</mods:identifier>
|
||
<mods:identifier id="48F4A97E62C2B14661060FCDEA801279" type="GBIF-Dataset">12d1122e-3201-4fb2-a139-cb58026f2146</mods:identifier>
|
||
<mods:identifier id="B411054815F4FD357B2165D7DA29DF87" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="20954AD2E6670C061F5B0355BD3BD6E0" type="Zenodo-Dep">234596</mods:identifier>
|
||
<mods:identifier id="0DF9A9B889E133072AD32FDDA6090E86" type="ZooBank">FBF20477-7DF3-4927-A415-D4BC98736B98</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="8B2F87A12C16DF7FFF42D7D7FC9CFB82" ID-DOI="http://doi.org/10.5281/zenodo.6094741" ID-GBIF-Taxon="119640306" ID-Zenodo-Dep="6094741" LSID="urn:lsid:plazi:treatment:8B2F87A12C16DF7FFF42D7D7FC9CFB82" httpUri="http://treatment.plazi.org/id/8B2F87A12C16DF7FFF42D7D7FC9CFB82" pageId="3">
|
||
<subSubSection id="4B9C653C2C16DF7FFF42D7D7FDA5FEC5" pageId="3" type="nomenclature">
|
||
<paragraph id="033936B72C16DF7FFF42D7D7FDE8FF40" blockId="3.[151,580,151,211]" box="[151,580,151,178]" pageId="3">
|
||
<heading id="587181DB2C16DF7FFF42D7D7FDE8FF40" bold="true" box="[151,580,151,178]" fontSize="11" level="1" pageId="3" reason="1">
|
||
<taxonomicName id="C4864D342C16DF7FFF42D7D7FDE8FF40" ID-CoL="47BZZ" authority="Meyrick, 1912" authorityName="Meyrick" authorityYear="1912" box="[151,580,151,178]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="3" phylum="Arthropoda" rank="species" species="hexaphala">
|
||
<emphasis id="31F2EAA52C16DF7FFF42D7D7FDE8FF40" bold="true" box="[151,580,151,178]" pageId="3">
|
||
<emphasis id="31F2EAA52C16DF7FFF42D7D7FEDBFF43" bold="true" box="[151,375,151,177]" italics="true" pageId="3">Nexosa hexaphala</emphasis>
|
||
(Meyrick, 1912)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="033936B72C16DF7FFF42D7FAFEE6FF21" blockId="3.[151,580,151,211]" box="[151,330,186,211]" pageId="3">
|
||
(
|
||
<figureCitation id="9BBD2A322C16DF7FFF4AD7FAFE8BFF21" box="[159,295,186,211]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 6" captionStartId-0="1.[151,264,1602,1625]" captionStartId-1="2.[151,264,1813,1836]" captionTargetBox-0="[160,1436,917,1565]" captionTargetBox-1="[150,1436,591,1791]" captionTargetId-0="figure@1.[151,1436,882,1578]" captionTargetId-1="figure@2.[147,1440,591,1791]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1 – 2. Nexosa species in Vietnam. 1. N. hexaphala tamdaoana, n. subsp., holotype male (13.8 mm wingspan). 2. N. tonkinensis, n. sp., holotype female (12.3 mm wingspan)." captionText-1="FIGURES 3 – 6. Nexosa species head close-ups (lateral and dorsal views). 3 – 4. N. hexaphala tamdaoana, n. subsp., male. 5 – 6. N. tonkinensis, n. sp., female." httpUri-0="https://zenodo.org/record/234597/files/figure.png" httpUri-1="https://zenodo.org/record/234598/files/figure.png" pageId="3">Figs. 1, 2–3</figureCitation>
|
||
,
|
||
<figureCitation id="9BBD2A322C16DF7FFEE7D7FBFEEDFF21" box="[306,321,187,211]" captionStart="FIGURE 7" captionStartId="3.[151,250,1976,1998]" captionTargetBox="[169,1408,1184,1955]" captionTargetId="figure@3.[159,1424,1175,1955]" captionTargetPageId="3" captionText="FIGURE 7. Nexosa hexaphala tamdaoana, n. subsp., male genitalia, with details a) aedeagus, b) abdominal pregenital sternal plate (slide JBH 2858)." httpUri="https://zenodo.org/record/234599/files/figure.png" pageId="3">7</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="033936B72C16DF7FFF42D642FDA5FEC5" blockId="3.[151,870,258,311]" pageId="3">
|
||
<taxonomicName id="C4864D342C16DF7FFF42D642FD84FEEA" authority="Meyrick, 1912" authorityName="Meyrick" authorityYear="1912" box="[151,552,258,280]" class="Insecta" family="Tortricidae" genus="Mictopsichia" kingdom="Animalia" order="Lepidoptera" pageId="3" phylum="Arthropoda" rank="species" species="hexaphala">
|
||
<emphasis id="31F2EAA52C16DF7FFF42D642FE27FEEA" box="[151,395,258,280]" italics="true" pageId="3">Mictopsichia hexaphala</emphasis>
|
||
Meyrick, 1912
|
||
</taxonomicName>
|
||
, Exot. Microlepidoptera, 1: 36.
|
||
<typeStatus id="DC3D88152C16DF7FFF42D662FF65FEC5" box="[151,201,290,311]" pageId="3">Type</typeStatus>
|
||
locality: Maskeliya,
|
||
<collectingCountry id="7B9176272C16DF7FFE4AD661FDAAFEC5" box="[415,518,289,311]" name="Sri Lanka" pageId="3">Sri Lanka</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4B9C653C2C16DF7FFF42D624FED5FCFA" pageId="3" type="diagnosis">
|
||
<paragraph id="033936B72C16DF7FFF42D624FF45FE1A" blockId="3.[151,1437,356,1136]" pageId="3">
|
||
<emphasis id="31F2EAA52C16DF7FFF42D624FEA6FE8F" bold="true" box="[151,266,356,381]" pageId="3">Diagnosis</emphasis>
|
||
. Wing expanse:
|
||
<quantity id="C47E9B522C16DF7FFE13D624FDB9FE8E" box="[454,533,356,380]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="3" unit="mm" value="14.0">14 mm</quantity>
|
||
(n = 1). This species is notable for the extensive white on the hindwings in both males and females, whereas the new species from
|
||
<collectingCountry id="7B9176272C16DF7FFD35D6C7FC93FE52" box="[736,831,391,416]" name="Vietnam" pageId="3">Vietnam</collectingCountry>
|
||
has darker hindwings in the female, with less white evident. The white hindwing of the female
|
||
<taxonomicName id="C4864D342C16DF7FFDA3D6EDFCA7FE37" box="[630,779,428,453]" class="Insecta" family="Tortricidae" genus="Mictopsichia" kingdom="Animalia" order="Lepidoptera" pageId="3" phylum="Arthropoda" rank="species" species="hexaphala">
|
||
<emphasis id="31F2EAA52C16DF7FFDA3D6EDFCA7FE37" box="[630,779,428,453]" italics="true" pageId="3">N. hexaphala</emphasis>
|
||
</taxonomicName>
|
||
can be noted in the figure of the
|
||
<typeStatus id="DC3D88152C16DF7FFBA2D6ECFB75FE37" box="[1143,1241,428,453]" pageId="3" type="holotype">holotype</typeStatus>
|
||
shown by
|
||
<bibRefCitation id="67174B462C16DF7FFA84D6ECFF48FE1A" author="Clarke" pageId="3" refString="Clarke, J. F. G. (1969) Catalogue of the Type Specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick. Vol. VI. Glyphipterigidae, Gelechiidae (A - C). British Museum (Natural History), London, 537 pp., 267 pls." type="book" year="1969">Clarke (1969)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="033936B72C16DF7FFF12D6B4FD83FD32" blockId="3.[151,1437,356,1136]" pageId="3">
|
||
<bibRefCitation id="67174B462C16DF7FFF12D6B4FE21FDFE" author="Diakonoff" box="[199,397,500,525]" pageId="3" refString="Diakonoff, A. N. (1977) Rearrangement of certain Glyphipterygidae sensu Meyrick, 1913, with descriptions of new taxa (Lepidoptera). Zoologische Verhandelingen, 158, 1 - 55." type="journal article" year="1977">Diakonoff (1977)</bibRefCitation>
|
||
erroneously stated that
|
||
<bibRefCitation id="67174B462C16DF7FFD74D6B4FC93FDFF" author="Clarke" box="[673,831,500,525]" pageId="3" refString="Clarke, J. F. G. (1969) Catalogue of the Type Specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick. Vol. VI. Glyphipterigidae, Gelechiidae (A - C). British Museum (Natural History), London, 537 pp., 267 pls." type="book" year="1969">Clarke (1969)</bibRefCitation>
|
||
had illustrated the female genitalia of
|
||
<taxonomicName id="C4864D342C16DF7FFB28D6B5FA38FDFF" box="[1277,1428,500,525]" class="Insecta" family="Tortricidae" genus="Mictopsichia" kingdom="Animalia" order="Lepidoptera" pageId="3" phylum="Arthropoda" rank="species" species="hexaphala">
|
||
<emphasis id="31F2EAA52C16DF7FFB28D6B5FA38FDFF" box="[1277,1428,500,525]" italics="true" pageId="3">N. hexaphala</emphasis>
|
||
</taxonomicName>
|
||
, but Diakonoff mistook a figure of the genitalia of a Neotropical species of
|
||
<taxonomicName id="C4864D342C16DF7FFC06D557FBC9FDC2" box="[979,1125,535,560]" class="Insecta" family="Tortricidae" genus="Mictopsichia" kingdom="Animalia" order="Lepidoptera" pageId="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C16DF7FFC06D557FBC9FDC2" box="[979,1125,535,560]" italics="true" pageId="3">Mictopsichia</emphasis>
|
||
</taxonomicName>
|
||
illustrated above that for
|
||
<taxonomicName id="C4864D342C16DF7FFA57D559FEA0FDA7" class="Insecta" family="Tortricidae" genus="Mictopsichia" kingdom="Animalia" order="Lepidoptera" pageId="3" phylum="Arthropoda" rank="species" species="hexaphala">
|
||
<emphasis id="31F2EAA52C16DF7FFA57D559FEA0FDA7" italics="true" pageId="3">N. hexaphala</emphasis>
|
||
</taxonomicName>
|
||
: the
|
||
<typeStatus id="DC3D88152C16DF7FFE9ED57CFE01FDA7" box="[331,429,572,597]" pageId="3" type="holotype">holotype</typeStatus>
|
||
female lacks an abdomen as Clarke noted. Because the species has not had a modern description, and none for the genitalia, a redescription is given in more detail herein based on the new
|
||
<collectingCountry id="7B9176272C16DF7FFAE8D51FFA30FD8A" box="[1341,1436,607,632]" name="Vietnam" pageId="3">Vietnam</collectingCountry>
|
||
subspecies which can be compared to the
|
||
<collectingCountry id="7B9176272C16DF7FFD55D5C4FD5DFD6F" box="[640,753,644,669]" name="Sri Lanka" pageId="3">Sri Lanka</collectingCountry>
|
||
population of the nominate subspecies, which otherwise is known from only a single specimen.
|
||
</paragraph>
|
||
<paragraph id="033936B72C16DF7FFF12D58CFED5FCFA" blockId="3.[151,1437,356,1136]" pageId="3">
|
||
<emphasis id="31F2EAA52C16DF7FFF12D58CFE10FD17" bold="true" box="[199,444,716,741]" pageId="3">Specimens recorded</emphasis>
|
||
.
|
||
<emphasis id="31F2EAA52C16DF7FFE1BD58CFD20FD17" box="[462,652,716,741]" italics="true" pageId="3">
|
||
<typeStatus id="DC3D88152C16DF7FFE1BD58CFD98FD17" box="[462,564,716,741]" pageId="3" type="holotype">Holotype</typeStatus>
|
||
female
|
||
</emphasis>
|
||
: Maskeliya, [Ratnapura Dist.],
|
||
<collectingCountry id="7B9176272C16DF7FFBD7D58CFBFFFD17" box="[1026,1107,716,741]" name="Sri Lanka" pageId="3">Ceylon</collectingCountry>
|
||
[
|
||
<collectingCountry id="7B9176272C16DF7FFBB3D58CFB4DFD17" box="[1126,1249,716,741]" name="Sri Lanka" pageId="3">Sri Lanka</collectingCountry>
|
||
],
|
||
<date id="773810772C16DF7FFB22D58DFADDFD17" box="[1271,1393,716,741]" pageId="3" value="1906-05">May 1906</date>
|
||
, de Mowbray (BMNH).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4B9C653C2C16DF7FFF12D454FAAAFCDF" box="[199,1286,788,813]" pageId="3" type="distribution">
|
||
<paragraph id="033936B72C16DF7FFF12D454FAAAFCDF" blockId="3.[151,1437,356,1136]" box="[199,1286,788,813]" pageId="3">
|
||
<emphasis id="31F2EAA52C16DF7FFF12D454FEF5FCDF" bold="true" box="[199,345,788,813]" pageId="3">Distribution</emphasis>
|
||
. The nominate subspecies,
|
||
<taxonomicName id="C4864D342C16DF7FFD5BD455FC33FCDF" box="[654,927,788,813]" class="Insecta" family="Tortricidae" genus="Mictopsichia" kingdom="Animalia" order="Lepidoptera" pageId="3" phylum="Arthropoda" rank="subSpecies" species="hexaphala" subSpecies="hexaphala">
|
||
<emphasis id="31F2EAA52C16DF7FFD5BD455FC33FCDF" box="[654,927,788,813]" italics="true" pageId="3">N. hexaphala hexaphala</emphasis>
|
||
</taxonomicName>
|
||
, is known only from
|
||
<collectingCountry id="7B9176272C16DF7FFB44D454FAAEFCDF" box="[1169,1282,788,813]" name="Sri Lanka" pageId="3">Sri Lanka</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4B9C653C2C16DF7FFF12D477FC9CFB82" pageId="3" type="discussion">
|
||
<paragraph id="033936B72C16DF7FFF12D477FC9CFB82" blockId="3.[151,1437,356,1136]" pageId="3">
|
||
<emphasis id="31F2EAA52C16DF7FFF12D477FEE9FCA2" bold="true" box="[199,325,823,848]" pageId="3">Discussion</emphasis>
|
||
. The description below for the
|
||
<collectingCountry id="7B9176272C16DF7FFD6CD477FCB4FCA2" box="[697,792,823,848]" name="Vietnam" pageId="3">Vietnam</collectingCountry>
|
||
subspecies is essentially the same as for the
|
||
<collectingCountry id="7B9176272C16DF7FFAFED477FA30FCA2" box="[1323,1436,823,848]" name="Sri Lanka" pageId="3">Sri Lanka</collectingCountry>
|
||
nominate subspecies, as far as can be determined at this time, except for the following points: forewing with fewer black-brown spots near base at 1/3 and the hindwing with much larger black patch on the termen and tornus and with two brown patches near the wing base (only one in the
|
||
<collectingCountry id="7B9176272C16DF7FFC87D4E4FC1DFC4F" box="[850,945,932,957]" name="Vietnam" pageId="3">Vietnam</collectingCountry>
|
||
specimens). The female
|
||
<typeStatus id="DC3D88152C16DF7FFB04D4E4FA9FFC4F" box="[1233,1331,932,957]" pageId="3" type="holotype">holotype</typeStatus>
|
||
from
|
||
<collectingCountry id="7B9176272C16DF7FFAA9D4E4FF71FC12" name="Sri Lanka" pageId="3">Sri Lanka</collectingCountry>
|
||
remains the single known female of this species and lacks an abdomen, thus the genitalia are not known and cannot be compared with the
|
||
<collectingCountry id="7B9176272C16DF7FFE3ED4ACFDE6FBF7" box="[491,586,1004,1029]" name="Vietnam" pageId="3">Vietnam</collectingCountry>
|
||
congeners. Although the female would be expected to differ somewhat in maculation from the male, some aspects of the hindwing, as well as the great distance from
|
||
<collectingCountry id="7B9176272C16DF7FFB7CD34FFAB4FBDA" box="[1193,1304,1039,1064]" name="Sri Lanka" pageId="3">Sri Lanka</collectingCountry>
|
||
to northern
|
||
<collectingCountry id="7B9176272C16DF7FFF42D374FF5AFBBF" box="[151,246,1076,1101]" name="Vietnam" pageId="3">Vietnam</collectingCountry>
|
||
and no intervening records in
|
||
<collectingCountry id="7B9176272C16DF7FFD9DD374FD28FBBF" box="[584,644,1076,1101]" name="India" pageId="3">India</collectingCountry>
|
||
,
|
||
<collectingCountry id="7B9176272C16DF7FFD5AD375FD72FBBE" box="[655,734,1077,1100]" name="Myanmar" pageId="3">Burma</collectingCountry>
|
||
,
|
||
<collectingCountry id="7B9176272C16DF7FFD3DD374FCE2FBBF" box="[744,846,1076,1101]" name="Thailand" pageId="3">Thailand</collectingCountry>
|
||
, or
|
||
<collectingCountry id="7B9176272C16DF7FFCA2D375FC1CFBBE" box="[887,944,1077,1100]" name="Laos" pageId="3">Laos</collectingCountry>
|
||
, suggest that the
|
||
<collectingCountry id="7B9176272C16DF7FFBBBD374FB61FBBF" box="[1134,1229,1076,1101]" name="Vietnam" pageId="3">Vietnam</collectingCountry>
|
||
population should be treated as a separate subspecies from those in
|
||
<collectingCountry id="7B9176272C16DF7FFD6ED317FC81FB82" box="[699,813,1111,1136]" name="Sri Lanka" pageId="3">Sri Lanka</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |