700 lines
126 KiB
XML
700 lines
126 KiB
XML
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<mods:title id="CFC8F66E349F4C4E242394487A8E7B2B">The Lace Web Spider Genus Vytfutia Deeleman-Reinhold (Araneae, Phyxelididae) in the Indo-Pacific Region</mods:title>
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<mods:namePart id="CB1D2E0C79502ADC8FAA8686A394A9EC">Griswold, Charles E.</mods:namePart>
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<mods:title id="F55226CBEFF3FB5040DA519EDECDAFBC">Proceedings of the California Academy of Sciences</mods:title>
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<mods:date id="E7B2DEAD76109AF7048B2AF55DE8F8EE">2022</mods:date>
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<mods:number id="264BE64311973ABBB5578E898592E31E">2022-07-29</mods:number>
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<treatment id="03B487984B6CFFBD28A2919EE6EEF3FD" ID-DOI="http://doi.org/10.5281/zenodo.12692105" ID-GBIF-Taxon="234981471" ID-Zenodo-Dep="12692105" LSID="urn:lsid:plazi:treatment:03B487984B6CFFBD28A2919EE6EEF3FD" httpUri="http://treatment.plazi.org/id/03B487984B6CFFBD28A2919EE6EEF3FD" lastPageId="6" lastPageNumber="335" pageId="4" pageNumber="333">
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<subSubSection id="C30765054B6CFFBF28A2919EE14AF77C" box="[486,811,508,534]" pageId="4" pageNumber="333" type="nomenclature">
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<paragraph id="8BA2368E4B6CFFBF28A2919EE14AF77C" blockId="4.[486,811,508,534]" box="[486,811,508,534]" pageId="4" pageNumber="333">
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<heading id="D0EA81E24B6CFFBF28A2919EE14AF77C" bold="true" box="[486,811,508,534]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="333" reason="6">
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<emphasis id="B969EA9C4B6CFFBF28A2919EE14AF77C" bold="true" box="[486,811,508,534]" pageId="4" pageNumber="333">
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<taxonomicName id="4C1D4D0D4B6CFFBF28A2919EE001F77C" authority="Griswold, 1990" authorityName="Griswold" authorityYear="1990" box="[486,608,508,534]" class="Arachnida" family="Agelenidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="329" phylum="Arthropoda" rank="tribe" tribe="Vytfutiini">Vytfutiini</taxonomicName>
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,
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<bibRefCitation id="EF8C4B7F4B6CFFBF2B29919EE14AF77C" author="GRISWOLD, C. E." box="[621,811,508,534]" pageId="4" pageNumber="333" pagination="1 - 206" refId="ref11937" refString="GRISWOLD, C. E. 1990. A revision and phylogenetic analysis of the spider subfamily Phyxelidinae (Araneae, Amaurobiidae). Bulletin of the American Museum of Natural History, 196: 1 - 206." type="journal article" year="1990">Griswold, 1990</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30765054B6CFFBF28D5925AE14CF700" pageId="4" pageNumber="333" type="type_taxon">
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<paragraph id="8BA2368E4B6CFFBF28D5925AE14CF700" blockId="4.[401,896,568,618]" pageId="4" pageNumber="333">
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Type genus
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<taxonomicName id="4C1D4D0D4B6CFFBF2B42925AE11DF727" authority="Deeleman-Reinhold, 1986: 34" authorityName="Deeleman-Reinhold" authorityPageNumber="34" authorityYear="1986" box="[518,892,568,589]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="genus">
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<emphasis id="B969EA9C4B6CFFBF2B42925AE031F727" box="[518,592,568,589]" italics="true" pageId="4" pageNumber="333">Vytfutia</emphasis>
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<bibRefCitation id="EF8C4B7F4B6CFFBF2B12925AE11DF727" author="DEELEMAN-REINHOLD, C. L." box="[598,892,568,589]" pageId="4" pageNumber="333" pagination="34 - 36" refId="ref11784" refString="DEELEMAN-REINHOLD, C. L. 1986. A new cribellate amaurobioid spider from Sumatra (Araneae: Agelenidae). Bulletin of the British Arachnological Society, 7: 34 - 36." type="journal article" year="1986">Deeleman-Reinhold, 1986: 34</bibRefCitation>
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</taxonomicName>
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||
. Comprise a single genus,
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<taxonomicName id="4C1D4D0D4B6CFFBF2B9A9237E146F700" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[734,807,597,618]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B969EA9C4B6CFFBF2B9A9237E146F700" box="[734,807,597,618]" italics="true" pageId="4" pageNumber="333">Vytfutia</emphasis>
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</taxonomicName>
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||
.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C30765054B6CFFBF28C8921CE173F7DF" pageId="4" pageNumber="333" type="multiple">
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<paragraph id="8BA2368E4B6CFFBF28C8921CE173F7DF" blockId="4.[396,900,638,693]" pageId="4" pageNumber="333">
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<emphasis id="B969EA9C4B6CFFBF28C8921CE1E5F7FF" bold="true" box="[396,900,638,661]" pageId="4" pageNumber="333">Diagnosis, Synapomorphies and Description</emphasis>
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See below under
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<taxonomicName id="4C1D4D0D4B6CFFBF2BFE92FCE16AF7DF" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[698,779,670,693]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="genus">
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<emphasis id="B969EA9C4B6CFFBF2BFE92FCE16AF7DF" box="[698,779,670,693]" italics="true" pageId="4" pageNumber="333">Vytfutia</emphasis>
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</taxonomicName>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30765054B6CFFBF293F92ABE3F2F65F" pageId="4" pageNumber="333" type="reference_group">
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<paragraph id="8BA2368E4B6CFFBF293F92ABE3F2F65F" blockId="4.[123,1174,713,1725]" pageId="4" pageNumber="333">
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<treatmentCitationGroup id="AB0D11A04B6CFFBF293F92ABE6F7F7B4" box="[123,1174,713,734]" pageId="4" pageNumber="333">
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<treatmentCitation id="0ABC109F4B6CFFBF293F92ABE062F7B4" author="DEELEMAN-REINHOLD, C. L." box="[123,515,713,734]" page="34" pageId="4" pageNumber="333" year="1986">
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<taxonomicName id="4C1D4D0D4B6CFFBF293F92ABE062F7B4" ID-CoL="87CS" authority="Deeleman-Reinhold, 1986: 34" authorityName="Deeleman-Reinhold" authorityPageNumber="34" authorityYear="1986" box="[123,515,713,734]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="genus">
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<emphasis id="B969EA9C4B6CFFBF293F92ABE2A4F7B4" box="[123,197,713,734]" italics="true" pageId="4" pageNumber="333">Vytfutia</emphasis>
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<bibRefCitation id="EF8C4B7F4B6CFFBF299692ABE062F7B4" author="DEELEMAN-REINHOLD, C. L." box="[210,515,713,734]" pageId="4" pageNumber="333" pagination="34 - 36" refId="ref11784" refString="DEELEMAN-REINHOLD, C. L. 1986. A new cribellate amaurobioid spider from Sumatra (Araneae: Agelenidae). Bulletin of the British Arachnological Society, 7: 34 - 36." type="journal article" year="1986">Deeleman-Reinhold, 1986: 34</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(
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||
<typeStatus id="54A6882C4B6CFFBF2B5392ABE023F7B4" box="[535,578,713,734]" pageId="4" pageNumber="333">type</typeStatus>
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||
species by monotypy
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||
<taxonomicName id="4C1D4D0D4B6CFFBF2A7292ABE6F3F7B4" ID-CoL="5BSNJ" authority="Deeleman-Reinhold" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[822,1170,713,734]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="species" species="bedel">
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||
<emphasis id="B969EA9C4B6CFFBF2A7292ABE1A0F7B4" box="[822,961,713,734]" italics="true" pageId="4" pageNumber="333">Vytfutia bedel</emphasis>
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Deeleman-Reinhold
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</taxonomicName>
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,
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</treatmentCitationGroup>
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<typeStatus id="54A6882C4B6CFFBF29DF9284E28EF791" box="[155,239,742,763]" pageId="4" pageNumber="333" type="holotype">holotype</typeStatus>
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male in
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<collectionCode id="ED0CAE4B4B6CFFBF28069284E3EDF791" box="[322,396,742,763]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="333" type="Museum">RMNH</collectionCode>
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, examined).
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<collectionCode id="ED0CAE4B4B6CFFBF2B429284E076F791" box="[518,535,742,763]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="333" type="Museum">V</collectionCode>
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.
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||
<bibRefCitation id="EF8C4B7F4B6CFFBF2B659284E125F791" author="DEELEMAN-REINHOLD, C. L." box="[545,836,742,763]" pageId="4" pageNumber="333" pagination="34 - 36" refId="ref11784" refString="DEELEMAN-REINHOLD, C. L. 1986. A new cribellate amaurobioid spider from Sumatra (Araneae: Agelenidae). Bulletin of the British Arachnological Society, 7: 34 - 36." type="journal article" year="1986">Deeleman-Reinhold, 1986: 34</bibRefCitation>
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, type
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<taxonomicName id="4C1D4D0D4B6CFFBF2A399284E2AFF672" authority="Deeleman-Reinhold, 1986" authorityName="Deeleman-Reinhold" authorityYear="1986" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="species" species="bedel">
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<emphasis id="B969EA9C4B6CFFBF2A399284E1A9F791" box="[893,968,742,763]" italics="true" pageId="4" pageNumber="333">V. bedel</emphasis>
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||
<bibRefCitation id="EF8C4B7F4B6CFFBF2A8A9284E6F7F791" author="DEELEMAN-REINHOLD, C. L." box="[974,1174,742,763]" pageId="4" pageNumber="333" pagination="34 - 36" refId="ref11784" refString="DEELEMAN-REINHOLD, C. L. 1986. A new cribellate amaurobioid spider from Sumatra (Araneae: Agelenidae). Bulletin of the British Arachnological Society, 7: 34 - 36." type="journal article" year="1986">Deeleman-Reinhold,</bibRefCitation>
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1986
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</taxonomicName>
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; N.B.: transferred from
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<taxonomicName id="4C1D4D0D4B6CFFBF28FA9361E04AF672" baseAuthorityName="Deeleman-Reinhold" baseAuthorityYear="1986" box="[446,555,771,792]" class="Arachnida" family="Agelenidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="family">Agelenidae</taxonomicName>
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to
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<taxonomicName id="4C1D4D0D4B6CFFBF2B0B9361E0B9F672" authorityName="Thorell" authorityYear="1870" box="[591,728,771,792]" class="Arachnida" family="Amaurobiidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="family">Amaurobiidae</taxonomicName>
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||
by
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||
<bibRefCitation id="EF8C4B7F4B6CFFBF2A459361E1AEF672" author="GRISWOLD, C. E." box="[769,975,771,792]" pageId="4" pageNumber="333" pagination="1 - 206" refId="ref11937" refString="GRISWOLD, C. E. 1990. A revision and phylogenetic analysis of the spider subfamily Phyxelidinae (Araneae, Amaurobiidae). Bulletin of the American Museum of Natural History, 196: 1 - 206." type="journal article" year="1990">Griswold, 1990: 186</bibRefCitation>
|
||
, to
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||
<taxonomicName id="4C1D4D0D4B6CFFBF2AB39361E615F672" authorityName="Lehtinen" authorityYear="1967" box="[1015,1140,771,792]" class="Arachnida" family="Phyxelididae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="family">Phyxelididae</taxonomicName>
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||
by Griswold et al., 1999: 59.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30765054B6CFFBF29EF9326E113F151" pageId="4" pageNumber="333" type="diagnosis">
|
||
<paragraph id="8BA2368E4B6CFFBF29EF9326E0BCF6B1" blockId="4.[123,1174,713,1725]" pageId="4" pageNumber="333">
|
||
<emphasis id="B969EA9C4B6CFFBF29EF9326E378F631" bold="true" box="[171,281,836,859]" pageId="4" pageNumber="333">Diagnosis</emphasis>
|
||
:
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF286D9326E31AF631" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[297,379,836,859]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA9C4B6CFFBF286D9326E31AF631" box="[297,379,836,859]" italics="true" pageId="4" pageNumber="333">Vytfutia</emphasis>
|
||
</taxonomicName>
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||
may be distinguished from other
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||
<taxonomicName id="4C1D4D0D4B6CFFBF2BB19326E1E1F631" authorityName="Lehtinen" authorityYear="1967" box="[757,896,836,859]" class="Arachnida" family="Phyxelididae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="family">Phyxelididae</taxonomicName>
|
||
by having the epigynum (
|
||
<figureCitation id="13262A0B4B6CFFBF29C79306E2BCF611" box="[131,221,868,891]" captionStart="FIGURES" captionStartId="18.[155,167,1684,1703]" captionTargetBox="[224,1059,1157,1665]" captionTargetId="figure-120@18.[215,1081,1140,1672]" captionTargetPageId="18" captionText="FIGURES 24, 25. Vytfutia halandrefana new species, female genitalia drawings, Tsingy de Bemaraha, Madagascar, CASENT9008848. 24. Epigynum, ventral. 25. Vulva, left half, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513187" httpUri="https://zenodo.org/record/11513187/files/figure.png" pageId="4" pageNumber="333">Figs. 24</figureCitation>
|
||
,
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||
<figureCitation id="13262A0B4B6CFFBF29AD9306E367F611" box="[233,262,868,891]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="4" pageNumber="333">39</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6CFFBF28569306E34EF611" box="[274,303,868,891]" captionStart="FIGURES" captionStartId="22.[155,167,1009,1028]" captionTargetBox="[134,1160,233,982]" captionTargetId="figure-19@22.[120,1176,216,992]" captionTargetPageId="22" captionText="FIGURES 41–44. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842, female genitalia, scanning electron micrographs (SEM). 41. Epigynum, ventral. 42. Vulva, dorsal. 43. Bennet’s gland pore (BG), left. 44. Pores of spermathecal head (HS), left. Scale bars, 41 = 100µm, 42 = 30µm, 43, 44 = 10µm." figureDoi="http://doi.org/10.5281/zenodo.11513199" httpUri="https://zenodo.org/record/11513199/files/figure.png" pageId="4" pageNumber="333">41</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6CFFBF287F9306E336F611" box="[315,343,868,891]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="4" pageNumber="333">48</figureCitation>
|
||
) with the posterior median lobe fused to the lateral lobes so that no suture is visible, by the presence of a retrolateral process (
|
||
<collectionCode id="ED0CAE4B4B6CFFBF2BD693E6E0D6F6F1" box="[658,695,900,923]" pageId="4" pageNumber="333">RL</collectionCode>
|
||
) on the male palpal tarsus and a sclerotized dorsal spur on male metatarsus I (
|
||
<figureCitation id="13262A0B4B6CFFBF28A993C6E026F6D1" box="[493,583,932,955]" captionStart="FIGURES" captionStartId="17.[155,167,1265,1284]" captionTargetBox="[120,1176,216,1249]" captionTargetId="figure-126@17.[120,1176,216,1249]" captionTargetPageId="17" captionText="FIGURES 15–19. Vytfutia bedel Deeleman-Reinhold, 1986, genitalia-drawings, Holotype male and paratype female collected from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. 15–17. Male left palp. 15. Tibia and tarsus, ventral. 16. Tibia, dorsal. 17. Tibia and tarsus, retrolateral. 18, 19. Female genitalia. 18. Epigynum, ventral. 19. Vulva, dorsal. C = conductor; D = Dorsal tibial apophysis; E = embolus; MA = median apophysis; RL = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.11513183" httpUri="https://zenodo.org/record/11513183/files/figure.png" pageId="4" pageNumber="333">Figs. 16</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6CFFBF2B1093C6E010F6D1" box="[596,625,932,955]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="4" pageNumber="333">37</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6CFFBF2B3A93C6E0FDF6D1" box="[638,668,932,955]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="4" pageNumber="333">46</figureCitation>
|
||
), by the entire chilum (
|
||
<figureCitation id="13262A0B4B6CFFBF2ADF93C6E1BCF6D1" box="[923,989,932,955]" captionStart="FIGURES" captionStartId="16.[155,167,1044,1063]" captionTargetBox="[121,1174,217,1025]" captionTargetId="figure-19@16.[120,1176,216,1026]" captionTargetPageId="16" captionText="FIGURES 1–11. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842, female, morphology photos. 1. Cephalothorax, anterior, showing chelicerae and palps. 2. PME, dorsal, showing canoe-shaped tapeta. 3. ALE and PLE, lateral, showing canoe-shaped tapeta. 4. Cephalothorax, anterior, showing AER, clypeus and chilum. 5. Trochanter I. 6. Trochanter IV. 7. Posterior margin of carapace and pedicel, dorsal. 8. Left metatarsus IV, retrolateral, showing calamistrum. 9. Left palp, prolateral, arrows to probasal thorns. 10. Left leg I, prolateral. 11. Left leg IV, retrolateral. Scale bars, 1, 8, 9, 10, 11 = 0.5 mm, 2, 3, 4 = 0.25 mm, 5, 6, 7 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11513179" httpUri="https://zenodo.org/record/11513179/files/figure.png" pageId="4" pageNumber="333">Fig. 4</figureCitation>
|
||
) and by the PLS lacking the stout, curved, apicolateral seta (
|
||
<figureCitation id="13262A0B4B6CFFBF2B0993A6E0B1F6B1" box="[589,720,964,987]" captionStart="FIGURES" captionStartId="16.[155,167,1550,1569]" captionTargetBox="[120,1176,1223,1532]" captionTargetId="figure-157@16.[120,1176,1223,1537]" captionTargetPageId="16" captionText="FIGURES 12–14. Spinnerets of female Phyxelididae. 12, 13. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842. 14. Ambohima ranohira Griswold, Wood and Carmichael, 2012, Ranohira, Madagascar, CASENT9019986. 12, 14. Ventral. 13. Lateral. Arrow and inset in 14 to stout, curved apicodorsal seta, which occurs in Phyxelidini and Vidoleini but not in Vytfutiini. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11513181" httpUri="https://zenodo.org/record/11513181/files/figure.png" pageId="4" pageNumber="333">Figs. 12, 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA2368E4B6CFFBF29EF9386E113F151" blockId="4.[123,1174,713,1725]" pageId="4" pageNumber="333">
|
||
<emphasis id="B969EA9C4B6CFFBF29EF9386E312F691" bold="true" box="[171,371,996,1019]" pageId="4" pageNumber="333">Synapomorphies:</emphasis>
|
||
Among the
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF2B4D9386E0F6F691" authorityName="Lehtinen" authorityYear="1967" box="[521,663,996,1019]" class="Arachnida" family="Phyxelididae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="family">Phyxelididae</taxonomicName>
|
||
, the dorsal spur on male metatarsus I (
|
||
<figureCitation id="13262A0B4B6CFFBF2D249386E2A0F171" captionStart="FIGURES" captionStartId="25.[155,167,1487,1506]" captionTargetBox="[153,1147,252,1446]" captionTargetId="figure-24@25.[120,1176,216,1475]" captionTargetPageId="25" captionText="FIGURES 56–61. Vytfutia spp., male leg I with clasping modification of metatarsus. 56, 58, 61. Dorsal. 57, 59, 60. Retrolateral. 56, 57. Vytfutia bedel Deeleman-Reinhold, 1986, Holotype male from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia, left. 58, 59. Vytfutia labalaba new species, Holotype male from the wall of a building at 1550m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842, right. 60, 61. Vytfutia pallens Deeleman-Reinhold, 1989, male paratype from Niah Cave, Sarawak, Malaysia, right. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.11513207" httpUri="https://zenodo.org/record/11513207/files/figure.png" pageId="4" pageNumber="333">Figs. 56–61</figureCitation>
|
||
), RTA on the male palpal tibia, and fusion of median and lateral sectors of the epigynum into a single plate (
|
||
<figureCitation id="13262A0B4B6CFFBF285C9446E313F151" box="[280,370,1060,1083]" captionStart="FIGURES" captionStartId="18.[155,167,1684,1703]" captionTargetBox="[224,1059,1157,1665]" captionTargetId="figure-120@18.[215,1081,1140,1672]" captionTargetPageId="18" captionText="FIGURES 24, 25. Vytfutia halandrefana new species, female genitalia drawings, Tsingy de Bemaraha, Madagascar, CASENT9008848. 24. Epigynum, ventral. 25. Vulva, left half, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513187" httpUri="https://zenodo.org/record/11513187/files/figure.png" pageId="4" pageNumber="333">Figs. 24</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6CFFBF283A9446E3FAF151" box="[382,411,1060,1083]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="4" pageNumber="333">39</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6CFFBF28E29446E3A2F151" box="[422,451,1060,1083]" captionStart="FIGURES" captionStartId="22.[155,167,1009,1028]" captionTargetBox="[134,1160,233,982]" captionTargetId="figure-19@22.[120,1176,216,992]" captionTargetPageId="22" captionText="FIGURES 41–44. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842, female genitalia, scanning electron micrographs (SEM). 41. Epigynum, ventral. 42. Vulva, dorsal. 43. Bennet’s gland pore (BG), left. 44. Pores of spermathecal head (HS), left. Scale bars, 41 = 100µm, 42 = 30µm, 43, 44 = 10µm." figureDoi="http://doi.org/10.5281/zenodo.11513199" httpUri="https://zenodo.org/record/11513199/files/figure.png" pageId="4" pageNumber="333">41</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6CFFBF288B9446E38AF151" box="[463,491,1060,1083]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="4" pageNumber="333">48</figureCitation>
|
||
) comprise
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF2B279446E0D4F151" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[611,693,1060,1083]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA9C4B6CFFBF2B279446E0D4F151" box="[611,693,1060,1083]" italics="true" pageId="4" pageNumber="333">Vytfutia</emphasis>
|
||
</taxonomicName>
|
||
synapomorphies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30765054B6CFFBD29EF9427E085F45E" lastPageId="6" lastPageNumber="335" pageId="4" pageNumber="333" type="description">
|
||
<paragraph id="8BA2368E4B6CFFBF29EF9427E0B3F196" blockId="4.[123,1174,713,1725]" pageId="4" pageNumber="333">
|
||
<emphasis id="B969EA9C4B6CFFBF29EF9427E292F136" bold="true" box="[171,243,1093,1116]" pageId="4" pageNumber="333">Notes:</emphasis>
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF29BD9427E380F136" authorityName="Griswold" authorityYear="2022" box="[249,481,1093,1116]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6CFFBF29BD9427E380F136" box="[249,481,1093,1116]" italics="true" pageId="4" pageNumber="333">Vytfutia halandrefana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF2B579427E01CF136" authorityName="Deeleman-Reinhold" authorityYear="1989" box="[531,637,1093,1116]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="species" species="pallens">
|
||
<emphasis id="B969EA9C4B6CFFBF2B579427E042F136" box="[531,547,1093,1116]" italics="true" pageId="4" pageNumber="333">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6CFFBF2B749427E01CF136" box="[560,637,1093,1116]" italics="true" pageId="4" pageNumber="333">pallens</emphasis>
|
||
</taxonomicName>
|
||
are the most morphologically divergent species of
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF293F9407E2ACF116" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[123,205,1125,1148]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA9C4B6CFFBF293F9407E2ACF116" box="[123,205,1125,1148]" italics="true" pageId="4" pageNumber="333">Vytfutia</emphasis>
|
||
</taxonomicName>
|
||
, and also very different to each other.
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF2B229407E12FF116" authorityName="Griswold" authorityYear="2022" box="[614,846,1125,1148]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6CFFBF2B229407E12FF116" box="[614,846,1125,1148]" italics="true" pageId="4" pageNumber="333">Vytfutia halandrefana</emphasis>
|
||
</taxonomicName>
|
||
is small, with a broad thoracic fovea, few leg spines and complex female genitalia with convoluted ducts. Unfortunately, the male of
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF29DC94C7E327F1D6" authorityName="Griswold" authorityYear="2022" box="[152,326,1189,1212]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6CFFBF29DC94C7E2C9F1D6" box="[152,168,1189,1212]" italics="true" pageId="4" pageNumber="333">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6CFFBF29F394C7E327F1D6" box="[183,326,1189,1212]" italics="true" pageId="4" pageNumber="333">halandrefana</emphasis>
|
||
</taxonomicName>
|
||
remains unknown. Vulval morphology suggests that the
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF2AEA94C7E63BF1D6" authorityName="Griswold" authorityYear="2022" box="[942,1114,1189,1212]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6CFFBF2AEA94C7E1DCF1D6" box="[942,957,1189,1212]" italics="true" pageId="4" pageNumber="333">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6CFFBF2A8F94C7E63BF1D6" box="[971,1114,1189,1212]" italics="true" pageId="4" pageNumber="333">halandrefana</emphasis>
|
||
</taxonomicName>
|
||
male palp has an elongate embolus.
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF288794A7E009F1B6" authorityName="Deeleman-Reinhold" authorityYear="1989" box="[451,616,1221,1244]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="species" species="pallens">
|
||
<emphasis id="B969EA9C4B6CFFBF288794A7E009F1B6" box="[451,616,1221,1244]" italics="true" pageId="4" pageNumber="333">Vytfutia pallens</emphasis>
|
||
</taxonomicName>
|
||
is much larger than other
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF2A3F94A7E1ACF1B6" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[891,973,1221,1244]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA9C4B6CFFBF2A3F94A7E1ACF1B6" box="[891,973,1221,1244]" italics="true" pageId="4" pageNumber="333">Vytfutia</emphasis>
|
||
</taxonomicName>
|
||
, with longer limbs and pale coloration, as befits a cave-dwelling organism.
|
||
</paragraph>
|
||
<paragraph id="8BA2368E4B6CFFBD29EF9567E0A6F479" blockId="4.[123,1174,713,1725]" lastBlockId="6.[123,1173,220,341]" lastPageId="6" lastPageNumber="335" pageId="4" pageNumber="333">
|
||
<emphasis id="B969EA9C4B6CFFBF29EF9567E359F076" bold="true" box="[171,312,1285,1308]" pageId="4" pageNumber="333">Description:</emphasis>
|
||
(
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF280E9567E3FDF076" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[330,412,1285,1308]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA9C4B6CFFBF280E9567E3FDF076" box="[330,412,1285,1308]" italics="true" pageId="4" pageNumber="333">Vytfutia</emphasis>
|
||
</taxonomicName>
|
||
) Total length 2.12–7.70. Markings differ significantly among species and are described under each species description. Carapace length 1.19–1.43 times width, height 0.39–0.53 width; thoracic fovea typically linear, length 0.11–0.18 carapace length, narrow oval in
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF293F9507E349F016" authorityName="Griswold" authorityYear="2022" box="[123,296,1381,1404]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6CFFBF293F9507E349F016" box="[123,296,1381,1404]" italics="true" pageId="4" pageNumber="333">V. halandrefana</emphasis>
|
||
</taxonomicName>
|
||
, length 0.07 carapace length.
|
||
<collectionCode id="ED0CAE4B4B6CFFBF2B3A9507E0CEF016" box="[638,687,1381,1404]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="4" pageNumber="333" type="Herbarium">PER</collectionCode>
|
||
straight to slightly recurved,
|
||
<collectionCode id="ED0CAE4B4B6CFFBF2ABA9507E652F016" box="[1022,1075,1381,1404]" pageId="4" pageNumber="333">AER</collectionCode>
|
||
straight, ocular area width 2.25–3.33 times length; clypeal height typically 1.20-1.70 times
|
||
<collectionCode id="ED0CAE4B4B6CFFBF2ABB95E7E64BF0F6" box="[1023,1066,1413,1436]" country="Australia" name="Australian Museum" pageId="4" pageNumber="333" type="Museum">AM</collectionCode>
|
||
diameter, but lower in
|
||
<taxonomicName id="4C1D4D0D4B6CFFBF284C95C7E3D4F0D6" authorityName="Griswold" authorityYear="2022" box="[264,437,1445,1468]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="333" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6CFFBF284C95C7E376F0D6" box="[264,279,1445,1468]" italics="true" pageId="4" pageNumber="333">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6CFFBF286295C7E3D4F0D6" box="[294,437,1445,1468]" italics="true" pageId="4" pageNumber="333">halandrefana</emphasis>
|
||
</taxonomicName>
|
||
, 0.60–0.80 times
|
||
<collectionCode id="ED0CAE4B4B6CFFBF2B3395C7E0C3F0D6" box="[631,674,1445,1468]" country="Australia" name="Australian Museum" pageId="4" pageNumber="333" type="Museum">AM</collectionCode>
|
||
diameter; chelicerae with small boss (
|
||
<figureCitation id="13262A0B4B6CFFBF2D0095C7E6EFF0D6" box="[1092,1166,1445,1468]" captionStart="FIGURES" captionStartId="16.[155,167,1044,1063]" captionTargetBox="[121,1174,217,1025]" captionTargetId="figure-19@16.[120,1176,216,1026]" captionTargetPageId="16" captionText="FIGURES 1–11. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842, female, morphology photos. 1. Cephalothorax, anterior, showing chelicerae and palps. 2. PME, dorsal, showing canoe-shaped tapeta. 3. ALE and PLE, lateral, showing canoe-shaped tapeta. 4. Cephalothorax, anterior, showing AER, clypeus and chilum. 5. Trochanter I. 6. Trochanter IV. 7. Posterior margin of carapace and pedicel, dorsal. 8. Left metatarsus IV, retrolateral, showing calamistrum. 9. Left palp, prolateral, arrows to probasal thorns. 10. Left leg I, prolateral. 11. Left leg IV, retrolateral. Scale bars, 1, 8, 9, 10, 11 = 0.5 mm, 2, 3, 4 = 0.25 mm, 5, 6, 7 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11513179" httpUri="https://zenodo.org/record/11513179/files/figure.png" pageId="4" pageNumber="333">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6CFFBF293F95A7E2F9F0B6" box="[123,152,1477,1500]" captionStart="FIGURES" captionStartId="20.[155,167,1187,1206]" captionTargetBox="[120,1176,216,1170]" captionTargetId="figure-19@20.[120,1176,216,1170]" captionTargetPageId="20" captionText="FIGURES 31–35. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842, female, habitus photos. 31. Dorsal. 32. Ventral. 33. Cephalothorax, ventral. 34. Cephalothorax, dorsal. 35. Cephalothorax, lateral. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11513193" httpUri="https://zenodo.org/record/11513193/files/figure.png" pageId="4" pageNumber="333">35</figureCitation>
|
||
), length 6.50–14.00 times clypeal height, pro- and retromargins of fang furrow with 3–5 heterogeneous teeth, retromargin with 3-4 escort setae at fang base, promargin with large group of fluffy whisker setae, rake setae not found (Ramirez 2014); cheliceral gland on flat cuticle, with numerous pores; labrum with flattened, anteriad pointing labral tongue, tongue apex free, deeply concave, with plumose setae dorsally and laterally on labrum, minute, bristle-like setae occur distad of tongue apex; sternum length 1.17–1.39 times width, apex a blunt point (
|
||
<figureCitation id="13262A0B4B6CFFBF2A8E9604E645F317" box="[970,1060,1638,1661]" captionStart="FIGURES" captionStartId="19.[155,167,1119,1138]" captionTargetBox="[120,1176,216,1100]" captionTargetId="figure-24@19.[120,1176,216,1100]" captionTargetPageId="19" captionText="FIGURES 26–30. Vytfutia labalaba new species, Holotype male, Mt. Kinabalu, CASENT9023842, habitus photos. 26. Carapace, dorsal. 27. Dorsal. 28. Ventral. 29. Cephalothorax, ventral. 30. Lateral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11513191" httpUri="https://zenodo.org/record/11513191/files/figure.png" pageId="4" pageNumber="333">Figs. 29</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6CFFBF2D749604E62FF317" box="[1072,1102,1638,1661]" captionStart="FIGURES" captionStartId="20.[155,167,1187,1206]" captionTargetBox="[120,1176,216,1170]" captionTargetId="figure-19@20.[120,1176,216,1170]" captionTargetPageId="20" captionText="FIGURES 31–35. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842, female, habitus photos. 31. Dorsal. 32. Ventral. 33. Cephalothorax, ventral. 34. Cephalothorax, dorsal. 35. Cephalothorax, lateral. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11513193" httpUri="https://zenodo.org/record/11513193/files/figure.png" pageId="4" pageNumber="333">33</figureCitation>
|
||
). Legs long, femur I 1.01–1.48 times carapace length; leg I always longest,
|
||
<collectionCode id="ED0CAE4B4B6CFFBF2A1296E4E110F3F7" box="[854,881,1670,1693]" pageId="4" pageNumber="333">III</collectionCode>
|
||
shortest, leg formula 1423, in some species 1243 or 1 [2=4] 3; male metatarsi I (
|
||
<figureCitation id="13262A0B4B6CFFBF2BAE96C4E117F3D7" box="[746,886,1702,1725]" captionStart="FIGURES" captionStartId="19.[155,167,1119,1138]" captionTargetBox="[120,1176,216,1100]" captionTargetId="figure-24@19.[120,1176,216,1100]" captionTargetPageId="19" captionText="FIGURES 26–30. Vytfutia labalaba new species, Holotype male, Mt. Kinabalu, CASENT9023842, habitus photos. 26. Carapace, dorsal. 27. Dorsal. 28. Ventral. 29. Cephalothorax, ventral. 30. Lateral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11513191" httpUri="https://zenodo.org/record/11513191/files/figure.png" pageId="4" pageNumber="333">Figs. 27, 28</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6CFFBF2AC396C4E1ABF3D7" box="[903,970,1702,1725]" captionStart="FIGURES" captionStartId="25.[155,167,1487,1506]" captionTargetBox="[153,1147,252,1446]" captionTargetId="figure-24@25.[120,1176,216,1475]" captionTargetPageId="25" captionText="FIGURES 56–61. Vytfutia spp., male leg I with clasping modification of metatarsus. 56, 58, 61. Dorsal. 57, 59, 60. Retrolateral. 56, 57. Vytfutia bedel Deeleman-Reinhold, 1986, Holotype male from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia, left. 58, 59. Vytfutia labalaba new species, Holotype male from the wall of a building at 1550m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842, right. 60, 61. Vytfutia pallens Deeleman-Reinhold, 1989, male paratype from Niah Cave, Sarawak, Malaysia, right. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.11513207" httpUri="https://zenodo.org/record/11513207/files/figure.png" pageId="4" pageNumber="333">56–61</figureCitation>
|
||
) with retrolateral, concavity at midsegment, concavity lacks spinules, at base of concavity a short, prolateral process surmounted by stout clasping spine, concavity arising at 0.21-0.47 length of metatarsus, concavity length 0.21–0.27 metatarsus, with short, dorsal spur at apex of concavity, spur height 0.06–0.95 of metatarsus length; legs otherwise unmodified; palpal claw and
|
||
<collectionCode id="ED0CAE4B4B6DFFBE2A5C915EE128F439" box="[792,841,316,339]" country="China" httpUri="http://grbio.org/cool/e5w9-0q9s" name="Sichuan Teacher's College" pageId="5" pageNumber="334">STC</collectionCode>
|
||
with numerous (up to 11) teeth (
|
||
<figureCitation id="13262A0B4B6DFFBE29C7913EE370F419" box="[131,273,348,371]" captionStart="FIGURES" captionStartId="24.[155,167,765,784]" captionTargetBox="[120,1173,216,746]" captionTargetId="figure-19@24.[120,1176,216,747]" captionTargetPageId="24" captionText="FIGURES 50–55. Vytfutia halandrefana new species, Berenty, CASENT9012420, female legs and palps, SEM. 50. Metatarsus IV, retrolateral, showing calamistrum. 51. Metatarsus IV, retrolateral, showing calamistral setae. 52. Tarsus IV, apical, showing STC and ITC. 53. Palpal femur, anterolateral, showing thorns (arrow). 54. Palpal femur, anterolateral thorn, close up. 55. Tarsus IV, retrolateral showing STC and ITC. Scale bars: 50, 300 µm; 51, 20 µm, 52, 55, 50 µm, 53, 200 µm, 54, 30 µm." figureDoi="http://doi.org/10.5281/zenodo.11513205" httpUri="https://zenodo.org/record/11513205/files/figure.png" pageId="5" pageNumber="334">Figs. 52, 55</figureCitation>
|
||
),
|
||
<collectionCode id="ED0CAE4B4B6DFFBE2868913EE336F419" box="[300,343,348,371]" country="0" httpUri="http://grbio.org/cool/hz43-zma8" name="International Transit Centre" pageId="5" pageNumber="334">ITC</collectionCode>
|
||
with 2-3 teeth or smooth (
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE2BD8913EE16CF419" authorityName="Deeleman-Reinhold" authorityYear="1989" box="[668,781,348,371]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="pallens">
|
||
<emphasis id="B969EA9C4B6DFFBE2BD8913EE0CDF419" box="[668,684,348,371]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE2B84913EE16CF419" box="[704,781,348,371]" italics="true" pageId="5" pageNumber="334">pallens</emphasis>
|
||
</taxonomicName>
|
||
); calamistrum subapical, origin at 0.22-0.29 distance from metatarsus base, length 0.33–0.46 metatarsus length (
|
||
<figureCitation id="13262A0B4B6DFFBE2D40911FE636F4FE" box="[1028,1111,381,404]" captionStart="FIGURES" captionStartId="16.[155,167,1044,1063]" captionTargetBox="[121,1174,217,1025]" captionTargetId="figure-19@16.[120,1176,216,1026]" captionTargetPageId="16" captionText="FIGURES 1–11. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842, female, morphology photos. 1. Cephalothorax, anterior, showing chelicerae and palps. 2. PME, dorsal, showing canoe-shaped tapeta. 3. ALE and PLE, lateral, showing canoe-shaped tapeta. 4. Cephalothorax, anterior, showing AER, clypeus and chilum. 5. Trochanter I. 6. Trochanter IV. 7. Posterior margin of carapace and pedicel, dorsal. 8. Left metatarsus IV, retrolateral, showing calamistrum. 9. Left palp, prolateral, arrows to probasal thorns. 10. Left leg I, prolateral. 11. Left leg IV, retrolateral. Scale bars, 1, 8, 9, 10, 11 = 0.5 mm, 2, 3, 4 = 0.25 mm, 5, 6, 7 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11513179" httpUri="https://zenodo.org/record/11513179/files/figure.png" pageId="5" pageNumber="334">Figs. 8</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6DFFBE2D2F911FE6E8F4FE" box="[1131,1161,381,404]" captionStart="FIGURES" captionStartId="24.[155,167,765,784]" captionTargetBox="[120,1173,216,746]" captionTargetId="figure-19@24.[120,1176,216,747]" captionTargetPageId="24" captionText="FIGURES 50–55. Vytfutia halandrefana new species, Berenty, CASENT9012420, female legs and palps, SEM. 50. Metatarsus IV, retrolateral, showing calamistrum. 51. Metatarsus IV, retrolateral, showing calamistral setae. 52. Tarsus IV, apical, showing STC and ITC. 53. Palpal femur, anterolateral, showing thorns (arrow). 54. Palpal femur, anterolateral thorn, close up. 55. Tarsus IV, retrolateral showing STC and ITC. Scale bars: 50, 300 µm; 51, 20 µm, 52, 55, 50 µm, 53, 200 µm, 54, 30 µm." figureDoi="http://doi.org/10.5281/zenodo.11513205" httpUri="https://zenodo.org/record/11513205/files/figure.png" pageId="5" pageNumber="334">50</figureCitation>
|
||
). Spination (following summary is based on males and females of
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE2A0191FFE19CF4DE" authorityName="Griswold" authorityYear="2022" box="[837,1021,413,436]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="labalaba">
|
||
<emphasis id="B969EA9C4B6DFFBE2A0191FFE19CF4DE" box="[837,1021,413,436]" italics="true" pageId="5" pageNumber="334">Vytfutia labalaba</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE2D4991FFE607F4DE" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[1037,1126,413,436]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="bedel">
|
||
<emphasis id="B969EA9C4B6DFFBE2D4991FFE67CF4DE" box="[1037,1053,413,436]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE2D6891FFE607F4DE" box="[1068,1126,413,436]" italics="true" pageId="5" pageNumber="334">bedel</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE293F91DFE287F4BE" authorityName="Deeleman-Reinhold" authorityYear="1989" box="[123,230,445,468]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="pallens">
|
||
<emphasis id="B969EA9C4B6DFFBE293F91DFE2EAF4BE" box="[123,139,445,468]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE29DD91DFE287F4BE" box="[153,230,445,468]" italics="true" pageId="5" pageNumber="334">pallens</emphasis>
|
||
</taxonomicName>
|
||
are very similar; the female of
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE2B7A91DFE08BF4BE" authorityName="Griswold" authorityYear="2022" box="[574,746,445,468]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6DFFBE2B7A91DFE02CF4BE" box="[574,589,445,468]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE2B1F91DFE08BF4BE" box="[603,746,445,468]" italics="true" pageId="5" pageNumber="334">halandrefana</emphasis>
|
||
</taxonomicName>
|
||
is very different, which is noted in that species’ description). Male: palp: femur d0-1-1, p0-1-0; leg I: femur d1-1-1, p0-0-0-1, r0-0-1, tibia d0-0-1, p0-1-1-0, r1-1-1-0, metatarsus p0-0-1, v2-0-0-2, r0-1(clasper)-0; leg II: femur d1-1-1, p0-0-1-0, r0-0-1-0, tibia p0-1-1-0, r0-1-2-0, metatarsus p0-1-1-2, r0-0-2; leg
|
||
<collectionCode id="ED0CAE4B4B6DFFBE2D6D927CE626F75F" box="[1065,1095,542,565]" pageId="5" pageNumber="334">III</collectionCode>
|
||
: femur d1-1-1, p0-0-1-0, r0-0-1-0, patella d0-1, p1, r1, tibia d1-0-0, p0-1-1-0, v0-0-1-0, r0-1-1-0, metatarsus p1-1-0-2, v2-0-0-2, r1-1-0-1; leg
|
||
<collectionCode id="ED0CAE4B4B6DFFBE2B5C923CE057F71F" box="[536,566,606,629]" pageId="5" pageNumber="334">IV</collectionCode>
|
||
: femur d1-1-1, p0-0-1-0, r0-0-1-0, patella d0-1-0, tibia d1-0-0, p0-1-1-0, v0-1-1-0, r0-1-1-0, metatarsus p1-1-0-2, v2-0-0-2, r1-1-0-1. Female: palp: femur d0-0-1-1, r0-0-1; leg I: femur d0-1-1-0, p0-0-0-1, tibia p0-1-1-0, r0-1-1-0, metatarsus p0-0-1, v2-0-0, r0-0-1; leg II: femur d1-1-1, p0-0-1, r0-0-1, tibia d1-1-0-2, v2-0-0-2, r1-1-0-2, p0-1-1-0, metatarsus d0-0-1, p0-1-1-0, v2-0-0-2, r0-0-1-0; leg
|
||
<collectionCode id="ED0CAE4B4B6DFFBE2A5992BDE15CF79C" box="[797,829,735,758]" pageId="5" pageNumber="334">III</collectionCode>
|
||
: femur d1-1-0-1, p0-0-0-1, r0-0-0-1, patella p1, r1, tibia p0-1-1-0, v1-1-0-0, r0-1-1-0, metatarsus p1-1-0-2, v2-0-2-1, r1-1-0-2; leg
|
||
<collectionCode id="ED0CAE4B4B6DFFBE28559342E34EF65D" box="[273,303,800,823]" pageId="5" pageNumber="334">IV</collectionCode>
|
||
: femur d1-0-0-1, p0-0-1, r0-0-1, patella d1, tibia p0-1-1-0, v1-1-0-0, r0-1-1-0, metatarsus d2-0-01, p1-1-0-2, v0-1-0-2, r0-0-0-1. Male epiandrum lacking spigots. Tracheae comprise four simple tubes that are limited to the abdomen (
|
||
<bibRefCitation id="EF8C4B7F4B6DFFBE2B889302E673F61D" author="DEELEMAN-REINHOLD, C. L." box="[716,1042,864,887]" pageId="5" pageNumber="334" pagination="34 - 36" refId="ref11784" refString="DEELEMAN-REINHOLD, C. L. 1986. A new cribellate amaurobioid spider from Sumatra (Araneae: Agelenidae). Bulletin of the British Arachnological Society, 7: 34 - 36." type="journal article" year="1986">Deeleman-Reinhold, 1986: 34</bibRefCitation>
|
||
). Spinnerets (based on scanning electron microscope [SEM] examination of three species: males and females of
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE293F93C3E3A7F6D2" authority="Deeleman-Reinhold" authorityName="Deeleman-Reinhold" authorityYear="1989" box="[123,454,929,952]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="pallens">
|
||
<emphasis id="B969EA9C4B6DFFBE293F93C3E2EAF6D2" box="[123,139,929,952]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE29DE93C3E286F6D2" box="[154,231,929,952]" italics="true" pageId="5" pageNumber="334">pallens</emphasis>
|
||
Deeleman-Reinhold
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF8C4B7F4B6DFFBE289193C3E0A1F6D2" author="GRISWOLD, C. E. & RAMIREZ, M. J. & CODDINGTON, J. A. & AND PLATNICK, N. I." box="[469,704,929,952]" pageId="5" pageNumber="334" pagination="1 - 324" refId="ref12029" refString="GRISWOLD, C. E., RAMIREZ, M. J., CODDINGTON, J. A. AND PLATNICK, N. I. 2005. Atlas of phylogenetic data for entelegyne spiders (Araneae: Araneomorphae: Entelegynae) with comments on their phylogeny. Proceedings of the California Academy of Sciences, ser. 4, 56 (Suppl. II): 1 - 324." type="journal article" year="2005">
|
||
Griswold
|
||
<emphasis id="B969EA9C4B6DFFBE2B0093C3E0E3F6D2" box="[580,642,929,952]" italics="true" pageId="5" pageNumber="334">et al.,</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
, figs. 46D, 47, 48) and of
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE2AA093C3E600F6D2" authority="Griswold, 2022" authorityName="Griswold" authorityYear="2022" box="[996,1121,929,952]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="labalaba" status="new species">
|
||
<emphasis id="B969EA9C4B6DFFBE2AA093C3E192F6D2" box="[996,1011,929,952]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE2D4693C3E600F6D2" box="[1026,1121,929,952]" italics="true" pageId="5" pageNumber="334">labalaba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A57E74B6DFFBE2D2D93C3E2A9F6B2" pageId="5" pageNumber="334" rank="species">new species</taxonomicNameLabel>
|
||
and a female of
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE28C093A3E050F6B2" authority="Griswold, 2022" authorityName="Griswold" authorityYear="2022" box="[388,561,961,984]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="halandrefana" status="new species">
|
||
<emphasis id="B969EA9C4B6DFFBE28C093A3E3F2F6B2" box="[388,403,961,984]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE28E693A3E050F6B2" box="[418,561,961,984]" italics="true" pageId="5" pageNumber="334">halandrefana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25A57E74B6DFFBE2B0493A3E0A6F6B2" box="[576,711,961,984]" pageId="5" pageNumber="334" rank="species">new species</taxonomicNameLabel>
|
||
).
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE2B9F93A3E14CF6B2" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[731,813,961,984]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA9C4B6DFFBE2B9F93A3E14CF6B2" box="[731,813,961,984]" italics="true" pageId="5" pageNumber="334">Vytfutia</emphasis>
|
||
</taxonomicName>
|
||
are typical phyxelidids in cribellum and paracribellar morphology, and universally differ from Vidoleini and Phyxelidini in lacking the large, stout, curved seta (black when viewed with light microscopy) at ectal margin of the PLS (compare
|
||
<figureCitation id="13262A0B4B6DFFBE28069443E38BF152" box="[322,490,1057,1080]" captionStart="FIGURES" captionStartId="16.[155,167,1550,1569]" captionTargetBox="[120,1176,1223,1532]" captionTargetId="figure-157@16.[120,1176,1223,1537]" captionTargetPageId="16" captionText="FIGURES 12–14. Spinnerets of female Phyxelididae. 12, 13. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842. 14. Ambohima ranohira Griswold, Wood and Carmichael, 2012, Ranohira, Madagascar, CASENT9019986. 12, 14. Ventral. 13. Lateral. Arrow and inset in 14 to stout, curved apicodorsal seta, which occurs in Phyxelidini and Vidoleini but not in Vytfutiini. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11513181" httpUri="https://zenodo.org/record/11513181/files/figure.png" pageId="5" pageNumber="334">Figs. 14 and 12</figureCitation>
|
||
). Cribellum width 0.43–0.67 spinnerets width (
|
||
<figureCitation id="13262A0B4B6DFFBE2AA09443E655F152" box="[996,1076,1057,1080]" captionStart="FIGURES" captionStartId="16.[155,167,1550,1569]" captionTargetBox="[120,1176,1223,1532]" captionTargetId="figure-157@16.[120,1176,1223,1537]" captionTargetPageId="16" captionText="FIGURES 12–14. Spinnerets of female Phyxelididae. 12, 13. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842. 14. Ambohima ranohira Griswold, Wood and Carmichael, 2012, Ranohira, Madagascar, CASENT9019986. 12, 14. Ventral. 13. Lateral. Arrow and inset in 14 to stout, curved apicodorsal seta, which occurs in Phyxelidini and Vidoleini but not in Vytfutiini. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11513181" httpUri="https://zenodo.org/record/11513181/files/figure.png" pageId="5" pageNumber="334">Fig. 12</figureCitation>
|
||
). Female ALS may have one mesal MAP spigot with a posterior MAP nubbin;
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE2A3A9420E64BF133" authorityName="Griswold" authorityYear="2022" box="[894,1066,1090,1113]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6DFFBE2A3A9420E64BF133" box="[894,1066,1090,1113]" italics="true" pageId="5" pageNumber="334">V. halandrefana</emphasis>
|
||
</taxonomicName>
|
||
differs in retaining two MAP spigots. Piriform gland (
|
||
<collectionCode id="ED0CAE4B4B6DFFBE2B309400E0EFF113" box="[628,654,1122,1145]" country="Russia" name="Paleontological Institute" pageId="5" pageNumber="334">PI</collectionCode>
|
||
) spigots in females range from as few as 18 (
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE29C094E0E352F1F3" authorityName="Griswold" authorityYear="2022" box="[132,307,1154,1177]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6DFFBE29C094E0E2F5F1F3" box="[132,148,1154,1177]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE29E094E0E352F1F3" box="[164,307,1154,1177]" italics="true" pageId="5" pageNumber="334">halandrefana</emphasis>
|
||
</taxonomicName>
|
||
) to more than 50 (
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE2B4294E0E012F1F3" authorityName="Deeleman-Reinhold" authorityYear="1989" box="[518,627,1154,1177]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="pallens">
|
||
<emphasis id="B969EA9C4B6DFFBE2B4294E0E077F1F3" box="[518,534,1154,1177]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE2B6294E0E012F1F3" box="[550,627,1154,1177]" italics="true" pageId="5" pageNumber="334">pallens</emphasis>
|
||
</taxonomicName>
|
||
); males have fewer; small tartipores are scattered throughout
|
||
<collectionCode id="ED0CAE4B4B6DFFBE29BE94C0E373F1D3" box="[250,274,1186,1209]" country="Russia" name="Paleontological Institute" pageId="5" pageNumber="334">PI</collectionCode>
|
||
spigot field; female
|
||
<collectionCode id="ED0CAE4B4B6DFFBE28B894C0E050F1D3" box="[508,561,1186,1209]" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" pageId="5" pageNumber="334">PMS</collectionCode>
|
||
with as few as 10 (
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE2A6494C0E1B1F1D3" authorityName="Griswold" authorityYear="2022" box="[800,976,1186,1209]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6DFFBE2A6494C0E151F1D3" box="[800,816,1186,1209]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE2A0594C0E1B1F1D3" box="[833,976,1186,1209]" italics="true" pageId="5" pageNumber="334">halandrefana</emphasis>
|
||
</taxonomicName>
|
||
) to more than 25 (
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE29C094A1E361F1B0" authorityName="Griswold" authorityYear="2022" box="[132,256,1219,1242]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="labalaba">
|
||
<emphasis id="B969EA9C4B6DFFBE29C094A1E2F5F1B0" box="[132,148,1219,1242]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE29E594A1E361F1B0" box="[161,256,1219,1242]" italics="true" pageId="5" pageNumber="334">labalaba</emphasis>
|
||
</taxonomicName>
|
||
) PC spigots; replaced by nubbins in the male; the
|
||
<collectionCode id="ED0CAE4B4B6DFFBE2A5294A1E12AF1B0" box="[790,843,1219,1242]" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" pageId="5" pageNumber="334">PMS</collectionCode>
|
||
of both sexes have one anteromedian mAP spigot and at least two (
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE2B589481E0AAF190" authorityName="Griswold" authorityYear="2022" box="[540,715,1251,1274]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6DFFBE2B589481E04DF190" box="[540,556,1251,1274]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE2B789481E0AAF190" box="[572,715,1251,1274]" italics="true" pageId="5" pageNumber="334">halandrefana</emphasis>
|
||
</taxonomicName>
|
||
) to six (
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE2A689481E1CDF190" authorityName="Griswold" authorityYear="2022" box="[812,940,1251,1274]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="labalaba">
|
||
<emphasis id="B969EA9C4B6DFFBE2A689481E15DF190" box="[812,828,1251,1274]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE2A099481E1CDF190" box="[845,940,1251,1274]" italics="true" pageId="5" pageNumber="334">labalaba</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<collectionCode id="ED0CAE4B4B6DFFBE2AF89481E180F190" box="[956,993,1251,1274]" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" pageId="5" pageNumber="334" type="University or college">AC</collectionCode>
|
||
spigots and one (
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE29C09561E350F070" authorityName="Griswold" authorityYear="2022" box="[132,305,1283,1306]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6DFFBE29C09561E2F5F070" box="[132,148,1283,1306]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE29E69561E350F070" box="[162,305,1283,1306]" italics="true" pageId="5" pageNumber="334">halandrefana</emphasis>
|
||
</taxonomicName>
|
||
) to four (
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE28DE9561E067F070" authorityName="Deeleman-Reinhold" authorityYear="1989" box="[410,518,1283,1306]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="pallens">
|
||
<emphasis id="B969EA9C4B6DFFBE28DE9561E3CBF070" box="[410,426,1283,1306]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE28FD9561E067F070" box="[441,518,1283,1306]" italics="true" pageId="5" pageNumber="334">pallens</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<collectionCode id="ED0CAE4B4B6DFFBE2B529561E05BF070" box="[534,570,1283,1306]" country="France" name="Centre des Yersinia" pageId="5" pageNumber="334">CY</collectionCode>
|
||
spigots; female PLS with large anterior MS spigot with cylindrical base and shaft, accompanied by a flanking
|
||
<collectionCode id="ED0CAE4B4B6DFFBE2B8A9541E097F050" box="[718,758,1315,1338]" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" pageId="5" pageNumber="334" type="University or college">AC</collectionCode>
|
||
, MS and flanking
|
||
<collectionCode id="ED0CAE4B4B6DFFBE2A859541E187F050" box="[961,998,1315,1338]" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" pageId="5" pageNumber="334" type="University or college">AC</collectionCode>
|
||
are replaced by nubbins in male; female with one (
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE28BC9526E0C7F031" authorityName="Griswold" authorityYear="2022" box="[504,678,1348,1371]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6DFFBE28BC9526E069F031" box="[504,520,1348,1371]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE2B539526E0C7F031" box="[535,678,1348,1371]" italics="true" pageId="5" pageNumber="334">halandrefana</emphasis>
|
||
</taxonomicName>
|
||
) to three (
|
||
<taxonomicName id="4C1D4D0D4B6DFFBE2A5F9526E1F8F031" authorityName="Griswold" authorityYear="2022" box="[795,921,1348,1371]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="334" phylum="Arthropoda" rank="species" species="labalaba">
|
||
<emphasis id="B969EA9C4B6DFFBE2A5F9526E14AF031" box="[795,811,1348,1371]" italics="true" pageId="5" pageNumber="334">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6DFFBE2A7E9526E1F8F031" box="[826,921,1348,1371]" italics="true" pageId="5" pageNumber="334">labalaba</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<collectionCode id="ED0CAE4B4B6DFFBE2AEE9526E1AFF031" box="[938,974,1348,1371]" country="France" name="Centre des Yersinia" pageId="5" pageNumber="334">CY</collectionCode>
|
||
spigots and fewer than eight
|
||
<collectionCode id="ED0CAE4B4B6DFFBE29AD9506E36FF011" box="[233,270,1380,1403]" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" pageId="5" pageNumber="334" type="University or college">AC</collectionCode>
|
||
spigots. Palpal femur with anterobasal row of four to eight stout setae set in enlarged bases, these setae shortened and enlarged as thorns (
|
||
<figureCitation id="13262A0B4B6DFFBE2BE795E6E08FF0F1" box="[675,750,1412,1435]" captionStart="FIGURES" captionStartId="16.[155,167,1044,1063]" captionTargetBox="[121,1174,217,1025]" captionTargetId="figure-19@16.[120,1176,216,1026]" captionTargetPageId="16" captionText="FIGURES 1–11. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842, female, morphology photos. 1. Cephalothorax, anterior, showing chelicerae and palps. 2. PME, dorsal, showing canoe-shaped tapeta. 3. ALE and PLE, lateral, showing canoe-shaped tapeta. 4. Cephalothorax, anterior, showing AER, clypeus and chilum. 5. Trochanter I. 6. Trochanter IV. 7. Posterior margin of carapace and pedicel, dorsal. 8. Left metatarsus IV, retrolateral, showing calamistrum. 9. Left palp, prolateral, arrows to probasal thorns. 10. Left leg I, prolateral. 11. Left leg IV, retrolateral. Scale bars, 1, 8, 9, 10, 11 = 0.5 mm, 2, 3, 4 = 0.25 mm, 5, 6, 7 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11513179" httpUri="https://zenodo.org/record/11513179/files/figure.png" pageId="5" pageNumber="334">Figs. 9</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6DFFBE2BBD95E6E15EF0F1" box="[761,831,1412,1435]" captionStart="FIGURES" captionStartId="24.[155,167,765,784]" captionTargetBox="[120,1173,216,746]" captionTargetId="figure-19@24.[120,1176,216,747]" captionTargetPageId="24" captionText="FIGURES 50–55. Vytfutia halandrefana new species, Berenty, CASENT9012420, female legs and palps, SEM. 50. Metatarsus IV, retrolateral, showing calamistrum. 51. Metatarsus IV, retrolateral, showing calamistral setae. 52. Tarsus IV, apical, showing STC and ITC. 53. Palpal femur, anterolateral, showing thorns (arrow). 54. Palpal femur, anterolateral thorn, close up. 55. Tarsus IV, retrolateral showing STC and ITC. Scale bars: 50, 300 µm; 51, 20 µm, 52, 55, 50 µm, 53, 200 µm, 54, 30 µm." figureDoi="http://doi.org/10.5281/zenodo.11513205" httpUri="https://zenodo.org/record/11513205/files/figure.png" pageId="5" pageNumber="334">53, 54</figureCitation>
|
||
); male palpal tibia (
|
||
<figureCitation id="13262A0B4B6DFFBE2D5595E6E60BF0F1" box="[1041,1130,1412,1435]" captionStart="FIGURES" captionStartId="17.[155,167,1265,1284]" captionTargetBox="[120,1176,216,1249]" captionTargetId="figure-126@17.[120,1176,216,1249]" captionTargetPageId="17" captionText="FIGURES 15–19. Vytfutia bedel Deeleman-Reinhold, 1986, genitalia-drawings, Holotype male and paratype female collected from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. 15–17. Male left palp. 15. Tibia and tarsus, ventral. 16. Tibia, dorsal. 17. Tibia and tarsus, retrolateral. 18, 19. Female genitalia. 18. Epigynum, ventral. 19. Vulva, dorsal. C = conductor; D = Dorsal tibial apophysis; E = embolus; MA = median apophysis; RL = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.11513183" httpUri="https://zenodo.org/record/11513183/files/figure.png" pageId="5" pageNumber="334">Figs. 16</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6DFFBE2D3095E6E2F6F0D1" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="5" pageNumber="334">46, 47</figureCitation>
|
||
) with short to long, convex, simple
|
||
<collectionCode id="ED0CAE4B4B6DFFBE2B6D95C6E03DF0D1" box="[553,604,1444,1467]" pageId="5" pageNumber="334">DTA</collectionCode>
|
||
extending apically, RTA developed as a trapezoidal flange; cymbium with retrolateral paracribellar projection (
|
||
<figureCitation id="13262A0B4B6DFFBE2BB195A6E12EF0B1" box="[757,847,1476,1499]" captionStart="FIGURES" captionStartId="17.[155,167,1265,1284]" captionTargetBox="[120,1176,216,1249]" captionTargetId="figure-126@17.[120,1176,216,1249]" captionTargetPageId="17" captionText="FIGURES 15–19. Vytfutia bedel Deeleman-Reinhold, 1986, genitalia-drawings, Holotype male and paratype female collected from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. 15–17. Male left palp. 15. Tibia and tarsus, ventral. 16. Tibia, dorsal. 17. Tibia and tarsus, retrolateral. 18, 19. Female genitalia. 18. Epigynum, ventral. 19. Vulva, dorsal. C = conductor; D = Dorsal tibial apophysis; E = embolus; MA = median apophysis; RL = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.11513183" httpUri="https://zenodo.org/record/11513183/files/figure.png" pageId="5" pageNumber="334">Figs. 15</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6DFFBE2A1F95A6E119F0B1" box="[859,888,1476,1499]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="5" pageNumber="334">38</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6DFFBE2AC095A6E1AAF0B1" box="[900,971,1476,1499]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="5" pageNumber="334">46, 47</figureCitation>
|
||
); palpal bulb with a small, oval petiole, anneli of subtegulum weakly developed, tegulum central, convex, course of reservoir a simple curve within bulb, without switchbacks, median apophysis present, fleshy, arising at mid-bulb near embolic base; embolus a slender, narrow, tapering blade to a slender spine, pars pendula fused basally with truncus of embolus, embolus flexibly attached to tegulum; conductor apical, fleshy, grooved to receive apex of embolus (
|
||
<figureCitation id="13262A0B4B6DFFBE2A6E9604E1E7F317" box="[810,902,1638,1661]" captionStart="FIGURES" captionStartId="17.[155,167,1265,1284]" captionTargetBox="[120,1176,216,1249]" captionTargetId="figure-126@17.[120,1176,216,1249]" captionTargetPageId="17" captionText="FIGURES 15–19. Vytfutia bedel Deeleman-Reinhold, 1986, genitalia-drawings, Holotype male and paratype female collected from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. 15–17. Male left palp. 15. Tibia and tarsus, ventral. 16. Tibia, dorsal. 17. Tibia and tarsus, retrolateral. 18, 19. Female genitalia. 18. Epigynum, ventral. 19. Vulva, dorsal. C = conductor; D = Dorsal tibial apophysis; E = embolus; MA = median apophysis; RL = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.11513183" httpUri="https://zenodo.org/record/11513183/files/figure.png" pageId="5" pageNumber="334">Figs. 15</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6DFFBE2AD79604E1BAF317" box="[915,987,1638,1661]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="5" pageNumber="334">36, 37</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6DFFBE2AAD9604E650F317" box="[1001,1073,1638,1661]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="5" pageNumber="334">45, 46</figureCitation>
|
||
). Female genitalia with epigynum (
|
||
<figureCitation id="13262A0B4B6DFFBE28D496E4E38AF3F7" box="[400,491,1670,1693]" captionStart="FIGURES" captionStartId="17.[155,167,1265,1284]" captionTargetBox="[120,1176,216,1249]" captionTargetId="figure-126@17.[120,1176,216,1249]" captionTargetPageId="17" captionText="FIGURES 15–19. Vytfutia bedel Deeleman-Reinhold, 1986, genitalia-drawings, Holotype male and paratype female collected from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. 15–17. Male left palp. 15. Tibia and tarsus, ventral. 16. Tibia, dorsal. 17. Tibia and tarsus, retrolateral. 18, 19. Female genitalia. 18. Epigynum, ventral. 19. Vulva, dorsal. C = conductor; D = Dorsal tibial apophysis; E = embolus; MA = median apophysis; RL = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.11513183" httpUri="https://zenodo.org/record/11513183/files/figure.png" pageId="5" pageNumber="334">Figs. 18</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6DFFBE28B396E4E075F3F7" box="[503,532,1670,1693]" captionStart="FIGURES" captionStartId="18.[155,167,1684,1703]" captionTargetBox="[224,1059,1157,1665]" captionTargetId="figure-120@18.[215,1081,1140,1672]" captionTargetPageId="18" captionText="FIGURES 24, 25. Vytfutia halandrefana new species, female genitalia drawings, Tsingy de Bemaraha, Madagascar, CASENT9008848. 24. Epigynum, ventral. 25. Vulva, left half, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513187" httpUri="https://zenodo.org/record/11513187/files/figure.png" pageId="5" pageNumber="334">24</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6DFFBE2B6496E4E05CF3F7" box="[544,573,1670,1693]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="5" pageNumber="334">39</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6DFFBE2B0D96E4E004F3F7" box="[585,613,1670,1693]" captionStart="FIGURES" captionStartId="22.[155,167,1009,1028]" captionTargetBox="[134,1160,233,982]" captionTargetId="figure-19@22.[120,1176,216,992]" captionTargetPageId="22" captionText="FIGURES 41–44. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842, female genitalia, scanning electron micrographs (SEM). 41. Epigynum, ventral. 42. Vulva, dorsal. 43. Bennet’s gland pore (BG), left. 44. Pores of spermathecal head (HS), left. Scale bars, 41 = 100µm, 42 = 30µm, 43, 44 = 10µm." figureDoi="http://doi.org/10.5281/zenodo.11513199" httpUri="https://zenodo.org/record/11513199/files/figure.png" pageId="5" pageNumber="334">41</figureCitation>
|
||
) a simple, undivided plate, copulatory openings slit like, transverse, oblique to longitudinal; vulva with capsulate structures(s) with internal chambers, with 2-3 lobes, at least
|
||
<taxonomicName id="4C1D4D0D4B6EFFBD282B90BEE38AF599" authorityName="Griswold" authorityYear="2022" box="[367,491,220,243]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="335" phylum="Arthropoda" rank="species" species="labalaba">
|
||
<emphasis id="B969EA9C4B6EFFBD282B90BEE31EF599" box="[367,383,220,243]" italics="true" pageId="6" pageNumber="335">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6EFFBD28C890BEE38AF599" box="[396,491,220,243]" italics="true" pageId="6" pageNumber="335">labalaba</emphasis>
|
||
</taxonomicName>
|
||
with sessile HS and posterdorsal Bennett’s gland pore (
|
||
<figureCitation id="13262A0B4B6EFFBD2D7E90BEE6E8F599" box="[1082,1161,220,243]" captionStart="FIGURES" captionStartId="22.[155,167,1009,1028]" captionTargetBox="[134,1160,233,982]" captionTargetId="figure-19@22.[120,1176,216,992]" captionTargetPageId="22" captionText="FIGURES 41–44. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842, female genitalia, scanning electron micrographs (SEM). 41. Epigynum, ventral. 42. Vulva, dorsal. 43. Bennet’s gland pore (BG), left. 44. Pores of spermathecal head (HS), left. Scale bars, 41 = 100µm, 42 = 30µm, 43, 44 = 10µm." figureDoi="http://doi.org/10.5281/zenodo.11513199" httpUri="https://zenodo.org/record/11513199/files/figure.png" pageId="6" pageNumber="335">Fig. 42</figureCitation>
|
||
), may have convoluted ducts (
|
||
<taxonomicName id="4C1D4D0D4B6EFFBD28F4909EE03CF479" authorityName="Griswold" authorityYear="2022" box="[432,605,252,275]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="335" phylum="Arthropoda" rank="species" species="halandrefana">
|
||
<emphasis id="B969EA9C4B6EFFBD28F4909EE3A1F479" box="[432,448,252,275]" italics="true" pageId="6" pageNumber="335">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6EFFBD288A909EE03CF479" box="[462,605,252,275]" italics="true" pageId="6" pageNumber="335">halandrefana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="13262A0B4B6EFFBD2B2F909EE0DBF479" box="[619,698,252,275]" captionStart="FIGURES" captionStartId="18.[155,167,1684,1703]" captionTargetBox="[224,1059,1157,1665]" captionTargetId="figure-120@18.[215,1081,1140,1672]" captionTargetPageId="18" captionText="FIGURES 24, 25. Vytfutia halandrefana new species, female genitalia drawings, Tsingy de Bemaraha, Madagascar, CASENT9008848. 24. Epigynum, ventral. 25. Vulva, left half, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513187" httpUri="https://zenodo.org/record/11513187/files/figure.png" pageId="6" pageNumber="335">Fig. 25</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA2368E4B6EFFBD29EF917FE085F45E" blockId="6.[123,1173,220,341]" box="[171,740,285,308]" pageId="6" pageNumber="335">
|
||
<emphasis id="B969EA9C4B6EFFBD29EF917FE35DF45E" bold="true" box="[171,316,285,308]" pageId="6" pageNumber="335">Composition</emphasis>
|
||
: Four species, all in the genus
|
||
<taxonomicName id="4C1D4D0D4B6EFFBD2BC8917FE0BCF45E" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[652,733,285,308]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EA9C4B6EFFBD2BC8917FE0BCF45E" box="[652,733,285,308]" italics="true" pageId="6" pageNumber="335">Vytfutia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30765054B6EFFBD29EF915CE627F43F" box="[171,1094,318,341]" pageId="6" pageNumber="335" type="distribution">
|
||
<paragraph id="8BA2368E4B6EFFBD29EF915CE627F43F" blockId="6.[123,1173,220,341]" box="[171,1094,318,341]" pageId="6" pageNumber="335">
|
||
<emphasis id="B969EA9C4B6EFFBD29EF915CE357F43F" bold="true" box="[171,310,318,341]" pageId="6" pageNumber="335">Distribution</emphasis>
|
||
: The Indo-Pacific islands of
|
||
<collectingCountry id="F30A761E4B6EFFBD2B2B915CE093F43F" box="[623,754,318,341]" name="Madagascar" pageId="6" pageNumber="335">Madagascar</collectingCountry>
|
||
, Sumatra and Borneo (
|
||
<figureCitation id="13262A0B4B6EFFBD2AAD915CE658F43F" box="[1001,1081,318,341]" captionStart="FIGURE" captionStartId="26.[155,167,1071,1090]" captionTargetBox="[120,1176,216,1056]" captionTargetId="figure-19@26.[120,1176,216,1056]" captionTargetPageId="26" captionText="FIGURE 62. Vytfutia spp. Distributions on map of Eurasia and Australasia. Filled circle. Vytfutia bedel Deeleman-Reinhold, 1986, Holotype male from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. Triangle. Vytfutia labalaba new species, Holotype male from the wall of a building at 1550m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia. Diamond. Vytfutia halandrefana new species, Madagascar. Star. Vytfutia pallens Deeleman-Reinhold, 1989, male paratype from Niah Cave, Sarawak, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.11513211" httpUri="https://zenodo.org/record/11513211/files/figure.png" pageId="6" pageNumber="335">Fig. 62</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30765054B6EFFBD28A6911DE6EEF3FD" pageId="6" pageNumber="335" type="key">
|
||
<paragraph id="8BA2368E4B6EFFBD28A6911DE14FF4FC" blockId="6.[482,814,383,406]" box="[482,814,383,406]" pageId="6" pageNumber="335">
|
||
<heading id="D0EA81E24B6EFFBD28A6911DE14FF4FC" bold="true" box="[482,814,383,406]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="335" reason="5">
|
||
<emphasis id="B969EA9C4B6EFFBD28A6911DE14FF4FC" bold="true" box="[482,814,383,406]" pageId="6" pageNumber="335">
|
||
Key to the species of
|
||
<taxonomicName id="4C1D4D0D4B6EFFBD2B96911DE14FF4FC" ID-CoL="87CS" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[722,814,383,406]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="335" phylum="Arthropoda" rank="genus">Vytfutia</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<key id="FF8C61BF4B6EFFBD293F91C9E6EEF3FD" pageId="6" pageNumber="335">
|
||
<keyStep id="30E92E2B4B6EFFBD293F91C9E6EFF489" pageId="6" pageNumber="335">
|
||
<paragraph id="8BA2368E4B6EFFBD293F91C9E6EFF4A8" blockId="6.[123,1166,427,483]" box="[123,1166,427,450]" pageId="6" pageNumber="335">
|
||
<keyLead id="30EC95BB4B6EFFBD293F91C9E6EFF4A8" box="[123,1166,427,450]" pageId="6" pageNumber="335">
|
||
<specimenCount id="9D1BFD074B6EFFBD293F91C9E28FF4A8" box="[123,238,427,450]" count="1" pageId="6" pageNumber="335" type="male">
|
||
<emphasis id="B969EA9C4B6EFFBD293F91C9E2E9F4A8" bold="true" box="[123,136,427,450]" pageId="6" pageNumber="335">1</emphasis>
|
||
Males
|
||
</specimenCount>
|
||
....................................................................
|
||
<emphasis id="B969EA9C4B6EFFBD2DC591C9E6EFF4A8" bold="true" box="[1153,1166,427,450]" pageId="6" pageNumber="335">5</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA2368E4B6EFFBD293F91AEE6EFF489" blockId="6.[123,1166,427,483]" box="[123,1166,460,483]" pageId="6" pageNumber="335">
|
||
<keyLead id="30EC95BB4B6EFFBD293F91AEE6EFF489" box="[123,1166,460,483]" pageId="6" pageNumber="335">
|
||
— Females..................................................................
|
||
<emphasis id="B969EA9C4B6EFFBD2DC591AEE6EFF489" bold="true" box="[1153,1166,460,483]" pageId="6" pageNumber="335">2</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E92E2B4B6EFFBD293F919AE6EEF79F" pageId="6" pageNumber="335">
|
||
<paragraph id="8BA2368E4B6EFFBD293F919AE6EFF71B" blockId="6.[123,1174,504,757]" pageId="6" pageNumber="335">
|
||
<keyLead id="30EC95BB4B6EFFBD293F919AE6EFF71B" pageId="6" pageNumber="335">
|
||
<emphasis id="B969EA9C4B6EFFBD293F919AE2CFF765" bold="true" box="[123,174,504,527]" pageId="6" pageNumber="335">2 (1)</emphasis>
|
||
Epigynum deep, quadrangle less than 1.00, copulatory opening rims oblique to nearly longitudinal, posterior margin of epigynum convex or concave (
|
||
<figureCitation id="13262A0B4B6EFFBD2A74927BE1EAF75A" box="[816,907,537,560]" captionStart="FIGURES" captionStartId="17.[155,167,1265,1284]" captionTargetBox="[120,1176,216,1249]" captionTargetId="figure-126@17.[120,1176,216,1249]" captionTargetPageId="17" captionText="FIGURES 15–19. Vytfutia bedel Deeleman-Reinhold, 1986, genitalia-drawings, Holotype male and paratype female collected from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. 15–17. Male left palp. 15. Tibia and tarsus, ventral. 16. Tibia, dorsal. 17. Tibia and tarsus, retrolateral. 18, 19. Female genitalia. 18. Epigynum, ventral. 19. Vulva, dorsal. C = conductor; D = Dorsal tibial apophysis; E = embolus; MA = median apophysis; RL = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.11513183" httpUri="https://zenodo.org/record/11513183/files/figure.png" pageId="6" pageNumber="335">Figs. 18</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6EFFBD2ADC927BE1D4F75A" box="[920,949,537,560]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="6" pageNumber="335">39</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6EFFBD2A86927BE181F75A" box="[962,992,537,560]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="6" pageNumber="335">48</figureCitation>
|
||
), vulva with 2-3 simple lobes, without slender ducts, vulva width greater than twice length (
|
||
<figureCitation id="13262A0B4B6EFFBD2A979258E64CF73B" box="[979,1069,570,593]" captionStart="FIGURES" captionStartId="17.[155,167,1265,1284]" captionTargetBox="[120,1176,216,1249]" captionTargetId="figure-126@17.[120,1176,216,1249]" captionTargetPageId="17" captionText="FIGURES 15–19. Vytfutia bedel Deeleman-Reinhold, 1986, genitalia-drawings, Holotype male and paratype female collected from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. 15–17. Male left palp. 15. Tibia and tarsus, ventral. 16. Tibia, dorsal. 17. Tibia and tarsus, retrolateral. 18, 19. Female genitalia. 18. Epigynum, ventral. 19. Vulva, dorsal. C = conductor; D = Dorsal tibial apophysis; E = embolus; MA = median apophysis; RL = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.11513183" httpUri="https://zenodo.org/record/11513183/files/figure.png" pageId="6" pageNumber="335">Figs. 19</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6EFFBD2D7D9258E637F73B" box="[1081,1110,570,593]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="6" pageNumber="335">40</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6EFFBD2D259258E61CF73B" box="[1121,1149,570,593]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="6" pageNumber="335">49</figureCitation>
|
||
) .........................................................................
|
||
<emphasis id="B969EA9C4B6EFFBD2DC59238E6EFF71B" bold="true" box="[1153,1166,602,625]" pageId="6" pageNumber="335">3</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA2368E4B6EFFBD293F9219E6EEF79F" blockId="6.[123,1174,504,757]" pageId="6" pageNumber="335">
|
||
<keyLead id="30EC95BB4B6EFFBD293F9219E6EEF79F" pageId="6" pageNumber="335">
|
||
— Epigynum broad, quadrangle greater than 2.50, copulatory opening rims transverse, posterior margin of epigynum concave (
|
||
<figureCitation id="13262A0B4B6EFFBD2B5892FEE019F7D9" box="[540,632,668,691]" captionStart="FIGURES" captionStartId="18.[155,167,1051,1070]" captionTargetBox="[216,1080,216,1035]" captionTargetId="figure-19@18.[216,1080,216,1035]" captionTargetPageId="18" captionText="FIGURES 20–23. Vytfutia halandrefana new species, female, Berenty, Madagascar, CASENT9012513, habitus photos. 20. Dorsal. 21. Ventral. 22. Left metatarsus-tarsus IV and apex of abdomen with spinnerets, ventral. 23. Epigynal region, dorsal. Scale bars 20–22 = 0.5 mm, 23 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.11513185" httpUri="https://zenodo.org/record/11513185/files/figure.png" pageId="6" pageNumber="335">Figs. 23</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6EFFBD2BC292FEE0C5F7D9" box="[646,676,668,691]" captionStart="FIGURES" captionStartId="18.[155,167,1684,1703]" captionTargetBox="[224,1059,1157,1665]" captionTargetId="figure-120@18.[215,1081,1140,1672]" captionTargetPageId="18" captionText="FIGURES 24, 25. Vytfutia halandrefana new species, female genitalia drawings, Tsingy de Bemaraha, Madagascar, CASENT9008848. 24. Epigynum, ventral. 25. Vulva, left half, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513187" httpUri="https://zenodo.org/record/11513187/files/figure.png" pageId="6" pageNumber="335">24</figureCitation>
|
||
), vulva complex with large anterior lobe and slender ducts, vulva width less than 1.50 times length (
|
||
<figureCitation id="13262A0B4B6EFFBD2BBE92DFE128F7BE" box="[762,841,701,724]" captionStart="FIGURES" captionStartId="18.[155,167,1684,1703]" captionTargetBox="[224,1059,1157,1665]" captionTargetId="figure-120@18.[215,1081,1140,1672]" captionTargetPageId="18" captionText="FIGURES 24, 25. Vytfutia halandrefana new species, female genitalia drawings, Tsingy de Bemaraha, Madagascar, CASENT9008848. 24. Epigynum, ventral. 25. Vulva, left half, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513187" httpUri="https://zenodo.org/record/11513187/files/figure.png" pageId="6" pageNumber="335">Fig. 25</figureCitation>
|
||
),
|
||
<collectingCountry id="F30A761E4B6EFFBD2A1A92DFE1BEF7BE" box="[862,991,701,724]" name="Madagascar" pageId="6" pageNumber="335">Madagascar</collectingCountry>
|
||
...............................................................
|
||
<emphasis id="B969EA9C4B6EFFBD2A0392BFE6EEF79F" bold="true" box="[839,1167,733,757]" pageId="6" pageNumber="335">
|
||
<taxonomicName id="4C1D4D0D4B6EFFBD2A0392BFE19CF79F" ID-CoL="C8MQC" authority="Griswold, 2022" authorityName="Griswold" authorityYear="2022" box="[839,1021,733,757]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="335" phylum="Arthropoda" rank="species" species="halandrefana" status="new species">
|
||
<emphasis id="B969EA9C4B6EFFBD2A0392BFE138F79E" bold="true" box="[839,857,733,756]" italics="true" pageId="6" pageNumber="335">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6EFFBD2A2392BFE19CF79F" bold="true" box="[871,1021,733,757]" italics="true" pageId="6" pageNumber="335">halandrefana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25A57E74B6EFFBD2D4F92BCE6EEF79F" box="[1035,1167,734,757]" pageId="6" pageNumber="335" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E92E2B4B6EFFBD293F9368E6EFF6CE" pageId="6" pageNumber="335">
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<paragraph id="8BA2368E4B6EFFBD293F9368E6EFF628" blockId="6.[123,1174,778,932]" pageId="6" pageNumber="335">
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||
<keyLead id="30EC95BB4B6EFFBD293F9368E6EFF628" pageId="6" pageNumber="335">
|
||
<emphasis id="B969EA9C4B6EFFBD293F9368E2CEF64B" bold="true" box="[123,175,778,801]" pageId="6" pageNumber="335">3 (2)</emphasis>
|
||
Epigynum with copulatory opening rims oblique (
|
||
<figureCitation id="13262A0B4B6EFFBD2B929368E150F64B" box="[726,817,778,801]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="6" pageNumber="335">Figs. 39</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6EFFBD2A799368E13AF64B" box="[829,859,778,801]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="6" pageNumber="335">48</figureCitation>
|
||
), vulva with integrated lobes anteriad of copulatory openings (
|
||
<figureCitation id="13262A0B4B6EFFBD2B4A9349E009F628" box="[526,616,811,834]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="6" pageNumber="335">Figs. 40</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6EFFBD2B309349E0F1F628" box="[628,656,811,834]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="6" pageNumber="335">49</figureCitation>
|
||
)....................................
|
||
<emphasis id="B969EA9C4B6EFFBD2DC59349E6EFF628" bold="true" box="[1153,1166,811,834]" pageId="6" pageNumber="335">4</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA2368E4B6EFFBD293F9329E6EFF6CE" blockId="6.[123,1174,778,932]" pageId="6" pageNumber="335">
|
||
<keyLead id="30EC95BB4B6EFFBD293F9329E6EFF6CE" pageId="6" pageNumber="335">
|
||
— Epigynum with copulatory opening rims longitudinal, epigynum posterior margin convex, elongated posteriorly in center (
|
||
<figureCitation id="13262A0B4B6EFFBD28BA930EE02CF6E9" box="[510,589,876,899]" captionStart="FIGURES" captionStartId="17.[155,167,1265,1284]" captionTargetBox="[120,1176,216,1249]" captionTargetId="figure-126@17.[120,1176,216,1249]" captionTargetPageId="17" captionText="FIGURES 15–19. Vytfutia bedel Deeleman-Reinhold, 1986, genitalia-drawings, Holotype male and paratype female collected from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. 15–17. Male left palp. 15. Tibia and tarsus, ventral. 16. Tibia, dorsal. 17. Tibia and tarsus, retrolateral. 18, 19. Female genitalia. 18. Epigynum, ventral. 19. Vulva, dorsal. C = conductor; D = Dorsal tibial apophysis; E = embolus; MA = median apophysis; RL = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.11513183" httpUri="https://zenodo.org/record/11513183/files/figure.png" pageId="6" pageNumber="335">Fig. 18</figureCitation>
|
||
), vulva with spherical lobe laterad of copulatory openings (
|
||
<figureCitation id="13262A0B4B6EFFBD29A293EFE355F6CE" box="[230,308,909,932]" captionStart="FIGURES" captionStartId="17.[155,167,1265,1284]" captionTargetBox="[120,1176,216,1249]" captionTargetId="figure-126@17.[120,1176,216,1249]" captionTargetPageId="17" captionText="FIGURES 15–19. Vytfutia bedel Deeleman-Reinhold, 1986, genitalia-drawings, Holotype male and paratype female collected from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. 15–17. Male left palp. 15. Tibia and tarsus, ventral. 16. Tibia, dorsal. 17. Tibia and tarsus, retrolateral. 18, 19. Female genitalia. 18. Epigynum, ventral. 19. Vulva, dorsal. C = conductor; D = Dorsal tibial apophysis; E = embolus; MA = median apophysis; RL = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.11513183" httpUri="https://zenodo.org/record/11513183/files/figure.png" pageId="6" pageNumber="335">Fig. 19</figureCitation>
|
||
)...................................
|
||
<taxonomicName id="4C1D4D0D4B6EFFBD2A5D93EFE6EFF6CE" ID-CoL="5BSNJ" authority="Deeleman-Reinhold, 1986" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[793,1166,909,932]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="335" phylum="Arthropoda" rank="species" species="bedel">
|
||
<emphasis id="B969EA9C4B6EFFBD2A5D93EFE10FF6CE" box="[793,878,909,932]" italics="true" pageId="6" pageNumber="335">V. bedel</emphasis>
|
||
<bibRefCitation id="EF8C4B7F4B6EFFBD2A3193EFE6EFF6CE" author="DEELEMAN-REINHOLD, C. L." box="[885,1166,909,932]" pageId="6" pageNumber="335" pagination="34 - 36" refId="ref11784" refString="DEELEMAN-REINHOLD, C. L. 1986. A new cribellate amaurobioid spider from Sumatra (Araneae: Agelenidae). Bulletin of the British Arachnological Society, 7: 34 - 36." type="journal article" year="1986">Deeleman-Reinhold, 1986</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E92E2B4B6EFFBD293F93DBE6EFF11E" pageId="6" pageNumber="335">
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<paragraph id="8BA2368E4B6EFFBD293F93DBE6EEF178" blockId="6.[123,1174,953,1140]" pageId="6" pageNumber="335">
|
||
<keyLead id="30EC95BB4B6EFFBD293F93DBE6EEF178" pageId="6" pageNumber="335">
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<emphasis id="B969EA9C4B6EFFBD293F93DBE2CCF6BA" bold="true" box="[123,173,953,976]" pageId="6" pageNumber="335">4 (3)</emphasis>
|
||
Epigynum with copulatory opening rims converging anteriorly, posterior margin slightly concave (
|
||
<figureCitation id="13262A0B4B6EFFBD29AF93B8E324F69B" box="[235,325,986,1009]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="6" pageNumber="335">Figs. 39</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6EFFBD281493B8E30CF69B" box="[336,365,986,1009]" captionStart="FIGURES" captionStartId="22.[155,167,1009,1028]" captionTargetBox="[134,1160,233,982]" captionTargetId="figure-19@22.[120,1176,216,992]" captionTargetPageId="22" captionText="FIGURES 41–44. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842, female genitalia, scanning electron micrographs (SEM). 41. Epigynum, ventral. 42. Vulva, dorsal. 43. Bennet’s gland pore (BG), left. 44. Pores of spermathecal head (HS), left. Scale bars, 41 = 100µm, 42 = 30µm, 43, 44 = 10µm." figureDoi="http://doi.org/10.5281/zenodo.11513199" httpUri="https://zenodo.org/record/11513199/files/figure.png" pageId="6" pageNumber="335">41</figureCitation>
|
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), vulva with paired lobes laterad of copulatory openings (
|
||
<figureCitation id="13262A0B4B6EFFBD2A9F93B8E654F69B" box="[987,1077,986,1009]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="6" pageNumber="335">Figs. 40</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6EFFBD2D0593B8E63CF69B" box="[1089,1117,986,1009]" captionStart="FIGURES" captionStartId="22.[155,167,1009,1028]" captionTargetBox="[134,1160,233,982]" captionTargetId="figure-19@22.[120,1176,216,992]" captionTargetPageId="22" captionText="FIGURES 41–44. Vytfutia labalaba new species, Mt. Kinabalu, Borneo, CASENT9023842, female genitalia, scanning electron micrographs (SEM). 41. Epigynum, ventral. 42. Vulva, dorsal. 43. Bennet’s gland pore (BG), left. 44. Pores of spermathecal head (HS), left. Scale bars, 41 = 100µm, 42 = 30µm, 43, 44 = 10µm." figureDoi="http://doi.org/10.5281/zenodo.11513199" httpUri="https://zenodo.org/record/11513199/files/figure.png" pageId="6" pageNumber="335">42</figureCitation>
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||
).........................................................
|
||
<emphasis id="B969EA9C4B6EFFBD2A3A9398E6EEF178" bold="true" box="[894,1167,1018,1042]" pageId="6" pageNumber="335">
|
||
<taxonomicName id="4C1D4D0D4B6EFFBD2A3A9398E19CF178" ID-CoL="C8MQD" authority="Griswold, 2022" authorityName="Griswold" authorityYear="2022" box="[894,1021,1018,1042]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="335" phylum="Arthropoda" rank="species" species="labalaba" status="new species">
|
||
<emphasis id="B969EA9C4B6EFFBD2A3A9398E1F1F17B" bold="true" box="[894,912,1018,1041]" italics="true" pageId="6" pageNumber="335">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6EFFBD2ADA9398E19CF178" bold="true" box="[926,1021,1018,1042]" italics="true" pageId="6" pageNumber="335">labalaba</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25A57E74B6EFFBD2D4F9399E6EEF178" box="[1035,1167,1019,1042]" pageId="6" pageNumber="335" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA2368E4B6EFFBD293F9479E6EFF11E" blockId="6.[123,1174,953,1140]" pageId="6" pageNumber="335">
|
||
<keyLead id="30EC95BB4B6EFFBD293F9479E6EFF11E" pageId="6" pageNumber="335">
|
||
— Epigynum with copulatory opening rims diverging anteriorly, posterior margin straight to slightly convex (
|
||
<figureCitation id="13262A0B4B6EFFBD2829945EE3A4F139" box="[365,453,1084,1107]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="6" pageNumber="335">Fig. 48</figureCitation>
|
||
), vulva with paired lobes far anteriad of copulatory openings (
|
||
<figureCitation id="13262A0B4B6EFFBD29F7943FE360F11E" box="[179,257,1117,1140]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="6" pageNumber="335">Fig. 49</figureCitation>
|
||
)......................................
|
||
<taxonomicName id="4C1D4D0D4B6EFFBD2A41943FE6EFF11E" ID-CoL="5BSNK" authority="Deeleman-Reinhold, 1989" authorityName="Deeleman-Reinhold" authorityYear="1989" box="[773,1166,1117,1140]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="335" phylum="Arthropoda" rank="species" species="pallens">
|
||
<emphasis id="B969EA9C4B6EFFBD2A41943FE10CF11E" box="[773,877,1117,1140]" italics="true" pageId="6" pageNumber="335">V. pallens</emphasis>
|
||
<bibRefCitation id="EF8C4B7F4B6EFFBD2A31943FE6EFF11E" author="DEELEMAN-REINHOLD, C. L." box="[885,1166,1117,1140]" pageId="6" pageNumber="335" pagination="619 - 627" refId="ref11818" refString="DEELEMAN-REINHOLD, C. L. 1989. Spiders from Niah Cave, Sarawak, East Malaysia, collected by P. Strinati. Revue Suisse de Zoologie, 96: 619 - 627." type="journal article" year="1989">Deeleman-Reinhold, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E92E2B4B6EFFBD293F94EBE6EFF0EC" pageId="6" pageNumber="335">
|
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<paragraph id="8BA2368E4B6EFFBD293F94EBE6EFF069" blockId="6.[123,1174,1161,1414]" pageId="6" pageNumber="335">
|
||
<keyLead id="30EC95BB4B6EFFBD293F94EBE6EFF069" pageId="6" pageNumber="335">
|
||
<emphasis id="B969EA9C4B6EFFBD293F94EBE2CFF1CA" bold="true" box="[123,174,1161,1184]" pageId="6" pageNumber="335">5 (1)</emphasis>
|
||
Male palpal tibia with
|
||
<collectionCode id="ED0CAE4B4B6EFFBD28ED94EBE3BDF1CA" box="[425,476,1161,1184]" pageId="6" pageNumber="335">DTA</collectionCode>
|
||
short, less than 0.25 length tibia (
|
||
<figureCitation id="13262A0B4B6EFFBD2A0C94EBE1ABF1CA" box="[840,970,1161,1184]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="6" pageNumber="335">Figs. 37, 38</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6EFFBD2A9294EBE67DF1CA" box="[982,1052,1161,1184]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="6" pageNumber="335">46, 49</figureCitation>
|
||
), tibia with subapical spine or cuspule, embolus short, arising near base of bulb (past 6 o’clock) and making turn of less than 180º, median apophysis long, length greater than 5.00 times width (
|
||
<figureCitation id="13262A0B4B6EFFBD29F7948EE354F069" box="[179,309,1260,1283]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="6" pageNumber="335">Figs. 36, 37</figureCitation>
|
||
,
|
||
<figureCitation id="13262A0B4B6EFFBD2805948EE3E4F069" box="[321,389,1260,1283]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="6" pageNumber="335">45, 46</figureCitation>
|
||
)........................................................
|
||
<emphasis id="B969EA9C4B6EFFBD2DC5948EE6EFF069" bold="true" box="[1153,1166,1260,1283]" pageId="6" pageNumber="335">6</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA2368E4B6EFFBD293F956EE6EFF0EC" blockId="6.[123,1174,1161,1414]" pageId="6" pageNumber="335">
|
||
<keyLead id="30EC95BB4B6EFFBD293F956EE6EFF0EC" pageId="6" pageNumber="335">
|
||
— Male palpal tibia with
|
||
<collectionCode id="ED0CAE4B4B6EFFBD28EC956EE3BAF049" box="[424,475,1292,1315]" pageId="6" pageNumber="335">DTA</collectionCode>
|
||
elongate, greater than 0.50 length tibia (
|
||
<figureCitation id="13262A0B4B6EFFBD2AD6956EE674F049" box="[914,1045,1292,1315]" captionStart="FIGURES" captionStartId="17.[155,167,1265,1284]" captionTargetBox="[120,1176,216,1249]" captionTargetId="figure-126@17.[120,1176,216,1249]" captionTargetPageId="17" captionText="FIGURES 15–19. Vytfutia bedel Deeleman-Reinhold, 1986, genitalia-drawings, Holotype male and paratype female collected from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. 15–17. Male left palp. 15. Tibia and tarsus, ventral. 16. Tibia, dorsal. 17. Tibia and tarsus, retrolateral. 18, 19. Female genitalia. 18. Epigynum, ventral. 19. Vulva, dorsal. C = conductor; D = Dorsal tibial apophysis; E = embolus; MA = median apophysis; RL = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.11513183" httpUri="https://zenodo.org/record/11513183/files/figure.png" pageId="6" pageNumber="335">Figs. 16, 17</figureCitation>
|
||
), tibia lacking subapical enlarged seta, embolus long, arising on retrolateral side of bulb (4 o’clock) and making turn of more than 225º, median apophysis short, length less than 4.00 times width (
|
||
<figureCitation id="13262A0B4B6EFFBD29F7950DE352F0EC" box="[179,307,1391,1414]" captionStart="FIGURES" captionStartId="17.[155,167,1265,1284]" captionTargetBox="[120,1176,216,1249]" captionTargetId="figure-126@17.[120,1176,216,1249]" captionTargetPageId="17" captionText="FIGURES 15–19. Vytfutia bedel Deeleman-Reinhold, 1986, genitalia-drawings, Holotype male and paratype female collected from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. 15–17. Male left palp. 15. Tibia and tarsus, ventral. 16. Tibia, dorsal. 17. Tibia and tarsus, retrolateral. 18, 19. Female genitalia. 18. Epigynum, ventral. 19. Vulva, dorsal. C = conductor; D = Dorsal tibial apophysis; E = embolus; MA = median apophysis; RL = retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.11513183" httpUri="https://zenodo.org/record/11513183/files/figure.png" pageId="6" pageNumber="335">Figs. 15, 17</figureCitation>
|
||
)...................................
|
||
<taxonomicName id="4C1D4D0D4B6EFFBD2A5D950DE6EFF0EC" ID-CoL="5BSNJ" authority="Deeleman-Reinhold, 1986" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[793,1166,1391,1414]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="335" phylum="Arthropoda" rank="species" species="bedel">
|
||
<emphasis id="B969EA9C4B6EFFBD2A5D950DE10FF0EC" box="[793,878,1391,1414]" italics="true" pageId="6" pageNumber="335">V. bedel</emphasis>
|
||
<bibRefCitation id="EF8C4B7F4B6EFFBD2A31950DE6EFF0EC" author="DEELEMAN-REINHOLD, C. L." box="[885,1166,1391,1414]" pageId="6" pageNumber="335" pagination="34 - 36" refId="ref11784" refString="DEELEMAN-REINHOLD, C. L. 1986. A new cribellate amaurobioid spider from Sumatra (Araneae: Agelenidae). Bulletin of the British Arachnological Society, 7: 34 - 36." type="journal article" year="1986">Deeleman-Reinhold, 1986</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E92E2B4B6EFFBD293F95F9E6EEF3FD" pageId="6" pageNumber="335">
|
||
<paragraph id="8BA2368E4B6EFFBD293F95F9E6EFF37E" blockId="6.[123,1174,1435,1687]" pageId="6" pageNumber="335">
|
||
<keyLead id="30EC95BB4B6EFFBD293F95F9E6EFF37E" pageId="6" pageNumber="335">
|
||
<emphasis id="B969EA9C4B6EFFBD293F95F9E2CEF0D8" bold="true" box="[123,175,1435,1458]" pageId="6" pageNumber="335">6 (5)</emphasis>
|
||
Palpal tibia with elongate apical spine (
|
||
<figureCitation id="13262A0B4B6EFFBD2B2795F9E089F0D8" box="[611,744,1435,1458]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="6" pageNumber="335">Figs. 46, 47</figureCitation>
|
||
), embolus short, straight, arises on prolateral side of bulb (near 8 o’clock), median apophysis hooked at apex (
|
||
<figureCitation id="13262A0B4B6EFFBD2AE395DEE648F0B9" box="[935,1065,1468,1491]" captionStart="FIGURES" captionStartId="23.[155,167,1021,1040]" captionTargetBox="[120,1164,216,976]" captionTargetId="figure-24@23.[120,1176,216,1004]" captionTargetPageId="23" captionText="FIGURES 45–49. Vytfutia pallens Deeleman-Reinhold, 1989, genitalia-drawings, male and female paratypes from Niah Cave, Sarawak, Malaysia. 45–47. Male left palp, tibia and tarsus. 45. Ventral. 46. Retrolateral. 47. Dorsal. 48. 49. Female genitalia. 48. Epigynum, ventral. 49. Vulva, dorsal. C: conductor, D: dorsal tibial apophysis, MA: median apophysis, PC: paracymbium, RL: retrolateral lobe of tibial apex. Grey arrow to tibial apical spine." figureDoi="http://doi.org/10.5281/zenodo.11513203" httpUri="https://zenodo.org/record/11513203/files/figure.png" pageId="6" pageNumber="335">Figs. 45, 46</figureCitation>
|
||
); metatarsus I long, base of concavity originates at more than 0.45 metatarsus length, spur high, height great than 0.09 metatarsus length (
|
||
<figureCitation id="13262A0B4B6EFFBD2B5A959FE0FFF37E" box="[542,670,1533,1556]" captionStart="FIGURES" captionStartId="25.[155,167,1487,1506]" captionTargetBox="[153,1147,252,1446]" captionTargetId="figure-24@25.[120,1176,216,1475]" captionTargetPageId="25" captionText="FIGURES 56–61. Vytfutia spp., male leg I with clasping modification of metatarsus. 56, 58, 61. Dorsal. 57, 59, 60. Retrolateral. 56, 57. Vytfutia bedel Deeleman-Reinhold, 1986, Holotype male from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia, left. 58, 59. Vytfutia labalaba new species, Holotype male from the wall of a building at 1550m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842, right. 60, 61. Vytfutia pallens Deeleman-Reinhold, 1989, male paratype from Niah Cave, Sarawak, Malaysia, right. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.11513207" httpUri="https://zenodo.org/record/11513207/files/figure.png" pageId="6" pageNumber="335">Figs. 60, 61</figureCitation>
|
||
)......
|
||
<taxonomicName id="4C1D4D0D4B6EFFBD2A41959FE6EFF37E" ID-CoL="5BSNK" authority="Deeleman-Reinhold, 1989" authorityName="Deeleman-Reinhold" authorityYear="1989" box="[773,1166,1533,1556]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="335" phylum="Arthropoda" rank="species" species="pallens">
|
||
<emphasis id="B969EA9C4B6EFFBD2A41959FE10CF37E" box="[773,877,1533,1556]" italics="true" pageId="6" pageNumber="335">V. pallens</emphasis>
|
||
<bibRefCitation id="EF8C4B7F4B6EFFBD2A31959FE6EFF37E" author="DEELEMAN-REINHOLD, C. L." box="[885,1166,1533,1556]" pageId="6" pageNumber="335" pagination="619 - 627" refId="ref11818" refString="DEELEMAN-REINHOLD, C. L. 1989. Spiders from Niah Cave, Sarawak, East Malaysia, collected by P. Strinati. Revue Suisse de Zoologie, 96: 619 - 627." type="journal article" year="1989">Deeleman-Reinhold, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA2368E4B6EFFBD293F967CE6EEF3FD" blockId="6.[123,1174,1435,1687]" pageId="6" pageNumber="335">
|
||
<keyLead id="30EC95BB4B6EFFBD293F967CE6EEF3FD" pageId="6" pageNumber="335">
|
||
— Palpal tibia with short subapical cuspule (
|
||
<figureCitation id="13262A0B4B6EFFBD2B30967CE099F35F" box="[628,760,1566,1589]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="6" pageNumber="335">Figs. 37, 38</figureCitation>
|
||
), embolus long, curved, arises at base of bulb (near 6 o’clock), median apophysis flattened, spoon shaped at apex (
|
||
<figureCitation id="13262A0B4B6EFFBD2D44965DE6E9F33C" box="[1024,1160,1599,1622]" captionStart="FIGURES" captionStartId="21.[155,167,1160,1179]" captionTargetBox="[120,1172,217,1122]" captionTargetId="figure-24@21.[120,1176,216,1144]" captionTargetPageId="21" captionText="FIGURES 36–40. Vytfutia labalaba new species, genitalia-drawings, Holotype male and paratype female collected on the wall of a building at 1550 m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842. 36–38. Male left palp, tibia and tarsus. 36. Ventral. 37. Retrolateral. 38. Dorsal. 39, 40. Female genitalia. 39. Epigynum, ventral. 40. Vulva, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513197" httpUri="https://zenodo.org/record/11513197/files/figure.png" pageId="6" pageNumber="335">Figs. 36, 37</figureCitation>
|
||
); metatarsus I shorter, base of concavity originates at less than 0.35 metatarsus length, spur low, height greater than 0.08 metatarsus length (
|
||
<figureCitation id="13262A0B4B6EFFBD2B3996E2E09FF3FD" box="[637,766,1664,1687]" captionStart="FIGURES" captionStartId="25.[155,167,1487,1506]" captionTargetBox="[153,1147,252,1446]" captionTargetId="figure-24@25.[120,1176,216,1475]" captionTargetPageId="25" captionText="FIGURES 56–61. Vytfutia spp., male leg I with clasping modification of metatarsus. 56, 58, 61. Dorsal. 57, 59, 60. Retrolateral. 56, 57. Vytfutia bedel Deeleman-Reinhold, 1986, Holotype male from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia, left. 58, 59. Vytfutia labalaba new species, Holotype male from the wall of a building at 1550m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia, CASENT9023842, right. 60, 61. Vytfutia pallens Deeleman-Reinhold, 1989, male paratype from Niah Cave, Sarawak, Malaysia, right. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.11513207" httpUri="https://zenodo.org/record/11513207/files/figure.png" pageId="6" pageNumber="335">Figs. 58, 59</figureCitation>
|
||
).........
|
||
<emphasis id="B969EA9C4B6EFFBD2A3A96E2E6EEF3FD" bold="true" box="[894,1167,1663,1687]" pageId="6" pageNumber="335">
|
||
<taxonomicName id="4C1D4D0D4B6EFFBD2A3A96E2E19CF3FD" ID-CoL="C8MQD" authority="Griswold, 2022" authorityName="Griswold" authorityYear="2022" box="[894,1021,1663,1687]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="335" phylum="Arthropoda" rank="species" species="labalaba" status="new species">
|
||
<emphasis id="B969EA9C4B6EFFBD2A3A96E2E1F1F3FD" bold="true" box="[894,912,1664,1687]" italics="true" pageId="6" pageNumber="335">V</emphasis>
|
||
.
|
||
<emphasis id="B969EA9C4B6EFFBD2ADA961DE19CF3FD" bold="true" box="[926,1021,1663,1687]" italics="true" pageId="6" pageNumber="335">labalaba</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25A57E74B6EFFBD2D4F96E2E6EEF3FD" box="[1035,1167,1664,1687]" pageId="6" pageNumber="335" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |