141 lines
18 KiB
XML
141 lines
18 KiB
XML
<document id="BF52E9F02EBB747BED765B030ABABAB7" ID-DOI="10.11646/zootaxa.4020.1.7" ID-GBIF-Dataset="82b2f539-5d25-42d6-95c5-e8b16a353e12" ID-ISSN="1175-5326" ID-Zenodo-Dep="237970" ID-ZooBank="BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461269068152" checkinUser="plazi" docAuthor="Bickel, Daniel J." docDate="2015" docId="172D87FDFFBFFF8DFF3DA27D05F8FC02" docLanguage="en" docName="zt04020p182.pdf" docOrigin="Zootaxa 4020 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus tenorio Bickel, sp. nov." docType="treatment" docVersion="8" lastPageNumber="172" masterDocId="EB14FF85FFBDFF8EFFAAA70A0459FFCC" masterDocTitle="The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus" masterLastPageNumber="182" masterPageNumber="169" pageNumber="171" updateTime="1698611248703" updateUser="plazi">
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<mods:mods id="828CF6DC45A4A04C34A595A029A22052" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FA9FCC877AB71A1714690CF094299669">
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<mods:title id="E4682401CC49352C8679D8CD5B0479EE">The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus</mods:title>
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<mods:name id="DAA5ECE513EBBB97B2447F3323328B2E" type="personal">
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<mods:namePart id="49FBDD4CD8CD54CD1B48C0D8EBABB6B6">Bickel, Daniel J.</mods:namePart>
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<mods:typeOfResource id="0EB339FEE2D342DB8C5018C6E649915D">text</mods:typeOfResource>
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<mods:title id="439CEE10E44E464CECA24ADBC89149B7">Zootaxa</mods:title>
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<mods:part id="FCB6EA41D2C8A2D369B179E7A6FF944F">
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<mods:date id="46E9E70848F46F712D866B26A6508C69">2015</mods:date>
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<mods:detail id="CC4A1218E0C3E56EC674F168DF7354CF" type="volume">
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<mods:number id="A2C7902026234089978B3903E0FF0493">4020</mods:number>
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<mods:detail id="B1EC71920343EBC25579CC68DE65BE95" type="issue">
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<mods:number id="B74F2005A4266338968491CC59344328">1</mods:number>
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<mods:end id="82FE46D59E5BEF965FBDDA03BBB05560">182</mods:end>
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<mods:classification id="ACEE4861BFEC7E66E79F532D60DFDD92">journal article</mods:classification>
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<mods:identifier id="E7798136F88C34CABEBD2C2CDAFF8238" type="DOI">10.11646/zootaxa.4020.1.7</mods:identifier>
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<mods:identifier id="FE1A6D6EAF01F4E29FD519692347606F" type="GBIF-Dataset">82b2f539-5d25-42d6-95c5-e8b16a353e12</mods:identifier>
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<mods:identifier id="E46E4510C03E6CDC52A11D97C958608C" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="70A3C1394CAAA572BBBC46E3664CD55F" type="Zenodo-Dep">237970</mods:identifier>
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<mods:identifier id="A18ADC52F3353A2C82365CBE1EF5D481" type="ZooBank">BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE</mods:identifier>
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</mods:mods>
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<treatment id="172D87FDFFBFFF8DFF3DA27D05F8FC02" ID-DOI="http://doi.org/10.5281/zenodo.6121799" ID-GBIF-Taxon="119638515" ID-Zenodo-Dep="6121799" LSID="urn:lsid:plazi:treatment:172D87FDFFBFFF8DFF3DA27D05F8FC02" httpUri="http://treatment.plazi.org/id/172D87FDFFBFFF8DFF3DA27D05F8FC02" lastPageId="3" lastPageNumber="172" pageId="2" pageNumber="171">
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<subSubSection id="D79E6560FFBFFF8CFF3DA27D054EFA7F" pageId="2" pageNumber="171" type="nomenclature">
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<paragraph id="9F3B36EBFFBFFF8CFF3DA27D0640FA5D" blockId="2.[151,537,1399,1459]" box="[151,537,1399,1425]" pageId="2" pageNumber="171">
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<heading id="C4738187FFBFFF8CFF3DA27D0640FA5D" bold="true" box="[151,537,1399,1425]" fontSize="11" level="1" pageId="2" pageNumber="171" reason="1">
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<emphasis id="ADF0EAF9FFBFFF8CFF3DA27D0640FA5D" bold="true" box="[151,537,1399,1425]" pageId="2" pageNumber="171">
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<taxonomicName id="58844D68FFBFFF8CFF3DA27D05EAFA5D" ID-CoL="9HZLJ" authority="Bickel" authorityName="Bickel" box="[151,435,1399,1425]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="tenorio" status="sp. nov.">
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<emphasis id="ADF0EAF9FFBFFF8CFF3DA27D0507FA5D" bold="true" box="[151,350,1399,1425]" italics="true" pageId="2" pageNumber="171">Systenus tenorio</emphasis>
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Bickel
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFBFFF8CFE11A27D0640FA5D" box="[443,537,1399,1425]" pageId="2" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="9F3B36EBFFBFFF8CFF3DA290054EFA7F" blockId="2.[151,537,1399,1459]" box="[151,279,1434,1459]" pageId="2" pageNumber="171">
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(
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<figureCitation id="07BF2A6EFFBFFF8CFF35A29004BBFA7F" box="[159,226,1434,1459]" captionStart="FIGURE 1" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[238,1352,195,1838]" captionTargetId="figure@4.[234,1352,195,1839]" captionTargetPageId="4" captionText="FIGURE 1. Systenus tenorio sp. nov. (holotype): a. male habitus, left lateral (terminalia removed); b. hypopygium, left lateral. S. flavifemoratus sp. nov. (holotype): c. male habitus, right lateral; d. hypopygium, left lateral. S. maculipennis sp. nov.: e. male habitus, left lateral (terminalia removed); f. hypopygium, left lateral. Abbreviations: cer, cercus; epan, epandrium; hyp, hypandrium; ph, phallus; st, sternite; sur, surstylus; tg, tergite; 6, 7, 8, abdominal segments." httpUri="https://zenodo.org/record/237971/files/figure.png" pageId="2" pageNumber="171">Figs 1</figureCitation>
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a, b)
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFBFFF8CFF3DA2D60636F9D5" pageId="2" pageNumber="171" type="materials_examined">
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<paragraph id="9F3B36EBFFBFFF8CFF3DA2D60636F9D5" blockId="2.[151,1437,1500,2030]" pageId="2" pageNumber="171">
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<emphasis id="ADF0EAF9FFBFFF8CFF3DA2D60519FA39" bold="true" box="[151,320,1500,1525]" pageId="2" pageNumber="171">
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<typeStatus id="403F8849FFBFFF8CFF3DA2D6048BFA39" box="[151,210,1500,1525]" pageId="2" pageNumber="171">Type</typeStatus>
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material
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</emphasis>
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.
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<emphasis id="ADF0EAF9FFBFFF8CFEFAA2D605B6FA39" bold="true" box="[336,495,1500,1525]" pageId="2" pageNumber="171">
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<typeStatus id="403F8849FFBFFF8CFEFAA2D605B6FA39" box="[336,495,1500,1525]" pageId="2" pageNumber="171" type="holotype">HOLOTYPE</typeStatus>
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</emphasis>
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♂,
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<collectingCountry id="E793767BFFBFFF8CFDB7A2D7069BFA39" box="[541,706,1501,1525]" name="Costa Rica" pageId="2" pageNumber="171">COSTA RICA</collectingCountry>
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:
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<emphasis id="ADF0EAF9FFBFFF8CFD7FA2D60738FA39" bold="true" box="[725,865,1500,1525]" pageId="2" pageNumber="171">Guanacaste</emphasis>
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:
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<quantity id="587C9B0EFFBFFF8CFCD8A2D707F4FA39" box="[882,941,1500,1525]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="171" unit="km" value="3.0">3 km</quantity>
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SE Rio Naranjo,
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<date id="EB3A102BFFBFFF8CFBD6A2D70155FA39" box="[1148,1292,1501,1525]" pageId="2" pageNumber="171" value="1992-05-03" valueMax="1992-05-08" valueMin="1992-05-03">3—8.V.1992</date>
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, F.D. Parker (
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<collectionCode id="F995AE2EFFBFFF8CFF34A10B04AAF9D5" LSID="urn:lsid:biocol.org:col:34802" box="[158,243,1537,1561]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="2" pageNumber="171">LACM</collectionCode>
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, ex.
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<collectionCode id="F995AE2EFFBFFF8CFE8DA10B0520F9D5" box="[295,377,1537,1561]" pageId="2" pageNumber="171">EMUS</collectionCode>
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;
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<collectionCode id="F995AE2EFFBFFF8CFE2EA10B058CF9D5" LSID="urn:lsid:biocol.org:col:34802" box="[388,469,1537,1561]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="2" pageNumber="171">LACM</collectionCode>
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Ent 329994).
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFBFFF8DFF6DA12F00C7FD3A" lastPageId="3" lastPageNumber="172" pageId="2" pageNumber="171" type="description">
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<paragraph id="9F3B36EBFFBFFF8DFF6DA12F00C7FD3A" blockId="2.[151,1437,1500,2030]" lastBlockId="3.[151,1437,151,974]" lastPageId="3" lastPageNumber="172" pageId="2" pageNumber="171">
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<emphasis id="ADF0EAF9FFBFFF8CFF6DA12F0508F9F2" bold="true" box="[199,337,1573,1598]" pageId="2" pageNumber="171">Description</emphasis>
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.
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<emphasis id="ADF0EAF9FFBFFF8CFECAA12F05C5F9F2" bold="true" box="[352,412,1573,1598]" pageId="2" pageNumber="171">Male</emphasis>
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: body length:
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<quantity id="587C9B0EFFBFFF8CFDEAA12F06C0F9F1" box="[576,665,1573,1598]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="171" unit="mm" value="2.5">2.5 mm</quantity>
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; wing 2.4 x
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<quantity id="587C9B0EFFBFFF8CFC8EA12F0722F9F1" box="[804,891,1573,1597]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="171" unit="mm" value="0.7">0.7 mm</quantity>
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(
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<figureCitation id="07BF2A6EFFBFFF8CFC20A12F0793F9F1" box="[906,970,1573,1598]" captionStart="FIGURE 1" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[238,1352,195,1838]" captionTargetId="figure@4.[234,1352,195,1839]" captionTargetPageId="4" captionText="FIGURE 1. Systenus tenorio sp. nov. (holotype): a. male habitus, left lateral (terminalia removed); b. hypopygium, left lateral. S. flavifemoratus sp. nov. (holotype): c. male habitus, right lateral; d. hypopygium, left lateral. S. maculipennis sp. nov.: e. male habitus, left lateral (terminalia removed); f. hypopygium, left lateral. Abbreviations: cer, cercus; epan, epandrium; hyp, hypandrium; ph, phallus; st, sternite; sur, surstylus; tg, tergite; 6, 7, 8, abdominal segments." httpUri="https://zenodo.org/record/237971/files/figure.png" pageId="2" pageNumber="171">Fig. 1</figureCitation>
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a).
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<emphasis id="ADF0EAF9FFBFFF8CFC5BA12F0074F9F2" box="[1009,1069,1573,1598]" italics="true" pageId="2" pageNumber="171">Head</emphasis>
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: head almost circular in anterior view; postcranium dorsally concave; single row of postorbitals, black dorsally, and white ventrally, with dorsalmost pair behind ocellar triangle strong; frons, face, dark metallic green with thick brownish pruinosity; strong vertical and ocellar setae present; eyes distinctly separated across face; ventral eye facets slightly larger than dorsal facets; palpus brown with short black setae and strong apical seta; proboscis dark brown; antenna dark brown; scape and pedicel short; postpedicel triangular and tapering gradually, more than twice as long as basal width, covered with short pubescence, and with short subapical arista.
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<emphasis id="ADF0EAF9FFBFFF8CFC6FA1F7004FF8DA" box="[965,1046,1789,1814]" italics="true" pageId="2" pageNumber="171">Thorax</emphasis>
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: dorsum dark metallic green with bronze reflections over ac band, and covered with grey pruinosity; pleura including metepimeron green with dense grey pruinosity; posterior third of mesonotum distinctly flattened; thoracic setae black except where noted; some 12 pairs of ac with posterior pair bordering mesonotal depression larger and offset laterally; 6 strong dc, decreasing in size anteriorly.
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<emphasis id="ADF0EAF9FFBFFF8CFEF5A08405CDF869" box="[351,404,1934,1957]" italics="true" pageId="2" pageNumber="171">Legs</emphasis>
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: CI brown basally but pale yellow on distal half; CII and CIII mostly dark brown but distal third pale yellow; all trochanters pale yellow; FI and FII mostly dark brown but with distal quarter yellow; FIII basally yellow and distal two-thirds to quarter brown to apex; TI and It tarsus ivory-colored; TII and IIt yellow, but distal tarsomeres infuscated; TIII brown but yellowish basally; IIIt mostly brown; CI and CII with white anterior setae and shorter vestiture, CIII with strong brown lateral seta at ½; major leg setae black; I: 3.1; 3.0; 1.3/ 0.7/ 0.6/ 0.3/ 0.2; leg I bare of major setation; TI and tarsus I covered with short white (not black) vestiture (MSSC); II: 3.4; 3.8; 2.0/ 1.1/ 0.8/ 0.5/ 0.4; TII with strong ad seta and weaker pd seta at ¼, ad seta at 2/3, with apical ring of ad, av and pv seta; III: 3.8; 4.7; 0.9/ 1.5/ 0.9/ 0.5/ 0.4; TIII with 5 evenly spaced black dorsal setae from 1/5 to 4/5, and with apical ring of dorsal, ad and av seta.
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<emphasis id="ADF0EAF9FFBEFF8DFDD0A64706EBFEA8" box="[634,690,333,356]" italics="true" pageId="3" pageNumber="172">Wing</emphasis>
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: hyaline; R2+3 and R4+5 diverging, R2+3 joining costa near 7/8, and R4+5 joining subapically; vein M diverging from R4+5 with two veins very slightly bowed half way between dm-cu crossvein and apex, and with M joining costa just behind wing apex; CuAx ratio: 0.5; lower calypter pale yellow with yellowish marginal setae; halter yellow.
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<emphasis id="ADF0EAF9FFBEFF8DFD3EA6B706A7FE1A" box="[660,766,445,470]" italics="true" pageId="3" pageNumber="172">Abdomen</emphasis>
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: metallic bronze-green and with dusting of grey pruinosity; posterior margin of tergite 1 with posterior row of long black setae, tergites otherwise covered with short black vestiture; segment 7 forming elongate peduncle; hypopygium (
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<figureCitation id="07BF2A6EFFBEFF8DFCDCA50F07E3FDD2" box="[886,954,517,542]" captionStart="FIGURE 1" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[238,1352,195,1838]" captionTargetId="figure@4.[234,1352,195,1839]" captionTargetPageId="4" captionText="FIGURE 1. Systenus tenorio sp. nov. (holotype): a. male habitus, left lateral (terminalia removed); b. hypopygium, left lateral. S. flavifemoratus sp. nov. (holotype): c. male habitus, right lateral; d. hypopygium, left lateral. S. maculipennis sp. nov.: e. male habitus, left lateral (terminalia removed); f. hypopygium, left lateral. Abbreviations: cer, cercus; epan, epandrium; hyp, hypandrium; ph, phallus; st, sternite; sur, surstylus; tg, tergite; 6, 7, 8, abdominal segments." httpUri="https://zenodo.org/record/237971/files/figure.png" pageId="3" pageNumber="172">Fig. 1</figureCitation>
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b) dark brown with yellow surstylus and cercus; hypandrium as large hood over epandrium with basally projecting extensions and bearing lateral row of 3 setae; phallus hooked and enclosed by hypandrium; short epandrial lobes present, with strong apical and weaker seta arising midlength; surstylus not solidly fused to epandrium, but with line of weakness at base; in lateral view, surstylus subrectangular and distally forked with two elongate digitiform projections separated by broad U-shaped excavation, and each projection bearing apical setae, and with curved digitiform medio-dorsal digitiform projection with short apical seta; cercus elongate and digitiform, with yellow setae.
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<emphasis id="ADF0EAF9FFBEFF8DFC62A5D70046FD3A" bold="true" box="[968,1055,733,758]" pageId="3" pageNumber="172">Female</emphasis>
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: unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFBEFF8DFF6DA40B061AFC4A" pageId="3" pageNumber="172" type="discussion">
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<paragraph id="9F3B36EBFFBEFF8DFF6DA40B061AFC4A" blockId="3.[151,1437,151,974]" pageId="3" pageNumber="172">
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<emphasis id="ADF0EAF9FFBEFF8DFF6DA40B056DFCD6" bold="true" box="[199,308,769,794]" pageId="3" pageNumber="172">Remarks</emphasis>
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.
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<taxonomicName id="58844D68FFBEFF8DFEE9A40805A5FCD5" box="[323,508,770,793]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="172" phylum="Arthropoda" rank="species" species="tenorio">
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<emphasis id="ADF0EAF9FFBEFF8DFEE9A40805A5FCD5" box="[323,508,770,793]" italics="true" pageId="3" pageNumber="172">Systenus tenorio</emphasis>
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</taxonomicName>
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is known from a single male taken at the Rio Naranjo site,
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<collectingCountry id="E793767BFFBEFF8DFB09A4080179FCD6" box="[1187,1312,769,794]" name="Costa Rica" pageId="3" pageNumber="172">Costa Rica</collectingCountry>
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, in May. It has the diagnostic male character of black tapering postpedicel (similar to
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<taxonomicName id="58844D68FFBEFF8DFC5EA42C00D9FCF2" box="[1012,1152,805,830]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="172" phylum="Arthropoda" rank="species" species="eboritibia">
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<emphasis id="ADF0EAF9FFBEFF8DFC5EA42C00D9FCF2" box="[1012,1152,805,830]" italics="true" pageId="3" pageNumber="172">S. eboritibia</emphasis>
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</taxonomicName>
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), femora I and II brown basally but yellow distally, whereas femur III is yellow in the basal third and brown distally. Tibia and basitarsus I are ivory colored with white vestiture.
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFBEFF8DFF6DA49B05F8FC02" pageId="3" pageNumber="172" type="etymology">
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<paragraph id="9F3B36EBFFBEFF8DFF6DA49B05F8FC02" blockId="3.[151,1437,151,974]" pageId="3" pageNumber="172">
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<emphasis id="ADF0EAF9FFBEFF8DFF6DA49B051FFC66" bold="true" box="[199,326,913,938]" pageId="3" pageNumber="172">Etymology</emphasis>
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. The specific epithet “
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<taxonomicName id="58844D68FFBEFF8DFDE9A49806CDFC65" box="[579,660,914,937]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="172" phylum="Arthropoda" rank="species" species="tenorio">
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<emphasis id="ADF0EAF9FFBEFF8DFDE9A49806CDFC65" box="[579,660,914,937]" italics="true" pageId="3" pageNumber="172">tenorio</emphasis>
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</taxonomicName>
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” is the name of the nearby Vulcan
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<taxonomicName id="58844D68FFBEFF8DFBB6A49B002FFC66" box="[1052,1142,913,938]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="179" phylum="Arthropoda" rank="species" species="tenorio">Tenorio</taxonomicName>
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, and should be considered as a noun in apposition.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |