266 lines
29 KiB
XML
266 lines
29 KiB
XML
<document id="C24EBDDB46FE376A814B9B5FDCEAB696" ID-DOI="10.11646/zootaxa.4020.1.7" ID-GBIF-Dataset="82b2f539-5d25-42d6-95c5-e8b16a353e12" ID-ISSN="1175-5326" ID-Zenodo-Dep="237970" ID-ZooBank="BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461269068152" checkinUser="plazi" docAuthor="Bickel, Daniel J." docDate="2015" docId="172D87FDFFBCFF8CFF3DA5BD004BFE1C" docLanguage="en" docName="zt04020p182.pdf" docOrigin="Zootaxa 4020 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus Loew" docType="treatment" docVersion="8" lastPageNumber="171" masterDocId="EB14FF85FFBDFF8EFFAAA70A0459FFCC" masterDocTitle="The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus" masterLastPageNumber="182" masterPageNumber="169" pageNumber="170" updateTime="1698611248703" updateUser="plazi">
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<mods:title id="D366576D1228BD718A38A2097B8068CB">The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus</mods:title>
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<mods:namePart id="CAC861A93EE826B5BCE1DD02AE1FAFFB">Bickel, Daniel J.</mods:namePart>
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<mods:typeOfResource id="8D5C08D2DED174BE936F30F3EEFE9741">text</mods:typeOfResource>
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<mods:title id="11B03712A9FF6B8F7926C93144926280">Zootaxa</mods:title>
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<mods:date id="DAFBC824D1DE7B8F21B6A14EFCB5FC29">2015</mods:date>
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<mods:number id="D3A7A782D5CDDCF74C5A2677A714C97F">4020</mods:number>
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<mods:number id="EB46A1341DC6F9ED85FF3D9A4C891A61">1</mods:number>
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<mods:identifier id="5233AB5CB8ECFEE0D3564C526FFA57FB" type="DOI">10.11646/zootaxa.4020.1.7</mods:identifier>
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<treatment id="172D87FDFFBCFF8CFF3DA5BD004BFE1C" ID-DOI="http://doi.org/10.5281/zenodo.6121793" ID-GBIF-Taxon="119638514" ID-Zenodo-Dep="6121793" LSID="urn:lsid:plazi:treatment:172D87FDFFBCFF8CFF3DA5BD004BFE1C" httpUri="http://treatment.plazi.org/id/172D87FDFFBCFF8CFF3DA5BD004BFE1C" lastPageId="2" lastPageNumber="171" pageId="1" pageNumber="170">
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<subSubSection id="D79E6560FFBCFF8FFF3DA5BD05FAFD1D" box="[151,419,694,721]" pageId="1" pageNumber="170" type="nomenclature">
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<paragraph id="9F3B36EBFFBCFF8FFF3DA5BD05FAFD1D" blockId="1.[151,419,694,721]" box="[151,419,694,721]" pageId="1" pageNumber="170">
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<heading id="C4738187FFBCFF8FFF3DA5BD05FAFD1D" bold="true" box="[151,419,694,721]" fontSize="11" level="1" pageId="1" pageNumber="170" reason="1">
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<emphasis id="ADF0EAF9FFBCFF8FFF3DA5BD05FAFD1D" bold="true" box="[151,419,694,721]" pageId="1" pageNumber="170">
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Genus
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<taxonomicName id="58844D68FFBCFF8FFF44A5BC05FAFD1D" ID-CoL="7RNG" ID-ENA="1000637" authority="Loew" authorityName="Loew" box="[238,419,694,721]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus">
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<emphasis id="ADF0EAF9FFBCFF8FFF44A5BC0501FD1C" bold="true" box="[238,344,694,720]" italics="true" pageId="1" pageNumber="170">Systenus</emphasis>
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Loew
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFBCFF8FFF3DA5F40534FCFE" pageId="1" pageNumber="170" type="reference_group">
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<paragraph id="9F3B36EBFFBCFF8FFF3DA5F40534FCFE" blockId="1.[151,1436,765,818]" pageId="1" pageNumber="170">
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<treatmentCitationGroup id="BF9411C5FFBCFF8FFF3DA5F40534FCFE" pageId="1" pageNumber="170">
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<taxonomicName id="58844D68FFBCFF8FFF3DA5F405CEFCDF" ID-CoL="7RNG" authority="Loew, 1857: 34" authorityName="Loew" authorityPageNumber="34" authorityYear="1857" box="[151,407,765,787]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus">
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<emphasis id="ADF0EAF9FFBCFF8FFF3DA5F404B7FCDF" box="[151,238,766,787]" italics="true" pageId="1" pageNumber="170">Systenus</emphasis>
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<treatmentCitation id="1E2510FAFFBCFF8FFF5FA5F405CEFCDF" author="Loew" box="[245,407,765,787]" page="34" pageId="1" pageNumber="170" year="1857">
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<bibRefCitation id="FB154B1AFFBCFF8FFF5FA5F40528FCDF" author="Loew" box="[245,369,765,787]" pageId="1" pageNumber="170" refString="Loew, H. (1857) Neue Beitrage zur Kenntniss der Dipteren. Funfter Beitrage. Programme der Koniglichen Realschule zu Meseritz, 1857, 1 - 56." type="journal article" year="1857">Loew, 1857</bibRefCitation>
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: 34
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</treatmentCitation>
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</taxonomicName>
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.
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<typeStatus id="403F8849FFBCFF8FFE08A5F4058DFCDF" box="[418,468,766,787]" pageId="1" pageNumber="170">Type</typeStatus>
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species,
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<taxonomicName id="58844D68FFBCFF8FFD98A5F707E1FCDF" ID-CoL="BB2MC" authority="Loew, 1857" authorityName="Loew" authorityYear="1857" box="[562,952,765,787]" class="Insecta" family="Dolichopodidae" genus="Rhaphium" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="adpropinquans">
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<emphasis id="ADF0EAF9FFBCFF8FFD98A5F70760FCDE" box="[562,825,765,786]" italics="true" pageId="1" pageNumber="170">Rhaphium adpropinquans</emphasis>
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<bibRefCitation id="FB154B1AFFBCFF8FFCEAA5F407E1FCDF" author="Loew" box="[832,952,765,787]" pageId="1" pageNumber="170" refString="Loew, H. (1857) Neue Beitrage zur Kenntniss der Dipteren. Funfter Beitrage. Programme der Koniglichen Realschule zu Meseritz, 1857, 1 - 56." type="journal article" year="1857">Loew, 1857</bibRefCitation>
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</taxonomicName>
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(=
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<taxonomicName id="58844D68FFBCFF8FFC77A5F70178FCDF" ID-CoL="8DML5" authority="Roser, 1840" authorityName="Roser" authorityYear="1840" box="[989,1313,765,787]" class="Insecta" family="Dolichopodidae" genus="Rhaphium" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="pallipes">
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<emphasis id="ADF0EAF9FFBCFF8FFC77A5F700C5FCDE" box="[989,1180,765,786]" italics="true" pageId="1" pageNumber="170">Rhaphium pallipes</emphasis>
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Roser, 1840
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</taxonomicName>
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), des. Foote
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<emphasis id="ADF0EAF9FFBCFF8FFF6DA41704AAFCFD" box="[199,243,796,818]" italics="true" pageId="1" pageNumber="170">et al</emphasis>
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. 1966: 517.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFBCFF8FFF3DA4540751F9C9" pageId="1" pageNumber="170" type="diagnosis">
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<paragraph id="9F3B36EBFFBCFF8FFF3DA4540751F9C9" blockId="1.[151,1437,862,2009]" pageId="1" pageNumber="170">
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<emphasis id="ADF0EAF9FFBCFF8FFF3DA4540553FCBB" bold="true" box="[151,266,862,887]" pageId="1" pageNumber="170">Diagnosis</emphasis>
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.
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<emphasis id="ADF0EAF9FFBCFF8FFEB6A4540500FCBB" box="[284,345,862,887]" italics="true" pageId="1" pageNumber="170">Head</emphasis>
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: head almost circular in anterior view; eyes with tiny setulae (ommatrichia) between facets; frontoclypeal suture complete; proboscis projecting anteriorly, keel-like, each labellar lobe with 6 unsclerotized pseudotracheae; single row of postorbitals present; scape without dorsal setae; postpedicel with marked sexual dimorphism: in male lancet shaped, elongate, usually 2-3 times as long as basal width, bearing short apical arista; in female subovate, about as long as wide, bearing long apical arista; postcranium dorsally concave.
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<emphasis id="ADF0EAF9FFBCFF8FFAA8A4E4010AFBCB" box="[1282,1363,1006,1031]" italics="true" pageId="1" pageNumber="170">Thorax</emphasis>
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: often with bronze medial stripe over ac band; posterior mesonotum distinctly flattened; ac biserial, posteriormost pair larger and laterally offset; 6 strong pairs dc; anterior slope of mesonotum laterad of dc rows with field of short setulae; 1 postalar, 2 postsutural and 2 presutural supraalars, 2 notopleurals, 1 humeral, and 1 posthumeral present per side;. 2 pairs scutellar setae per side, lateral setae less than less half length of medians; strong ventrally projecting white propleural seta present above base of CI.
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<emphasis id="ADF0EAF9FFBCFF8FFC9EA3AE072AFB77" box="[820,883,1188,1211]" italics="true" pageId="1" pageNumber="170">Legs:</emphasis>
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coxa III with strong lateral seta; femora lacking anterior preapical setae; femora mostly bare of major setae; tibia I mostly bare; tibiae II and III variously with dorsal, and ventral, usually with ad-pd setal pair near ¼.
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<emphasis id="ADF0EAF9FFBCFF8FFC9FA3E60734FACF" box="[821,877,1260,1283]" italics="true" pageId="1" pageNumber="170">Wing</emphasis>
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: R4+5 and M subparallel but bowed subapically; flexion (
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<emphasis id="ADF0EAF9FFBCFF8FFF5DA21B05DEFAE6" box="[247,391,1297,1322]" italics="true" pageId="1" pageNumber="170">bosse alaire)</emphasis>
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present in distal sector of M; A1 present.
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<emphasis id="ADF0EAF9FFBCFF8FFCF0A21B0069FAE6" box="[858,1072,1297,1322]" italics="true" pageId="1" pageNumber="170">Male postabdomen</emphasis>
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: segment 7 forming an elongate peduncle, with tergite and sternite 7 incompletely fused, separated by partially sclerotized pleural membrane; hypopygial peduncle, formed of tergite and sternite 7 often held withdrawn between sternites 5 and 6 at rest, and sternite 7 is glabrous while tergite 7 is setose (
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<figureCitation id="07BF2A6EFFBCFF8FFD6BA28A075DFA55" box="[705,772,1408,1433]" captionStart="FIGURE 1" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[238,1352,195,1838]" captionTargetId="figure@4.[234,1352,195,1839]" captionTargetPageId="4" captionText="FIGURE 1. Systenus tenorio sp. nov. (holotype): a. male habitus, left lateral (terminalia removed); b. hypopygium, left lateral. S. flavifemoratus sp. nov. (holotype): c. male habitus, right lateral; d. hypopygium, left lateral. S. maculipennis sp. nov.: e. male habitus, left lateral (terminalia removed); f. hypopygium, left lateral. Abbreviations: cer, cercus; epan, epandrium; hyp, hypandrium; ph, phallus; st, sternite; sur, surstylus; tg, tergite; 6, 7, 8, abdominal segments." httpUri="https://zenodo.org/record/237971/files/figure.png" pageId="1" pageNumber="170">Fig. 1</figureCitation>
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b); hypopygial foramen left lateral; hypandrium fused to epandrium, immovable; two seta-bearing epandrial lobes separate but short, each with only one strong seta present; surstylus usually bifurcate; cercus various.
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<emphasis id="ADF0EAF9FFBCFF8FFD03A2C20733FA2C" box="[681,874,1480,1505]" italics="true" pageId="1" pageNumber="170">Female oviscapt</emphasis>
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: fused tergites 9+10 divided medially into 2 hemitergites, each bearing 4 spine-like acanthophorites.
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFBCFF8CFF6DA11A004BFE1C" lastPageId="2" lastPageNumber="171" pageId="1" pageNumber="170" type="discussion">
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<paragraph id="9F3B36EBFFBCFF8FFF6DA11A0750F975" blockId="1.[151,1437,862,2009]" pageId="1" pageNumber="170">
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<emphasis id="ADF0EAF9FFBCFF8FFF6DA11A056DF9E5" bold="true" box="[199,308,1552,1577]" pageId="1" pageNumber="170">Remarks</emphasis>
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. Including the nine new species described here, 36 species are currently placed in the genus
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<taxonomicName id="58844D68FFBCFF8FFF3DA13F05CBF981" authority="Afrotropical" authorityName="Afrotropical" box="[151,402,1588,1613]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus">
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<emphasis id="ADF0EAF9FFBCFF8FFF3DA13F04AEF980" box="[151,247,1589,1612]" italics="true" pageId="1" pageNumber="170">Systenus</emphasis>
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: Afrotropical
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</taxonomicName>
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(
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<specimenCount id="8982FD62FFBCFF8FFE0EA13F0583F981" box="[420,474,1589,1613]" pageId="1" pageNumber="170" type="generic">1 sp.</specimenCount>
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), Australasian (2 spp.), Nearctic (6 spp.), Neotropical (16 spp.), Oriental (
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<specimenCount id="8982FD62FFBCFF8FFA8DA13F0105F981" box="[1319,1372,1589,1613]" pageId="1" pageNumber="170" type="generic">1 sp.</specimenCount>
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), and Palearctic (10 spp.) (
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<bibRefCitation id="FB154B1AFFBCFF8FFED4A1530663F9BD" author="Yang" box="[382,570,1624,1649]" pageId="1" pageNumber="170" refString="Yang, D., Zhu, Y. J., Wang, M. Q. & Zhang, L. L. (2006) World Catalog of Dolichopodidae (Insecta: Diptera). China Agricultural University Press, Beijing, 704 pp." type="book" year="2006">
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Yang
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<emphasis id="ADF0EAF9FFBCFF8FFE6AA15305ABF9BD" box="[448,498,1624,1649]" italics="true" pageId="1" pageNumber="170">et al</emphasis>
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. 2006
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</bibRefCitation>
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;
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<bibRefCitation id="FB154B1AFFBCFF8FFDECA15207C3F9BD" author="Grichanov" box="[582,922,1624,1649]" pageId="1" pageNumber="170" refString="Grichanov, I. Ya. & Mostovski, M. B. (2009) Discovery of Systenus in the Afrotropical Region with a description of a new species (Diptera: Dolichopodidae). Zoosystematica Rossica, 18, 285 - 290." type="journal article" year="2009">Grichanov & Mostovski 2009</bibRefCitation>
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). In addition, I have seen a female
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<taxonomicName id="58844D68FFBCFF8FFAB5A1530126F9BC" box="[1311,1407,1625,1648]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus">
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<emphasis id="ADF0EAF9FFBCFF8FFAB5A1530126F9BC" box="[1311,1407,1625,1648]" italics="true" pageId="1" pageNumber="170">Systenus</emphasis>
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</taxonomicName>
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in amber from Chiapas,
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<collectingCountry id="E793767BFFBCFF8FFE25A17605BDF959" box="[399,484,1660,1685]" name="Mexico" pageId="1" pageNumber="170">Mexico</collectingCountry>
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(Bickel & Solórzano Kraemer, in press), suggesting the genus has been part of the Mesoamerican fauna since at least the Middle Miocene.
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</paragraph>
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<paragraph id="9F3B36EBFFBCFF8FFF6DA1CE0092F8A1" blockId="1.[151,1437,862,2009]" pageId="1" pageNumber="170">
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The general morphology and taxonomic placement of
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<taxonomicName id="58844D68FFBCFF8FFCE2A1CF07F1F910" box="[840,936,1733,1756]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus">
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<emphasis id="ADF0EAF9FFBCFF8FFCE2A1CF07F1F910" box="[840,936,1733,1756]" italics="true" pageId="1" pageNumber="170">Systenus</emphasis>
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</taxonomicName>
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within the
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<taxonomicName id="58844D68FFBCFF8FFB9FA1CE0098F911" box="[1077,1217,1732,1757]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="subFamily" subFamily="Medeterinae">Medeterinae</taxonomicName>
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were discussed in
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<bibRefCitation id="FB154B1AFFBCFF8FFF3DA1E2056FF8CD" author="Bickel" box="[151,310,1768,1793]" pageId="1" pageNumber="170" refString="Bickel, D. J. (1986) Australian species of Systenus (Diptera: Dolichopodidae). Records of the Australian Museum, 38 (5), 263 - 270. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.38.1986.350" type="journal article" year="1986">Bickel (1986)</bibRefCitation>
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. An additional observation on the male postabdomen of
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<taxonomicName id="58844D68FFBCFF8FFC17A1E30044F8CC" box="[957,1053,1769,1792]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus">
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<emphasis id="ADF0EAF9FFBCFF8FFC17A1E30044F8CC" box="[957,1053,1769,1792]" italics="true" pageId="1" pageNumber="170">Systenus</emphasis>
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</taxonomicName>
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should be noted. The hypopygial peduncle, formed of tergite and sternite 7 is often held withdrawn between sternites 5 and 6 at rest, leaving only tergite 7 and the genital capsule exposed. Correspondingly, sternite 7 is glabrous to facilitate retraction within the preabdomen, while tergite 7 is setose, like the adjoining tergite 6, as clearly shown in
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<figureCitation id="07BF2A6EFFBCFF8FFBFDA05E00EBF8A1" box="[1111,1202,1876,1901]" captionStart="FIGURE 1" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[238,1352,195,1838]" captionTargetId="figure@4.[234,1352,195,1839]" captionTargetPageId="4" captionText="FIGURE 1. Systenus tenorio sp. nov. (holotype): a. male habitus, left lateral (terminalia removed); b. hypopygium, left lateral. S. flavifemoratus sp. nov. (holotype): c. male habitus, right lateral; d. hypopygium, left lateral. S. maculipennis sp. nov.: e. male habitus, left lateral (terminalia removed); f. hypopygium, left lateral. Abbreviations: cer, cercus; epan, epandrium; hyp, hypandrium; ph, phallus; st, sternite; sur, surstylus; tg, tergite; 6, 7, 8, abdominal segments." httpUri="https://zenodo.org/record/237971/files/figure.png" pageId="1" pageNumber="170">Figure 1</figureCitation>
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b.
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</paragraph>
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<paragraph id="9F3B36EBFFBCFF8FFF6DA0720786F815" blockId="1.[151,1437,862,2009]" pageId="1" pageNumber="170">
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The Costa Rican
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<taxonomicName id="58844D68FFBCFF8FFE26A07305B5F85C" box="[396,492,1913,1936]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="ADF0EAF9FFBCFF8FFE26A07305B5F85C" box="[396,492,1913,1936]" italics="true" pageId="1" pageNumber="170">Systenus</emphasis>
|
|
</taxonomicName>
|
|
species treated here appear similar in overall hypopygial structure and variation to the six Brazilian Amazonas species as described and figured by
|
|
<bibRefCitation id="FB154B1AFFBCFF8FFCDAA0960056F879" author="Naglis" box="[880,1039,1948,1973]" pageId="1" pageNumber="170" refString="Naglis S. (2000) Six new species of Systenus (Diptera: Dolichopodidae) from Brazil, with a key to the Neotropical species. Studia dipterologica, 7 (1), 59 - 68." type="journal article" year="2000">Naglis (2000)</bibRefCitation>
|
|
. The following species of
|
|
<taxonomicName id="58844D68FFBCFF8FFA96A09701C5F878" box="[1340,1436,1949,1972]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="ADF0EAF9FFBCFF8FFA96A09701C5F878" box="[1340,1436,1949,1972]" italics="true" pageId="1" pageNumber="170">Systenus</emphasis>
|
|
</taxonomicName>
|
|
are known from
|
|
<collectingCountry id="E793767BFFBCFF8FFEFBA0CB0594F815" box="[337,461,1984,2009]" name="Costa Rica" pageId="1" pageNumber="170">Costa Rica</collectingCountry>
|
|
, with the Province and collection month noted.
|
|
</paragraph>
|
|
<paragraph id="9F3B36EBFFBFFF8CFF3DA7920721FF7C" blockId="2.[151,1042,151,464]" box="[151,888,151,176]" pageId="2" pageNumber="171">
|
|
<taxonomicName id="58844D68FFBFFF8CFF3DA79205DFFF7C" authority="Bickel." authorityName="Bickel." box="[151,390,151,176]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="divericatus" status="sp. nov.">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFF3DA7920576FF7C" box="[151,303,151,176]" italics="true" pageId="2" pageNumber="171">S. divericatus</emphasis>
|
|
Bickel.
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="B6C35782FFBFFF8CFE27A79D05BEFF7C" box="[397,487,151,176]" pageId="2" pageNumber="171" rank="species">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFE27A79D05BEFF7C" bold="true" box="[397,487,151,176]" pageId="2" pageNumber="171">sp. nov.</emphasis>
|
|
</taxonomicNameLabel>
|
|
,
|
|
<collectingCountry id="E793767BFFBFFF8CFE5FA792062DFF7C" box="[501,628,151,176]" name="Costa Rica" pageId="2" pageNumber="171">Costa Rica</collectingCountry>
|
|
: Guanacaste, February.
|
|
</paragraph>
|
|
<paragraph id="9F3B36EBFFBFFF8CFF3DA7B70709FF18" blockId="2.[151,1042,151,464]" box="[151,848,188,213]" pageId="2" pageNumber="171">
|
|
<taxonomicName id="58844D68FFBFFF8CFF3DA7B70528FF19" authority="Bickel" authorityName="Bickel" box="[151,369,188,213]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="eboritibia" status="sp. nov.">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFF3DA7B70578FF19" box="[151,289,188,213]" italics="true" pageId="2" pageNumber="171">S. eboritibia</emphasis>
|
|
Bickel
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="B6C35782FFBFFF8CFED2A7B6058BFF19" box="[376,466,188,213]" pageId="2" pageNumber="171" rank="species">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFED2A7B6058BFF19" bold="true" box="[376,466,188,213]" pageId="2" pageNumber="171">sp. nov.</emphasis>
|
|
</taxonomicNameLabel>
|
|
,
|
|
<collectingCountry id="E793767BFFBFFF8CFE75A7B60604FF19" box="[479,605,188,213]" name="Costa Rica" pageId="2" pageNumber="171">Costa Rica</collectingCountry>
|
|
: Guanacaste, August.
|
|
</paragraph>
|
|
<paragraph id="9F3B36EBFFBFFF8CFF3DA7EA0716FF34" blockId="2.[151,1042,151,464]" box="[151,847,223,248]" pageId="2" pageNumber="171">
|
|
<taxonomicName id="58844D68FFBFFF8CFF3DA7EA052EFF34" authority="Bickel" authorityName="Bickel" box="[151,375,223,248]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="emusorum" status="sp. nov.">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFF3DA7EA0571FF34" box="[151,296,224,248]" italics="true" pageId="2" pageNumber="171">S. emusorum</emphasis>
|
|
Bickel
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="B6C35782FFBFFF8CFED4A7D50581FF34" box="[382,472,223,248]" pageId="2" pageNumber="171" rank="species">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFED4A7D50581FF34" bold="true" box="[382,472,223,248]" pageId="2" pageNumber="171">sp. nov.</emphasis>
|
|
</taxonomicNameLabel>
|
|
,
|
|
<collectingCountry id="E793767BFFBFFF8CFE4CA7EA063CFF34" box="[486,613,223,248]" name="Costa Rica" pageId="2" pageNumber="171">Costa Rica</collectingCountry>
|
|
: Guanacaste, March.
|
|
</paragraph>
|
|
<paragraph id="9F3B36EBFFBFFF8CFF3DA60F072EFEA9" blockId="2.[151,1042,151,464]" pageId="2" pageNumber="171">
|
|
<taxonomicName id="58844D68FFBFFF8CFF3DA60F05FCFED1" authority="Bickel" authorityName="Bickel" box="[151,421,260,285]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="flavifemoratus" status="sp. nov.">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFF3DA60F050CFED1" box="[151,341,260,285]" italics="true" pageId="2" pageNumber="171">S. flavifemoratus</emphasis>
|
|
Bickel
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="B6C35782FFBFFF8CFE06A60E065CFED1" box="[428,517,260,285]" pageId="2" pageNumber="171" rank="species">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFE06A60E065CFED1" bold="true" box="[428,517,260,285]" pageId="2" pageNumber="171">sp. nov.</emphasis>
|
|
</taxonomicNameLabel>
|
|
,
|
|
<collectingCountry id="E793767BFFBFFF8CFDB9A60E06CBFED1" box="[531,658,260,285]" name="Costa Rica" pageId="2" pageNumber="171">Costa Rica</collectingCountry>
|
|
: Guanacaste, January.
|
|
<taxonomicName id="58844D68FFBFFF8CFF3DA62205C1FE8C" authority="Bickel" authorityName="Bickel" box="[151,408,295,320]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="maculipennis" status="sp. nov.">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFF3DA6220511FE8C" box="[151,328,295,320]" italics="true" pageId="2" pageNumber="171">S. maculipennis</emphasis>
|
|
Bickel
|
|
</taxonomicName>
|
|
<emphasis id="ADF0EAF9FFBFFF8CFE35A62D0659FE8C" bold="true" box="[415,512,295,320]" pageId="2" pageNumber="171">
|
|
<taxonomicNameLabel id="B6C35782FFBFFF8CFE35A62D05A3FE8C" box="[415,506,295,320]" pageId="2" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
|
|
,
|
|
</emphasis>
|
|
<collectingCountry id="E793767BFFBFFF8CFDADA62206DCFE8C" box="[519,645,295,320]" name="Costa Rica" pageId="2" pageNumber="171">Costa Rica</collectingCountry>
|
|
: Guanacaste, April, July.
|
|
<taxonomicName id="58844D68FFBFFF8CFF3DA64705DCFEA9" authority="Bickel" authorityName="Bickel" box="[151,389,332,357]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="naranjensis" status="sp. nov.">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFF3DA647056FFEA8" box="[151,310,333,356]" italics="true" pageId="2" pageNumber="171">S. naranjensis</emphasis>
|
|
Bickel
|
|
</taxonomicName>
|
|
<emphasis id="ADF0EAF9FFBFFF8CFE26A64605B7FEA9" bold="true" box="[396,494,332,357]" pageId="2" pageNumber="171">
|
|
<taxonomicNameLabel id="B6C35782FFBFFF8CFE26A64605BEFEA9" box="[396,487,332,357]" pageId="2" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
|
|
,
|
|
</emphasis>
|
|
<collectingCountry id="E793767BFFBFFF8CFE5EA646062BFEA9" box="[500,626,332,357]" name="Costa Rica" pageId="2" pageNumber="171">Costa Rica</collectingCountry>
|
|
: Guanacaste, February.
|
|
</paragraph>
|
|
<paragraph id="9F3B36EBFFBFFF8CFF3DA67A074AFE61" blockId="2.[151,1042,151,464]" pageId="2" pageNumber="171">
|
|
<taxonomicName id="58844D68FFBFFF8CFF3DA67A050CFE44" authority="Bickel" authorityName="Bickel" box="[151,341,367,392]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="parkeri" status="sp. nov.">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFF3DA67A055CFE44" box="[151,261,367,392]" italics="true" pageId="2" pageNumber="171">S. parkeri</emphasis>
|
|
Bickel
|
|
</taxonomicName>
|
|
<emphasis id="ADF0EAF9FFBFFF8CFEF6A66505E4FE44" bold="true" box="[348,445,367,392]" pageId="2" pageNumber="171">
|
|
<taxonomicNameLabel id="B6C35782FFBFFF8CFEF6A66505EEFE44" box="[348,439,367,392]" pageId="2" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
|
|
,
|
|
</emphasis>
|
|
<collectingCountry id="E793767BFFBFFF8CFE6EA67A061AFE44" box="[452,579,367,392]" name="Costa Rica" pageId="2" pageNumber="171">Costa Rica</collectingCountry>
|
|
: Guanacaste, November & January.
|
|
<taxonomicName id="58844D68FFBFFF8CFF3DA69F050DFE61" authority="Bickel" authorityName="Bickel" box="[151,340,404,429]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="tenorio" status="sp. nov.">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFF3DA69F055DFE60" box="[151,260,405,428]" italics="true" pageId="2" pageNumber="171">S. tenorio</emphasis>
|
|
Bickel
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="B6C35782FFBFFF8CFEF1A69E05EDFE61" box="[347,436,404,429]" pageId="2" pageNumber="171" rank="species">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFEF1A69E05EDFE61" bold="true" box="[347,436,404,429]" pageId="2" pageNumber="171">sp. nov.</emphasis>
|
|
</taxonomicNameLabel>
|
|
,
|
|
<collectingCountry id="E793767BFFBFFF8CFE68A69E0618FE61" box="[450,577,404,429]" name="Costa Rica" pageId="2" pageNumber="171">Costa Rica</collectingCountry>
|
|
: Guanacaste, May.
|
|
</paragraph>
|
|
<paragraph id="9F3B36EBFFBFFF8CFF3DA6B2004BFE1C" blockId="2.[151,1042,151,464]" box="[151,1042,439,464]" pageId="2" pageNumber="171">
|
|
<taxonomicName id="58844D68FFBFFF8CFF3DA6B20510FE1C" authority="Bickel" authorityName="Bickel" box="[151,329,439,464]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="zurqui" status="sp. nov.">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFF3DA6B204A0FE1C" box="[151,249,440,464]" italics="true" pageId="2" pageNumber="171">S. zurqui</emphasis>
|
|
Bickel
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="B6C35782FFBFFF8CFEFAA6BD05F3FE1C" box="[336,426,439,464]" pageId="2" pageNumber="171" rank="species">
|
|
<emphasis id="ADF0EAF9FFBFFF8CFEFAA6BD05F3FE1C" bold="true" box="[336,426,439,464]" pageId="2" pageNumber="171">sp. nov.</emphasis>
|
|
</taxonomicNameLabel>
|
|
,
|
|
<collectingCountry id="E793767BFFBFFF8CFE12A6B2066FFE1C" box="[440,566,439,464]" name="Costa Rica" pageId="2" pageNumber="171">Costa Rica</collectingCountry>
|
|
: San José, August; Puntarenas, December.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |