166 lines
18 KiB
XML
166 lines
18 KiB
XML
<document id="DAE5CF87FEA718913EEA161BAD333953" ID-DOI="10.11646/zootaxa.4020.1.7" ID-GBIF-Dataset="82b2f539-5d25-42d6-95c5-e8b16a353e12" ID-ISSN="1175-5326" ID-Zenodo-Dep="237970" ID-ZooBank="BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461269068152" checkinUser="plazi" docAuthor="Bickel, Daniel J." docDate="2015" docId="172D87FDFFBAFF87FF3DA1C3017DFE8E" docLanguage="en" docName="zt04020p182.pdf" docOrigin="Zootaxa 4020 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus divericatus Bickel, sp. nov." docType="treatment" docVersion="8" lastPageNumber="178" masterDocId="EB14FF85FFBDFF8EFFAAA70A0459FFCC" masterDocTitle="The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus" masterLastPageNumber="182" masterPageNumber="169" pageNumber="176" updateTime="1698611248703" updateUser="plazi">
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<mods:mods id="EE0FE8AF86FD146556A9CD08CF6D10AB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5BE1B3C67E044F71D15BC1FD3290C26C">
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<mods:title id="9EDA5491F4F8EE7AD7ABFD459797021C">The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus</mods:title>
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</mods:titleInfo>
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<mods:name id="076A15DFAD938102C3E0C32570663A69" type="personal">
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<mods:role id="CD9312F28E88AAC1358C5B98AA01EA34">
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<mods:namePart id="62A91EC716681D89D27B4B3EBA36ED00">Bickel, Daniel J.</mods:namePart>
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<mods:typeOfResource id="C89F382C70C69B24F5A54B45242385AC">text</mods:typeOfResource>
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<mods:title id="B85FD6622BA917949ACF93BCCC8A99CC">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9F3B1D2A28652392410F149D923D40FE">
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<mods:date id="912C42C2B89690BB76C4919EB1B411EB">2015</mods:date>
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<mods:detail id="57D32A9FFC5039384CCFFB676E1CAF4D" type="volume">
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<mods:number id="019AC721E7093F62101D3BC08D378488">4020</mods:number>
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</mods:detail>
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<mods:detail id="42ABCD06AE8884218434493D1BB5D3C2" type="issue">
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<mods:number id="C633C734AA35A4C336E6288460021851">1</mods:number>
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<mods:classification id="7CA33D8651933122D3DC7562BB403824">journal article</mods:classification>
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<mods:identifier id="2D0DB3A1E00531648306E0AAD6FFF433" type="DOI">10.11646/zootaxa.4020.1.7</mods:identifier>
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<mods:identifier id="7FFE0BB824D73F5226B2219B62225D29" type="GBIF-Dataset">82b2f539-5d25-42d6-95c5-e8b16a353e12</mods:identifier>
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<mods:identifier id="A930BD9B20C596378E1B9160D2ED6AA1" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5C0498C106E0CE04BE8779F50BED5381" type="Zenodo-Dep">237970</mods:identifier>
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<mods:identifier id="5507ECBF72F15F9C667C0950D9BA1A03" type="ZooBank">BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE</mods:identifier>
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</mods:mods>
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<treatment id="172D87FDFFBAFF87FF3DA1C3017DFE8E" ID-DOI="http://doi.org/10.5281/zenodo.6121809" ID-GBIF-Taxon="119638519" ID-Zenodo-Dep="6121809" LSID="urn:lsid:plazi:treatment:172D87FDFFBAFF87FF3DA1C3017DFE8E" httpUri="http://treatment.plazi.org/id/172D87FDFFBAFF87FF3DA1C3017DFE8E" lastPageId="9" lastPageNumber="178" pageId="7" pageNumber="176">
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<subSubSection id="D79E6560FFBAFF89FF3DA1C3054EF8C9" pageId="7" pageNumber="176" type="nomenclature">
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<paragraph id="9F3B36EBFFBAFF89FF3DA1C3061FF92F" blockId="7.[151,582,1737,1797]" box="[151,582,1737,1763]" pageId="7" pageNumber="176">
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<heading id="C4738187FFBAFF89FF3DA1C3061FF92F" bold="true" box="[151,582,1737,1763]" fontSize="11" level="1" pageId="7" pageNumber="176" reason="1">
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<emphasis id="ADF0EAF9FFBAFF89FF3DA1C3061FF92F" bold="true" box="[151,582,1737,1763]" pageId="7" pageNumber="176">
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<taxonomicName id="58844D68FFBAFF89FF3DA1C305B8F92F" ID-CoL="9HYSV" authority="Bickel" authorityName="Bickel" box="[151,481,1737,1763]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="176" phylum="Arthropoda" rank="species" species="divericatus" status="sp. nov.">
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<emphasis id="ADF0EAF9FFBAFF89FF3DA1C305D5F92F" bold="true" box="[151,396,1737,1763]" italics="true" pageId="7" pageNumber="176">Systenus divericatus</emphasis>
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Bickel
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFBAFF89FE43A1C3061FF92F" box="[489,582,1737,1763]" pageId="7" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="9F3B36EBFFBAFF89FF3DA1E6054EF8C9" blockId="7.[151,582,1737,1797]" box="[151,279,1772,1797]" pageId="7" pageNumber="176">
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(
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<figureCitation id="07BF2A6EFFBAFF89FF35A1E604BCF8C8" box="[159,229,1772,1797]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[166,1436,813,1836]" captionTargetId="figure@8.[151,1436,813,1836]" captionTargetPageId="8" captionText="FIGURE 3. Systenus divericatus sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. zurqui sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral." httpUri="https://zenodo.org/record/237973/files/figure.png" pageId="7" pageNumber="176">Figs 3</figureCitation>
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a, b)
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFBAFF89FF3DA03E0636F8BC" pageId="7" pageNumber="176" type="materials_examined">
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<paragraph id="9F3B36EBFFBAFF89FF3DA03E0636F8BC" blockId="7.[151,1437,1844,2013]" pageId="7" pageNumber="176">
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<emphasis id="ADF0EAF9FFBAFF89FF3DA03E0567F881" bold="true" box="[151,318,1844,1869]" pageId="7" pageNumber="176">
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<typeStatus id="403F8849FFBAFF89FF3DA03E048BF881" box="[151,210,1844,1869]" pageId="7" pageNumber="176">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="403F8849FFBAFF89FEE6A03E05B2F881" box="[332,491,1844,1869]" pageId="7" pageNumber="176" type="holotype">
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<emphasis id="ADF0EAF9FFBAFF89FEE6A03E05B2F881" bold="true" box="[332,491,1844,1869]" pageId="7" pageNumber="176">HOLOTYPE</emphasis>
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</typeStatus>
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♂,
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<collectingCountry id="E793767BFFBAFF89FDBEA03E06E4F880" box="[532,701,1844,1868]" name="Costa Rica" pageId="7" pageNumber="176">COSTA RICA</collectingCountry>
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:
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<emphasis id="ADF0EAF9FFBAFF89FD6DA03E070AF881" bold="true" box="[711,851,1844,1869]" pageId="7" pageNumber="176">Guanacaste</emphasis>
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:
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<quantity id="587C9B0EFFBAFF89FCC8A03E07C2F881" box="[866,923,1844,1869]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="176" unit="km" value="3.0">3 km</quantity>
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SE Rio Naranjo,
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<date id="EB3A102BFFBAFF89FBCBA03E0148F880" box="[1121,1297,1844,1868]" pageId="7" pageNumber="176" value="1992-02-21" valueMax="1992-02-28" valueMin="1992-02-21">21—28.II.1992</date>
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, F.D. Parker (
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<collectionCode id="F995AE2EFFBAFF89FF34A05204AAF8BC" LSID="urn:lsid:biocol.org:col:34802" box="[158,243,1880,1904]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="176">LACM</collectionCode>
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, ex.
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<collectionCode id="F995AE2EFFBAFF89FE8DA0520520F8BC" box="[295,377,1880,1904]" pageId="7" pageNumber="176">EMUS</collectionCode>
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;
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<collectionCode id="F995AE2EFFBAFF89FE2EA052058CF8BC" LSID="urn:lsid:biocol.org:col:34802" box="[388,469,1880,1904]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="176">LACM</collectionCode>
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Ent 320000).
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFBAFF86FF6DA0760732FD35" lastPageId="8" lastPageNumber="177" pageId="7" pageNumber="176" type="description">
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<paragraph id="9F3B36EBFFBAFF86FF6DA0760732FD35" blockId="7.[151,1437,1844,2013]" lastBlockId="8.[151,1437,151,761]" lastPageId="8" lastPageNumber="177" pageId="7" pageNumber="176">
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<emphasis id="ADF0EAF9FFBAFF89FF6DA0760508F859" bold="true" box="[199,337,1916,1941]" pageId="7" pageNumber="176">Description</emphasis>
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.
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<emphasis id="ADF0EAF9FFBAFF89FECAA07605C5F859" bold="true" box="[352,412,1916,1941]" pageId="7" pageNumber="176">Male</emphasis>
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: body length:
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<quantity id="587C9B0EFFBAFF89FDE9A07606C4F858" box="[579,669,1916,1940]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="176" unit="mm" value="2.3">2.3 mm</quantity>
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; wing 2.2 x
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<quantity id="587C9B0EFFBAFF89FC81A07607DBF858" box="[811,898,1916,1940]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="176" unit="mm" value="0.8">0.8 mm</quantity>
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(
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<figureCitation id="07BF2A6EFFBAFF89FC38A076078FF858" box="[914,982,1916,1941]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[166,1436,813,1836]" captionTargetId="figure@8.[151,1436,813,1836]" captionTargetPageId="8" captionText="FIGURE 3. Systenus divericatus sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. zurqui sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral." httpUri="https://zenodo.org/record/237973/files/figure.png" pageId="7" pageNumber="176">Fig. 3</figureCitation>
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a); similar to
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<taxonomicName id="58844D68FFBAFF89FBC5A0770084F858" box="[1135,1245,1917,1940]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="176" phylum="Arthropoda" rank="species" species="tenorio">
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<emphasis id="ADF0EAF9FFBAFF89FBC5A0770084F858" box="[1135,1245,1917,1940]" italics="true" pageId="7" pageNumber="176">S. tenorio</emphasis>
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</taxonomicName>
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except as noted:
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<emphasis id="ADF0EAF9FFBAFF89FF3DA0AA048DF875" box="[151,212,1952,1977]" italics="true" pageId="7" pageNumber="176">Head</emphasis>
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: palpus yellow with short black setae and strong apical seta; proboscis yellowish; antenna mostly dark brown but postpedicel yellowish laterally near base; postpedicel elongate, triangular tapering gradually, covered with short pubescence, more than 3 times as long as basal width, and with short apical arista.
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<emphasis id="ADF0EAF9FFB5FF86FB28A79300EEFF7C" box="[1154,1207,153,176]" italics="true" pageId="8" pageNumber="177">Legs</emphasis>
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: CI yellow; CII and CIII mostly dark brown, but distal eighth yellow; all trochanters yellow; femora mostly yellow but distal quarter of FIII infuscated; remainder of legs mostly yellow; TIII and tarsomeres III mostly with white cuticle (MSSC): CI and CII with white anterior setae and shorter vestiture, CIII with strong while lateral seta at ½; major leg setae black; I: 3.1; 3.0; 1.6/ 0.6/ 0.4/ 0.3/ 0.2; leg I bare of major setation; TI and tarsus I only covered with ivory colored vestiture (MSSC); II: 3.6; 3.8; 2.0/ 1.3/distal tarsomeres missing; TII with strong ad seta and weaker pd seta at ¼, and short anterior, long ad and short pd seta near 2/3, and with apical ring of ad, pd, av and ventral seta; III: 3.8; 4.6; 0.8/ 1.8/ 1.0/ 0.7/ 0.5; TIII with 4 spaced short black dorsal setae, and with apical ring of dorsal, ad and av seta.
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<emphasis id="ADF0EAF9FFB5FF86FAF1A69F01CAFE60" box="[1371,1427,405,428]" italics="true" pageId="8" pageNumber="177">Wing</emphasis>
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: hyaline; R2+3 and R4+5 diverging from base, R2+3 joining costa near 7/8, and R4+5 joining costa almost at wing apex; vein M diverging from R4+5 from base, and not becoming subparallel, but with M continuing to diverge to join costa well behind wing apex, and with distal outward bend causing small bump on posterior wing margin (MSSC); costa between apex of R4+5 and M almost straight (MSSC); CuAx ratio: 0.7; lower calypter pale yellow with yellowish marginal setae; halter yellow.
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<emphasis id="ADF0EAF9FFB5FF86FE42A55A060BFDA5" box="[488,594,592,617]" italics="true" pageId="8" pageNumber="177">Abdomen</emphasis>
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: segment 7 forming elongate peduncle, with sternite 7 glabrous and tergite 7 covered with same black vestiture as preabdomen; sternite 8 forming cap-like cover over hypopygial foramen; hypopygium (
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<figureCitation id="07BF2A6EFFB5FF86FE9EA592052FFD7D" box="[308,374,664,689]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[166,1436,813,1836]" captionTargetId="figure@8.[151,1436,813,1836]" captionTargetPageId="8" captionText="FIGURE 3. Systenus divericatus sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. zurqui sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral." httpUri="https://zenodo.org/record/237973/files/figure.png" pageId="8" pageNumber="177">Fig. 3</figureCitation>
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b) dark brown with yellow surstylus and cercus; epandrium ventroapically truncated; surstylus as elongate digitiform projection which is apically swollen, and bearing apical seta; cercus elongate and flagellate, and bearing elongate pale setae along length.
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<emphasis id="ADF0EAF9FFB5FF86FD3FA5EA06B5FD35" bold="true" box="[661,748,736,761]" pageId="8" pageNumber="177">Female</emphasis>
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: unknown.
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</paragraph>
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</subSubSection>
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<caption id="CBFB6663FFB5FF86FF3DA0480759F8BA" httpUri="https://zenodo.org/record/237973/files/figure.png" pageId="8" pageNumber="177" targetBox="[166,1436,813,1836]" targetPageId="8">
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<paragraph id="9F3B36EBFFB5FF86FF3DA0480759F8BA" blockId="8.[151,1436,1858,1910]" pageId="8" pageNumber="177">
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<emphasis id="ADF0EAF9FFB5FF86FF3DA0480556F894" bold="true" box="[151,271,1858,1880]" pageId="8" pageNumber="177">FIGURE 3</emphasis>
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.
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<taxonomicName id="58844D68FFB5FF86FEB4A04805B6F89B" box="[286,495,1858,1879]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="177" phylum="Arthropoda" rank="species" species="divericatus" status="sp. nov.">
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<emphasis id="ADF0EAF9FFB5FF86FEB4A04805B6F89B" box="[286,495,1858,1879]" italics="true" pageId="8" pageNumber="177">Systenus divericatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFB5FF86FE5DA0480613F894" box="[503,586,1858,1880]" pageId="8" pageNumber="177" rank="species">
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<emphasis id="ADF0EAF9FFB5FF86FE5DA0480613F894" bold="true" box="[503,586,1858,1880]" pageId="8" pageNumber="177">sp. nov.</emphasis>
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</taxonomicNameLabel>
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(holotype):
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<emphasis id="ADF0EAF9FFB5FF86FD66A0480681F894" bold="true" box="[716,728,1858,1880]" pageId="8" pageNumber="177">a</emphasis>
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. male habitus, left lateral;
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<emphasis id="ADF0EAF9FFB5FF86FC5EA048005BF894" bold="true" box="[1012,1026,1858,1880]" pageId="8" pageNumber="177">b</emphasis>
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. hypopygium, left lateral.
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<taxonomicName id="58844D68FFB5FF86FAB3A048012CF894" box="[1305,1397,1858,1880]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="177" phylum="Arthropoda" rank="species" species="zurqui" status="sp. nov.">
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<emphasis id="ADF0EAF9FFB5FF86FAB3A048012CF894" box="[1305,1397,1858,1880]" italics="true" pageId="8" pageNumber="177">S. zurqui</emphasis>
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</taxonomicName>
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<emphasis id="ADF0EAF9FFB5FF86FAD7A048049AF8BA" bold="true" pageId="8" pageNumber="177">
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<taxonomicNameLabel id="B6C35782FFB5FF86FAD7A048049AF8BA" pageId="8" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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:
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<emphasis id="ADF0EAF9FFB5FF86FF7AA06A0482F8BA" bold="true" box="[208,219,1888,1910]" pageId="8" pageNumber="177">c</emphasis>
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. male habitus, left lateral;
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<emphasis id="ADF0EAF9FFB5FF86FE40A06A05A1F8BA" bold="true" box="[490,504,1888,1910]" pageId="8" pageNumber="177">d</emphasis>
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. hypopygium, left lateral.
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</paragraph>
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</caption>
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<subSubSection id="D79E6560FFB4FF87FF6DA79D017DFE8E" pageId="9" pageNumber="178" type="discussion">
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<paragraph id="9F3B36EBFFB4FF87FF6DA79D017DFE8E" blockId="9.[151,1436,151,327]" pageId="9" pageNumber="178">
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<emphasis id="ADF0EAF9FFB4FF87FF6DA79D056DFF7C" bold="true" box="[199,308,151,176]" pageId="9" pageNumber="178">Remarks</emphasis>
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.
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<taxonomicName id="58844D68FFB4FF87FEECA7920677FF7C" box="[326,558,151,176]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="divericatus">
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<emphasis id="ADF0EAF9FFB4FF87FEECA7920677FF7C" box="[326,558,151,176]" italics="true" pageId="9" pageNumber="178">Systenus divericatus</emphasis>
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</taxonomicName>
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is known from a single male taken at the Rio Naranjo site,
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<collectingCountry id="E793767BFFB4FF87FB57A7920122FF7C" box="[1277,1403,151,176]" name="Costa Rica" pageId="9" pageNumber="178">Costa Rica</collectingCountry>
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in February. It has the diagnostic male character with veins R4+5 diverging distally with vein M and apically bent (MSSC), a black tapering postpedicel, and coxa I and remainder of legs mostly yellow except femur III infuscated along the distal quarter. Like many Costa Rican
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<taxonomicName id="58844D68FFB4FF87FD19A60D074AFED2" box="[691,787,263,286]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="genus">
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<emphasis id="ADF0EAF9FFB4FF87FD19A60D074AFED2" box="[691,787,263,286]" italics="true" pageId="9" pageNumber="178">Systenus</emphasis>
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</taxonomicName>
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species, tibia and tarsus I have ivory colored vestiture.
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<emphasis id="ADF0EAF9FFB4FF87FF6DA620051FFE8F" bold="true" box="[199,326,298,323]" pageId="9" pageNumber="178">Etymology</emphasis>
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. The specific epithet “
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<emphasis id="ADF0EAF9FFB4FF87FDEEA6200689FE8F" box="[580,720,298,323]" italics="true" pageId="9" pageNumber="178">
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<taxonomicName id="58844D68FFB4FF87FDEEA6200691FE8F" box="[580,712,298,323]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="divericatus">divericatus</taxonomicName>
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”
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</emphasis>
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is Latin and refers to the diverging veins R4+5 and M.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |