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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1187.104536" ID-Pensoft-Pub="1313-2970-1187-1" ID-Pensoft-UUID="7E263DF0EDD652A281B2CBBCDAC8B95B" ID-ZooBank="226A24ABB4BE4EFDBF11D6CA719AB601" ModsDocID="1313-2970-1187-1" checkinTime="1703142022244" checkinUser="pensoft" docAuthor="Herrera-Alva, Valia, Catenazzi, Alessandro &amp; Aguilar-Puntriano, Cesar" docDate="2023" docId="106D7E7B730C5CACB3E922BD2BD5E452" docLanguage="en" docName="ZooKeys 1187: 1-29" docOrigin="ZooKeys 1187" docPubDate="2023-12-20" docSource="http://dx.doi.org/10.3897/zookeys.1187.104536" docTitle="Pristimantis similaris Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano 2023, sp. nov." docType="treatment" docUuid="BC56FD8A-6EBD-43C9-A446-689FC3253576" docUuidSource="ZooBank" docVersion="2" id="7E263DF0EDD652A281B2CBBCDAC8B95B" lastPageNumber="1" masterDocId="7E263DF0EDD652A281B2CBBCDAC8B95B" masterDocTitle="A new cryptic species of terrestrial breeding frog of the Pristimantis danae Group (Anura, Strabomantidae) from montane forests in Ayacucho, Peru" masterLastPageNumber="29" masterPageNumber="1" pageNumber="1" updateTime="1703142324792" updateUser="ExternalLinkService">
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<mods:titleInfo id="0F0EE946C9423DEE92128EE0844E782D">
<mods:title id="F910C2E43814FC737E9577E28991B707">A new cryptic species of terrestrial breeding frog of the Pristimantis danae Group (Anura, Strabomantidae) from montane forests in Ayacucho, Peru</mods:title>
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<mods:name id="0DDD3804430AE2FE252968BFAA485EEA" type="personal">
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<mods:namePart id="69BD87881A552421444A59378484355A">Herrera-Alva, Valia</mods:namePart>
<mods:nameIdentifier id="FA8E38CEBE107B04E25F32B0F455DCBB" type="ORCID">https://orcid.org/0000-0001-7858-8279</mods:nameIdentifier>
<mods:affiliation id="B412BB1065A4173397B656C9F4351A7C">Departamento de Herpetologia, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Lima, Peru &amp; Laboratorio de Sistematica y Ecologia de Vertebrados, Facultad de Ciencias Biologicas, Universidad Nacional Mayor de San Marcos, Lima, Peru</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="47AC588B65629A4FED1E29AF87D78DE5">Catenazzi, Alessandro</mods:namePart>
<mods:nameIdentifier id="A321BF9344FA7E6C2DEE6C7D003387FE" type="ORCID">https://orcid.org/0000-0002-3650-4783</mods:nameIdentifier>
<mods:affiliation id="B14AAE7A812109A90CE28D87DA00BE46">Florida International University, Department of Biological Sciences, 11200 SW 8 th Street, Miami, FL 33199, USA</mods:affiliation>
</mods:name>
<mods:name id="93190C8F01B99D11E2262CE27603BE0E" type="personal">
<mods:role id="1E9AD00E1E7E823BBD90DB5B4957E3A0">
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<mods:namePart id="66CEE208D920E09BA79F470BF9645E19">Aguilar-Puntriano, Cesar</mods:namePart>
<mods:nameIdentifier id="B96DF4A58F867C77B76B0AAF95C2B08D" type="ORCID">https://orcid.org/0000-0001-6372-7926</mods:nameIdentifier>
<mods:affiliation id="5A38ACE5E98345B9F8B636902CF97139">Departamento de Herpetologia, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Lima, Peru &amp; Laboratorio de Sistematica y Ecologia de Vertebrados, Facultad de Ciencias Biologicas, Universidad Nacional Mayor de San Marcos, Lima, Peru</mods:affiliation>
<mods:nameIdentifier id="B397779F6F98D10D0DCF885D32438840" type="email">caguilarp@unmsm.edu.pe</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A146E4957FF9D13BF329305839BFA04D">text</mods:typeOfResource>
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<mods:title id="E1D2F6F2FD6084848F4A5B0A2FA22DC7">ZooKeys</mods:title>
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<mods:part id="79323519C2545D436B628FDE8E959217">
<mods:date id="2E2468D71A7F5E92392B01737CC47E05">2023</mods:date>
<mods:detail id="B3B91F265E685EFA67B07FF82F039074" type="pubDate">
<mods:number id="DBBE939B46A3EDAA375F1D62EED152B5">2023-12-20</mods:number>
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<mods:number id="AE3815BF3F7C40FB7EEFD297AE549C01">1187</mods:number>
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<mods:url id="15E8F52EE2DD65A90B8608FD3ABE9FE4">http://dx.doi.org/10.3897/zookeys.1187.104536</mods:url>
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<mods:classification id="86244AEA83B388B4F3667F9BCD0C844C">journal article</mods:classification>
<mods:identifier id="404F74C7DA78A03FBA26E9940F01F3C5" type="DOI">http://dx.doi.org/10.3897/zookeys.1187.104536</mods:identifier>
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</mods:mods>
<treatment id="106D7E7B730C5CACB3E922BD2BD5E452" LSID="urn:lsid:zoobank.org:act:BC56FD8A-6EBD-43C9-A446-689FC3253576" httpUri="http://treatment.plazi.org/id/106D7E7B730C5CACB3E922BD2BD5E452" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="CECB259D9B4179CC2DB87AF21E4461E0" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="7EE3205ED36F3E09D792EB7DFC60687C" pageId="0" pageNumber="1">
<taxonomicName id="4D096A4147D91669E5D37AFB200EE90A" LSID="https://zoobank.org/BC56FD8A-6EBD-43C9-A446-689FC3253576" authority="Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano, 2023" authorityName="Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano" authorityYear="2023" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis similaris" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="similaris" status="sp. nov.">Pristimantis similaris</taxonomicName>
<taxonomicNameLabel id="1D275C1978861A241417AB1FA991B999" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C39570E336A0A2CFCA2FB10C0F3FCE3A" pageId="0" pageNumber="1" type="description">
<paragraph id="3467E919B2CCD6C2E6EEBBCD3E4945DE" pageId="0" pageNumber="1">
<figureCitation id="B58DBE29C2A8726B352A57CD62C330F2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pristimantis similaris sp. nov. (A-D) male. SVL: 17.0 mm. Holotype. MUSM 41030. Photos by Vladimir Diaz-Vargas." figureDoi="10.3897/zookeys.1187.104536.figure4" httpUri="https://binary.pensoft.net/fig/955206" pageId="0" pageNumber="1">Figs 4</figureCitation>
<figureCitation id="136D1DD46E4EE9C8816769C904FE8B0A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pristimantis similaris sp. nov. A hand B toe. Male. Holotype. MUSM 41030. Photos by VHA." figureDoi="10.3897/zookeys.1187.104536.figure5" httpUri="https://binary.pensoft.net/fig/955207" pageId="0" pageNumber="1">, 5</figureCitation>
<figureCitation id="0CFC7F39F29C238321EAF5F117272345" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-H colour and pattern variation of Pristimantis similaris sp. nov. Specimen from A-F collected in Cajadela: A, B male MUSM 41029 C, D female MUSM 41031 E, F female MUSM 41032. Specimen G, H male MUSM 41326 collected in Machente. Photos by V. Diaz-Vargas and E. Castillo-Urbina." figureDoi="10.3897/zookeys.1187.104536.figure6" httpUri="https://binary.pensoft.net/fig/955208" pageId="0" pageNumber="1">, 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="87DE3D721FCBF728BB464A6D51794839" pageId="0" pageNumber="1" type="common name">
<paragraph id="D7A68500A056EFA9A95B3C066937C841" pageId="0" pageNumber="1">Common name.</paragraph>
<paragraph id="3E2B7E80B22AA90D4FE2C2C0D7480E8A" pageId="0" pageNumber="1">
English: Similar Rubber Frog. Spanish: Rana
<normalizedToken id="F007BC86022253ABED0AA0DDE4F53EE2" originalValue="cutín">cutin</normalizedToken>
similar.
</paragraph>
</subSubSection>
<subSubSection id="4F192370574EEA6ACDB3D74712AA31F0" pageId="0" pageNumber="1" type="generic placement">
<paragraph id="FDF17960A5829A85DB31881C9C835FAA" pageId="0" pageNumber="1">Generic placement.</paragraph>
<paragraph id="11E16F0A8A8A5EB7946470AF8D9290FC" pageId="0" pageNumber="1">
We assign this species to the genus
<taxonomicName id="6A2C03DFF1579AC429683DFA8C696818" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">
<emphasis id="F730BE1F91334CE7AF9117CBAF376867" italics="true" pageId="0" pageNumber="1">Pristimantis</emphasis>
</taxonomicName>
, based on morphology and molecular data (Figs
<figureCitation id="C46ED64BBDA8BEC7AED2038BC1F79B73" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Maximum Likelihood tree of concatenated genes 16 S rRNA, COI, RAG 1 and TYR taken from GenBank and novel sequences. Numbers on nodes are bootstrap values (see Materials and Methods section for details). Green shadow corresponds to the ingroup. Pristimantis similaris sp. nov. in red, Pristimantis sp. 3 from Bolivia in purple and Pristimantis sp. from Cusco in blue. Maximum Likelihood optimal tree with bootstrap node values from the analysis of a concatenated dataset of 591 bp (16 S), 685 bp (COI), 624 bp (RAG 1) and 545 bp (TYR) of 128 species aligned by MAFFT and node support assessed using 10,000 replicates in IQTREE." figureDoi="10.3897/zookeys.1187.104536.figure2" httpUri="https://binary.pensoft.net/fig/955204" pageId="0" pageNumber="1">2</figureCitation>
,
<figureCitation id="158CA75A1A1E449EFBA623700C831506" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pristimantis similaris sp. nov. (A-D) male. SVL: 17.0 mm. Holotype. MUSM 41030. Photos by Vladimir Diaz-Vargas." figureDoi="10.3897/zookeys.1187.104536.figure4" httpUri="https://binary.pensoft.net/fig/955206" pageId="0" pageNumber="1">4</figureCitation>
,
<figureCitation id="5F0FE2615D0BEB7F51A4AC52BA248941" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-H colour and pattern variation of Pristimantis similaris sp. nov. Specimen from A-F collected in Cajadela: A, B male MUSM 41029 C, D female MUSM 41031 E, F female MUSM 41032. Specimen G, H male MUSM 41326 collected in Machente. Photos by V. Diaz-Vargas and E. Castillo-Urbina." figureDoi="10.3897/zookeys.1187.104536.figure6" httpUri="https://binary.pensoft.net/fig/955208" pageId="0" pageNumber="1">6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B9C2036FD0DD5AD7D1BFAF6790921A45" pageId="0" pageNumber="1" type="type material">
<paragraph id="7C1DCF09B5546D997CE3DC9FC4A9985C" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="DB423413788321C4DA20349D7AB0E257" pageId="0" pageNumber="1">
<emphasis id="9DFBD9D70DDBC2E343F824ED32A8E9F1" bold="true" pageId="0" pageNumber="1">
<emphasis id="AC075D5645C28CF8D821BC87693AD53D" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
MUSM 41030, adult male (Figs
<figureCitation id="981BA091166F96565D50B99E682A1B4B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pristimantis similaris sp. nov. (A-D) male. SVL: 17.0 mm. Holotype. MUSM 41030. Photos by Vladimir Diaz-Vargas." figureDoi="10.3897/zookeys.1187.104536.figure4" httpUri="https://binary.pensoft.net/fig/955206" pageId="0" pageNumber="1">4</figureCitation>
,
<figureCitation id="DF4695715D20485492D86D4DFB74478D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pristimantis similaris sp. nov. A hand B toe. Male. Holotype. MUSM 41030. Photos by VHA." figureDoi="10.3897/zookeys.1187.104536.figure5" httpUri="https://binary.pensoft.net/fig/955207" pageId="0" pageNumber="1">5</figureCitation>
) from Comunidad Cajadela (
<geoCoordinate id="9A5FC07D72C7050F0395B934C1D990E0" degrees="12" direction="south" minutes="57" orientation="latitude" precision="1" seconds="16.50" value="-12.954583">12°57'16.50&quot;S</geoCoordinate>
, 73°35'0.70
<normalizedToken id="D3690E8BBE981090C3F4D2F8D8CDB185" originalValue="”’">&quot;'</normalizedToken>
W, 1460 m a.s.l.), Distrito Anco, Provincia La Mar, Departamento Ayacucho, Peru, collected on 15 November 2018 by V. Herrera-Alva, E. Castillo-Urbina, V.
<normalizedToken id="5FBC87C3EB451274ED03006579B14A7D" originalValue="Díaz">Diaz</normalizedToken>
, M. Fernandez, and J. Gamboa.
</paragraph>
<caption id="B0986B05C21A156769252650945FA22B" doi="10.3897/zookeys.1187.104536.figure4" httpUri="https://binary.pensoft.net/fig/955206" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph id="9D0CEE3EE63838AE3AFCD2519EBCACB6" pageId="0" pageNumber="1">
<emphasis id="D86408E6AF1DB3CA57959E354A1A21DB" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
<taxonomicName id="B0F828B7B0E4ECAD72652B48893F9E38" authorityName="Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano" authorityYear="2023" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis similaris" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="similaris">
<emphasis id="ADCE24F0403758B15EE7F99BB96293DB" italics="true" pageId="0" pageNumber="1">Pristimantis similaris</emphasis>
</taxonomicName>
sp. nov. (
<emphasis id="38BC46034BCB7DA05E5586DA713370ED" bold="true" pageId="0" pageNumber="1">A-D</emphasis>
) male. SVL: 17.0 mm. Holotype. MUSM 41030. Photos by Vladimir Diaz-Vargas.
</paragraph>
</caption>
<caption id="0433C5153BF6889E00BAA1E0A738D6FF" doi="10.3897/zookeys.1187.104536.figure5" httpUri="https://binary.pensoft.net/fig/955207" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph id="BA1A838C2599D6352858D8838E721DBF" pageId="0" pageNumber="1">
<emphasis id="0D2BF2AA6E726E77A54A061F6159D7DB" bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
<taxonomicName id="665F3F4F5D132AA48B4D15E0B36A3EF7" authorityName="Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano" authorityYear="2023" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis similaris" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="similaris">
<emphasis id="7324BA809A9031F7F923058DCD4C782C" italics="true" pageId="0" pageNumber="1">Pristimantis similaris</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="7B0313B10128A90F738032BDF34731C6" bold="true" pageId="0" pageNumber="1">A</emphasis>
hand
<emphasis id="0DBCB8E529C3661851C7FB27C7870375" bold="true" pageId="0" pageNumber="1">B</emphasis>
toe. Male. Holotype. MUSM 41030. Photos by VHA.
</paragraph>
</caption>
<paragraph id="6AA07DC6CA4821264281EA1A9A200C2B" pageId="0" pageNumber="1">
<emphasis id="2C40E59376BF709899B2D6B781C4C464" bold="true" pageId="0" pageNumber="1">
<emphasis id="B1DB1AA5E961F8B95D79DA4B7C99FFC8" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
.
</emphasis>
Nine specimens. Five adult females (MUSM 41031, 41032, 41035, 41036 and MUSM 41037). Four adult males (MUSM 41029, 41028, 41033 and MUSM 41034). All the specimens were collected at the type locality, except MUSM 41037, which was collected in Comunidad Machente (
<geoCoordinate id="042993916F63BDA3A0D702442C180E1D" degrees="12" direction="south" minutes="41" orientation="latitude" precision="1" seconds="31.70" value="-12.692139">12°41'31.70&quot;S</geoCoordinate>
,
<geoCoordinate id="C68422D77146C9017A72A6CD65C8E6E7" degrees="73" direction="west" minutes="51" orientation="longitude" precision="1" seconds="0.30" value="-73.85008">73°51'0.30&quot;W</geoCoordinate>
, 1640 m a.s.l.), Distrito Ayna, Provincia La Mar, Departamento Ayacucho, Peru, on 11 November 2021 by V. Herrera-Alva, E. Castillo-Urbina, V.
<normalizedToken id="2FCE76A7E8BC763E963096DC1B0CB281" originalValue="Díaz">Diaz</normalizedToken>
and K.
<normalizedToken id="59E736D3847586DC5EFA5115E0B1EEE5" originalValue="Ñaccha">Naccha</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="3B7226F3B91759DB2994F62A521812E8" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="3C6C22B04D59E28BD3975526690520BD" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="B8EF1D71B6B0B095E0546ED161369115" pageId="0" pageNumber="1">
A new species of
<taxonomicName id="BC9ABD5D80F70B8CEEEB9DB110AEE2FE" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="genus">
<emphasis id="3D83B0E670782EC578091A2B1E73FA83" italics="true" pageId="0" pageNumber="1">Pristimantis</emphasis>
</taxonomicName>
assigned to the
<taxonomicName id="D81A1387D49DEF4DBB381F8FD3669BD6" lsidName="P. danae" pageId="0" pageNumber="1" rank="species" species="danae">
<emphasis id="6E6CA98393BC26721CD8DE96DB335962" italics="true" pageId="0" pageNumber="1">P. danae</emphasis>
</taxonomicName>
species Group having the following combination of characters: (1) Skin on dorsum shagreen, skin on venter areolate; discoidal and dorsolateral folds present, weak; thoracic fold present; (2) tympanic membrane and tympanic annulus present, distinct, visible externally; (3) snout subaccuminated in dorsal view, round in lateral view; (4) upper eyelid lacking tubercles; EW smaller than IOD; cranial crest absent; two small and flat tubercles above the snout near the eyes; (5) dentigerous processes of vomers low, oblique in five of the paratypes, absent in four paratypes and the holotype; (6) males with vocal slits, subgular vocal sac large extending on to chest and without nuptial pads; (7) Finger I slightly shorter than Finger II; discs of digits expanded, flat and truncated; (8) fingers without lateral fringes; (9) ulnar tubercles present, but diffuse; (10) heel with two to three small and flat tubercles; inner tarsal fold present, small; (11) inner metatarsal tubercle ovoid, 2-3 times larger than outer one; outer metatarsal tubercle small, ovoid; numerous and flat supernumerary tubercles; (12) toes without lateral fringes; basal toe webbing absent; toe V is slightly longer than toe III; toe discs about as large as those on fingers; (13) in life, dorsum varies from blackish to dark brown with three conspicuous chevrons (not very visible in some cases) (Fig.
<figureCitation id="6BA032E428D8F7A21AA7256EC3638E2B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-H colour and pattern variation of Pristimantis similaris sp. nov. Specimen from A-F collected in Cajadela: A, B male MUSM 41029 C, D female MUSM 41031 E, F female MUSM 41032. Specimen G, H male MUSM 41326 collected in Machente. Photos by V. Diaz-Vargas and E. Castillo-Urbina." figureDoi="10.3897/zookeys.1187.104536.figure6" httpUri="https://binary.pensoft.net/fig/955208" pageId="0" pageNumber="1">6</figureCitation>
); in most of the adults, the anterior surfaces of thighs reddish-orange, posterior surfaces of thighs brown; flanks cream without tubercles; groin same pattern as flanks mostly, some specimens with orange-reddish colouration (Figs
<figureCitation id="96087C7A22BA4DAA06E3517FE08C5D69" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pristimantis similaris sp. nov. (A-D) male. SVL: 17.0 mm. Holotype. MUSM 41030. Photos by Vladimir Diaz-Vargas." figureDoi="10.3897/zookeys.1187.104536.figure4" httpUri="https://binary.pensoft.net/fig/955206" pageId="0" pageNumber="1">4</figureCitation>
,
<figureCitation id="3E2ADD1670AE324C32375C8509622807" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-H colour and pattern variation of Pristimantis similaris sp. nov. Specimen from A-F collected in Cajadela: A, B male MUSM 41029 C, D female MUSM 41031 E, F female MUSM 41032. Specimen G, H male MUSM 41326 collected in Machente. Photos by V. Diaz-Vargas and E. Castillo-Urbina." figureDoi="10.3897/zookeys.1187.104536.figure6" httpUri="https://binary.pensoft.net/fig/955208" pageId="0" pageNumber="1">6</figureCitation>
); venter cream to yellow with black conspicuous reticulations in the throat and black marks in the chest, males present yellow throat with black or white longitudinal reticulations not as conspicuous as in females (Figs
<figureCitation id="EDE315BAD6BB7A8A7B59170AC0317881" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pristimantis similaris sp. nov. (A-D) male. SVL: 17.0 mm. Holotype. MUSM 41030. Photos by Vladimir Diaz-Vargas." figureDoi="10.3897/zookeys.1187.104536.figure4" httpUri="https://binary.pensoft.net/fig/955206" pageId="0" pageNumber="1">4</figureCitation>
,
<figureCitation id="842B2886BB8A3F59C88C1BABE70028B7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-H colour and pattern variation of Pristimantis similaris sp. nov. Specimen from A-F collected in Cajadela: A, B male MUSM 41029 C, D female MUSM 41031 E, F female MUSM 41032. Specimen G, H male MUSM 41326 collected in Machente. Photos by V. Diaz-Vargas and E. Castillo-Urbina." figureDoi="10.3897/zookeys.1187.104536.figure6" httpUri="https://binary.pensoft.net/fig/955208" pageId="0" pageNumber="1">6</figureCitation>
); iris dark copper-coloured with fine black vermiculations; (14) SVL in adult females 20.8-25.2 mm (mean = 23.4
<normalizedToken id="25D65FAA9A782DDE674DAA6A921BCD8A" originalValue="±">+/-</normalizedToken>
1.8 SE, n = 5), in adult males 17.0-18.6 mm (mean = 18.1 mm
<normalizedToken id="6DE5C3425DBCBED30A90AA957B880740" originalValue="±">+/-</normalizedToken>
0.7 SE, n = 5).
</paragraph>
<caption id="13DD1A9D83B1FE964016FD724F9B573F" doi="10.3897/zookeys.1187.104536.figure6" httpUri="https://binary.pensoft.net/fig/955208" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
<paragraph id="3D2DF34EF40AF1AB22592BD776F88AC7" pageId="0" pageNumber="1">
<emphasis id="41CFF0FFF25C5D46A422D7DE8CFC3E10" bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
<emphasis id="C631F590370B02018DCF0206D6486D06" bold="true" pageId="0" pageNumber="1">A-H</emphasis>
colour and pattern variation of
<taxonomicName id="64E35065772169078EB504E471E87250" authorityName="Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano" authorityYear="2023" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis similaris" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="similaris">
<emphasis id="49241C1D1D05BA77F764CE5A204E3BBE" italics="true" pageId="0" pageNumber="1">Pristimantis similaris</emphasis>
</taxonomicName>
sp. nov. Specimen from
<emphasis id="20B2651A0D23D0266CD738DD18B0A59C" bold="true" pageId="0" pageNumber="1">A-F</emphasis>
collected in Cajadela:
<emphasis id="D4B473BAF768E01CEABF0AF2BC318C5E" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
male MUSM 41029
<emphasis id="7D6621D03FDD70DF6633FE8F122EE2E2" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
female MUSM 41031
<emphasis id="ABF9322563E98F14B4E41E664EB9458E" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
female MUSM 41032. Specimen
<emphasis id="EC38C11937988A5DF224E4724196916A" bold="true" pageId="0" pageNumber="1">G, H</emphasis>
male MUSM 41326 collected in Machente. Photos by V. Diaz-Vargas and E. Castillo-Urbina.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DE26CE334F49D7CA070630C2FB5602A0" pageId="0" pageNumber="1" type="comparisons">
<paragraph id="21AF64E5329A65ECDB7FEA1BCE814103" pageId="0" pageNumber="1">Comparisons.</paragraph>
<paragraph id="A058669032CD9CA249922112A92EDC51" pageId="0" pageNumber="1">
<taxonomicName id="87266C7D0FDF1E5B795C71F2635604D4" authorityName="Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano" authorityYear="2023" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis similaris" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="similaris">
<emphasis id="8D256F16051DABE73BE739912DB67723" italics="true" pageId="0" pageNumber="1">Pristimantis similaris</emphasis>
</taxonomicName>
is distinguished from its congeners in Peru and Bolivia by the following combination of characters: skin on dorsum areolate, tympanum and tympanic annulus distinct, weakly-defined discoidal and dorsolateral folds, two small and flat tubercles above the snout near the eyes (not conspicuous in preservative), dorsum dark brown with three darker chevrons, anterior surface of thighs usually orange-reddish and posterior surface of thighs dark brown.
<taxonomicName id="D561BA23E1A4A648885BF3032E6CE2BE" authorityName="Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano" authorityYear="2023" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis similaris" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="similaris">
<emphasis id="36684EC6CEAD734FB1816342C5E2BDEF" italics="true" pageId="0" pageNumber="1">Pristimantis similaris</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="83FECA5BFE7045134A40CCC175369E9D" lsidName="P. rhabdolaemus" pageId="0" pageNumber="1" rank="species" species="rhabdolaemus">
<emphasis id="25E3421B614A00790BA3FF4806F0A15F" italics="true" pageId="0" pageNumber="1">P. rhabdolaemus</emphasis>
</taxonomicName>
and
<taxonomicName id="7A4CCDCED47AE72194B5D4B2BFD344AB" lsidName="P. pharangobates" pageId="0" pageNumber="1" rank="species" species="pharangobates">
<emphasis id="FEDC1BCE8ABD8F3FA3E32B4958943BA3" italics="true" pageId="0" pageNumber="1">P. pharangobates</emphasis>
</taxonomicName>
by the following characters (characters in parenthesis): smaller SVL of 20.8-25.8 mm in ten females and 15.2-18.9 mm in eight males (
<taxonomicName id="83743561525BB8913DABE27AD0088791" lsidName="P. pharangobates" pageId="0" pageNumber="1" rank="species" species="pharangobates">
<emphasis id="456F9AF02F5A0708F54F6EFAFD4934C7" italics="true" pageId="0" pageNumber="1">P. pharangobates</emphasis>
</taxonomicName>
23.1-27.8 mm in females and 15.2-18.2 mm in males;
<taxonomicName id="C72C02045C14C31C2D251249246021E7" lsidName="P. rhabdolaemus" pageId="0" pageNumber="1" rank="species" species="rhabdolaemus">
<emphasis id="7B00F36A68C05F63CDFA03491B3096EE" italics="true" pageId="0" pageNumber="1">P. rhabdolaemus</emphasis>
</taxonomicName>
25.5-31.9 mm in females and 24.1-26.3 mm in males); absence of scapular tubercles (present in both species); presence of conspicuous longitudinal black and white or yellow marks on the throat and chest (less conspicuous in
<taxonomicName id="836AC174E90880E8C00F8444B44E7CA7" lsidName="P. pharangobates" pageId="0" pageNumber="1" rank="species" species="pharangobates">
<emphasis id="5BAF638D2D9C59A1B196F683DDEF4789" italics="true" pageId="0" pageNumber="1">P. pharangobates</emphasis>
</taxonomicName>
and
<taxonomicName id="8714B80D4B06E469FC0C18EC65511AAF" lsidName="P. rhabdolaemus" pageId="0" pageNumber="1" rank="species" species="rhabdolaemus">
<emphasis id="BCC5569E026B0F70ECFEAD85319AA592" italics="true" pageId="0" pageNumber="1">P. rhabdolaemus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8C2D945A333DD674DACE41E0968FCCC6" pageId="0" pageNumber="1">
Other species in the
<taxonomicName id="62F27D9F1BAB5F91E1AC72CD6D0884AE" baseAuthorityName="Duellman" baseAuthorityYear="1978" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis danae" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="danae">
<emphasis id="EDB2B78E1E445535144DAA8EB9134933" italics="true" pageId="0" pageNumber="1">Pristimantis danae</emphasis>
</taxonomicName>
species Group that are similar to
<taxonomicName id="D21B073A0F0CAF365BCDC64A34806F8E" lsidName="P. similaris" pageId="0" pageNumber="1" rank="species" species="similaris">
<emphasis id="9C2122E7626721E2BF5D43A8348F953D" italics="true" pageId="0" pageNumber="1">P. similaris</emphasis>
</taxonomicName>
include
<taxonomicName id="F9B3203D8071669C169E4B4B8A8D7A55" lsidName="P. danae" pageId="0" pageNumber="1" rank="species" species="danae">
<emphasis id="FE469F013C82E6DFDDF8D256002E8625" italics="true" pageId="0" pageNumber="1">P. danae</emphasis>
</taxonomicName>
,
<taxonomicName id="88DEDD219230F7D34F42271A6F1B0406" lsidName="P. reichlei" pageId="0" pageNumber="1" rank="species" species="reichlei">
<emphasis id="85BEF089E2176A4CF2D9D8B940EE1D09" italics="true" pageId="0" pageNumber="1">P. reichlei</emphasis>
</taxonomicName>
,
<taxonomicName id="4542EDA9197AD4F1AB8C72542D3DF8CB" lsidName="P. scitulus" pageId="0" pageNumber="1" rank="species" species="scitulus">
<emphasis id="698F86B82D1C6CD6EB919A64AA7F994B" italics="true" pageId="0" pageNumber="1">P. scitulus</emphasis>
</taxonomicName>
and
<taxonomicName id="2FF4C3EB399CD7357C118694288C50B2" lsidName="P. toftae" pageId="0" pageNumber="1" rank="species" species="toftae">
<emphasis id="26E0332058FAEF91BA5346C4CA6A0207" italics="true" pageId="0" pageNumber="1">P. toftae</emphasis>
</taxonomicName>
.
<taxonomicName id="0B6B0857C8413B1C6F37691B03897363" baseAuthorityName="Duellman" baseAuthorityYear="1978" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis danae" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="danae">
<emphasis id="42A28E7468926ED340CBA08A8C30F491" italics="true" pageId="0" pageNumber="1">Pristimantis danae</emphasis>
</taxonomicName>
and
<taxonomicName id="C277DAC3FAB7B17B354DD27D9F0400DD" authorityName="Padial &amp; De la Riva" authorityYear="2009" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis reichlei" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="reichlei">
<emphasis id="CC4A8A51B53B92090BEDA219766C37EA" italics="true" pageId="0" pageNumber="1">Pristimantis reichlei</emphasis>
</taxonomicName>
also have brown chevrons in the dorsum and differ from the new species by the combination of the following characters: males nuptial pads absent (present in
<taxonomicName id="302C6077DC2C059A9598534772757F32" lsidName="P. danae" pageId="0" pageNumber="1" rank="species" species="danae">
<emphasis id="217FB00C26C9D09760C89EE820F5953E" italics="true" pageId="0" pageNumber="1">P. danae</emphasis>
</taxonomicName>
and
<taxonomicName id="C8D91F330CF7D852746615ACBD676D3A" lsidName="P. reichlei" pageId="0" pageNumber="1" rank="species" species="reichlei">
<emphasis id="72CC8C77948B8BB11A5A61F46404755F" italics="true" pageId="0" pageNumber="1">P. reichlei</emphasis>
</taxonomicName>
), dorsolateral folds present (weak in
<taxonomicName id="A3F8A53B4AD2F01D4A20BC3011D43A54" lsidName="P. danae" pageId="0" pageNumber="1" rank="species" species="danae">
<emphasis id="8953C8B0CF4F1D30D85CC925C04FC33A" italics="true" pageId="0" pageNumber="1">P. danae</emphasis>
</taxonomicName>
and absent
<taxonomicName id="6720FE6242229E3A8B5FDD4EB78AA31E" lsidName="P. reichlei" pageId="0" pageNumber="1" rank="species" species="reichlei">
<emphasis id="4D806FE6D4625B72FF1D5DBC802F9355" italics="true" pageId="0" pageNumber="1">P. reichlei</emphasis>
</taxonomicName>
), small pale spots in the posterior surfaces of the thighs absent (present in
<taxonomicName id="91F776122836B8C381FA9C24688DF32C" lsidName="P. danae" pageId="0" pageNumber="1" rank="species" species="danae">
<emphasis id="8EB8E227A9961231084A69B811C51047" italics="true" pageId="0" pageNumber="1">P. danae</emphasis>
</taxonomicName>
and
<taxonomicName id="AB2526589E8AA9C51718DE20E677ABC6" lsidName="P. reichlei" pageId="0" pageNumber="1" rank="species" species="reichlei">
<emphasis id="8C0EF427623EF05EA4C90A451A142C34" italics="true" pageId="0" pageNumber="1">P. reichlei</emphasis>
</taxonomicName>
) and smaller size in
<taxonomicName id="CE44D3ED5F0A135E0E469DA3E8DA0F9F" lsidName="P. similaris" pageId="0" pageNumber="1" rank="species" species="similaris">
<emphasis id="BDF3BC0ECD376AF92F85BF21C12B9745" italics="true" pageId="0" pageNumber="1">P. similaris</emphasis>
</taxonomicName>
.
<taxonomicName id="1AD5973E315288D53185D9B3E4A61392" baseAuthorityName="Duellman" baseAuthorityYear="1978" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis scitulus" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="scitulus">
<emphasis id="60E98B3D4BE314EFF142F70BB7437B23" italics="true" pageId="0" pageNumber="1">Pristimantis scitulus</emphasis>
</taxonomicName>
is morphologically similar to
<taxonomicName id="842FD279A8EFDCB2EE5FA29F2C863459" lsidName="P. similaris" pageId="0" pageNumber="1" rank="species" species="similaris">
<emphasis id="4136951C5231EBF0B570192A72E07B72" italics="true" pageId="0" pageNumber="1">P. similaris</emphasis>
</taxonomicName>
and has a parapatric distribution (Yuraccyacu, in the Piene Valley, Ayacucho Region).
<taxonomicName id="8272F4972CA783DDE3F6C3ADEA45B149" authorityName="Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano" authorityYear="2023" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis similaris" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="similaris">
<emphasis id="4792C2E00F9A1A4A2FFB8B7727F67B47" italics="true" pageId="0" pageNumber="1">Pristimantis similaris</emphasis>
</taxonomicName>
can be distinguished by lacking a conical tubercle in the upper eyelid and heels (present in
<taxonomicName id="247B8965C6B48DB3E21889C0712FAC0A" lsidName="P. scitulus" pageId="0" pageNumber="1" rank="species" species="scitulus">
<emphasis id="46DE345CC129ADC4B41F3629F80AA7A8" italics="true" pageId="0" pageNumber="1">P. scitulus</emphasis>
</taxonomicName>
), mid-ventral line absent (present in
<taxonomicName id="02D3168B3E0AB123D95127056973AD56" lsidName="P. scitulus" pageId="0" pageNumber="1" rank="species" species="scitulus">
<emphasis id="8AD4EDC12C76C37ABC39041EDB534701" italics="true" pageId="0" pageNumber="1">P. scitulus</emphasis>
</taxonomicName>
) and absence of marks in the groin or thighs (conspicuous dark spots in the groin that is continuous as marks on the posterior surfaces of the thighs in
<taxonomicName id="93A2459DF896EEC73BFB339E54850EAA" lsidName="P. scitulus" pageId="0" pageNumber="1" rank="species" species="scitulus">
<emphasis id="A7B73272C3ACA8B2C298E042C2DDCDA0" italics="true" pageId="0" pageNumber="1">P. scitulus</emphasis>
</taxonomicName>
).
<taxonomicName id="5FA1DB1E268447FC5054CB47EA80C90A" baseAuthorityName="Duellman" baseAuthorityYear="1978" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis toftae" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="toftae">
<emphasis id="0B080CC6F28B70B8EB50AFFF73AAB72F" italics="true" pageId="0" pageNumber="1">Pristimantis toftae</emphasis>
</taxonomicName>
is a smaller species that is superficially similar to
<taxonomicName id="980FFA8FB67F0B03F38CC807D08A28B4" lsidName="P. similaris" pageId="0" pageNumber="1" rank="species" species="similaris">
<emphasis id="CD3682D9146B7FBBFF0DCAA19AD4A52F" italics="true" pageId="0" pageNumber="1">P. similaris</emphasis>
</taxonomicName>
. The new species can be distinghished by the absence of coloured marks or spots in the groin or other parts of its body (yellow spot in the groins of
<taxonomicName id="8D9D377492F134114DD799D22831E5DC" lsidName="P. toftae" pageId="0" pageNumber="1" rank="species" species="toftae">
<emphasis id="4452F84F93398AA236E46DC59AB2426E" italics="true" pageId="0" pageNumber="1">P. toftae</emphasis>
</taxonomicName>
), absence of labial bar (presence of a white labial bar in
<taxonomicName id="8864F365A8496370F7853B3B77D22268" lsidName="P. toftae" pageId="0" pageNumber="1" rank="species" species="toftae">
<emphasis id="D7BE3D23AF7E8BEC5924AEBE90F76B87" italics="true" pageId="0" pageNumber="1">P. toftae</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="94E3C97071612D82A72A690515D21C57" pageId="0" pageNumber="1">
<taxonomicName id="5C625413A6ECD4BE8B27EA37A01B6717" authorityName="Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano" authorityYear="2023" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis similaris" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="similaris">
<emphasis id="80D78910343AE8990F0AC3176BFACCA6" italics="true" pageId="0" pageNumber="1">Pristimantis similaris</emphasis>
</taxonomicName>
is also similar to some species in the
<taxonomicName id="302CE35FEE9F4B3B03F821D785B4DEB0" baseAuthorityName="Gunther" baseAuthorityYear="1858" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis conspicillatus" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="conspicillatus">
<emphasis id="39B5C44ADDFC6248C019132DF1C5E70C" italics="true" pageId="0" pageNumber="1">Pristimantis conspicillatus</emphasis>
</taxonomicName>
species Group, which includes
<taxonomicName id="FA5C65DCE2ACB63B7E67294C216DBE01" lsidName="P. bipunctatus" pageId="0" pageNumber="1" rank="species" species="bipunctatus">
<emphasis id="3AF927151794739EBBB323DC2A4334EA" italics="true" pageId="0" pageNumber="1">P. bipunctatus</emphasis>
</taxonomicName>
,
<taxonomicName id="65D5C7992ABB1F7007C532B9734F87A4" lsidName="P. skydmainos" pageId="0" pageNumber="1" rank="species" species="skydmainos">
<emphasis id="2DCC7BA9480F0E3521356CC5B6DBCC2B" italics="true" pageId="0" pageNumber="1">P. skydmainos</emphasis>
</taxonomicName>
and
<taxonomicName id="BD00BEB095FB6C3298FF6A22CB597800" lsidName="P. iiap" pageId="0" pageNumber="1" rank="species" species="iiap">
<emphasis id="197C33843EC6D13C5872EA78DEA80C0C" italics="true" pageId="0" pageNumber="1">P. iiap</emphasis>
</taxonomicName>
. The parapatric
<taxonomicName id="93FA2E94DDA2FAD4F1F40354F2235E3A" baseAuthorityName="Duellman &amp; Hedges" baseAuthorityYear="2005" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis bipunctatus" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="bipunctatus">
<emphasis id="0167136B6FA4FD925CBD411B0CDF78D7" italics="true" pageId="0" pageNumber="1">Pristimantis bipunctatus</emphasis>
</taxonomicName>
(found in Calicanto at 1940 m. a.s.l. in the Piene Valley, Ayacucho Region), has dorsum and ventral skin shagreen and areolate, snout long, upper eyelids without tubercules similar to
<taxonomicName id="73CFF859D06DF50485C384CF4A847A37" lsidName="P. similaris" pageId="0" pageNumber="1" rank="species" species="similaris">
<emphasis id="63F1AC7E78627EDEF9007C172832FC99" italics="true" pageId="0" pageNumber="1">P. similaris</emphasis>
</taxonomicName>
, but the latter differs by having finger I slightly shorter than finger II (finger I and finger II about equal length in
<taxonomicName id="06A4B7FA6727D71EF820CFB16C2DDF92" lsidName="P. bipunctatus" pageId="0" pageNumber="1" rank="species" species="bipunctatus">
<emphasis id="1A7622396ECF89AC74CCEABE432F0CE3" italics="true" pageId="0" pageNumber="1">P. bipunctatus</emphasis>
</taxonomicName>
), discs on outer fingers truncated (broadly rounded in
<taxonomicName id="44EAEEA94229AC8BABA564A1B751DD96" lsidName="P. bipunctatus" pageId="0" pageNumber="1" rank="species" species="bipunctatus">
<emphasis id="1A3CBD84780B4E84326C99F825A33580" italics="true" pageId="0" pageNumber="1">P. bipunctatus</emphasis>
</taxonomicName>
), scapular tubercules absent (present in
<taxonomicName id="23961A170FD8811BE12565B66143EC39" lsidName="P. bipunctatus" pageId="0" pageNumber="1" rank="species" species="bipunctatus">
<emphasis id="9661CCBA28D04203BDF7EAED2EFB882D" italics="true" pageId="0" pageNumber="1">P. bipunctatus</emphasis>
</taxonomicName>
) and by its smaller size (22.6-28.8 mm in males and 32.4-41.5 mm in females in
<taxonomicName id="434C38BB5C2C875CE7FBD5647A1B9E19" lsidName="P. bipunctatus" pageId="0" pageNumber="1" rank="species" species="bipunctatus">
<emphasis id="02D450B52EAF79FACD60C6AE11AA7157" italics="true" pageId="0" pageNumber="1">P. bipunctatus</emphasis>
</taxonomicName>
).
<taxonomicName id="20933B21493373F5DF6092646B56176D" lsidName="P. similaris" pageId="0" pageNumber="1" rank="species" species="similaris">
<emphasis id="7841F6D6C2569F85B18BEEDDE78571D3" italics="true" pageId="0" pageNumber="1">P. similaris</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="450BE230B08D8E0288B9909E6F788A0A" lsidName="P. skydmainos" pageId="0" pageNumber="1" rank="species" species="skydmainos">
<emphasis id="F907E01062CDF622B85DEB5D1F2A3D99" italics="true" pageId="0" pageNumber="1">P. skydmainos</emphasis>
</taxonomicName>
by the absence of a mid-dorsal tubercle (present in
<taxonomicName id="F9998A762DF2F46CAAEDCF1582B03655" lsidName="P. skydmainos" pageId="0" pageNumber="1" rank="species" species="skydmainos">
<emphasis id="1726B4F259BEF7D2FFCD14F4C260BF7C" italics="true" pageId="0" pageNumber="1">P. skydmainos</emphasis>
</taxonomicName>
), absence of nuptial pads (present in
<taxonomicName id="F99C647092B4984486B38AD04C797140" lsidName="P. skydmainos" pageId="0" pageNumber="1" rank="species" species="skydmainos">
<emphasis id="7B3486B388B2062D39FA3CFE957981E6" italics="true" pageId="0" pageNumber="1">P. skydmainos</emphasis>
</taxonomicName>
), finger I smaller than finger II (finger I longer than finger II in
<taxonomicName id="A72099D842416C1BDBE82EC282382F46" lsidName="P. skydmainos" pageId="0" pageNumber="1" rank="species" species="skydmainos">
<emphasis id="CC51FD35FF55B07EE239720D8FB3FC91" italics="true" pageId="0" pageNumber="1">P. skydmainos</emphasis>
</taxonomicName>
), absence of spots or marks in the posterior surfaces of the thighs (minute cream flecks on the posterior surfaces of the thighs in
<taxonomicName id="400EDFEBF7A34861FFDE80B742E69461" lsidName="P. skydmainos" pageId="0" pageNumber="1" rank="species" species="skydmainos">
<emphasis id="7912521ADAB80EDE632A619C2C9E4742" italics="true" pageId="0" pageNumber="1">P. skydmainos</emphasis>
</taxonomicName>
) and the absence of W-shaped marks (present in the scapular region in
<taxonomicName id="9460F6B4740504B784AF8B3DEBAC0905" lsidName="P. skydmainos" pageId="0" pageNumber="1" rank="species" species="skydmainos">
<emphasis id="E17A98C4ED93B24CADC6D6B51678B183" italics="true" pageId="0" pageNumber="1">P. skydmainos</emphasis>
</taxonomicName>
).
<taxonomicName id="BD4F955896324651F18FC48BE586541D" authorityName="Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano" authorityYear="2023" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis similaris" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="similaris">
<emphasis id="61E8D364B187A9C8C176D598A994BF5A" italics="true" pageId="0" pageNumber="1">Pristimantis similaris</emphasis>
</taxonomicName>
differs from
<taxonomicName id="5589435C5B18834B0C1FF272EBE73A01" lsidName="P. iiap" pageId="0" pageNumber="1" rank="species" species="iiap">
<emphasis id="BF0C550201601C6EA521D46CD0B80B07" italics="true" pageId="0" pageNumber="1">P. iiap</emphasis>
</taxonomicName>
from the lowland Amazon of the Ucayali Region by lacking large granules on flanks (present in
<taxonomicName id="B2659EF4474DBB0732BC4B404C572074" lsidName="P. iiap" pageId="0" pageNumber="1" rank="species" species="iiap">
<emphasis id="72D5F08A1C57F890513F90E3BA858BBD" italics="true" pageId="0" pageNumber="1">P. iiap</emphasis>
</taxonomicName>
), lacking granules on the upper eyelids (present in
<taxonomicName id="9E0C904AB98B638047D553AC9F1C3799" lsidName="P. iiap" pageId="0" pageNumber="1" rank="species" species="iiap">
<emphasis id="5A6774D3725437CD0AEF02E7E7D2C158" italics="true" pageId="0" pageNumber="1">P. iiap</emphasis>
</taxonomicName>
) and by having finger I shorter than finger II (finger I and II about the same length in
<taxonomicName id="12B6E9C6A6370BADAE3AF9C378D3A7F1" lsidName="P. iiap" pageId="0" pageNumber="1" rank="species" species="iiap">
<emphasis id="450AF5A9B92E382EB001EC431312451F" italics="true" pageId="0" pageNumber="1">P. iiap</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="E41E61D9CAF1AA3E7410854C70B53F75" pageId="0" pageNumber="1">
Another species with some resemblance (mainly on the throat in ventral view) to the new species is
<taxonomicName id="805112D0780E24E9B5DF52907525B58A" authorityName="Lehr" authorityYear="2007" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis tanyrhynchus" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="tanyrhynchus">
<emphasis id="76D3F55E3BE332D4D66F6CCF95194556" italics="true" pageId="0" pageNumber="1">Pristimantis tanyrhynchus</emphasis>
</taxonomicName>
.
<taxonomicName id="56A882658FA74E33EB5B695DD547D80D" authorityName="Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano" authorityYear="2023" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis similaris" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="similaris">
<emphasis id="DA436EDAF0CCD4E7DCFE323FFA2B46F0" italics="true" pageId="0" pageNumber="1">Pristimantis similaris</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="FD705D645D6C3612B4094B994DB0EEBC" lsidName="P. tanyrhynchus" pageId="0" pageNumber="1" rank="species" species="tanyrhynchus">
<emphasis id="B4856B21795F2BDFB9CBF79895888432" italics="true" pageId="0" pageNumber="1">P. tanyrhynchus</emphasis>
</taxonomicName>
by the absence of nuptial pads in males (present in
<taxonomicName id="4A5C2D572151B7187319F9EFC8BBBF37" lsidName="P. tanyrhynchus" pageId="0" pageNumber="1" rank="species" species="tanyrhynchus">
<emphasis id="A3E9AC8427BB26148060A159D18FB6D5" italics="true" pageId="0" pageNumber="1">P. tanyrhynchus</emphasis>
</taxonomicName>
) and absence of tubercles on the heel (heel with a conical and large tubercle in
<taxonomicName id="0557DAD72708A9B032B5836DE1D889E1" lsidName="P. tanyrhynchus" pageId="0" pageNumber="1" rank="species" species="tanyrhynchus">
<emphasis id="ADC4E40334311196314716E4A42A41D0" italics="true" pageId="0" pageNumber="1">P. tanyrhynchus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="7FA15F9E6ECAFC899C554B709666FD55" pageId="0" pageNumber="1" type="description">
<paragraph id="5DF61B1465C135BABE9481B67F709B1E" pageId="0" pageNumber="1">Description of the holotype.</paragraph>
<paragraph id="BB5FF34E928F9BDA16659795AE8F950A" pageId="0" pageNumber="1">
Head longer than wide; head length 43% of SVL; head width 35% of SVL; cranial crests absent; snout subaccuminated in dorsal view and in lateral view (Fig.
<figureCitation id="C084E7FCE5292CC924977D87463C4861" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pristimantis similaris sp. nov. (A-D) male. SVL: 17.0 mm. Holotype. MUSM 41030. Photos by Vladimir Diaz-Vargas." figureDoi="10.3897/zookeys.1187.104536.figure4" httpUri="https://binary.pensoft.net/fig/955206" pageId="0" pageNumber="1">4A, B, D</figureCitation>
); eye-nostril distance same as the eye diameter; nostrils slightly protuberant, directed dorsolaterally; canthus rostralis long, straight in lateral and in dorsal views; loreal region concave; upper eyelid without tubercles, width 90% of IOD (see photo in life Fig.
<figureCitation id="3172A59AD9C34A82D40103E8451D87C5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pristimantis similaris sp. nov. (A-D) male. SVL: 17.0 mm. Holotype. MUSM 41030. Photos by Vladimir Diaz-Vargas." figureDoi="10.3897/zookeys.1187.104536.figure4" httpUri="https://binary.pensoft.net/fig/955206" pageId="0" pageNumber="1">4</figureCitation>
); supratympanic fold short and narrow, extending from posterior margin of upper eyelid slightly curved to insertion of arm; tympanic membrane and annulus present; distinct conical postrictal tubercles present bilaterally. Choanae small, ovoid, not concealed by palatal shelf of maxilla; dentigerous processes of vomers absent; vocal slits present; tongue longer than short, oval, about a quarter times as long as wide, notched posteriorly, half of the tongue posteriorly free; one large vocal sac extending on to chest.
</paragraph>
<paragraph id="01EC6A5D28E9F91E6F84F22905C4018B" pageId="0" pageNumber="1">Skin on dorsum and flanks shagreen, continuous dorsolateral folds present extending from posterior level of tympanic area to level of hind limb insertion; skin on throat, chest and belly areolate; discoidal fold present; thoracic fold present.</paragraph>
<paragraph id="461D819306B90F9F62363C6882574185" pageId="0" pageNumber="1">
Outer ulnar surface without tubercles; palmar tubercle bifid; thenar tubercle ovoid; subarticular tubercles well defined, most prominent on base of fingers, ovoid in ventral view, subconical in lateral view; supernumerary tubercles indistinct; fingers long and thin lacking lateral fringes, Finger I shorter than Finger II; tips of digits of fingers expanded, truncated, with circumferential grooves; nuptial pads absent (Fig.
<figureCitation id="C753C8394D62544F9291ECE2364CBAE1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pristimantis similaris sp. nov. A hand B toe. Male. Holotype. MUSM 41030. Photos by VHA." figureDoi="10.3897/zookeys.1187.104536.figure5" httpUri="https://binary.pensoft.net/fig/955207" pageId="0" pageNumber="1">5A</figureCitation>
).
</paragraph>
<paragraph id="08C27CCCAD16FED8B1B4D9E0DA929BB1" pageId="0" pageNumber="1">
Hind limbs long, slender, tibia length 58% of SVL; foot length 49% of SVL; dorsal surfaces of hind limbs tuberculate; inner surface of thighs smooth, posterior surfaces of thighs shagreen, ventral surfaces of thighs smooth; heels each with three small conical tubercles; outer surface of tarsus with one minute low tubercle; inner tarsal fold present; inner metatarsal tubercle ovoid, two times the size of round outer metatarsal tubercle; subarticular tubercles well defined, ovoid in ventral view, subconical in lateral view; few plantar supernumerary tubercles, about one quarter the size of subarticular tubercles; toes without lateral fringes; basal webbing absent; tips of digits expanded, truncated, less expanded than those on fingers, with circumferential grooves; relative length of toes: 1 &lt;2 &lt;3 &lt;5 &lt;4; Toe V slightly longer than Toe III (tip of digit of Toe III and Toe V not reaching distal subarticular tubercle on Toe IV; Fig.
<figureCitation id="547913AE41F1836C2807EC0CA2D2E937" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pristimantis similaris sp. nov. A hand B toe. Male. Holotype. MUSM 41030. Photos by VHA." figureDoi="10.3897/zookeys.1187.104536.figure5" httpUri="https://binary.pensoft.net/fig/955207" pageId="0" pageNumber="1">5B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7D00E895BEABDB9460DC8C5814A08910" pageId="0" pageNumber="1" type="description">
<paragraph id="5C486CE4B0F8BA8316D3CBE00BC968D4" pageId="0" pageNumber="1">Measurements (in mm) of the holotype.</paragraph>
<paragraph id="BCE9E1E442F53C08EB19FB750118EAD0" pageId="0" pageNumber="1">SVL 17.0; tibia length 9.9; foot length 8.4; head length 7.3; head width 5.9; eye diameter 2.3; inter orbital distance 1.9; upper eyelid width 1.7; internarial distance 2.0; eye-nostril distance 2.3; tympanum length 1.0; tympanum height 1.1; forearm length 4.3.</paragraph>
</subSubSection>
<subSubSection id="46DDC536430FA67501DCB3B333381BE1" pageId="0" pageNumber="1" type="colouration of the holotype in life">
<paragraph id="3D28D4FB2A39C9179674C1D1E390FBA4" pageId="0" pageNumber="1">Colouration of the holotype in life</paragraph>
<paragraph id="561EF88C4E5A5A207A22089F73434D75" pageId="0" pageNumber="1">
<emphasis id="BC3C69B76D51780938742840CEB0FDF9" bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation id="51CB31E642FE1BE12C97AAAD4012262E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pristimantis similaris sp. nov. (A-D) male. SVL: 17.0 mm. Holotype. MUSM 41030. Photos by Vladimir Diaz-Vargas." figureDoi="10.3897/zookeys.1187.104536.figure4" httpUri="https://binary.pensoft.net/fig/955206" pageId="0" pageNumber="1">4</figureCitation>
).
</emphasis>
In life, dorsum dark brown with three conspicuous chevrons; flanks cream without tubercles, groin same pattern as flanks with orange-reddish colouration; anterior surfaces of thighs reddish-orange, posterior surfaces of thighs brown; venter cream to yellow with black conspicuous reticulations in the throat and black marks in the chest; iris dark copper-coloured with fine black vermiculations (Fig.
<figureCitation id="A5AC6D9AEB13DB0F2B3A860F742E1A37" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pristimantis similaris sp. nov. (A-D) male. SVL: 17.0 mm. Holotype. MUSM 41030. Photos by Vladimir Diaz-Vargas." figureDoi="10.3897/zookeys.1187.104536.figure4" httpUri="https://binary.pensoft.net/fig/955206" pageId="0" pageNumber="1">4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A41BD02365BED06684C30169361DBF61" pageId="0" pageNumber="1" type="colouration of the holotype in preservative">
<paragraph id="F2429087B45CE5A70BED6B3829AD9A1E" pageId="0" pageNumber="1">Colouration of the holotype in preservative.</paragraph>
<paragraph id="0FD93FF4A9BEFFD07D2D9CEF95DFA1E2" pageId="0" pageNumber="1">The dorsal ground colouration is pale brown with three browner chevrons; narrow blackish canthal and supratympanic stripes; flanks pale brown with dark brown and cream flecks forming irregularly-shaped diagonal bars; groin and anterior surfaces of thighs brown with dark brown flecks; chest, belly and ventral surfaces of thighs pale cream, throat pale cream and grey-striped; palmar and plantar surfaces and fingers and toes dark brown; iris pale grey.</paragraph>
</subSubSection>
<subSubSection id="CE4B0935B7C9C1C51B5CF9832381EE34" pageId="0" pageNumber="1" type="variation">
<paragraph id="69048009BA63B236968570664DEBBD56" pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph id="8EDF7650FE0692DE59EB11A226562A47" pageId="0" pageNumber="1">
All specimens have the same general appearance, with three chevrons on the dorsum. MUSM 41029 is completely yellow and lacks marks on the chest or throat. MUSM 41032 has two brown-yellowish longitudinal bars on the dorsolateral folds. MUSM 41341 is blackish-brown and the three chevrons are not very visible (Fig.
<figureCitation id="F8F5446C865EA06D2E8EEAF9F4632521" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-H colour and pattern variation of Pristimantis similaris sp. nov. Specimen from A-F collected in Cajadela: A, B male MUSM 41029 C, D female MUSM 41031 E, F female MUSM 41032. Specimen G, H male MUSM 41326 collected in Machente. Photos by V. Diaz-Vargas and E. Castillo-Urbina." figureDoi="10.3897/zookeys.1187.104536.figure6" httpUri="https://binary.pensoft.net/fig/955208" pageId="0" pageNumber="1">6</figureCitation>
). Some individuals (MUSM 41030-2, 41036-7) lack the dentigerous processes of vomers. Morphological measurements ranges and proportions of types are included in Tables
<tableCitation id="BF219152E7CE43BEB7C9393E30B8E5FE" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological measurements (mm) of nine paratype specimens of Pristimantis similaris sp. nov. For abbreviations, see Materials and methods." httpUri="http://table.plazi.org/id/BED4CBB6EDB4228D532F1107578556A0" pageId="0" pageNumber="1" tableUuid="BED4CBB6EDB4228D532F1107578556A0">2</tableCitation>
,
<tableCitation id="770BEE17DCD4762FAFD8743D50D61FE7" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Measurements (in mm) and proportions of adult male and female type specimens of Pristimantis similaris sp. nov.; ranges followed by means and one standard deviation in parentheses. For abbreviations, see Materials and methods." httpUri="http://table.plazi.org/id/C6E77729A15B49B9C8BA7BADA2469F1E" pageId="0" pageNumber="1" tableUuid="C6E77729A15B49B9C8BA7BADA2469F1E">3</tableCitation>
.
</paragraph>
<caption id="BED4CBB6EDB4228D532F1107578556A0" ID-Table-UUID="BED4CBB6EDB4228D532F1107578556A0" httpUri="http://table.plazi.org/id/BED4CBB6EDB4228D532F1107578556A0" pageId="0" pageNumber="1" start="Table 2" startId="T2">
<paragraph id="4E90880F1E85BC35F25D5EF7DD87AE89" pageId="0" pageNumber="1">
<emphasis id="295511F9287B1B98F806E15D8046765D" bold="true" pageId="0" pageNumber="1">Table 2.</emphasis>
Morphological measurements (mm) of nine paratype specimens of
<taxonomicName id="C395960FD1D3D514EB127DD33C55B004" authorityName="Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano" authorityYear="2023" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis similaris" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="similaris">
<emphasis id="ACC458A6BE2773C16BDD35C4B701303D" italics="true" pageId="0" pageNumber="1">Pristimantis similaris</emphasis>
</taxonomicName>
sp. nov. For abbreviations, see Materials and methods.
</paragraph>
</caption>
<paragraph id="B318A29E0C42774F6B79AAA878E423CA" pageId="0" pageNumber="1">
<table id="2026D7309E32C5239B082DD74CACB9D9" pageId="0" pageNumber="1">
<tr id="16DE73769BAAE1E724DD4FEB54658003" pageId="0" pageNumber="1">
<th id="9C336C79D71D6B41B166F51886B5FD09" colspan="1" pageId="0" pageNumber="1" rowspan="1">Character</th>
<th id="01A459331FAEDAA0A6BFAE9B76F8D727" colspan="1" pageId="0" pageNumber="1" rowspan="1">MUSM 41028</th>
<th id="C2320C524D52C2C27B2C4FC4B0893147" colspan="1" pageId="0" pageNumber="1" rowspan="1">MUSM 41029</th>
<th id="56049D2171643CB28FCB3584D6727645" colspan="1" pageId="0" pageNumber="1" rowspan="1">MUSM 41031</th>
<th id="E37A7A54FEFF96266F46E06D98C65134" colspan="1" pageId="0" pageNumber="1" rowspan="1">MUSM 41032</th>
<th id="BCCE5E053C9DF1BB9D8D67943CBD9603" colspan="1" pageId="0" pageNumber="1" rowspan="1">MUSM 41033</th>
<th id="4B70B114106C19663B0CAD441F25C686" colspan="1" pageId="0" pageNumber="1" rowspan="1">MUSM 41034</th>
<th id="3E8CAB942737B4B7F5DAB1A52B10A509" colspan="1" pageId="0" pageNumber="1" rowspan="1">MUSM 41035</th>
<th id="4E79E6F5EBC9EDC7C3B5FAD5D8EA72FF" colspan="1" pageId="0" pageNumber="1" rowspan="1">MUSM 41036</th>
<th id="AAEF2604C32350F0A7EBCCFFEC440F73" colspan="1" pageId="0" pageNumber="1" rowspan="1">MUSM 41037</th>
</tr>
<tr id="364ECDFE87EE5C995872D872D10F2645" pageId="0" pageNumber="1">
<td id="044CEB554E0EC7CAEDC416D92B13065A" colspan="1" pageId="0" pageNumber="1" rowspan="1">Sex</td>
<td id="2ED9A3D60D30FF47E958CBFFE4D28253" colspan="1" pageId="0" pageNumber="1" rowspan="1">Male</td>
<td id="F813A74FEFB3BA5FC6CD01C9CD347469" colspan="1" pageId="0" pageNumber="1" rowspan="1">Male</td>
<td id="5C8E86FBA963D1C1FC01624B9DA11B7D" colspan="1" pageId="0" pageNumber="1" rowspan="1">Female</td>
<td id="AD52DB2F67C7B3F0C4C7062D14618C8D" colspan="1" pageId="0" pageNumber="1" rowspan="1">Female</td>
<td id="8550C62EA402EC3756F6A8C2A974E108" colspan="1" pageId="0" pageNumber="1" rowspan="1">Male</td>
<td id="7236B20E5C93322A23EEB67D463A9FBA" colspan="1" pageId="0" pageNumber="1" rowspan="1">Male</td>
<td id="F4D71F13036F61249CE2D5AB52861B53" colspan="1" pageId="0" pageNumber="1" rowspan="1">Female</td>
<td id="1196F6F8BB2E883D8773DCA15C1426B9" colspan="1" pageId="0" pageNumber="1" rowspan="1">Female</td>
<td id="859420FE5467FCB55080A3C7976D8C9C" colspan="1" pageId="0" pageNumber="1" rowspan="1">Female</td>
</tr>
<tr id="D97A01BADA5BF9CCF328C52A7BA62D1A" pageId="0" pageNumber="1">
<td id="F72828FF648FE27EFA572675C4A08150" colspan="1" pageId="0" pageNumber="1" rowspan="1">SVL</td>
<td id="9BAEAF5F5108AAA9DB7E1D8F399D8A2E" colspan="1" pageId="0" pageNumber="1" rowspan="1">18.1</td>
<td id="FB4BAC1701274684692E1B37B05CDCF3" colspan="1" pageId="0" pageNumber="1" rowspan="1">17.7</td>
<td id="3154BB68B178407164C00A50E7139518" colspan="1" pageId="0" pageNumber="1" rowspan="1">25.2</td>
<td id="A24BC4AC0AE2BE9E464EFDBFF9EA709D" colspan="1" pageId="0" pageNumber="1" rowspan="1">22.7</td>
<td id="4A7E0EB29A0C52EE2A945A166AE740F4" colspan="1" pageId="0" pageNumber="1" rowspan="1">18.8</td>
<td id="0FFE38276FE6E92576876360ED4F2809" colspan="1" pageId="0" pageNumber="1" rowspan="1">18.6</td>
<td id="92C4D915CCAF085628BEFAC76802CE13" colspan="1" pageId="0" pageNumber="1" rowspan="1">24.7</td>
<td id="A24669CBDCE565FF00F162FBEC59DA3B" colspan="1" pageId="0" pageNumber="1" rowspan="1">23.4</td>
<td id="73EB8AAF3DEC7A5F9E2A1584B1757D44" colspan="1" pageId="0" pageNumber="1" rowspan="1">20.8</td>
</tr>
<tr id="F20DEFC1366A5EF4747A0E617E1ACAB4" pageId="0" pageNumber="1">
<td id="FF8101ED4C9FC11E033A76AC4D82F239" colspan="1" pageId="0" pageNumber="1" rowspan="1">TL</td>
<td id="57CEBF562B381C620536422265BD2840" colspan="1" pageId="0" pageNumber="1" rowspan="1">10.3</td>
<td id="C88C83449354C0C66C5126C0888931FD" colspan="1" pageId="0" pageNumber="1" rowspan="1">10.4</td>
<td id="DC9E256A70DC633DDA6D50D761C31658" colspan="1" pageId="0" pageNumber="1" rowspan="1">14.8</td>
<td id="B62F4F980442172EF1B0594AD8D16A7E" colspan="1" pageId="0" pageNumber="1" rowspan="1">13.5</td>
<td id="C0B7001C7B5C996846774EDC74A50E10" colspan="1" pageId="0" pageNumber="1" rowspan="1">10.5</td>
<td id="185FEE6240EDC639F58616B29442F026" colspan="1" pageId="0" pageNumber="1" rowspan="1">10.8</td>
<td id="0090EE9FC109ED5DE1500380D7DAA3A4" colspan="1" pageId="0" pageNumber="1" rowspan="1">14.0</td>
<td id="879180815E6F780561544603B8F81E96" colspan="1" pageId="0" pageNumber="1" rowspan="1">14.1</td>
<td id="81696C4BDB7AFE2DBEFEC4ABFA40FD93" colspan="1" pageId="0" pageNumber="1" rowspan="1">12.1</td>
</tr>
<tr id="5876B0C4875D79F83386FBAF49C00962" pageId="0" pageNumber="1">
<td id="CE6E0BE116A6C0D53AADC1EE2AE58A4D" colspan="1" pageId="0" pageNumber="1" rowspan="1">FL</td>
<td id="ACB617232C5CCB9AF65F323BFF7629F1" colspan="1" pageId="0" pageNumber="1" rowspan="1">9.0</td>
<td id="8F4E47FB3F259FE00F9A8416D13FC176" colspan="1" pageId="0" pageNumber="1" rowspan="1">8.4</td>
<td id="69AE23B5B44BDE46F91D51E55047776A" colspan="1" pageId="0" pageNumber="1" rowspan="1">12.4</td>
<td id="21609DBFC0911268B8BCA62853BAFDC3" colspan="1" pageId="0" pageNumber="1" rowspan="1">11.3</td>
<td id="9E33E650DD21E2B164672C410DDC95B4" colspan="1" pageId="0" pageNumber="1" rowspan="1">9.0</td>
<td id="FC47E6B66515275E50330CD20DB7BC5F" colspan="1" pageId="0" pageNumber="1" rowspan="1">9.5</td>
<td id="205D47E13E60C14282FD50C11B5F088D" colspan="1" pageId="0" pageNumber="1" rowspan="1">11.6</td>
<td id="CB1D26E24712C2346777215AF5E5CEF1" colspan="1" pageId="0" pageNumber="1" rowspan="1">12.4</td>
<td id="6C4E8A7C9E3A1A3BD7ACDF3BFC7097B2" colspan="1" pageId="0" pageNumber="1" rowspan="1">9.4</td>
</tr>
<tr id="631F4404C5F285CD9155D4EED6044D2A" pageId="0" pageNumber="1">
<td id="0D07251A0A4BEAC1D98A3D5A3DF45121" colspan="1" pageId="0" pageNumber="1" rowspan="1">HL</td>
<td id="00DF5B138F1DAACD96A6717D350A52F3" colspan="1" pageId="0" pageNumber="1" rowspan="1">7.2</td>
<td id="1834500AE507B3AA6F6E540DF9D0D667" colspan="1" pageId="0" pageNumber="1" rowspan="1">7.2</td>
<td id="74F9A8F10C103E348ECB7CF754A9D823" colspan="1" pageId="0" pageNumber="1" rowspan="1">9.6</td>
<td id="860ED72258BF3F2983B239DDF67FE206" colspan="1" pageId="0" pageNumber="1" rowspan="1">8.9</td>
<td id="29503B47C698855554F63402E3442ED3" colspan="1" pageId="0" pageNumber="1" rowspan="1">7.3</td>
<td id="6E2A0692E151624D49396F46239631F2" colspan="1" pageId="0" pageNumber="1" rowspan="1">7.9</td>
<td id="27C6DE9E94348417132A61294335F53E" colspan="1" pageId="0" pageNumber="1" rowspan="1">9.8</td>
<td id="7265C779DD79574C0AB911F57421441A" colspan="1" pageId="0" pageNumber="1" rowspan="1">9.2</td>
<td id="8B83FEA99997E297DC0A957A888B883E" colspan="1" pageId="0" pageNumber="1" rowspan="1">8.9</td>
</tr>
<tr id="E0BE412B58C98DAD9D40AC0957F3920E" pageId="0" pageNumber="1">
<td id="05B46DC47753AF108C6607978E204354" colspan="1" pageId="0" pageNumber="1" rowspan="1">HW</td>
<td id="554394237F400D3ECF10843754C838B4" colspan="1" pageId="0" pageNumber="1" rowspan="1">5.9</td>
<td id="5EDE48A530A2B9226D8C9E42C3151B46" colspan="1" pageId="0" pageNumber="1" rowspan="1">6.5</td>
<td id="E4F983B86B8CC8E26BA893F5805A390F" colspan="1" pageId="0" pageNumber="1" rowspan="1">8.5</td>
<td id="1DA26B429D5883685163EB5391B36059" colspan="1" pageId="0" pageNumber="1" rowspan="1">7.5</td>
<td id="1AD1237540E87E33AE248888DC51BF20" colspan="1" pageId="0" pageNumber="1" rowspan="1">6.6</td>
<td id="BC152BEF77F60861213F1C2C80B4B513" colspan="1" pageId="0" pageNumber="1" rowspan="1">6.9</td>
<td id="4F95DD31C1A5FEF33ADA7413D51C1BD7" colspan="1" pageId="0" pageNumber="1" rowspan="1">8.2</td>
<td id="149151A758E2CFB3188824996FE6869A" colspan="1" pageId="0" pageNumber="1" rowspan="1">8.0</td>
<td id="25E264DAD3520158F933138D64A95F18" colspan="1" pageId="0" pageNumber="1" rowspan="1">7.5</td>
</tr>
<tr id="EF147A922FC610B82413A929913D93A3" pageId="0" pageNumber="1">
<td id="33276EECA18CD42966D9CEBB9A0ABED6" colspan="1" pageId="0" pageNumber="1" rowspan="1">ED</td>
<td id="D81667B90C0ADF691F516DF9FF434EF6" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.9</td>
<td id="1E9F3EA4507093AC3D19D70F86F1383E" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.4</td>
<td id="A9F4E2F3457FA099E8344EF311EA2A84" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.8</td>
<td id="BF5960E66FAEBC6E81930A84D41B17FD" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.5</td>
<td id="71CD34025622428182E10681D2C81816" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.3</td>
<td id="023D28D19F4738DA5B287F697D9C5694" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.3</td>
<td id="98F059E3F609F6A4C61F47B731B6CC8A" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.9</td>
<td id="A119C31016E946A64430DE27780BF5C0" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.6</td>
<td id="7BC97D584A766A084991F0EADACB92A8" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.4</td>
</tr>
<tr id="F3AFF4D700F0D11472F4EDF13F97CBFA" pageId="0" pageNumber="1">
<td id="D75351A23908B8BC8EB05BC1CEC6BA8B" colspan="1" pageId="0" pageNumber="1" rowspan="1">TY</td>
<td id="63C9DB98DA101D089167F97091FE6B08" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.9</td>
<td id="B609586B301399BE9D870FBB1ED7F29A" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.0</td>
<td id="4D54426C469AE3EE1D2BF061D4660EA4" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.1</td>
<td id="C5693DEDF51E80258E395C236CC9171C" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.1</td>
<td id="537562300E6F3A7908CF6B23E620C034" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.0</td>
<td id="3CE7C2474A84AEB0B8459E7B10944555" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.1</td>
<td id="73AD2154308DE0347761F54161471C98" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.3</td>
<td id="BF77E5077D317A9A80913A5F04553F45" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.1</td>
<td id="5FFF285D3F4AD4E5AB3143E78A4A690D" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.2</td>
</tr>
<tr id="E08C47E72B963AD0FF3A87D4F7A46CF2" pageId="0" pageNumber="1">
<td id="E9106025762B50C880E83D536F55E423" colspan="1" pageId="0" pageNumber="1" rowspan="1">IOD</td>
<td id="35A6FDC3D6A8A06BF6263D132B520842" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.1</td>
<td id="8159AEBE3A627C9B0FD9F633F8A27FC5" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.0</td>
<td id="1D603ABA0E1A7590D15818C2F6894079" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.4</td>
<td id="22540818F6F2867C489D09FCEAAB7159" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.2</td>
<td id="A4FC9A8E48ADFF94AA0FD1A416546237" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.1</td>
<td id="1CB027548574F0A3AAEBF3341B3F44EF" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.0</td>
<td id="357EF22DB9A8817D85A190BC2884511A" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.4</td>
<td id="9C8885CD948036E86BBD48AD5A826089" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.3</td>
<td id="F146E178B4749BD849207AE4F300DFEC" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.2</td>
</tr>
<tr id="EAA264D2BCBD281EA4B104B8D0C82476" pageId="0" pageNumber="1">
<td id="EB279A376F5E86D81ABE0EDB5877D466" colspan="1" pageId="0" pageNumber="1" rowspan="1">EW</td>
<td id="B3A7FF193591904765B53B4CD25915BB" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.8</td>
<td id="A0109141367963DB29F95CC01B71A388" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.7</td>
<td id="DE6E36E25A7A1099C1F0E41C6A19BF9C" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.3</td>
<td id="5C2F0EE05FA6232F178C5C07D0CC3388" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.9</td>
<td id="BA4155CE7206425230F69DA41C11392A" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.8</td>
<td id="7498D35F90C2BB3D761BE4E5A295304B" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.8</td>
<td id="8E28B36E676F816CDD398E2EA0E27BF3" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.3</td>
<td id="7E7544D028F4ADF91CB21222C44FBF81" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.2</td>
<td id="D238578467E2FC18B8C6C7C69B12F8BB" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.1</td>
</tr>
<tr id="ECB9DB0ADE2319B243F81071B991A147" pageId="0" pageNumber="1">
<td id="9BE8ECF19461E415713BD420874B1753" colspan="1" pageId="0" pageNumber="1" rowspan="1">IND</td>
<td id="BD51BED4DF8BD22EA732A02DD0150A63" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.0</td>
<td id="06699A663950DED89C114D7B686F8E7A" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.3</td>
<td id="1EE3D1480E7E3F1E2204CE966E23C519" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.9</td>
<td id="9FFF95BA3C9F118CB416A96CF098C0D9" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.6</td>
<td id="3807906A1E3EFCE483F9AB7816606588" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.2</td>
<td id="256486FBB3A265DCC13D9AA928018C11" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.2</td>
<td id="790C8DF3AC067CAE92CB004B42069EA6" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.7</td>
<td id="14281037D181E4A1C3A9AEBED2C636DD" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.5</td>
<td id="213A6D4CA03020E574734DC8727B04CB" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.5</td>
</tr>
<tr id="DD1C0EEE9F5643533AFC0CBF16936B47" pageId="0" pageNumber="1">
<td id="E0A09DC5698192F1F9FE94EE1B50CAD2" colspan="1" pageId="0" pageNumber="1" rowspan="1">EN</td>
<td id="BFBA6C432EEA015962EB79E3213169F0" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.3</td>
<td id="BF5E45F7E39B4F4D841EB08245A146CA" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.3</td>
<td id="979790DA3916B229FE1A091FD0FAE373" colspan="1" pageId="0" pageNumber="1" rowspan="1">3.1</td>
<td id="B3761562F98CA4F47A8A43F60DD84B3E" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.8</td>
<td id="24D7D16C58CB5140C8351AADA03224D2" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.5</td>
<td id="8E26F139558CA5218F1984D55CDBA214" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.6</td>
<td id="98A2691353BABCE5E8549088543F83FE" colspan="1" pageId="0" pageNumber="1" rowspan="1">3.0</td>
<td id="29BEB164AED6C05C45A663957D9435FB" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.7</td>
<td id="6EA3D64121A3B4CBFC9B5CE131C9396F" colspan="1" pageId="0" pageNumber="1" rowspan="1">2.8</td>
</tr>
<tr id="00FE62B886A90DE979B7D8E56FE6B76E" pageId="0" pageNumber="1">
<td id="DC85AF72A349F97E9828CF7FB6BD13D5" colspan="1" pageId="0" pageNumber="1" rowspan="1">F4</td>
<td id="98F7C361656C7A7CDAE65C6B19FFCA4C" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.7</td>
<td id="607544953A0469AC4C3B471F1B34B2D7" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.7</td>
<td id="6674EE7DE8F35B4D9913C9F5B9130905" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.0</td>
<td id="37B0A21CB0FF3A3A95A82BBB1A4135EA" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.8</td>
<td id="17DF6BF634F22276B9503A2732796A5B" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.9</td>
<td id="C606CAD02913A632478C56CEC39A8C39" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.7</td>
<td id="6CB1197F968631ADCCA0938E2629D6AF" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.9</td>
<td id="37713D4EF84BB3844C484E8B5731D2A7" colspan="1" pageId="0" pageNumber="1" rowspan="1">1.0</td>
<td id="0364A1D46BEC084D0169D937E3A7F9F3" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.8</td>
</tr>
</table>
</paragraph>
<caption id="C6E77729A15B49B9C8BA7BADA2469F1E" ID-Table-UUID="C6E77729A15B49B9C8BA7BADA2469F1E" httpUri="http://table.plazi.org/id/C6E77729A15B49B9C8BA7BADA2469F1E" pageId="0" pageNumber="1" start="Table 3" startId="T3">
<paragraph id="E4A41A38AE0AE99BD5B7F9046BA96CDD" pageId="0" pageNumber="1">
<emphasis id="43A58025ACD316C755B6AC46E0ADC671" bold="true" pageId="0" pageNumber="1">Table 3.</emphasis>
Measurements (in mm) and proportions of adult male and female type specimens of
<taxonomicName id="6D67F2370ABA8177D7E2C06F5E37E91F" authorityName="Herrera-Alva &amp; Catenazzi &amp; Aguilar-Puntriano" authorityYear="2023" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis similaris" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="similaris">
<emphasis id="E49985C853FB5908884A31729DDA223F" italics="true" pageId="0" pageNumber="1">Pristimantis similaris</emphasis>
</taxonomicName>
sp. nov.; ranges followed by means and one standard deviation in parentheses. For abbreviations, see Materials and methods.
</paragraph>
</caption>
<paragraph id="37A583C43F220FC2DBB23DBC846DFA7C" pageId="0" pageNumber="1">
<table id="705F98A8B70EC3FBDD86D84419E8C865" pageId="0" pageNumber="1">
<tr id="F6D05F630847260BA6AC78649325E259" pageId="0" pageNumber="1">
<th id="93B7C0E95E057BCBB71800B8262795BA" colspan="1" pageId="0" pageNumber="1" rowspan="1">Character</th>
<th id="D0C93F20FD8C7F5E69C13422F3CDD01E" colspan="1" pageId="0" pageNumber="1" rowspan="1">Males (n = 5)</th>
<th id="474BF3ACB213314B6FBDCF2943B520E3" colspan="1" pageId="0" pageNumber="1" rowspan="1">Females (n = 5)</th>
</tr>
<tr id="6F8BE987E3A0FD9018045F43CB2C6F52" pageId="0" pageNumber="1">
<td id="B3F75694DF2FDBC369BC266D0E9FF593" colspan="1" pageId="0" pageNumber="1" rowspan="1">SVL</td>
<td id="BA3ED53B00D91E3249A6FD04B30693B9" colspan="1" pageId="0" pageNumber="1" rowspan="1">
17.0-18.6 (18.1
<normalizedToken id="EF44CE29CE0ACA1B2064D8C84DBCB4D4" originalValue="±">+/-</normalizedToken>
0.7)
</td>
<td id="2CC02166228FAD7025FE5FE387EE73C8" colspan="1" pageId="0" pageNumber="1" rowspan="1">
20.8-25.2 (23.4
<normalizedToken id="2A779FC7385ADCFA7CF6207C67A6404D" originalValue="±">+/-</normalizedToken>
1.8)
</td>
</tr>
<tr id="A2A83B61C013FECDFC6BEBDAC5155D02" pageId="0" pageNumber="1">
<td id="E178DA64D7508DACADB9C2F0B09FE1E5" colspan="1" pageId="0" pageNumber="1" rowspan="1">TL</td>
<td id="96E9C7E0AA3700C17B8FBB23A564C9A1" colspan="1" pageId="0" pageNumber="1" rowspan="1">
9.9-10.8 (10.4
<normalizedToken id="18E4A2637CF1360312E095A8773BF102" originalValue="±">+/-</normalizedToken>
0.3)
</td>
<td id="D65913C12DB46D973C54C1803ADAB2F7" colspan="1" pageId="0" pageNumber="1" rowspan="1">
12.1-14.8 (13.7
<normalizedToken id="E88D812A3CBE7E4189045148E92F8E97" originalValue="±">+/-</normalizedToken>
1.0)
</td>
</tr>
<tr id="193B114C1F735DF749EE8FCA2297F3BE" pageId="0" pageNumber="1">
<td id="3E2BB3497284540E7018AE102982258C" colspan="1" pageId="0" pageNumber="1" rowspan="1">FL</td>
<td id="06B52D90010B8B622772BCE4AD0DA535" colspan="1" pageId="0" pageNumber="1" rowspan="1">
8.4-9.5 (8.9
<normalizedToken id="237D199EFDC788AB4ED9125C31665C05" originalValue="±">+/-</normalizedToken>
0.5)
</td>
<td id="DFE2B945379A9DFE3A96301BBF318820" colspan="1" pageId="0" pageNumber="1" rowspan="1">
9.4-12.4 (11.4
<normalizedToken id="24996D5FCC90F2CA3A13EACF904AD93D" originalValue="±">+/-</normalizedToken>
1.2)
</td>
</tr>
<tr id="5613F842BCD7865E8AC0F53925EF2FAD" pageId="0" pageNumber="1">
<td id="70CFBB87EE8E300709E1FC015D97B01D" colspan="1" pageId="0" pageNumber="1" rowspan="1">HL</td>
<td id="5781DFB06B5D69D728303AC5C71DB5C2" colspan="1" pageId="0" pageNumber="1" rowspan="1">
7.2-7.9 (7.4
<normalizedToken id="7FBFA83ADECF42A080AB02E49B604F0C" originalValue="±">+/-</normalizedToken>
0.7)
</td>
<td id="0C4B89B7C684966FE38193673635706B" colspan="1" pageId="0" pageNumber="1" rowspan="1">
8.9-9.8 (9.3
<normalizedToken id="0F6605D441EEC72D729A692379C1264B" originalValue="±">+/-</normalizedToken>
0.4)
</td>
</tr>
<tr id="BE6C4DA2F198E84C5F05F13325015242" pageId="0" pageNumber="1">
<td id="31E6AFC7DE20506F6A2F1301ED871C9E" colspan="1" pageId="0" pageNumber="1" rowspan="1">HW</td>
<td id="FE5400CE575987A5A5E8B8408B46CA7F" colspan="1" pageId="0" pageNumber="1" rowspan="1">
5.9-6.9 (6.5
<normalizedToken id="AE3EC17DAE171233AEDDF7E5775B7089" originalValue="±">+/-</normalizedToken>
1.0)
</td>
<td id="F322CBA51AA2C3D2075152DB918C65AC" colspan="1" pageId="0" pageNumber="1" rowspan="1">
7.5-8.5 (7.9
<normalizedToken id="3F6E41F02CCB5B08AF9394E84C14D5AD" originalValue="±">+/-</normalizedToken>
0.4)
</td>
</tr>
<tr id="C3A2EBFFC357BBD94FABF31D3576B8B5" pageId="0" pageNumber="1">
<td id="39A5EB1C351C95E9B13FDE0884B31D43" colspan="1" pageId="0" pageNumber="1" rowspan="1">ED</td>
<td id="91896FF2CDD53D6D414A21BB24755B85" colspan="1" pageId="0" pageNumber="1" rowspan="1">
1.9-2.4 (2.2
<normalizedToken id="8C6F620D604E7B97714272A0363D4F4F" originalValue="±">+/-</normalizedToken>
0.5)
</td>
<td id="4EFE66ECCED666CF074A86D83E2AACA4" colspan="1" pageId="0" pageNumber="1" rowspan="1">
2.4-2.9 (2.6
<normalizedToken id="ABB80ED0DF6781E9A99F8621D1E9B7B3" originalValue="±">+/-</normalizedToken>
0.2)
</td>
</tr>
<tr id="CDE004E61AFD38C77237711791E57780" pageId="0" pageNumber="1">
<td id="0EE4F0119DD22C05BC849C6D00964C25" colspan="1" pageId="0" pageNumber="1" rowspan="1">TY</td>
<td id="C408A8E26DCF334C4FE6276E4271F7CB" colspan="1" pageId="0" pageNumber="1" rowspan="1">
1.0-1.1 (1.0
<normalizedToken id="17A37220472E01CE7A6117D6691C37BA" originalValue="±">+/-</normalizedToken>
0.1)
</td>
<td id="6CF8C4450272A7618A5106D7E82C4835" colspan="1" pageId="0" pageNumber="1" rowspan="1">
1.0-1.3 (1.2
<normalizedToken id="593F892B5AFA6B83CCF4306D52DF0C25" originalValue="±">+/-</normalizedToken>
0.1)
</td>
</tr>
<tr id="67D4A7AAA4BEC35F5D9CD16D37F13AD4" pageId="0" pageNumber="1">
<td id="37143EAF9746B75223B8CE94F12AE0FF" colspan="1" pageId="0" pageNumber="1" rowspan="1">IOD</td>
<td id="E173916F50EF782CA98355508481381D" colspan="1" pageId="0" pageNumber="1" rowspan="1">
1.9-2.1 (2.0
<normalizedToken id="2F0F5464C89BAD6E8C24A69DD9ABEFF8" originalValue="±">+/-</normalizedToken>
0.1)
</td>
<td id="3D1639CC8BFC7D75E27CD491D7BF919F" colspan="1" pageId="0" pageNumber="1" rowspan="1">
2.2-2.4 (2.3
<normalizedToken id="63D1C673ED7B5044C108BDBB0BC9B6F3" originalValue="±">+/-</normalizedToken>
0.1)
</td>
</tr>
<tr id="2B61752DD91A8E47F00650FB3F6CEC18" pageId="0" pageNumber="1">
<td id="4730F789DC73987F26F1BF95F248CBE5" colspan="1" pageId="0" pageNumber="1" rowspan="1">EW</td>
<td id="4B5AB3C0590E4E25866A6800DD09B71A" colspan="1" pageId="0" pageNumber="1" rowspan="1">
1.7-1.8 (1.75
<normalizedToken id="1DD2369147540E3F30C69834CA814856" originalValue="±">+/-</normalizedToken>
0.05)
</td>
<td id="83C18490B4F2EE80A3A1642866FF75B8" colspan="1" pageId="0" pageNumber="1" rowspan="1">
1.9-2.3 (2.1
<normalizedToken id="4B7A622811879F3379043C73674A3772" originalValue="±">+/-</normalizedToken>
0.2)
</td>
</tr>
<tr id="EF5200ACAEDF315F65B2669C6A769C61" pageId="0" pageNumber="1">
<td id="55ADCBC89488C4710DAC2BE85E07B500" colspan="1" pageId="0" pageNumber="1" rowspan="1">IND</td>
<td id="776E659966B02BD4EEC05EAC6E328E2B" colspan="1" pageId="0" pageNumber="1" rowspan="1">
2.0-2.3 (2.1
<normalizedToken id="F600F8BBEE5714A2D10B8DBF2049ACB3" originalValue="±">+/-</normalizedToken>
0.1)
</td>
<td id="AC39CF14FE88201EEAEE415410DCD72A" colspan="1" pageId="0" pageNumber="1" rowspan="1">
2.5-2.9 (2.6
<normalizedToken id="AF76445D617AAA2886006611D7D62F33" originalValue="±">+/-</normalizedToken>
0.2)
</td>
</tr>
<tr id="B6085D84762BB59740C023906EA533E9" pageId="0" pageNumber="1">
<td id="4D6EBA31CC2FB15F76E8850DD8266146" colspan="1" pageId="0" pageNumber="1" rowspan="1">EN</td>
<td id="EEB05CA5F88E72B53AD2CA025F85D77D" colspan="1" pageId="0" pageNumber="1" rowspan="1">
2.3-2.6 (2.4
<normalizedToken id="05B0C04E4FE15FD500B5F0FF74D623F7" originalValue="±">+/-</normalizedToken>
0.3)
</td>
<td id="BBFFC4E16F6BBFA7D774918998F795A9" colspan="1" pageId="0" pageNumber="1" rowspan="1">
2.8-3.1 (2.9
<normalizedToken id="DF9C5673AE990D56A3F48331741A8566" originalValue="±">+/-</normalizedToken>
0.2)
</td>
</tr>
<tr id="DD325A045C6F95CD81FF10E87721668F" pageId="0" pageNumber="1">
<td id="A1B41932B7CBD8E91A3152383266FC36" colspan="1" pageId="0" pageNumber="1" rowspan="1">F4</td>
<td id="CDA52FC5B4135A7F2469D8122C93B4FC" colspan="1" pageId="0" pageNumber="1" rowspan="1">
0.7-0.9 (0.8
<normalizedToken id="EF148BC70917DE051A416CFB551B6AA6" originalValue="±">+/-</normalizedToken>
0.1)
</td>
<td id="E752330CEA873ECBB3BFF324CC8BBA2D" colspan="1" pageId="0" pageNumber="1" rowspan="1">
0.8-1.0 (0.9
<normalizedToken id="ED571B068D724F2DBA7FB217B6C073DB" originalValue="±">+/-</normalizedToken>
0.1)
</td>
</tr>
<tr id="0E7AE72E197234A7B2C609F05FCC6D4D" pageId="0" pageNumber="1">
<td id="E3136DEA3EBD4F3679B52DFC29EFC692" colspan="1" pageId="0" pageNumber="1" rowspan="1">TL/SVL</td>
<td id="458A455456DF917BE91C9D3DEC935C4D" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.56-0.59</td>
<td id="4698B5C30EC2652E506439975C49482E" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.57-0.60</td>
</tr>
<tr id="71372D88454F476A3FC29399295D04B1" pageId="0" pageNumber="1">
<td id="B56290344FA4EF0B0556E0EF7DB4AA19" colspan="1" pageId="0" pageNumber="1" rowspan="1">HL/SVL</td>
<td id="9E81AE73914911E0FDA31A1E1309A7CA" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.39-0.43</td>
<td id="2CD3F2D11FC6B6FD0F96868B4F3B33E0" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.38-0.43</td>
</tr>
<tr id="9D26CEE4D779B5D5A4C830E32687305B" pageId="0" pageNumber="1">
<td id="A6AB9BB27DA39AE72EBB195CDDD1323E" colspan="1" pageId="0" pageNumber="1" rowspan="1">EN/HL</td>
<td id="3AC76F5649E4AC2ED2267E74295FAC76" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.32-0.34</td>
<td id="9F14E13077BB308F21B540138EA9FD67" colspan="1" pageId="0" pageNumber="1" rowspan="1">0.30-0.33</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="CE11D5284EC06EFCE9AB3444621A4374" pageId="0" pageNumber="1" type="etymology">
<paragraph id="67232DD6669B131CEE623FE833F269E6" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="1AE4B48AC9FAC6852795290B0192D4FD" pageId="0" pageNumber="1">
The specific name corresponds to the Latin word
<normalizedToken id="2C8E5C19DB49DB3A82EB239B2B02158A" originalValue="“similar”">&quot;similar&quot;</normalizedToken>
. This refers to the similarity of the new species and its close phylogenetic relationship with
<taxonomicName id="D88ACE1A024FB324F0088E1ABE25AE29" lsidName="P. rhabdolaemus" pageId="0" pageNumber="1" rank="species" species="rhabdolaemus">
<emphasis id="6CED9D8C0210D6F672E263D20AF97115" italics="true" pageId="0" pageNumber="1">P. rhabdolaemus</emphasis>
</taxonomicName>
and
<taxonomicName id="5F9994291BAF5297121D18094C171024" lsidName="P. pharangobates" pageId="0" pageNumber="1" rank="species" species="pharangobates">
<emphasis id="69E24BD13F389DA4C991B724F9900CA5" italics="true" pageId="0" pageNumber="1">P. pharangobates</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="47DA85B4516E20FA961B5448ABB68971" pageId="0" pageNumber="1" type="distribution">
<paragraph id="CB6E4438B9CA61849296DD725DE21009" pageId="0" pageNumber="1">Distribution and natural history.</paragraph>
<paragraph id="98C35091AF1AA47F94F4E7FD265B4EB5" pageId="0" pageNumber="1">
The new species is only known from montane forests of Ayna and Anco in Departamento Ayacucho at elevations from 1200-2000 m a.s.l. in secondary forests (Figs
<figureCitation id="922C03E5BFCC55DD4DD9EE4EECBA17B5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Montane forest of Cajadela (A) and Machente (B), Ayacucho Department. Type locality of Pristimantis similaris sp. nov. in Cajadela (A). Note the presence of secondary forest in both localities. Photo A taken on 8 November 2021 and B, on 24 October 2022. Arrow marks the type locality." figureDoi="10.3897/zookeys.1187.104536.figure1" httpUri="https://binary.pensoft.net/fig/955203" pageId="0" pageNumber="1">1</figureCitation>
,
<figureCitation id="9293D63E9279B5EAF34D148F7E780BB0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution map of some species of the P. danae species Group in Ayacucho and Cusco Departments. Raster of altitude from 500 to 4500 m. a. s. l. (white to black). Locality of new species in red circles." figureDoi="10.3897/zookeys.1187.104536.figure3" httpUri="https://binary.pensoft.net/fig/955205" pageId="0" pageNumber="1">3</figureCitation>
). This species was found only at night after 18:00 hours, usually perching on wet leaves 0.5-1.5 m above the ground. Males call rarely and their calls are overshadowed by other male species (
<taxonomicName id="01189B56791D0352D4E4D7D1519B0E8B" baseAuthorityName="Jimenez de la Espada" baseAuthorityYear="1875" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis lacrimosus" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="lacrimosus">
<emphasis id="8117E067BCECE0D45D67E0D2D01B77EB" italics="true" pageId="0" pageNumber="1">Pristimantis lacrimosus</emphasis>
</taxonomicName>
species Group) calling louder. The species is common and appears to tolerate some human disturbance, because it was found near abandoned farms, less frequented roads and in the sourroudings of abandoned houses. Syntopic species included candidate new species in the
<taxonomicName id="D5FDA484A5FEF30283A8AD929DDC1A5F" baseAuthorityName="Jimenez de la Espada" baseAuthorityYear="1875" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis lacrimosus" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="lacrimosus">
<emphasis id="DB291FDEBBC5DC0B9994DE340D2016CB" italics="true" pageId="0" pageNumber="1">Pristimantis lacrimosus</emphasis>
</taxonomicName>
species Group and candidate new species in the
<taxonomicName id="636A033E6F9A8EF56543F9FAB623C1F7" baseAuthorityName="Boulenger" baseAuthorityYear="1903" class="Amphibia" family="Strabomantidae" genus="Pristimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristimantis platydactylus" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="platydactylus">
<emphasis id="637EEB97A16D7C198C0C8A964936E70F" italics="true" pageId="0" pageNumber="1">Pristimantis platydactylus</emphasis>
</taxonomicName>
species Group, which were more abundant than the new species. Sympatric species included frogs and toads
<taxonomicName id="3B1EEEBFE262F71AC41204C435ABF0A4" authorityName="Duellman" authorityYear="1987" class="Amphibia" family="Hemiphractidae" genus="Gastrotheca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gastrotheca pacchamama" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="pacchamama">
<emphasis id="CDDC082574638F3A4B73D05477AEF9FC" italics="true" pageId="0" pageNumber="1">Gastrotheca pacchamama</emphasis>
</taxonomicName>
,
<taxonomicName id="748EE45E8B99CE037DD88956B85332F4" baseAuthorityName="Cannatella &amp; Duellman" baseAuthorityYear="1982" class="Amphibia" family="Centrolenidae" genus="Nymphargus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nymphargus pluvialis" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="33AF6C83C69005D1C4FC88957905A833" italics="true" pageId="0" pageNumber="1">Nymphargus pluvialis</emphasis>
</taxonomicName>
,
<taxonomicName id="CDD81A9F7095CC0001D0106DC3F22171" baseAuthorityName="Duellman, De la Riva &amp; Wild" baseAuthorityYear="1997" class="Amphibia" family="Hylidae" genus="Boana" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Boana palaestes" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="palaestes">
<emphasis id="10B5C5DA4E4BE43C923531B002068726" italics="true" pageId="0" pageNumber="1">Boana palaestes</emphasis>
</taxonomicName>
,
<taxonomicName id="2DB54139CF05E736BFC366C30AC7B059" baseAuthorityName="Stejneger" baseAuthorityYear="1913" class="Amphibia" family="Bufonidae" genus="Rhinella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhinella inca" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="inca">
<emphasis id="80B5A84E285929B41FDB229CBBEDA398" italics="true" pageId="0" pageNumber="1">Rhinella inca</emphasis>
</taxonomicName>
,
<taxonomicName id="066C0E20263190BF8816B0BC86CEE458" authorityName="Caminer, Mila, Jansen, Fouquet, Venegas, Chavez, Lougheed &amp; Ron" authorityYear="2017" class="Amphibia" family="Dendrobatidae" genus="Dendropsophus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dendropsophus vraemi" order="Anura" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="vraemi">
<emphasis id="3E2EECC542ED2C23A5ACBB531D8A8110" italics="true" pageId="0" pageNumber="1">Dendropsophus vraemi</emphasis>
</taxonomicName>
and
<taxonomicName id="E209BBB271B0E9026F3A12437D5F5E5C" genus="Hyalinobactrachium" lsidName="Hyalinobactrachium bergeri" pageId="0" pageNumber="1" rank="species" species="bergeri">Hyalinobactrachium aff. bergeri</taxonomicName>
; lizards
<taxonomicName id="1DF57B1C3A61BB7333F12871008E9FCF" authorityName="O'Shaughnessy" authorityYear="1881" class="Squamata" family="Gymnophthalmidae" genus="Cercosaura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercosaura manicata" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="manicata">
<emphasis id="E44CC4E1AABDCB2DF5BA9EB2DCC73F00" italics="true" pageId="0" pageNumber="1">Cercosaura manicata</emphasis>
</taxonomicName>
,
<taxonomicName id="C399D1AA8F43D16E29FA9C3D2D866351" baseAuthorityName="Tschudi" baseAuthorityYear="1845" class="Squamata" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus crassicaudatus" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="crassicaudatus">
<emphasis id="DDCE2440ECA665C34ADB602CEC1539BA" italics="true" pageId="0" pageNumber="1">Stenocercus crassicaudatus</emphasis>
</taxonomicName>
and
<taxonomicName id="9C9FF08D1E1EAC8BA686022113F37072" authorityName="Chavez &amp; Vasquez" authorityYear="2012" class="Squamata" family="Gymnophthalmidae" genus="Potamites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamites montanicola" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="montanicola">
<emphasis id="8AE473A54D76A3FE549202EDC7B68BF7" italics="true" pageId="0" pageNumber="1">Potamites montanicola</emphasis>
</taxonomicName>
; and snakes
<taxonomicName id="0F17899FB2B1EE0B59F18148EE54F24C" authorityName="Boettger" authorityYear="1898" baseAuthorityName="Boettger" baseAuthorityYear="1898" class="Amphibia" family="Colubridae" genus="Dipsas" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dipsas peruana" order="Squamata" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="peruana">Dipsas cf. peruana</taxonomicName>
,
<taxonomicName id="27EB2BCD946D8204B75235756071F044" authorityName="Linnaeus" authorityYear="1758" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Squamata" family="Colubridae" genus="Leptodeira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptodeira annulata" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="annulata">
<emphasis id="F9D4653BB259C2E660F6C211EFF8C5F3" italics="true" pageId="0" pageNumber="1">Leptodeira annulata</emphasis>
</taxonomicName>
and
<taxonomicName id="8C1FC6D01CED4734DD5FB40D562B5F46" baseAuthorityName="Orejas-Miranda" baseAuthorityYear="1969" class="Reptilia" family="Boidae" genus="Epictia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Epictia peruviana" order="Squamata" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="peruviana">Epictia cf. peruviana</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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