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<document id="143CD184B5BDFBFCF2B4E907D8C92735" ID-DOI="10.5852/ejt.2024.962.2683" ID-ISSN="2118-9773" ID-ZooBank="40171C5B-90EB-4641-91FF-EA8CA2C60D23" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729183243575" checkinUser="plazi" docAuthor="Ramos-Sánchez, Mariela" docDate="2024" docId="03F2DF38026EFF9EFEE74D45FDE07B99" docLanguage="en" docName="EJT.2024.962.1-67.pdf" docOrigin="European Journal of Taxonomy 962 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2683/12419" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Boninia oaxaquensis Ramos-Sanchez, Bahia &amp; Bastida-Zavala 2020" docType="treatment" docVersion="1" lastPageNumber="38" masterDocId="FFCBA740024AFFBBFF994B10FFC27830" masterDocTitle="The polyclads (Platyhelminthes: Polycladida) from the Tropical Eastern Pacific: Commented checklist and description of a new species" masterLastPageNumber="67" masterPageNumber="1" pageNumber="37" updateTime="1729183522411" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="93CC7A988E948577B8688C45B93BF87A">The polyclads (Platyhelminthes: Polycladida) from the Tropical Eastern Pacific: Commented checklist and description of a new species</mods:title>
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<mods:namePart id="CEFC2ACC10007196444D8E244273B5D3">Ramos-Sánchez, Mariela</mods:namePart>
<mods:affiliation id="61C27008EE4D253A0A974204D46FD74A">Doctorado en Ciencias Biológico Agropecuarias, Área Ciencias Biológico Agropecuaria y Pesquera, Ciudad de la cultura “ Amado Nervo ”, Tepic, Nayarit, 63000, Mexico. &amp; Laboratorio de Sistemática de Invertebrados Marinos (LABSIM), Universidad del Mar, campus Puerto Ángel, Ciudad Universitaria, Puerto Ángel, Oaxaca, 70902, Mexico.</mods:affiliation>
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<mods:title id="6FCCA111AEE4AA4D3821578677C3306A">European Journal of Taxonomy</mods:title>
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<mods:date id="32AC668048D3D9AE372DFA7EA3A69C40">2024</mods:date>
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<treatment id="03F2DF38026EFF9EFEE74D45FDE07B99" LSID="urn:lsid:plazi:treatment:03F2DF38026EFF9EFEE74D45FDE07B99" httpUri="http://treatment.plazi.org/id/03F2DF38026EFF9EFEE74D45FDE07B99" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<subSubSection id="C3413DA5026EFF9FFEE74D45FB747E5F" box="[382,1206,1621,1647]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="8BE46E2E026EFF9FFEE74D45FB747E5F" blockId="36.[382,1206,1621,1647]" box="[382,1206,1621,1647]" pageId="36" pageNumber="37">
<heading id="D0ACD942026EFF9FFEE74D45FB747E5F" box="[382,1206,1621,1647]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="37" reason="2">
<taxonomicName id="4C5B15AD026EFF9FFEE74D45FB747E5F" authority="Ramos-Sanchez, Bahia &amp; Bastida-Zavala, 2020" authorityName="Ramos-Sanchez, Bahia &amp; Bastida-Zavala" authorityYear="2020" box="[382,1206,1621,1647]" family="Boniniidae" genus="Boninia" kingdom="Animalia" order="Polycladida" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="species" species="oaxaquensis">
<emphasis id="B92FB23C026EFF9FFEE74D45FDBF7E5F" bold="true" box="[382,637,1621,1647]" italics="true" pageId="36" pageNumber="37">Boninia oaxaquensis</emphasis>
<bibRefCitation id="EFCA13DF026EFF9FFD1C4D45FB747E5F" author="Ramos-Sanchez M. &amp; Bahia J. &amp; Bastida-Zavala J. R." box="[645,1206,1621,1647]" pageId="36" pageNumber="37" pagination="49 - 83" refId="ref32124" refString="Ramos-Sanchez M., Bahia J. &amp; Bastida-Zavala J. R. 2020. Five new species of cotylean flatworms (Platyhelminthes: Polycladida: Cotylea) from Oaxaca, southern Mexican Pacific. Zootaxa 4819 (1): 49 - 83. https: // doi. org / 10.11646 / zootaxa. 4819.1.3" type="journal article" year="2020">Ramos-Sánchez, Bahia &amp; Bastida-Zavala, 2020</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3413DA5026EFF9FFF244D87FC4E7EE3" pageId="36" pageNumber="37" type="reference_group">
<paragraph id="8BE46E2E026EFF9FFF244D87FC4E7EE3" blockId="36.[189,1398,1686,1750]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB4B4900026EFF9FFF244D87FC4E7EE3" pageId="36" pageNumber="37">
<treatmentCitation id="0AFA483F026EFF9FFF244D87FBFE7E81" author="Ramos-Sanchez M. &amp; Bahia J. &amp; Bastida-Zavala J. R." box="[189,1084,1686,1713]" page="56" pageId="36" pageNumber="37" year="2020">
<taxonomicName id="4C5B15AD026EFF9FFF244D87FBFE7E81" authority="Ramos-Sanchez, Bahia &amp; Bastida-Zavala, 2020: 51 - 56" authorityName="Ramos-Sanchez, Bahia &amp; Bastida-Zavala" authorityYear="2020" box="[189,1084,1686,1713]" family="Boniniidae" genus="Boninia" kingdom="Animalia" order="Polycladida" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="species" species="oaxaquensis">
<emphasis id="B92FB23C026EFF9FFF244D87FE737E80" box="[189,433,1687,1712]" italics="true" pageId="36" pageNumber="37">Boninia oaxaquensis</emphasis>
<bibRefCitation id="EFCA13DF026EFF9FFE2E4D86FBFE7E81" author="Ramos-Sanchez M. &amp; Bahia J. &amp; Bastida-Zavala J. R." box="[439,1084,1686,1713]" pageId="36" pageNumber="37" pagination="49 - 83" refId="ref32124" refString="Ramos-Sanchez M., Bahia J. &amp; Bastida-Zavala J. R. 2020. Five new species of cotylean flatworms (Platyhelminthes: Polycladida: Cotylea) from Oaxaca, southern Mexican Pacific. Zootaxa 4819 (1): 49 - 83. https: // doi. org / 10.11646 / zootaxa. 4819.1.3" type="journal article" year="2020">Ramos-Sánchez, Bahia &amp; Bastida-Zavala, 2020: 51–56</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 2a–i, 3a–b, 4a–e (
<typeStatus id="54E0D08C026EFF9FFADB4D86FAB47E80" box="[1346,1398,1686,1712]" pageId="36" pageNumber="37">type</typeStatus>
locality: Panteón Beach, Puerto Ángel,
<collectingRegion id="499FA0CC026EFF9FFD234DA9FCD77EE3" box="[698,789,1721,1747]" country="Mexico" name="Oaxaca" pageId="36" pageNumber="37">Oaxaca</collectingRegion>
,
<collectingCountry id="F34C2EBE026EFF9FFCB84DA9FC427EE3" box="[801,896,1721,1747]" name="Mexico" pageId="36" pageNumber="37">Mexico</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3413DA5026EFF9EFF244DEAFAEB7944" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="distribution">
<paragraph id="8BE46E2E026EFF9FFF244DEAFE947F24" blockId="36.[189,342,1786,1812]" box="[189,342,1786,1812]" pageId="36" pageNumber="37">
<heading id="D0ACD942026EFF9FFF244DEAFE947F24" bold="true" box="[189,342,1786,1812]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B92FB23C026EFF9FFF244DEAFE947F24" bold="true" box="[189,342,1786,1812]" pageId="36" pageNumber="37">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46E2E026EFF9FFF244C39FC127F56" blockId="36.[188,1399,1833,1895]" pageId="36" pageNumber="37">
Central coast of
<collectingRegion id="499FA0CC026EFF9FFEE34C39FE157F73" box="[378,471,1833,1859]" country="Mexico" name="Oaxaca" pageId="36" pageNumber="37">Oaxaca</collectingRegion>
: Agua Blanca, San Agustinillo, Panteón, Estacahuite Beaches and Cacaluta Bay. This is the first record of the genus for the Tropical Eastern Pacific.
</paragraph>
<paragraph id="8BE46E2E026FFF9EFF244A18FEDF7912" blockId="37.[189,285,264,290]" box="[189,285,264,290]" pageId="37" pageNumber="38">
<heading id="D0ACD942026FFF9EFF244A18FEDF7912" bold="true" box="[189,285,264,290]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B92FB23C026FFF9EFF244A18FEDF7912" bold="true" box="[189,285,264,290]" pageId="37" pageNumber="38">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46E2E026FFF9EFF244A27FAEB7944" blockId="37.[188,1399,311,373]" pageId="37" pageNumber="38">
Benthic, littoral to sublittoral (
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). Specimens of
<taxonomicName id="4C5B15AD026FFF9EFCAC4A28FC507961" authorityName="Bock" authorityYear="1923" box="[821,914,312,337]" family="Boniniidae" genus="Boninia" kingdom="Animalia" order="Polycladida" pageId="37" pageNumber="38" phylum="Platyhelminthes" rank="genus">
<emphasis id="B92FB23C026FFF9EFCAC4A28FC507961" box="[821,914,312,337]" italics="true" pageId="37" pageNumber="38">Boninia</emphasis>
</taxonomicName>
were collected under rocks, frequently in tide pools, in groups of three to
<specimenCount id="9D5DA5A7026FFF9EFDAA4A4AFD157944" box="[563,727,346,372]" count="6" pageId="37" pageNumber="38" type="generic">six specimens</specimenCount>
on each rock and alongside chitons and seaweeds.
</paragraph>
</subSubSection>
<subSubSection id="C3413DA5026FFF9EFF244A8CFDE07B99" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8BE46E2E026FFF9EFF244A8CFEED7986" blockId="37.[189,303,412,438]" box="[189,303,412,438]" pageId="37" pageNumber="38">
<heading id="D0ACD942026FFF9EFF244A8CFEED7986" bold="true" box="[189,303,412,438]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B92FB23C026FFF9EFF244A8CFEED7986" bold="true" box="[189,303,412,438]" pageId="37" pageNumber="38">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46E2E026FFF9EFF244ADBFE3E7A41" blockId="37.[189,1399,458,625]" pageId="37" pageNumber="38">
<taxonomicName id="4C5B15AD026FFF9EFF244ADBFE7B79D4" authorityName="Ramos-Sanchez, Bahia &amp; Bastida-Zavala" authorityYear="2020" box="[189,441,459,484]" family="Boniniidae" genus="Boninia" kingdom="Animalia" order="Polycladida" pageId="37" pageNumber="38" phylum="Platyhelminthes" rank="species" species="oaxaquensis">
<emphasis id="B92FB23C026FFF9EFF244ADBFE7B79D4" box="[189,441,459,484]" italics="true" pageId="37" pageNumber="38">Boninia oaxaquensis</emphasis>
</taxonomicName>
is assigned to the genus
<taxonomicName id="4C5B15AD026FFF9EFC9E4ADBFCA679D4" authorityName="Bock" authorityYear="1923" box="[775,868,459,484]" family="Boniniidae" genus="Boninia" kingdom="Animalia" order="Polycladida" pageId="37" pageNumber="38" phylum="Platyhelminthes" rank="genus">
<emphasis id="B92FB23C026FFF9EFC9E4ADBFCA679D4" box="[775,868,459,484]" italics="true" pageId="37" pageNumber="38">Boninia</emphasis>
</taxonomicName>
based on prostatoid organs symmetrically surrounding the male gonopore.
<taxonomicName id="4C5B15AD026FFF9EFDA14AFEFCEF7A37" authorityName="Ramos-Sanchez, Bahia &amp; Bastida-Zavala" authorityYear="2020" box="[568,813,494,519]" family="Boniniidae" genus="Boninia" kingdom="Animalia" order="Polycladida" pageId="37" pageNumber="38" phylum="Platyhelminthes" rank="species" species="oaxaquensis">
<emphasis id="B92FB23C026FFF9EFDA14AFEFCEF7A37" box="[568,813,494,519]" italics="true" pageId="37" pageNumber="38">Boninia oaxaquensis</emphasis>
</taxonomicName>
presents 14–66 (n = 24, μ = 36, SD = 26) cerebral eyes and 36–126 (n = 24, μ = 78, SD = 45) marginal eyes, has between 16–24 (n = 24) prostatoid organs arranged in one girdle, and its habitat is in the littoral to sublittoral zone (
<quantity id="4CA3C3CB026FFF9EFBBD4924FB407A7E" box="[1060,1154,564,590]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="10.0" metricValueMin="2.0" pageId="37" pageNumber="38" unit="m" value="6.0" valueMax="10.0" valueMin="2.0">2–10 m</quantity>
), under rocks in tide pools and sublittoral rocks.
</paragraph>
<paragraph id="8BE46E2E026FFF9EFF244988FDE07B99" blockId="37.[188,1399,664,937]" pageId="37" pageNumber="38">
Currently, the genus comprises seven valid species
<taxonomicName id="4C5B15AD026FFF9EFCBD4989FB457A83" authority="(Hyman 1955 c)" baseAuthorityName="Hyman" baseAuthorityYear="1955" box="[804,1159,664,691]" family="Boniniidae" genus="Boninia" kingdom="Animalia" order="Polycladida" pageId="37" pageNumber="38" phylum="Platyhelminthes" rank="species" species="antillarum">
<emphasis id="B92FB23C026FFF9EFCBD4989FC007A82" box="[804,962,664,690]" italics="true" pageId="37" pageNumber="38">B. antillarum</emphasis>
(
<bibRefCitation id="EFCA13DF026FFF9EFC484989FBBC7A83" author="Hyman L. H." box="[977,1150,664,691]" pageId="37" pageNumber="38" pagination="115 - 150" refId="ref29842" refString="Hyman L. H. 1955 c. Some polyclad flatworms from the west Indies and Florida. Proceedings of the United States National Museum 104 (3341): 115 - 150. https: // doi. org / 10.5479 / si. 00963801.104 - 3341.115" type="journal article" year="1955">Hyman 1955c</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C5B15AD026FFF9EFB0F4989FEA37AE5" authority="Marcus &amp; Marcus, 1968" authorityName="Marcus &amp; Marcus" authorityYear="1968" family="Boniniidae" genus="Boninia" kingdom="Animalia" order="Polycladida" pageId="37" pageNumber="38" phylum="Platyhelminthes" rank="species" species="divae">
<emphasis id="B92FB23C026FFF9EFB0F4989FB3A7A82" box="[1174,1272,664,690]" italics="true" pageId="37" pageNumber="38">B. divae</emphasis>
<bibRefCitation id="EFCA13DF026FFF9EFB664989FEA37AE5" author="Marcus E. &amp; Marcus E." pageId="37" pageNumber="38" pagination="1 - 11" refId="ref30889" refString="Marcus E. &amp; Marcus E. 1968. Polycladida from Curacao and faunistically related regions. Studies on the Fauna Curacao and other Caribbean Islands 26: 1 - 11. Available from https: // repository. naturalis. nl / pub / 506072 [accessed 2 Aug. 2023]." type="journal article" year="1968">Marcus &amp; Marcus, 1968</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5B15AD026FFF9EFEF049ACFD417AE6" authority="Bock, 1923" authorityName="Bock" authorityYear="1923" box="[361,643,700,726]" family="Boniniidae" genus="Boninia" kingdom="Animalia" order="Polycladida" pageId="37" pageNumber="38" phylum="Platyhelminthes" rank="species" species="mirabilis">
<emphasis id="B92FB23C026FFF9EFEF049ACFE317AE6" box="[361,499,700,726]" italics="true" pageId="37" pageNumber="38">B. mirabilis</emphasis>
<bibRefCitation id="EFCA13DF026FFF9EFE6349ACFD417AE6" author="Bock S." box="[506,643,700,726]" pageId="37" pageNumber="38" pagination="341 - 372" refId="ref28091" refString="Bock S. 1923. Polycladen aus Juan Fernandez. In: Skottsberg C. (ed.) The Natural History of Juan Fernandez and Easter Island. Zoology 3: 341 - 372. Almqvist &amp; Wiksells Boktryckeri, Uppsala. https: // doi. org / 10.5962 / bhl. title. 41367" type="journal article" year="1923">Bock, 1923</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5B15AD026FFF9EFD1449ACFB767AE6" authority="Curini-Galletti &amp; Campus, 2007" authorityName="Curini-Galletti &amp; Campus" authorityYear="2007" box="[653,1204,700,726]" family="Boniniidae" genus="Boninia" kingdom="Animalia" order="Polycladida" pageId="37" pageNumber="38" phylum="Platyhelminthes" rank="species" species="neotethydis">
<emphasis id="B92FB23C026FFF9EFD1449ACFCF07AE6" box="[653,818,700,726]" italics="true" pageId="37" pageNumber="38">B. neotethydis</emphasis>
<bibRefCitation id="EFCA13DF026FFF9EFCA049ACFB767AE6" author="Curini-Galletti M. &amp; Campus P." box="[825,1204,700,726]" pageId="37" pageNumber="38" pagination="435 - 442" refId="ref28540" refString="Curini-Galletti M. &amp; Campus P. 2007. Boninia neotethydis sp. nov. (Platyhelminthes: Polycladida: Cotylea) the first lessepsian flatworm. Journal of the Marine Biological Association of the United Kingdom 87: 435 - 442. https: // doi. org / 10.1017 / S 0025315407055014" type="journal article" year="2007">Curini-Galletti &amp; Campus, 2007</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5B15AD026FFF9EFB2449ACFAAC7AE5" authorityName="Ramos-Sanchez, Bahia &amp; Bastida-Zavala" authorityYear="2020" box="[1213,1390,700,725]" family="Boniniidae" genus="Boninia" kingdom="Animalia" order="Polycladida" pageId="37" pageNumber="38" phylum="Platyhelminthes" rank="species" species="oaxaquensis">
<emphasis id="B92FB23C026FFF9EFB2449ACFAAC7AE5" box="[1213,1390,700,725]" italics="true" pageId="37" pageNumber="38">B. oaxaquensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5B15AD026FFF9EFF2449F0FD4F7AC9" authority="Tsuyuki, Oya &amp; Kajihara 2022" authorityName="Tsuyuki, Oya &amp; Kajihara" authorityYear="2022" box="[189,653,735,761]" family="Boniniidae" genus="Boninia" kingdom="Animalia" order="Polycladida" pageId="37" pageNumber="38" phylum="Platyhelminthes" rank="species" species="uru">
<emphasis id="B92FB23C026FFF9EFF2449F0FEC87AC9" box="[189,266,736,761]" italics="true" pageId="37" pageNumber="38">B. uru</emphasis>
Tsuyuki, Oya &amp; Kajihara 2022
</taxonomicName>
, and
<taxonomicName id="4C5B15AD026FFF9EFD4949F0FAC97AC9" authority="Tsuyuki, Oya &amp; Kajihara 2022" authorityName="Tsuyuki, Oya &amp; Kajihara" authorityYear="2022" box="[720,1291,735,761]" family="Boniniidae" genus="Boninia" kingdom="Animalia" order="Polycladida" pageId="37" pageNumber="38" phylum="Platyhelminthes" rank="species" species="yambarensis">
<emphasis id="B92FB23C026FFF9EFD4949F0FC457AC9" box="[720,903,735,761]" italics="true" pageId="37" pageNumber="38">B. yambarensis</emphasis>
Tsuyuki, Oya &amp; Kajihara 2022
</taxonomicName>
. Authors such as
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,
<bibRefCitation id="EFCA13DF026FFF9EFE504813FD4A7B2D" author="Hyman L. H." box="[457,648,770,797]" pageId="37" pageNumber="38" pagination="65 - 82" refId="ref29742" refString="Hyman L. H. 1955 a. Some polyclad flatworms from Polynesia and Micronesia. Proceedings of the United States National Museum 105 (3352): 65 - 82. https: // doi. org / 10.5479 / si. 00963801.105 - 3352.65 Hyman L. H. 1955 b. The polyclad flatworms of the Pacific coast of North America. Additions and corrections. American Museum Novitates 1704: 1 - 11. Available from http: // hdl. handle. net / 2246 / 4856 [accessed 2 Aug. 2023]." type="journal article" year="1955">Hyman (1955a)</bibRefCitation>
,
<bibRefCitation id="EFCA13DF026FFF9EFD0E4813FC0C7B2C" author="Marcus E. &amp; Marcus E." box="[663,974,770,797]" pageId="37" pageNumber="38" pagination="1 - 11" refId="ref30889" refString="Marcus E. &amp; Marcus E. 1968. Polycladida from Curacao and faunistically related regions. Studies on the Fauna Curacao and other Caribbean Islands 26: 1 - 11. Available from https: // repository. naturalis. nl / pub / 506072 [accessed 2 Aug. 2023]." type="journal article" year="1968">Marcus &amp; Marcus (1968)</bibRefCitation>
,
<bibRefCitation id="EFCA13DF026FFF9EFC444812FAB07B2C" author="Curini-Galletti M. &amp; Campus P." box="[989,1394,770,796]" pageId="37" pageNumber="38" pagination="435 - 442" refId="ref28540" refString="Curini-Galletti M. &amp; Campus P. 2007. Boninia neotethydis sp. nov. (Platyhelminthes: Polycladida: Cotylea) the first lessepsian flatworm. Journal of the Marine Biological Association of the United Kingdom 87: 435 - 442. https: // doi. org / 10.1017 / S 0025315407055014" type="journal article" year="2007">Curini-Galletti &amp; Campus (2007)</bibRefCitation>
, and
<bibRefCitation id="EFCA13DF026FFF9EFF764835FD817B0F" author="Ramos-Sanchez M. &amp; Bahia J. &amp; Bastida-Zavala J. R." box="[239,579,805,831]" pageId="37" pageNumber="38" pagination="49 - 83" refId="ref32124" refString="Ramos-Sanchez M., Bahia J. &amp; Bastida-Zavala J. R. 2020. Five new species of cotylean flatworms (Platyhelminthes: Polycladida: Cotylea) from Oaxaca, southern Mexican Pacific. Zootaxa 4819 (1): 49 - 83. https: // doi. org / 10.11646 / zootaxa. 4819.1.3" type="journal article" year="2020">
Ramos-Sánchez
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have suggested that the key diagnostic characters for defining species within the genus are: the number of prostatoid organs, the arrangement and number of their girdles, which are distributed on each side of the penis papillae, the distribution and number of cerebral eyes and marginal eyes, and the habitat.
</paragraph>
</subSubSection>
</treatment>
</document>
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