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<document id="DCBB6DCC45009F8373930EB7C18A80D0" ID-DOI="10.11646/zootaxa.4039.3.3" ID-GBIF-Dataset="0ad500ca-d193-4331-90af-70a6018a5cfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="238528" ID-ZooBank="87A108DC-6967-4CAD-8F6C-DAF7ADD622CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461331934834" checkinUser="plazi" docAuthor="Irmler, Ulrich" docDate="2015" docId="07009178FF86BB2FFF4882C13AD92AE6" docLanguage="en" docName="zt04039p444.pdf" docOrigin="Zootaxa 4039 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thoracophorus perplexus Irmler, 2015, n. sp." docType="treatment" docVersion="7" lastPageNumber="441" masterDocId="FB39E900FF8FBB25FFDF840938402C7E" masterDocTitle="Four new species of the genus Thoracophorus Motschulsky, 1857 for the Neotropical region (Coleoptera: Staphylinidae: Osoriinae)" masterLastPageNumber="444" masterPageNumber="431" pageNumber="440" updateTime="1698618579127" updateUser="plazi">
<mods:mods id="226DC6D2FEB679ABF6F6A6752159A317" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="396C364BF0A3A51298AA99C72714DE11">
<mods:title id="BDF7BE69A86600591F76A140A45B5BCB">Four new species of the genus Thoracophorus Motschulsky, 1857 for the Neotropical region (Coleoptera: Staphylinidae: Osoriinae)</mods:title>
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<mods:name id="52B72CCDD1F67BEEE0E744B0A6F56EB1" type="personal">
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<mods:roleTerm id="7B6DD8F439145CF8F8854FC5ACFF8A9D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B1AFC5EC2C72D83B72AEC3A5547B0990">Irmler, Ulrich</mods:namePart>
</mods:name>
<mods:typeOfResource id="0329BA6764DD4B13222867FF2C44DDD3">text</mods:typeOfResource>
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<mods:title id="0E030062D0947CE72CFBCEF4EA4ADF9E">Zootaxa</mods:title>
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<mods:part id="21349180FCBFF81005C2D3D8BCE8DE3D">
<mods:date id="A31198D7AB2B85C23BF96F02CDC5335F">2015</mods:date>
<mods:detail id="4260A35517EE9B5BB6FE422CF7D45CE7" type="volume">
<mods:number id="D911EB798A6DD008B51ADC6B2D7581D0">4039</mods:number>
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<mods:classification id="F7F46ECDB4161D3EBAEA2E7DC629801A">journal article</mods:classification>
<mods:identifier id="3EC90BE6C6D4678ABDFCDC02AAD3F7C1" type="DOI">10.11646/zootaxa.4039.3.3</mods:identifier>
<mods:identifier id="FDC4EC54629DD3FC5F12E94E7781694F" type="GBIF-Dataset">0ad500ca-d193-4331-90af-70a6018a5cfc</mods:identifier>
<mods:identifier id="272B02E9B0D224C7BF08FA49CCA722EF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4625C1C553D8BA10B557A402221F855F" type="Zenodo-Dep">238528</mods:identifier>
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<treatment id="07009178FF86BB2FFF4882C13AD92AE6" ID-DOI="http://doi.org/10.5281/zenodo.6107945" ID-GBIF-Taxon="119644045" ID-Zenodo-Dep="6107945" LSID="urn:lsid:plazi:treatment:07009178FF86BB2FFF4882C13AD92AE6" httpUri="http://treatment.plazi.org/id/07009178FF86BB2FFF4882C13AD92AE6" lastPageId="10" lastPageNumber="441" pageId="9" pageNumber="440">
<subSubSection id="C7B373E5FF86BB2CFF4882C139372B7D" pageId="9" pageNumber="440" type="nomenclature">
<paragraph id="8F16206EFF86BB2CFF4882C13A582A9C" blockId="9.[151,536,1736,1795]" box="[151,536,1736,1762]" pageId="9" pageNumber="440">
<heading id="D45E9702FF86BB2CFF4882C13A582A9C" bold="true" box="[151,536,1736,1762]" fontSize="11" level="1" pageId="9" pageNumber="440" reason="1">
<emphasis id="BDDDFC7CFF86BB2CFF4882C13A582A9C" bold="true" box="[151,536,1736,1762]" pageId="9" pageNumber="440">
<taxonomicName id="48A95BEDFF86BB2CFF4882C1398E2A9C" ID-CoL="56KDX" box="[151,462,1736,1762]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="440" phylum="Arthropoda" rank="species" species="perplexus" status="sp. nov.">
<emphasis id="BDDDFC7CFF86BB2CFF4882C1398E2A9C" bold="true" box="[151,462,1736,1762]" italics="true" pageId="9" pageNumber="440">Thoracophorus perplexus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6EE4107FF86BB2CFE0982C13A582A9C" box="[470,536,1736,1762]" pageId="9" pageNumber="440" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8F16206EFF86BB2CFF4882E339372B7D" blockId="9.[151,536,1736,1795]" box="[151,375,1770,1795]" pageId="9" pageNumber="440">
<figureCitation id="17923CEBFF86BB2CFF4882E338A22B7D" box="[151,226,1770,1795]" captionStart="FIGURES 4 – 8" captionStartId="4.[151,264,1958,1981]" captionTargetBox="[171,1410,614,1931]" captionTargetId="figure@4.[162,1422,603,1945]" captionTargetPageId="4" captionText="FIGURES 4 – 8. Thoracophorus struyvei (4); T. perplexus (5); T. proximus (6); T. exilis (7); T. aequalis (8); aedeagus in lateral aspect (A), antenna (B), shape of temples (C); scale bar: 0.1 mm." httpUri="https://zenodo.org/record/238530/files/figure.png" pageId="9" pageNumber="440">Figs. 5</figureCitation>
A, B, C, 10D
</paragraph>
</subSubSection>
<subSubSection id="C7B373E5FF86BB2CFF48833B3DDC2B11" pageId="9" pageNumber="440" type="materials_examined">
<paragraph id="8F16206EFF86BB2CFF48833B3DDC2B11" blockId="9.[151,1436,1842,2011]" pageId="9" pageNumber="440">
<emphasis id="BDDDFC7CFF86BB2CFF48833B39062B35" bold="true" box="[151,326,1842,1867]" pageId="9" pageNumber="440">
<typeStatus id="50129ECCFF86BB2CFF48833B38922B35" box="[151,210,1842,1867]" pageId="9" pageNumber="440">Type</typeStatus>
material.
</emphasis>
<typeStatus id="50129ECCFF86BB2CFE91833A39A62B35" box="[334,486,1843,1867]" pageId="9" pageNumber="440" type="holotype">HOLOTYPE</typeStatus>
. male,
<emphasis id="BDDDFC7CFF86BB2CFDE9833B3ABB2B35" bold="true" box="[566,763,1842,1867]" pageId="9" pageNumber="440">
French
<collectingCountry id="F7BE60FEFF86BB2CFD4B833B3AB72B35" box="[660,759,1842,1867]" name="Guyana" pageId="9" pageNumber="440">Guyana</collectingCountry>
:
</emphasis>
Coralie (
<geoCoordinate id="EA9D46A9FF86BB2CFCB9833B3BAD2B35" box="[870,1005,1842,1867]" direction="west" orientation="longitude" pageId="9" pageNumber="440" precision="9" value="-52.356335">52°21.38'W</geoCoordinate>
,
<geoCoordinate id="EA9D46A9FF86BB2CFC25833A3C2F2B35" box="[1018,1135,1843,1867]" direction="north" orientation="latitude" pageId="9" pageNumber="440" precision="9" value="4.506833">4°30.41'N</geoCoordinate>
), male,
<date id="FB1706AEFF86BB2CFB16833B3D082B35" box="[1225,1352,1842,1867]" pageId="9" pageNumber="440" value="2011-11-15">15.11.2011</date>
, leg. T. Struyve (
<collectionCode id="E9B8B8ABFF86BB2CFF23835E39722B11" box="[252,306,1879,1903]" pageId="9" pageNumber="440">UIC</collectionCode>
).
<typeStatus id="50129ECCFF86BB2CFE9A835E39A92B11" box="[325,489,1879,1903]" pageId="9" pageNumber="440" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="99AFEBE7FF86BB2CFE2B835E3A022B11" box="[500,578,1878,1903]" pageId="9" pageNumber="440" type="male">1 male</specimenCount>
,
<specimenCount id="99AFEBE7FF86BB2CFD92835E3AF92B11" box="[589,697,1878,1903]" pageId="9" pageNumber="440" type="female">2 females</specimenCount>
from same location and at same date as
<typeStatus id="50129ECCFF86BB2CFBA1835F3CA02B11" box="[1150,1248,1878,1903]" pageId="9" pageNumber="440" type="holotype">holotype</typeStatus>
(1
<collectionCode id="E9B8B8ABFF86BB2CFADA835E3D7B2B11" box="[1285,1339,1879,1903]" httpUri="http://grbio.org/cool/vspn-j4n9" name="Tarleton State University, Tarleton State Collection" pageId="9" pageNumber="440">TSC</collectionCode>
, 2
<collectionCode id="E9B8B8ABFF86BB2CFA84835E3DD02B11" box="[1371,1424,1879,1903]" pageId="9" pageNumber="440">UIC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C7B373E5FF86BB2FFF1883733A772E1E" lastPageId="10" lastPageNumber="441" pageId="9" pageNumber="440" type="diagnosis">
<paragraph id="8F16206EFF86BB2FFF1883733A772E1E" blockId="9.[151,1436,1842,2011]" lastBlockId="10.[151,1437,151,1688]" lastPageId="10" lastPageNumber="441" pageId="9" pageNumber="440">
<emphasis id="BDDDFC7CFF86BB2CFF18837339012BED" bold="true" box="[199,321,1914,1939]" pageId="9" pageNumber="440">Diagnosis.</emphasis>
Among the species of the genus with the lateral margin of the pronotum not denticulate,
<taxonomicName id="48A95BEDFF86BB2CFA58837539442BC9" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="440" phylum="Arthropoda" rank="species" species="perplexus" status="sp. nov.">
<emphasis id="BDDDFC7CFF86BB2CFA58837539442BC9" italics="true" pageId="9" pageNumber="440">T. perplexus</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF86BB2CFED1839739112BC9" bold="true" box="[270,337,1950,1975]" pageId="9" pageNumber="440">
<taxonomicNameLabel id="A6EE4107FF86BB2CFED1839739112BC9" box="[270,337,1950,1975]" pageId="9" pageNumber="440" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
represents a species that seems to be intermediate between
<taxonomicName id="48A95BEDFF86BB2CFBCB83963D432BC9" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" box="[1044,1283,1950,1975]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="440" phylum="Arthropoda" rank="species" species="filum">
<emphasis id="BDDDFC7CFF86BB2CFBCB83963C2D2BC9" box="[1044,1133,1950,1975]" italics="true" pageId="9" pageNumber="440">T. filum</emphasis>
<bibRefCitation id="EB385D9FFF86BB2CFBA783973D432BC9" author="Sharp" box="[1144,1283,1950,1975]" pageId="9" pageNumber="440" refString="Sharp, D. (1887) Fam. Staphylinidae. In: Sharp, D. (Ed.), Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor &amp; Francis, London, pp. 145 - 824. [plates. 5 - 19]" type="journal article" year="1887">Sharp, 1887</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="48A95BEDFF86BB2CFA9E839639162BA5" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="440" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="BDDDFC7CFF86BB2CFA9E83963DDC2BC9" box="[1345,1436,1950,1975]" italics="true" pageId="9" pageNumber="440">T. exilis</emphasis>
(
<bibRefCitation id="EB385D9FFF86BB2CFF4183CB390D2BA5" author="Erichson" box="[158,333,1986,2011]" pageId="9" pageNumber="440" refString="Erichson, W. F. (1840) Genera et species Staphylinorum Insectorum Coleopterorum Familiae. Morin, Berlin, 954 pp. http: // dx. doi. org / 10.5962 / bhl. title. 59644" type="book" year="1840">Erichson, 1840</bibRefCitation>
)
</taxonomicName>
. The elytral carinae and swellings on the head are more distinctly developed than in the species of
<taxonomicName id="48A95BEDFF85BB2FFF488490393A2CCE" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" box="[151,378,151,176]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="filum">
<emphasis id="BDDDFC7CFF85BB2FFF48849038AC2CCE" box="[151,236,151,176]" italics="true" pageId="10" pageNumber="441">T. filum</emphasis>
<bibRefCitation id="EB385D9FFF85BB2FFF2C849E393A2CCE" author="Sharp" box="[243,378,151,176]" pageId="10" pageNumber="441" refString="Sharp, D. (1887) Fam. Staphylinidae. In: Sharp, D. (Ed.), Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor &amp; Francis, London, pp. 145 - 824. [plates. 5 - 19]" type="journal article" year="1887">Sharp, 1887</bibRefCitation>
</taxonomicName>
group, but less developed than in
<taxonomicName id="48A95BEDFF85BB2FFD2684903C552CCE" authority="Erichson, 1887" authorityName="Erichson" authorityYear="1887" box="[761,1045,151,176]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="BDDDFC7CFF85BB2FFD2684903B0F2CCE" box="[761,847,151,176]" italics="true" pageId="10" pageNumber="441">T. exilis</emphasis>
(Erichson, 1887)
</taxonomicName>
. Furthermore, it resembles
<taxonomicName id="48A95BEDFF85BB2FFA98849039642CAA" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="filum">
<emphasis id="BDDDFC7CFF85BB2FFA9884903DDC2CCE" box="[1351,1436,151,176]" italics="true" pageId="10" pageNumber="441">T. filum</emphasis>
<bibRefCitation id="EB385D9FFF85BB2FFF4884B539642CAA" author="Sharp" box="[151,292,188,213]" pageId="10" pageNumber="441" refString="Sharp, D. (1887) Fam. Staphylinidae. In: Sharp, D. (Ed.), Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor &amp; Francis, London, pp. 145 - 824. [plates. 5 - 19]" type="journal article" year="1887">
Sharp,
<quantity id="48518D8BFF85BB2FFF3384B539642CAA" box="[236,292,188,212]" metricMagnitude="1" metricUnit="m" metricValue="4.79298" pageId="10" pageNumber="441" unit="in" value="1887.0">1887</quantity>
</bibRefCitation>
</taxonomicName>
in the orange colouration of the pronotum. In contrast to
<taxonomicName id="48A95BEDFF85BB2FFC3D84B43C982CAA" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" box="[994,1240,188,213]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="filum">
<emphasis id="BDDDFC7CFF85BB2FFC3D84B43C7C2CAB" box="[994,1084,188,213]" italics="true" pageId="10" pageNumber="441">T. filum</emphasis>
<bibRefCitation id="EB385D9FFF85BB2FFB9684B53C982CAA" author="Sharp" box="[1097,1240,188,213]" pageId="10" pageNumber="441" refString="Sharp, D. (1887) Fam. Staphylinidae. In: Sharp, D. (Ed.), Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor &amp; Francis, London, pp. 145 - 824. [plates. 5 - 19]" type="journal article" year="1887">Sharp, 1887</bibRefCitation>
</taxonomicName>
, the elytra of
<taxonomicName id="48A95BEDFF85BB2FFA5884B439442C86" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="perplexus" status="sp. nov.">
<emphasis id="BDDDFC7CFF85BB2FFA5884B439442C86" italics="true" pageId="10" pageNumber="441">T. perplexus</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF85BB2FFED284D6390F2C86" bold="true" box="[269,335,223,248]" pageId="10" pageNumber="441">
<taxonomicNameLabel id="A6EE4107FF85BB2FFED284D6390F2C86" box="[269,335,223,248]" pageId="10" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are only orange in the anterior half, whereas they are totally orange in
<taxonomicName id="48A95BEDFF85BB2FFB5384E83D3A2C86" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" box="[1164,1402,223,248]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="filum">
<emphasis id="BDDDFC7CFF85BB2FFB5384E83CA42C86" box="[1164,1252,223,248]" italics="true" pageId="10" pageNumber="441">T. filum</emphasis>
<bibRefCitation id="EB385D9FFF85BB2FFB3284D63D3A2C86" author="Sharp" box="[1261,1402,223,248]" pageId="10" pageNumber="441" refString="Sharp, D. (1887) Fam. Staphylinidae. In: Sharp, D. (Ed.), Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor &amp; Francis, London, pp. 145 - 824. [plates. 5 - 19]" type="journal article" year="1887">Sharp, 1887</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="48A95BEDFF85BB2FFA5884E839442D63" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="perplexus" status="sp. nov.">
<emphasis id="BDDDFC7CFF85BB2FFA5884E839442D63" italics="true" pageId="10" pageNumber="441">T. perplexus</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF85BB2FFED4850D390A2D63" bold="true" box="[267,330,260,285]" pageId="10" pageNumber="441">
<taxonomicNameLabel id="A6EE4107FF85BB2FFED4850D390A2D63" box="[267,330,260,285]" pageId="10" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is slightly longer than
<taxonomicName id="48A95BEDFF85BB2FFD92850C3B742D62" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" box="[589,820,260,285]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="filum">
<emphasis id="BDDDFC7CFF85BB2FFD92850C3AE32D63" box="[589,675,260,285]" italics="true" pageId="10" pageNumber="441">T. filum</emphasis>
<bibRefCitation id="EB385D9FFF85BB2FFD76850D3B742D62" author="Sharp" box="[681,820,260,285]" pageId="10" pageNumber="441" refString="Sharp, D. (1887) Fam. Staphylinidae. In: Sharp, D. (Ed.), Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor &amp; Francis, London, pp. 145 - 824. [plates. 5 - 19]" type="journal article" year="1887">Sharp, 1887</bibRefCitation>
</taxonomicName>
. The elytra are quadrate and not longer than wide such as in
<taxonomicName id="48A95BEDFF85BB2FFF0A852039802D3E" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" box="[213,448,295,320]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="filum">
<emphasis id="BDDDFC7CFF85BB2FFF0A8520396B2D3E" box="[213,299,295,320]" italics="true" pageId="10" pageNumber="441">T. filum</emphasis>
<bibRefCitation id="EB385D9FFF85BB2FFEEB852E39802D3E" author="Sharp" box="[308,448,295,320]" pageId="10" pageNumber="441" refString="Sharp, D. (1887) Fam. Staphylinidae. In: Sharp, D. (Ed.), Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor &amp; Francis, London, pp. 145 - 824. [plates. 5 - 19]" type="journal article" year="1887">Sharp, 1887</bibRefCitation>
</taxonomicName>
. In this respect it also resembles
<taxonomicName id="48A95BEDFF85BB2FFCE385203B8A2D3E" box="[828,970,295,320]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="verhaaghi" status="sp. nov.">
<emphasis id="BDDDFC7CFF85BB2FFCE385203B8A2D3E" box="[828,970,295,320]" italics="true" pageId="10" pageNumber="441">T. verhaaghi</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF85BB2FFC0C852E3C532D3E" bold="true" box="[979,1043,295,320]" pageId="10" pageNumber="441">
<taxonomicNameLabel id="A6EE4107FF85BB2FFC0C852E3C532D3E" box="[979,1043,295,320]" pageId="10" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
But
<taxonomicName id="48A95BEDFF85BB2FFB9285203C9B2D3E" box="[1101,1243,295,320]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="verhaaghi" status="sp. nov.">
<emphasis id="BDDDFC7CFF85BB2FFB9285203C9B2D3E" box="[1101,1243,295,320]" italics="true" pageId="10" pageNumber="441">T. verhaaghi</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF85BB2FFB3B852E3D642D3E" bold="true" box="[1252,1316,295,320]" pageId="10" pageNumber="441">
<taxonomicNameLabel id="A6EE4107FF85BB2FFB3B852E3D642D3E" box="[1252,1316,295,320]" pageId="10" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is slightly larger and nearly totally orange. On average, it is also slightly smaller than
<taxonomicName id="48A95BEDFF85BB2FFC3285443D4C2D1A" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" box="[1005,1292,332,357]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="BDDDFC7CFF85BB2FFC3285443C052D1B" box="[1005,1093,332,357]" italics="true" pageId="10" pageNumber="441">T. exilis</emphasis>
(
<bibRefCitation id="EB385D9FFF85BB2FFB8B85453D442D1A" author="Erichson" box="[1108,1284,332,357]" pageId="10" pageNumber="441" refString="Erichson, W. F. (1840) Genera et species Staphylinorum Insectorum Coleopterorum Familiae. Morin, Berlin, 954 pp. http: // dx. doi. org / 10.5962 / bhl. title. 59644" type="book" year="1840">Erichson, 1840</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="48A95BEDFF85BB2FFAC7854439622DF6" authority="Irmler, 1985" authorityName="Irmler" authorityYear="1985" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="BDDDFC7CFF85BB2FFAC785443DDC2D1A" box="[1304,1436,333,356]" italics="true" pageId="10" pageNumber="441">T. proximus</emphasis>
<bibRefCitation id="EB385D9FFF85BB2FFF48856639622DF6" author="Irmler" box="[151,290,367,392]" pageId="10" pageNumber="441" refString="Irmler, U. (1985) Neue Arten der Gattungen Aneucamptus und Thoracophorus (Col., Staphylinidae) aus der Neotropis. Entomologische Blatter, 81, 41 - 58." type="journal article" year="1985">Irmler, 1985</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="48A95BEDFF85BB2FFE878578398B2DF6" box="[344,459,369,392]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="struyvei" status="sp. nov.">
<emphasis id="BDDDFC7CFF85BB2FFE878578398B2DF6" box="[344,459,369,392]" italics="true" pageId="10" pageNumber="441">T. struyvei</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF85BB2FFE0C85663A522DF6" bold="true" box="[467,530,367,392]" pageId="10" pageNumber="441">
<taxonomicNameLabel id="A6EE4107FF85BB2FFE0C85663A522DF6" box="[467,530,367,392]" pageId="10" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
It is mainly differentiated from these species by the aedeagal characters. Such as in
<taxonomicName id="48A95BEDFF85BB2FFF6A859C39902DD3" authority="Irmler, 1985" authorityName="Irmler" authorityYear="1985" box="[181,464,404,429]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="BDDDFC7CFF85BB2FFF6A859C397A2DD2" box="[181,314,405,428]" italics="true" pageId="10" pageNumber="441">T. proximus</emphasis>
<bibRefCitation id="EB385D9FFF85BB2FFE9C859D39902DD3" author="Irmler" box="[323,464,404,429]" pageId="10" pageNumber="441" refString="Irmler, U. (1985) Neue Arten der Gattungen Aneucamptus und Thoracophorus (Col., Staphylinidae) aus der Neotropis. Entomologische Blatter, 81, 41 - 58." type="journal article" year="1985">Irmler, 1985</bibRefCitation>
</taxonomicName>
the paramera are distinctly longer than the central lobe, but a transparent lobe is not separated at the apex of the paramera and the inner edge of the apical lobe is slightly sinuate and not straight. Only a few sensillae occur at the base and the apex of the paramera, whereas sensillae are concentrated on the apical plate in
<taxonomicName id="48A95BEDFF85BB2FFF2C860839282E66" box="[243,360,513,536]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="struyvei" status="sp. nov.">
<emphasis id="BDDDFC7CFF85BB2FFF2C860839282E66" box="[243,360,513,536]" italics="true" pageId="10" pageNumber="441">T. struyvei</emphasis>
</taxonomicName>
<emphasis id="BDDDFC7CFF85BB2FFEAF85F639F02E66" bold="true" box="[368,432,511,536]" pageId="10" pageNumber="441">
<taxonomicNameLabel id="A6EE4107FF85BB2FFEAF85F639F02E66" box="[368,432,511,536]" pageId="10" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and at the base of the paramera in
<taxonomicName id="48A95BEDFF85BB2FFC9F86083C1A2E66" authority="Irmler, 1985" authorityName="Irmler" authorityYear="1985" box="[832,1114,511,536]" class="Insecta" family="Staphylinidae" genus="Thoracophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="441" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="BDDDFC7CFF85BB2FFC9F86083B832E66" box="[832,963,513,536]" italics="true" pageId="10" pageNumber="441">T. proximus</emphasis>
<bibRefCitation id="EB385D9FFF85BB2FFC1385F63C1A2E66" author="Irmler" box="[972,1114,511,536]" pageId="10" pageNumber="441" refString="Irmler, U. (1985) Neue Arten der Gattungen Aneucamptus und Thoracophorus (Col., Staphylinidae) aus der Neotropis. Entomologische Blatter, 81, 41 - 58." type="journal article" year="1985">Irmler, 1985</bibRefCitation>
</taxonomicName>
. In addition, the temples are more rounded than in the two other species and the posterior angles are only slightly obtuse with a short nearly parallel part directly behind the eyes.
</paragraph>
</subSubSection>
<subSubSection id="C7B373E5FF85BB2FFF1886653BC82A2E" pageId="10" pageNumber="441" type="description">
<paragraph id="8F16206EFF85BB2FFF18866539382ED6" blockId="10.[151,1437,151,1688]" pageId="10" pageNumber="441">
<emphasis id="BDDDFC7CFF85BB2FFF188665391B2EFB" bold="true" box="[199,347,620,645]" pageId="10" pageNumber="441">Description:</emphasis>
Length:
<quantity id="48518D8BFF85BB2FFE1E86653A5B2EFA" box="[449,539,620,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="441" unit="mm" value="1.6">1.6 mm</quantity>
. Colouration: black; clypeus, pronotum on lateral and posterior parts and elytra at shoulders orange.
</paragraph>
<paragraph id="8F16206EFF85BB2FFF1886BD39FA2F22" blockId="10.[151,1437,151,1688]" pageId="10" pageNumber="441">
Head:
<quantity id="48518D8BFF85BB2FFECE86BD39362EB2" box="[273,374,692,716]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="441" unit="mm" value="0.2">0.20 mm</quantity>
long;
<quantity id="48518D8BFF85BB2FFE6086BD3A632EB2" box="[447,547,692,716]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="441" unit="mm" value="0.27">0.27 mm</quantity>
wide; eyes large; slightly prominent; temples short; only one fourth as long as eyes; narrowed to neck in short convex curve; posterior angles of temples 120°; clypeus approximately semicircular; front edge margined; without punctation; pair of elongate swellings on vertex indistinct; with slight impression between them, but without impression between swellings and eyes; isodiametric microsculpture deep and coarse; surface matte.
</paragraph>
<paragraph id="8F16206EFF85BB2FFF1887613B22284B" blockId="10.[151,1437,151,1688]" pageId="10" pageNumber="441">Antennae as long as head and half of pronotum combined; slightly clavate; first and second antennomeres thick; first approximately rectangular; second oval; antennomeres three to six distinctly narrower than preceding two antennomeres; conical third antennomere slightly shorter than second; fourth to sixth antennomeres approximately quadrate; seventh antennomere slightly wider than preceding antennomeres; following antennomeres distinctly wider than preceding antennomeres; eighth antennomere considerably wider than long; tenth antennomere slightly wider than long and wider than first.</paragraph>
<paragraph id="8F16206EFF85BB2FFF1880493B6F2972" blockId="10.[151,1437,151,1688]" pageId="10" pageNumber="441">
Pronotum:
<quantity id="48518D8BFF85BB2FFE95803639F52829" box="[330,437,1087,1112]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="441" unit="mm" value="0.25">0.25 mm</quantity>
long,
<quantity id="48518D8BFF85BB2FFDD780493A322829" box="[520,626,1088,1112]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="441" unit="mm" value="0.3">0.30 mm</quantity>
wide; widest in anterior half; anterior half parallel; anterior angles approximately rectangular; posterior half deeply sinuate to posterior angles; posterior edge one third smaller than anterior edge; sides of anterior half indistinctly crenulate; central impression moderately deep; in anterior half deeper than in posterior half; central impression with oval swelling; moderately deep lateral impression elongate; extending from posterior angles to anterior angles; posterior edge with white short scales; without punctation; isodiametric microsculpture deep and dense; surface matte.
</paragraph>
<paragraph id="8F16206EFF85BB2FFF18811E3B6F2906" blockId="10.[151,1437,151,1688]" pageId="10" pageNumber="441">
Elytra:
<quantity id="48518D8BFF85BB2FFEC3811E39C3294E" box="[284,387,1303,1328]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="441" unit="mm" value="0.35">0.35 mm</quantity>
long,
<quantity id="48518D8BFF85BB2FFE11811E3A74294E" box="[462,564,1303,1328]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="441" unit="mm" value="0.35">0.35 mm</quantity>
wide; sides parallel; shoulders and posterior angles obtusely rounded; three indistinct carinae on each side of suture; without punctation; few white short scales at posterior margin; isodiametric microsculpture deep and dense; surface matte.
</paragraph>
<paragraph id="8F16206EFF85BB2FFF18818D3975299A" blockId="10.[151,1437,151,1688]" pageId="10" pageNumber="441">Abdomen parallel; segments III to V with weak transverse impression at base; with extremely fine punctation; punctures with short white scales; isodiametric microsculpture dense and deep; slightly less deep than on elytra; surface matte.</paragraph>
<paragraph id="8F16206EFF85BB2FFF1881E63BC82A2E" blockId="10.[151,1437,151,1688]" pageId="10" pageNumber="441">Aedeagus with obtuse angle between basal and apical lobe; apical lobe long; in lateral aspect, inner edge of apical lobe sinuate; endophallus with thick basal bulb and long ductus widened apically; paramera long and slender; projecting apical lobe; few sensillae at apex and at middle.</paragraph>
</subSubSection>
<subSubSection id="C7B373E5FF85BB2FFF1882553AD92AE6" pageId="10" pageNumber="441" type="etymology">
<paragraph id="8F16206EFF85BB2FFF1882553AD92AE6" blockId="10.[151,1437,151,1688]" pageId="10" pageNumber="441">
<emphasis id="BDDDFC7CFF85BB2FFF188255390E2A0B" bold="true" box="[199,334,1628,1653]" pageId="10" pageNumber="441">Etymology.</emphasis>
The specific name derives from the same Latin word meaning confused and refers to the high similarity between the species described here.
</paragraph>
</subSubSection>
</treatment>
</document>
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