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<document id="1618BA47D33CA7980D74BB7D6F87D2F4" ID-CLB-Dataset="2617" ID-DOI="10.5169/seals-787038" ID-GBIF-Dataset="736b1480-7963-499f-8ca5-19608edf08d1" ID-ISSN="2624-9170" ID-Zenodo-Dep="5846115" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637934267982" checkinUser="carolina" docAuthor="Herrmann, Manfred, Stossier, Günter &amp; Salisbury, Richard" docDate="2014" docId="61145817244DFFE4FF20045EFD12829A" docLanguage="en" docName="ContribNatHist.24.1-55.pdf.imf" docOrigin="Contributions to Natural History 24" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Vexillum (Protoelongata) corallinum Herrmann &amp; Stossier &amp; Salisbury 2014, comb. nov." docType="treatment" docVersion="7" lastPageNumber="16" masterDocId="9D2D206F2440FFEBFF970045FFF3807E" masterDocTitle="A new subgenus including three new species of the genus Vexillum (Gastropoda: Costellariidae) from the central Indo-Pacific with remarks on Vexillum (Pusia) semicostatum (ANTON, 1838)" masterLastPageNumber="55" masterPageNumber="1" pageNumber="14" updateTime="1699470458461" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="B696FE0B91AC7D79C48417A069AB43DC">A new subgenus including three new species of the genus Vexillum (Gastropoda: Costellariidae) from the central Indo-Pacific with remarks on Vexillum (Pusia) semicostatum (ANTON, 1838)</mods:title>
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<mods:namePart id="0B2F654A195442FC92CE45E98EDEC8AC">Herrmann, Manfred</mods:namePart>
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<mods:namePart id="2564DA72964510F272EEF986C75CB1C0">Stossier, Günter</mods:namePart>
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<mods:namePart id="A1D34299326C86B0798F26614140826F">Salisbury, Richard</mods:namePart>
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<mods:title id="16CE568C28A6C06F5F1E2FFB6046CD29">Contributions to Natural History</mods:title>
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<mods:date id="FE0AE9702402B123DFF15ED19A420924">2014</mods:date>
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<mods:number id="CAB2333C636FF2F7FFB0FAC99CEAC63D">24</mods:number>
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<mods:identifier id="7B68FEFC5A9008D333F4AC2E5BE50A01" type="DOI">10.5169/seals-787038</mods:identifier>
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<treatment id="61145817244DFFE4FF20045EFD12829A" ID-DOI="http://doi.org/10.5281/zenodo.5846179" ID-GBIF-Taxon="192068544" ID-Zenodo-Dep="5846179" LSID="urn:lsid:plazi:treatment:61145817244DFFE4FF20045EFD12829A" httpUri="http://treatment.plazi.org/id/61145817244DFFE4FF20045EFD12829A" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
<subSubSection id="A1A7BA8A244DFFE6FF20045EFC88844A" box="[183,891,1051,1076]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="E902E901244DFFE6FF20045EFC88844A" blockId="13.[183,891,1051,1113]" box="[183,891,1051,1076]" pageId="13" pageNumber="14">
<heading id="B24A5E6D244DFFE6FF20045EFC88844A" box="[183,891,1051,1076]" centered="true" fontSize="36" level="2" pageId="13" pageNumber="14" reason="4">
<emphasis id="DBC93513244DFFE6FF20045EFC88844A" bold="true" box="[183,891,1051,1076]" pageId="13" pageNumber="14">
<taxonomicName id="2EBD9282244DFFE6FF20045EFD08844A" ID-CoL="7FQN2" authority="(REEVE, 1845)" authorityName="Herrmann &amp; Stossier &amp; Salisbury" authorityYear="2014" baseAuthorityName="REEVE" baseAuthorityYear="1845" box="[183,763,1051,1076]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="corallinum" status="comb. nov." subGenus="Protoelongata">
<emphasis id="DBC93513244DFFE6FF20045EFEE8844A" bold="true" box="[183,283,1051,1076]" italics="true" pageId="13" pageNumber="14">Vexillum</emphasis>
(
<emphasis id="DBC93513244DFFE6FEBC045EFE3E844A" bold="true" box="[299,461,1051,1076]" italics="true" pageId="13" pageNumber="14">Protoelongata</emphasis>
)
<emphasis id="DBC93513244DFFE6FE4A045EFDA5844A" bold="true" box="[477,598,1051,1076]" italics="true" pageId="13" pageNumber="14">corallinum</emphasis>
(
<bibRefCitation id="8D2C94F0244DFFE6FDF2045EFD01844D" author="Reeve, L. A." box="[613,754,1051,1075]" pageId="13" pageNumber="14" refId="ref12476" refString="Reeve, L. A. (1845): Monograph of the genus Mitra. Vol. 2. - London." type="book" year="1845">REEVE, 1845</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="C0FA8868244DFFE6FC96045EFC88844A" box="[769,891,1051,1076]" pageId="13" pageNumber="14" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="A1A7BA8A244DFFE4FF200404FE858137" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
<paragraph id="E902E901244DFFE6FF200404FE5C8427" blockId="13.[183,891,1051,1113]" box="[183,431,1089,1113]" pageId="13" pageNumber="14">
(
<figureCitation id="7186F584244DFFE6FF2A0404FEF18427" box="[189,258,1089,1113]" captionStart="Plate 1" captionStartId="5.[184,229,1163,1181]" captionTargetId="figure-10@6.[221,1104,153,1624]" captionTargetPageId="6" captionText="Plate 1: Vexillum (Protoelongata) subgen. nov., all figures 4.5 x Fig. 1: Vexillum (Protoelongata) dekkersi sp. nov., paratype 1, Philippines, Bohol, Balicasag Island, collected 2000, 11.7 mm, MS 63760. Fig. 2: Vexillum (Protoelongata) bilineatum (REEVE, 1845) comb. nov., Philippines, Cebu, dived 15–20 m, 20.6 mm, MH. Fig. 3: Vexillum (Protoelongata) xerampelina (MELVILL, 1895) comb. nov., holotype, Arabian Sea, Yemen, Aden, 18.8 mm, ex coll. E.R. Shopland; J.C. Melvill, NMW.1955.158.00429 (coll. Melvill-Tomlin). Fig. 4: Vexillum (Protoelongata) rubrotaeniatum sp. nov., paratype 9, Solomon Islands, Guadalcanal, Bonegi, collected Aug 1987 by F. Lorenz on rocks at night at 21 m, 13.8 mm, MH. Fig. 5: Vexillum (Protoelongata) corallinum (REEVE, 1845) comb. nov., Philippines, Bohol, Balicasag Island, 73–88 m, 17.5 mm, ex coll. Steinke, MH. Fig. 6: Vexillum (Protoelongata) heleneae sp. nov., holotype, Guam, Orote Point, Orote Cliffs, collected by R. Salisbury under a pile of rocks at 18 m, 7 Jan 1978, 17.2 mm, MNHN 27071. Fig. 7: Vexillum (Protoelongata) loyaltyense (HERVIER, 1897) comb. nov., Philippines, Cebu, Mactan Island, Punta Engaño, collected 2009 by lumun lumun net at 25–50 m, 10.3 mm, MH. Photos: Fig. 3 Harriet Wood (NMW), Fig. 6 Richard Salisbury, other photos Manfred Herrmann." figureDoi="http://doi.org/10.5281/zenodo.5846119" httpUri="https://zenodo.org/record/5846119/files/figure.png" pageId="13" pageNumber="14">Figs 5</figureCitation>
,
<figureCitation id="7186F584244DFFE6FE870404FEA68427" box="[272,341,1089,1113]" captionStart="Plate 5" captionStartId="14.[221,266,1243,1261]" captionTargetBox="[221,1105,153,1219]" captionTargetId="figure-73@14.[221,1105,153,1221]" captionTargetPageId="14" captionText="Plate 5: Vexillum (Protoelongata) corallinum (REEVE, 1845) comb. nov. Figs 15–16: Philippines, Bohol, Panglao area, collected 2004. © MNHN." figureDoi="http://doi.org/10.5281/zenodo.5846129" httpUri="https://zenodo.org/record/5846129/files/figure.png" pageId="13" pageNumber="14">15–16</figureCitation>
,
<figureCitation id="7186F584244DFFE6FEF50404FE5A8427" box="[354,425,1089,1113]" captionStart="Plate 10" captionStartId="35.[184,229,1376,1394]" captionTargetId="figure-10@36.[221,1105,153,1627]" captionTargetPageId="36" captionText="Plate 10: Vexillum (Protoelongata) corallinum (REEVE, 1845) comb. nov., figures 4 x Fig. 31: Holotype, Philippines, Masbate, found on the sands, 14.8 mm, NHMUK 1967727. Figs 32–33: Juvenile Specimen, Philippines, Bohol, Balicasag, lumun lumun nets, collected 2006, 6.8 mm, MH (fig. 33 detail of first whorls with elongated protoconch, 12 x). Fig. 34: Specimen, Philippines, Bohol, Balicasag Island, 73–88 m, 17.5 mm, ex coll. Steinke, MH. Fig. 35: Specimen, Australia, Queensland, Credlin Reef, dived up Jul 1997 from reef slope at 8 m at night, 17.1 mm, MH. Photos: Fig. 31 John Wolff, other photos Manfred Herrmann." figureDoi="http://doi.org/10.5281/zenodo.5846141" httpUri="https://zenodo.org/record/5846141/files/figure.png" pageId="13" pageNumber="14">31–35</figureCitation>
)
</paragraph>
<paragraph id="E902E901244DFFE6FF2E04CEFECF84DA" blockId="13.[185,316,1163,1188]" box="[185,316,1163,1188]" pageId="13" pageNumber="14">
<emphasis id="DBC93513244DFFE6FF2E04CEFECF84DA" bold="true" box="[185,316,1163,1188]" pageId="13" pageNumber="14">Description</emphasis>
</paragraph>
<paragraph id="E902E901244DFFE6FF2F0493FEF68640" blockId="13.[183,1073,1238,1598]" pageId="13" pageNumber="14">
Slender, fusiform, nearly biconical shell, shiny, reaching about
<quantity id="2E4544E4244DFFE6FCE50493FC4D8490" box="[882,958,1238,1262]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="14" unit="mm" value="22.0">22 mm</quantity>
in length. Elongated protoconch consists of at least three whorls. Teleoconch consists of 7 slightly convex whorls. Spire outline straight, first two to three whorls with distinct narrow axial ribs, remaining whorls with broad but flat axial ribs. Last third to half of body whorl often smooth. Whole shell covered with fine spiral striae. Aperture shorter than half the entire shell, lirate inside. Lip straight in posterior half, then recurved towards the siphonal fasciole. Siphonal canal short, narrow and slightly recurved. Columella with 4 strong folds, decreasing in size anteriorly. Folds merge into weak spiral cords when leaving the columellar shield.
</paragraph>
<caption id="BDC2B989244EFFE5FF4A049EFCB8855D" ID-DOI="http://doi.org/10.5281/zenodo.5846129" ID-Zenodo-Dep="5846129" httpUri="https://zenodo.org/record/5846129/files/figure.png" pageId="14" pageNumber="15" startId="14.[221,266,1243,1261]" subCaptionStartIDs="14.[221,258,1297,1315]" subCaptionStarts="Figs 15" targetBox="[221,1105,153,1219]" targetPageId="14">
<paragraph id="E902E901244EFFE5FF4A049EFCB08493" blockId="14.[221,843,1243,1315]" box="[221,835,1243,1261]" pageId="14" pageNumber="15">
Plate 5:
<taxonomicName id="2EBD9282244EFFE5FEB2049EFD128493" authority="(REEVE, 1845)" authorityName="Herrmann &amp; Stossier &amp; Salisbury" authorityYear="2014" baseAuthorityName="REEVE" baseAuthorityYear="1845" box="[293,737,1243,1261]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="corallinum" status="comb. nov." subGenus="Protoelongata">
<emphasis id="DBC93513244EFFE5FEB2049EFE828493" bold="true" box="[293,369,1243,1261]" italics="true" pageId="14" pageNumber="15">Vexillum</emphasis>
(
<emphasis id="DBC93513244EFFE5FEE9049EFE098493" bold="true" box="[382,506,1243,1261]" italics="true" pageId="14" pageNumber="15">Protoelongata</emphasis>
)
<emphasis id="DBC93513244EFFE5FD91049EFD908493" bold="true" box="[518,611,1243,1261]" italics="true" pageId="14" pageNumber="15">corallinum</emphasis>
(
<bibRefCitation id="8D2C94F0244EFFE5FDF9049EFD2A8493" author="Reeve, L. A." box="[622,729,1243,1261]" pageId="14" pageNumber="15" refId="ref12476" refString="Reeve, L. A. (1845): Monograph of the genus Mitra. Vol. 2. - London." type="book" year="1845">REEVE, 1845</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="C0FA8868244EFFE5FD72049EFCB08493" box="[741,835,1243,1261]" pageId="14" pageNumber="15" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="E902E901244EFFE5FF4A0554FCB8855D" blockId="14.[221,843,1243,1315]" box="[221,843,1297,1315]" pageId="14" pageNumber="15">
<emphasis id="DBC93513244EFFE5FF4A0554FCB8855D" bold="true" box="[221,843,1297,1315]" pageId="14" pageNumber="15">
<collectingCounty id="0063918D244EFFE5FF4A0554FEF1855D" box="[221,258,1297,1315]" pageId="14" pageNumber="15">Figs</collectingCounty>
15–16:
<collectingCountry id="91AAA991244EFFE5FED10554FE5F855D" box="[326,428,1297,1315]" name="Philippines" pageId="14" pageNumber="15">Philippines</collectingCountry>
,
<collectingRegion id="2B7927E3244EFFE5FE230554FE19855D" box="[436,490,1297,1315]" country="Philippines" name="Bohol" pageId="14" pageNumber="15">Bohol</collectingRegion>
,
<collectingCounty id="0063918D244EFFE5FE650554FD94855D" box="[498,615,1297,1315]" pageId="14" pageNumber="15">Panglao area</collectingCounty>
, collected 2004. ©
<collectionCode id="8FAC71C4244EFFE5FC990554FCB5855D" box="[782,838,1297,1315]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="15">MNHN</collectionCode>
.
</emphasis>
</paragraph>
</caption>
<paragraph id="E902E901244EFFE5FE9405D4FBE98640" blockId="14.[222,1111,1425,1598]" pageId="14" pageNumber="15">Colour pattern: Shell colour deep red, seldom orange. Protoconch yellowish translucent, first two to three whorls changing from yellowish orange to deep red without white markings. All remaining whorls regularly covered with white dots on the ribs in the anterior part of the whorls, often faded in the last half of the body whorl. Columellar folds greyish, aperture reddish orange.</paragraph>
<paragraph id="E902E901244FFFE4FF4A00D9FE858137" blockId="15.[182,1073,156,740]" pageId="15" pageNumber="16">Animal: Siphon black with many, equal-sized white or yellow spots, animal body black, with white spots, foot underneath greyish black with yellowish white margin, base of eyestalk black with white spots, eyestalk thick with a large black eye at the base, upper portion of the eyestalk semitransparent with fine white spots.</paragraph>
</subSubSection>
<subSubSection id="A1A7BA8A244FFFE4FF490113FCBB81EA" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="E902E901244FFFE4FF490113FD7E8110" blockId="15.[182,1073,156,740]" box="[222,653,342,366]" pageId="15" pageNumber="16">
<materialsCitation id="59D5E35C244FFFE4FF490113FD7E8110" ID-GBIF-Occurrence="3436331334" box="[222,653,342,366]" collectionCode="NHMUK" pageId="15" pageNumber="16" specimenCode="NHMUK 1967727" specimenCount="1" typeStatus="holotype">
<typeStatus id="360657A3244FFFE4FF490113FEBF8110" box="[222,332,342,366]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
:
<specimenCode id="B91B417A244FFFE4FEC30113FDE68110" box="[340,533,342,366]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="15" pageNumber="16" type="Museum">NHMUK 1967727</specimenCode>
;
<quantity id="2E4544E4244FFFE4FDB40113FD7A8110" box="[547,649,342,366]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="15" pageNumber="16" unit="mm" value="14.8">14.8 mm</quantity>
.
</materialsCitation>
</paragraph>
<paragraph id="E902E901244FFFE4FF4B0139FCBB81EA" blockId="15.[182,1073,156,740]" box="[220,840,380,404]" pageId="15" pageNumber="16">
Type locality:
<collectingCountry id="91AAA991244FFFE4FEEE0139FE0881EA" box="[377,507,380,404]" name="Philippines" pageId="15" pageNumber="16">Philippines</collectingCountry>
,
<collectingRegion id="2B7927E3244FFFE4FD930139FD9981EA" box="[516,618,380,404]" country="Philippines" name="Masbate" pageId="15" pageNumber="16">Masbate</collectingRegion>
, found on the sand.
</paragraph>
</subSubSection>
<subSubSection id="A1A7BA8A244FFFE4FF4901E4FF0C8257" pageId="15" pageNumber="16" type="distribution">
<paragraph id="E902E901244FFFE4FF4901E4FF0C8257" blockId="15.[182,1073,156,740]" pageId="15" pageNumber="16">
Distribution: Western Pacific from
<collectingCountry id="91AAA991244FFFE4FDCB01E4FD6981C7" box="[604,666,417,441]" name="Japan" pageId="15" pageNumber="16">Japan</collectingCountry>
(
<bibRefCitation id="8D2C94F0244FFFE4FD3001E4FCB181C7" author="Okutani, T." box="[679,834,417,441]" pageId="15" pageNumber="16" refId="ref12357" refString="Okutani, T. (2000): Marine Mollusks in Japan. - 1173 pp., Tokyo." type="book" year="2000">Okutani 2000</bibRefCitation>
) via the
<typeStatus id="360657A3244FFFE4FC3501E4FC2781C7" box="[930,980,417,441]" pageId="15" pageNumber="16">type</typeStatus>
locality
<collectingCountry id="91AAA991244FFFE4FF2F0183FEC081A0" box="[184,307,454,478]" name="Philippines" pageId="15" pageNumber="16">Philippines</collectingCountry>
and
<collectingCountry id="91AAA991244FFFE4FEFA0183FE3C81A0" box="[365,463,454,478]" name="Malaysia" pageId="15" pageNumber="16">Malaysia</collectingCountry>
(
<bibRefCitation id="8D2C94F0244FFFE4FE490183FD3981A0" author="Coltro, J. &amp; Coltro, M." box="[478,714,454,478]" pageId="15" pageNumber="16" refId="ref12068" refString="Coltro, J. &amp; Coltro, M. (2013): Accessed through: Search for V. corallinum at: http: // www. femorale. com. br / search / index. asp on 2013 - 09 - 13." type="url" year="2013">Coltro &amp; Coltro 2013</bibRefCitation>
) to
<collectingCountry id="91AAA991244FFFE4FD6E0183FC3C81A0" box="[761,975,454,478]" name="Papua New Guinea" pageId="15" pageNumber="16">Papua New Guinea</collectingCountry>
(
<bibRefCitation id="8D2C94F0244FFFE4FC4A0183FEC3827A" author="Pechar, P. &amp; Prior, C. &amp; Parkinson, B." pageId="15" pageNumber="16" refId="ref12377" refString="Pechar, P., Prior, C. &amp; Parkinson, B. (1980): Mitre shells from the Pacific and Indian Oceans. - 126 pp., 56 plates, Bathurst N. S. W." type="book" year="1980">Pechar &amp; al. 1980</bibRefCitation>
),
<collectingRegion id="2B7927E3244FFFE4FEDE01A9FE26827A" box="[329,469,492,516]" country="Australia" name="Queensland" pageId="15" pageNumber="16">Queensland</collectingRegion>
,
<collectingCountry id="91AAA991244FFFE4FE7701A9FDB5827A" box="[480,582,492,516]" name="Australia" pageId="15" pageNumber="16">Australia</collectingCountry>
(this paper) and
<collectingCountry id="91AAA991244FFFE4FC8701A9FC9D827A" box="[784,878,492,516]" name="Vanuatu" pageId="15" pageNumber="16">Vanuatu</collectingCountry>
(
<bibRefCitation id="8D2C94F0244FFFE4FCEB01A9FF1C8257" author="Coltro, J. &amp; Coltro, M." pageId="15" pageNumber="16" refId="ref12068" refString="Coltro, J. &amp; Coltro, M. (2013): Accessed through: Search for V. corallinum at: http: // www. femorale. com. br / search / index. asp on 2013 - 09 - 13." type="url" year="2013">Coltro &amp; Coltro 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A1A7BA8A244FFFE4FF490273FD12829A" pageId="15" pageNumber="16" type="discussion">
<paragraph id="E902E901244FFFE4FF490273FD12829A" blockId="15.[182,1073,156,740]" pageId="15" pageNumber="16">
Remarks: This species was put into synonymy with
<taxonomicName id="2EBD9282244FFFE4FCA60273FE3E820A" authority="(ANTON, 1838)" baseAuthorityName="ANTON" baseAuthorityYear="1838" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="semicostatum" subGenus="Pusia">
<emphasis id="DBC93513244FFFE4FCA60273FC618230" box="[817,914,566,590]" italics="true" pageId="15" pageNumber="16">Vexillum</emphasis>
(
<emphasis id="DBC93513244FFFE4FC340273FC128230" box="[931,993,566,590]" italics="true" pageId="15" pageNumber="16">Pusia</emphasis>
)
<emphasis id="DBC93513244FFFE4FC640273FED2820A" italics="true" pageId="15" pageNumber="16">semicostatum</emphasis>
(ANTON, 1838)
</taxonomicName>
by different authors (
<bibRefCitation id="8D2C94F0244FFFE4FD290219FC7A820A" author="Pechar, P. &amp; Prior, C. &amp; Parkinson, B." box="[702,905,604,628]" pageId="15" pageNumber="16" refId="ref12377" refString="Pechar, P., Prior, C. &amp; Parkinson, B. (1980): Mitre shells from the Pacific and Indian Oceans. - 126 pp., 56 plates, Bathurst N. S. W." type="book" year="1980">Pechar &amp; al. 1980</bibRefCitation>
– plate 54, fig. 4;
<bibRefCitation id="8D2C94F0244FFFE4FF4102C4FDD982E7" author="Springsteen, F. J. &amp; Leobrera, F. M." box="[214,554,641,665]" pageId="15" pageNumber="16" refId="ref12761" refString="Springsteen, F. J. &amp; Leobrera, F. M. (1986): Shells of the Philippines. - 377 pp., 100 plates, Manila." type="book" year="1986">Springsteen &amp; Leobrera 1986</bibRefCitation>
;
<bibRefCitation id="8D2C94F0244FFFE4FDAB02C4FD2B82E7" author="Okutani, T." box="[572,728,641,665]" pageId="15" pageNumber="16" refId="ref12357" refString="Okutani, T. (2000): Marine Mollusks in Japan. - 1173 pp., Tokyo." type="book" year="2000">Okutani 2000</bibRefCitation>
; Turner 2001), but is easy to distinguish from the latter species by the red-orange colour and the regular white dots only in the anterior part of the whorls.
</paragraph>
</subSubSection>
</treatment>
</document>
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