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<mods:title id="6432729940D378C949CF2EC87D1DFF00">A world review of the bee fly tribe Usiini (Diptera, Bombyliidae) - Part 3: Parageron Paramonov s. lat.</mods:title>
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<treatment id="EE3F8791FFC04C62FD623BCED6F4E49A" ID-DOI="http://doi.org/10.5281/zenodo.8291089" ID-GBIF-Taxon="212472366" ID-Zenodo-Dep="8291089" LSID="urn:lsid:zoobank.org:act:4A91131A-1CA0-4BC3-AF37-39A8E764C56A" httpUri="http://treatment.plazi.org/id/EE3F8791FFC04C62FD623BCED6F4E49A" lastPageId="121" lastPageNumber="122" pageId="120" pageNumber="121">
<subSubSection id="2E8C650CFFC04C63FD623BCED1B1E40C" pageId="120" pageNumber="121" type="nomenclature">
<paragraph id="66293687FFC04C63FD623BCED6D0E5EF" blockId="120.[362,1226,1750,1811]" box="[648,939,1750,1776]" pageId="120" pageNumber="121">
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Genus
<taxonomicName id="A1964D04FFC04C63FD313BCED641E5EF" ID-CoL="BDNR8" authority="Gibbs, 2023" authorityName="Gibbs" authorityYear="2023" box="[731,826,1750,1776]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="54E2EA95FFC04C63FD313BCED641E5EF" bold="true" box="[731,826,1750,1776]" italics="true" pageId="120" pageNumber="121">Parusia</emphasis>
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<taxonomicNameLabel id="4FD157EEFFC04C63FCAB3BCED6D0E5EF" box="[833,939,1750,1776]" pageId="120" pageNumber="121" rank="genus">gen. nov.</taxonomicNameLabel>
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<heading id="3D6181EBFFC04C63FF573A22D423E44C" bold="true" box="[189,344,1849,1876]" fontSize="11" level="3" pageId="120" pageNumber="121" reason="3">
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<typeStatus id="B92D8825FFC04C63FF573A22D581E44B" box="[189,250,1850,1876]" pageId="120" pageNumber="121">Type</typeStatus>
species
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</paragraph>
<paragraph id="66293687FFC04C63FF573A7FD729E49D" blockId="120.[189,594,1895,1922]" box="[189,594,1895,1922]" pageId="120" pageNumber="121">
<taxonomicName id="A1964D04FFC04C63FF573A7FD735E49D" ID-CoL="8FLN9" authority="Fabricius, 1794" authorityName="Fabricius" authorityYear="1794" box="[189,590,1895,1922]" class="Insecta" family="Bombyliidae" genus="Voluccella" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="54E2EA95FFC04C63FF573A7FD4F6E49E" box="[189,397,1895,1921]" italics="true" pageId="120" pageNumber="121">Voluccella aurata</emphasis>
<bibRefCitation id="02074B76FFC04C63FE7E3A7FD735E49D" author="Fabricius J. C." box="[404,590,1895,1922]" pageId="120" pageNumber="121" refId="ref74393" refString="Fabricius J. C. 1794. Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tome IV. C. G. Proft, Hafniae [= Copenhagen]. https: // doi. org / 10.5962 / bhl. title. 125869" type="book" year="1794">Fabricius, 1794</bibRefCitation>
</taxonomicName>
.
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<subSubSection id="2E8C650CFFC14C62FF573C10D079E1B3" pageId="121" pageNumber="122" type="diagnosis">
<paragraph id="66293687FFC14C62FF573C10D44DE23D" blockId="121.[189,310,264,290]" box="[189,310,264,290]" pageId="121" pageNumber="122">
<heading id="3D6181EBFFC14C62FF573C10D44DE23D" bold="true" box="[189,310,264,290]" fontSize="11" level="3" pageId="121" pageNumber="122" reason="3">
<emphasis id="54E2EA95FFC14C62FF573C10D44DE23D" bold="true" box="[189,310,264,290]" pageId="121" pageNumber="122">Diagnosis</emphasis>
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<paragraph id="66293687FFC14C62FF573C2ED079E1B3" blockId="121.[188,1401,310,684]" pageId="121" pageNumber="122">
The combination of dichoptic males, very narrow gena plus oral margin, relatively short proboscis, entirely black legs, relatively weakly sclerotised matt-black tergites with yellow apical margins and large, globular genitalia in the male should serve to distinguish this genus from other
<taxonomicName id="A1964D04FFC14C62FB583C65D04BE289" authorityName="Paramonov" authorityYear="1929" box="[1202,1328,381,406]" class="Insecta" family="Bombyliidae" genus="Parageron" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="54E2EA95FFC14C62FB583C65D04BE289" box="[1202,1328,381,406]" italics="true" pageId="121" pageNumber="122">Parageron</emphasis>
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<taxonomicNameLabel id="4FD157EEFFC14C62FADD3C64D00CE289" box="[1335,1399,380,406]" pageId="121" pageNumber="122" sensu="lato">s. lat.</taxonomicNameLabel>
and
<taxonomicName id="A1964D04FFC14C62FF1B3C87D453E2A7" authorityName="Latreille" authorityYear="1802" box="[241,296,415,440]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="54E2EA95FFC14C62FF1B3C87D453E2A7" box="[241,296,415,440]" italics="true" pageId="121" pageNumber="122">Usia</emphasis>
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. However, all these characteristics are variable, and are found at least in part in other
<taxonomicName id="A1964D04FFC14C62FAD53C87D00DE2A7" authorityName="Latreille" authorityYear="1802" box="[1343,1398,415,440]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="54E2EA95FFC14C62FAD53C87D00DE2A7" box="[1343,1398,415,440]" italics="true" pageId="121" pageNumber="122">Usia</emphasis>
</taxonomicName>
and
<taxonomicName id="A1964D04FFC14C62FF053CDAD416E2C4" authorityName="Paramonov" authorityYear="1929" box="[239,365,450,475]" class="Insecta" family="Bombyliidae" genus="Parageron" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="54E2EA95FFC14C62FF053CDAD416E2C4" box="[239,365,450,475]" italics="true" pageId="121" pageNumber="122">Parageron</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFC14C62FE9E3CDAD4C8E2C4" box="[372,435,449,476]" pageId="121" pageNumber="122" sensu="lato">s. lat.</taxonomicNameLabel>
The most convincing characters are found in the male epiphallic complex and the female genitalia. Males have a strongly sclerotised epiphallus and very large basal ejaculatory apodeme enclosed within a very large, strongly sclerotised and shining black hypopygium. The female genital fork is strongly sclerotised, often flat, sometimes bent dorsally, sometimes strongly angled with the arms. Compared to other
<taxonomicName id="A1964D04FFC14C62FE0F3F54D74BE179" authorityName=", Becker" authorityYear="1913" box="[485,560,588,614]" class="Insecta" family="Bombyliidae" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="159" phylum="Arthropoda" rank="tribe" tribe="Usiini">Usiini</taxonomicName>
, there is no obviously separate vaginal plate, it appears to be contiguous with the genital fork body. In many
<taxonomicName id="A1964D04FFC14C62FDB43F68D7A5E196" authority="Gibbs, 2023" authorityName="Gibbs" authorityYear="2023" box="[606,734,624,649]" class="Insecta" family="Bombyliidae" genus="Protypusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="54E2EA95FFC14C62FDB43F68D7A5E196" box="[606,734,624,649]" italics="true" pageId="121" pageNumber="122">Protypusia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFC14C62FD0F3F68D634E195" box="[741,847,624,650]" pageId="121" pageNumber="122" rank="genus">gen. nov.</taxonomicNameLabel>
the genital fork is also strongly sclerotised, but the vaginal plate is always discernible, even when sclerotised and fused with the genital fork.
</paragraph>
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<subSubSection id="2E8C650CFFC14C62FF573FCBD654E004" box="[189,815,723,795]" pageId="121" pageNumber="122" type="etymology">
<paragraph id="66293687FFC14C62FF573FCBD43FE1F2" blockId="121.[189,324,723,749]" box="[189,324,723,749]" pageId="121" pageNumber="122">
<heading id="3D6181EBFFC14C62FF573FCBD43FE1F2" bold="true" box="[189,324,723,749]" fontSize="11" level="3" pageId="121" pageNumber="122" reason="3">
<emphasis id="54E2EA95FFC14C62FF573FCBD43FE1F2" bold="true" box="[189,324,723,749]" pageId="121" pageNumber="122">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="66293687FFC14C62FF573E19D654E004" blockId="121.[189,815,769,795]" box="[189,815,769,795]" pageId="121" pageNumber="122">
‘par(a)-’ and ‘
<emphasis id="54E2EA95FFC14C62FE8B3E19D4E9E005" box="[353,402,769,794]" italics="true" pageId="121" pageNumber="122">usia</emphasis>
’ meaning ‘near to the genus
<taxonomicName id="A1964D04FFC14C62FD0C3E19D666E005" authorityName="Latreille" authorityYear="1802" box="[742,797,769,794]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="54E2EA95FFC14C62FD0C3E19D666E005" box="[742,797,769,794]" italics="true" pageId="121" pageNumber="122">Usia</emphasis>
</taxonomicName>
’.
</paragraph>
</subSubSection>
<subSubSection id="2E8C650CFFC14C62FF573E5AD6F4E49A" pageId="121" pageNumber="122" type="discussion">
<paragraph id="66293687FFC14C62FF573E5AD454E043" blockId="121.[189,303,834,860]" box="[189,303,834,860]" pageId="121" pageNumber="122">
<heading id="3D6181EBFFC14C62FF573E5AD454E043" bold="true" box="[189,303,834,860]" fontSize="11" level="3" pageId="121" pageNumber="122" reason="3">
<emphasis id="54E2EA95FFC14C62FF573E5AD454E043" bold="true" box="[189,303,834,860]" pageId="121" pageNumber="122">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="66293687FFC14C62FF573E77D412E746" blockId="121.[188,1399,879,1113]" pageId="121" pageNumber="122">
This genus encompasses the former ‘
<taxonomicName id="A1964D04FFC14C62FD9A3E68D7C5E096" authorityName="Fabricius" authorityYear="1794" box="[624,702,880,905]" class="Insecta" family="Bombyliidae" genus="Voluccella" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="54E2EA95FFC14C62FD9A3E68D7C5E096" box="[624,702,880,905]" italics="true" pageId="121" pageNumber="122">aurata</emphasis>
</taxonomicName>
’-complex including
<emphasis id="54E2EA95FFC14C62FC593E68D156E096" box="[947,1069,879,905]" italics="true" pageId="121" pageNumber="122">
Pru.
<taxonomicName id="A1964D04FFC14C62FC073E77D156E096" box="[1005,1069,879,905]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="loewi">loewi</taxonomicName>
</emphasis>
. It is a very coherent group, all very closely related to each other and clearly monophyletic, but with no obvious affinities within the rest of the
<taxonomicName id="A1964D04FFC14C62FED63EADD4F3E0D0" authorityName=", Becker" authorityYear="1913" box="[316,392,949,975]" class="Insecta" family="Bombyliidae" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="159" phylum="Arthropoda" rank="tribe" tribe="Usiini">Usiini</taxonomicName>
. The group might arguably be closer to
<taxonomicName id="A1964D04FFC14C62FC8A3EAED6ECE0D0" authorityName="Latreille" authorityYear="1802" box="[864,919,950,975]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="54E2EA95FFC14C62FC8A3EAED6ECE0D0" box="[864,919,950,975]" italics="true" pageId="121" pageNumber="122">Usia</emphasis>
</taxonomicName>
, and it has often been placed here rather than in
<taxonomicName id="A1964D04FFC14C62FEF83EC0D4EBE0EE" authorityName="Paramonov" authorityYear="1929" box="[274,400,984,1009]" class="Insecta" family="Bombyliidae" genus="Parageron" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="54E2EA95FFC14C62FEF83EC0D4EBE0EE" box="[274,400,984,1009]" italics="true" pageId="121" pageNumber="122">Parageron</emphasis>
</taxonomicName>
. While there are some members of
<taxonomicName id="A1964D04FFC14C62FCC73EC0D6D0E0EE" authorityName="Paramonov" authorityYear="1929" box="[813,939,984,1009]" class="Insecta" family="Bombyliidae" genus="Parageron" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="54E2EA95FFC14C62FCC73EC0D6D0E0EE" box="[813,939,984,1009]" italics="true" pageId="121" pageNumber="122">Parageron</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFC14C62FC583EC0D68BE0EE" box="[946,1008,983,1010]" pageId="121" pageNumber="122" sensu="lato">s. lat.</taxonomicNameLabel>
that share some of the features of the ‘
<taxonomicName id="A1964D04FFC14C62FF183EE3D43BE70B" authorityName="Fabricius" authorityYear="1794" box="[242,320,1019,1044]" class="Insecta" family="Bombyliidae" genus="Voluccella" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="54E2EA95FFC14C62FF183EE3D43BE70B" box="[242,320,1019,1044]" italics="true" pageId="121" pageNumber="122">aurata</emphasis>
</taxonomicName>
’-complex, the
<taxonomicName id="A1964D04FFC14C62FE1F3EE3D708E70B" authorityName="Paramonov" authorityYear="1929" box="[501,627,1019,1044]" class="Insecta" family="Bombyliidae" genus="Parageron" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="54E2EA95FFC14C62FE1F3EE3D708E70B" box="[501,627,1019,1044]" italics="true" pageId="121" pageNumber="122">Parageron</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFC14C62FD903EE3D7C3E70A" box="[634,696,1019,1045]" pageId="121" pageNumber="122" sensu="stricto">s. str.</taxonomicNameLabel>
are really quite different, and it became apparent that if the separation of
<taxonomicName id="A1964D04FFC14C62FEB73906D4EFE728" authorityName="Latreille" authorityYear="1802" box="[349,404,1054,1079]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="54E2EA95FFC14C62FEB73906D4EFE728" box="[349,404,1054,1079]" italics="true" pageId="121" pageNumber="122">Usia</emphasis>
</taxonomicName>
and
<taxonomicName id="A1964D04FFC14C62FE243906D737E728" authorityName="Paramonov" authorityYear="1929" box="[462,588,1054,1079]" class="Insecta" family="Bombyliidae" genus="Parageron" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="54E2EA95FFC14C62FE243906D737E728" box="[462,588,1054,1079]" italics="true" pageId="121" pageNumber="122">Parageron</emphasis>
</taxonomicName>
were to be maintained, then
<taxonomicName id="A1964D04FFC14C62FC4A3906D165E728" authorityName="Paramonov" authorityYear="1929" box="[928,1054,1054,1079]" class="Insecta" family="Bombyliidae" genus="Parageron" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="54E2EA95FFC14C62FC4A3906D165E728" box="[928,1054,1054,1079]" italics="true" pageId="121" pageNumber="122">Parageron</emphasis>
</taxonomicName>
in its former sense needed to be subdivided.
</paragraph>
<paragraph id="66293687FFC14C62FF57399AD7DBE501" blockId="121.[189,1400,1153,1567]" pageId="121" pageNumber="122">
<taxonomicName id="A1964D04FFC14C62FF57399AD461E784" authority="Gibbs, 2023" authorityName="Gibbs" authorityYear="2023" box="[189,282,1154,1179]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="54E2EA95FFC14C62FF57399AD461E784" box="[189,282,1154,1179]" italics="true" pageId="121" pageNumber="122">Parusia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFC14C62FEC93999D4E9E784" box="[291,402,1153,1179]" pageId="121" pageNumber="122" rank="genus">gen. nov.</taxonomicNameLabel>
, like other
<taxonomicName id="A1964D04FFC14C62FDF03999D71EE784" authorityName=", Becker" authorityYear="1913" box="[538,613,1153,1179]" class="Insecta" family="Bombyliidae" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="159" phylum="Arthropoda" rank="tribe" tribe="Usiini">Usiini</taxonomicName>
, exhibits a strong tendency for endemism, in this case with many cryptic species superficially very similar and differing predominantly in the structure of male and female genitalia. During this study, it has been possible to show that the two formerly recognised species
<emphasis id="54E2EA95FFC14C62FAAE39D0D00BE7FE" box="[1348,1392,1224,1249]" italics="true" pageId="121" pageNumber="122">Pru</emphasis>
.
<taxonomicName id="A1964D04FFC14C62FF5739F3D470E61B" authorityName="Fabricius" authorityYear="1794" box="[189,267,1259,1284]" class="Insecta" family="Bombyliidae" genus="Voluccella" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="54E2EA95FFC14C62FF5739F3D470E61B" box="[189,267,1259,1284]" italics="true" pageId="121" pageNumber="122">aurata</emphasis>
</taxonomicName>
and
<emphasis id="54E2EA95FFC14C62FEA039F3D40DE61B" box="[330,374,1259,1284]" italics="true" pageId="121" pageNumber="122">Pru</emphasis>
.
<taxonomicName id="A1964D04FFC14C62FE6D39F2D4BCE61B" box="[391,455,1258,1284]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="loewi">
<emphasis id="54E2EA95FFC14C62FE6D39F2D4BCE61B" box="[391,455,1258,1284]" italics="true" pageId="121" pageNumber="122">loewi</emphasis>
</taxonomicName>
(with
<emphasis id="54E2EA95FFC14C62FDF339F3D73EE61B" box="[537,581,1259,1284]" italics="true" pageId="121" pageNumber="122">Pru</emphasis>
.
<taxonomicName id="A1964D04FFC14C62FDBD39F2D7B5E61B" authorityName="Gibbs" authorityYear="2023" baseAuthorityName="Costa" box="[599,718,1258,1284]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="taeniolata">
<emphasis id="54E2EA95FFC14C62FDBD39F2D7B5E61B" box="[599,718,1258,1284]" italics="true" pageId="121" pageNumber="122">taeniolata</emphasis>
</taxonomicName>
a synonym of the former) in fact comprise no less than eight species. All of these have been found within a small part of the previously published range of the ‘
<taxonomicName id="A1964D04FFC14C62FF1C382AD43FE654" authorityName="Fabricius" authorityYear="1794" box="[246,324,1330,1355]" class="Insecta" family="Bombyliidae" genus="Voluccella" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="54E2EA95FFC14C62FF1C382AD43FE654" box="[246,324,1330,1355]" italics="true" pageId="121" pageNumber="122">aurata</emphasis>
</taxonomicName>
’-complex in the western Mediterranean east to
<collectingCountry id="1E817617FFC14C62FC7A3829D6A3E654" box="[912,984,1329,1355]" name="Libya" pageId="121" pageNumber="122">Libya</collectingCountry>
. Records of
<emphasis id="54E2EA95FFC14C62FB9C382AD1D9E654" box="[1142,1186,1330,1355]" italics="true" pageId="121" pageNumber="122">Pru</emphasis>
.
<taxonomicName id="A1964D04FFC14C62FB5E382AD079E654" authorityName="Fabricius" authorityYear="1794" box="[1204,1282,1330,1355]" class="Insecta" family="Bombyliidae" genus="Voluccella" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="54E2EA95FFC14C62FB5E382AD079E654" box="[1204,1282,1330,1355]" italics="true" pageId="121" pageNumber="122">aurata</emphasis>
</taxonomicName>
and
<emphasis id="54E2EA95FFC14C62FAA9382AD014E654" box="[1347,1391,1330,1355]" italics="true" pageId="121" pageNumber="122">Pru</emphasis>
.
<taxonomicName id="A1964D04FFC14C62FF57384CD586E671" box="[189,253,1364,1390]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="loewi">
<emphasis id="54E2EA95FFC14C62FF57384CD586E671" box="[189,253,1364,1390]" italics="true" pageId="121" pageNumber="122">loewi</emphasis>
</taxonomicName>
come from many countries north to
<collectingCountry id="1E817617FFC14C62FD41384DD785E670" box="[683,766,1365,1391]" name="France" pageId="121" pageNumber="122">France</collectingCountry>
, east to
<collectingCountry id="1E817617FFC14C62FCB4384DD69EE670" box="[862,997,1365,1391]" name="Kyrgyzstan" pageId="121" pageNumber="122">Kyrgyzstan</collectingCountry>
and south to
<collectingCountry id="1E817617FFC14C62FB69384CD1A2E671" box="[1155,1241,1364,1390]" name="Kuwait" pageId="121" pageNumber="122">Kuwait</collectingCountry>
(
<bibRefCitation id="02074B76FFC14C62FB03384CD4F9E68D" author="Evenhuis N. L. &amp; Greathead D. J." pageId="121" pageNumber="122" refId="ref74346" refString="Evenhuis N. L. &amp; Greathead D. J. 2015. World Catalog of Bee Flies (Diptera: Bombyliidae) web site. Available from http: // hbs. bishopmuseum. org / bombcat / [accessed 25 Jan. 2023]." type="url" year="2015">Evenhuis &amp; Greathead 2015</bibRefCitation>
). During this study no specimens from the eastern Mediterranean or Central Asia have been seen so it seems likely that these are all misidentifications in the literature. The lack of any specimens from mainland
<collectingCountry id="1E817617FFC14C62FE1E38A6D751E6C7" box="[500,554,1470,1496]" name="Italy" pageId="121" pageNumber="122">Italy</collectingCountry>
or
<collectingRegion id="A452F865FFC14C62FDB838A6D7EDE6C7" box="[594,662,1470,1496]" country="Italy" name="Sicilia" pageId="121" pageNumber="122">Sicily</collectingRegion>
is somewhat surprising, so it could well occur there (probably
<emphasis id="54E2EA95FFC14C62FF5738FAD592E6E4" box="[189,233,1506,1531]" italics="true" pageId="121" pageNumber="122">Pru</emphasis>
.
<taxonomicName id="A1964D04FFC14C62FF1038F9D40AE6E4" authorityName="Gibbs" authorityYear="2023" baseAuthorityName="Costa" box="[250,369,1505,1531]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="taeniolata">
<emphasis id="54E2EA95FFC14C62FF1038F9D40AE6E4" box="[250,369,1505,1531]" italics="true" pageId="121" pageNumber="122">taeniolata</emphasis>
</taxonomicName>
). Given the number of cryptic species found during this study, it is highly likely that more local endemics remain to be found.
</paragraph>
<paragraph id="66293687FFC14C62FF573B5ED4F2E540" blockId="121.[189,393,1605,1632]" box="[189,393,1605,1632]" pageId="121" pageNumber="122">
<heading id="3D6181EBFFC14C62FF573B5ED4F2E540" bold="true" box="[189,393,1605,1632]" fontSize="11" level="3" pageId="121" pageNumber="122" reason="3">
<emphasis id="54E2EA95FFC14C62FF573B5ED4F2E540" bold="true" box="[189,393,1605,1632]" pageId="121" pageNumber="122">Included species</emphasis>
</heading>
</paragraph>
<paragraph id="66293687FFC14C62FF573B6DD74FE590" blockId="121.[189,911,1652,1925]" box="[189,564,1652,1679]" pageId="121" pageNumber="122">
<taxonomicName id="A1964D04FFC14C62FF573B6DD407E591" authority="Gibbs, 2023" authorityName="Gibbs" authorityYear="2023" box="[189,380,1652,1678]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="almeria" status="gen. et sp. nov.">
<emphasis id="54E2EA95FFC14C62FF573B6DD407E591" box="[189,380,1652,1678]" italics="true" pageId="121" pageNumber="122">Parusia almeria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFC14C62FE6E3B6DD74FE590" box="[388,564,1653,1679]" pageId="121" pageNumber="122" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="66293687FFC14C62FF573B80D664E5AD" blockId="121.[189,911,1652,1925]" box="[189,799,1687,1714]" pageId="121" pageNumber="122">
<taxonomicName id="A1964D04FFC14C62FF573B80D73AE5AE" authority="(Fabricius, 1794)" authorityName="Gibbs" authorityYear="2023" baseAuthorityName="Fabricius" baseAuthorityYear="1794" box="[189,577,1687,1713]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="aurata" status="gen. et comb. nov.">
<emphasis id="54E2EA95FFC14C62FF573B80D414E5AE" box="[189,367,1688,1713]" italics="true" pageId="121" pageNumber="122">Parusia aurata</emphasis>
(
<bibRefCitation id="02074B76FFC14C62FE953B8FD743E5AE" author="Fabricius J. C." box="[383,568,1687,1713]" pageId="121" pageNumber="122" refId="ref74393" refString="Fabricius J. C. 1794. Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tome IV. C. G. Proft, Hafniae [= Copenhagen]. https: // doi. org / 10.5962 / bhl. title. 125869" type="book" year="1794">Fabricius, 1794</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFC14C62FDA23B80D664E5AD" box="[584,799,1687,1714]" pageId="121" pageNumber="122" rank="species">gen. et comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="66293687FFC14C62FF573BA3D74DE5CA" blockId="121.[189,911,1652,1925]" box="[189,566,1722,1749]" pageId="121" pageNumber="122">
<taxonomicName id="A1964D04FFC14C62FF573BA3D405E5CB" authority="Gibbs, 2023" authorityName="Gibbs" authorityYear="2023" box="[189,382,1722,1748]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="benoisti" status="gen. et sp. nov.">
<emphasis id="54E2EA95FFC14C62FF573BA3D405E5CB" box="[189,382,1722,1748]" italics="true" pageId="121" pageNumber="122">Parusia benoisti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFC14C62FE6F3BA3D74DE5CA" box="[389,566,1723,1749]" pageId="121" pageNumber="122" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="66293687FFC14C62FF573BC6D730E5E7" blockId="121.[189,911,1652,1925]" box="[189,587,1758,1784]" pageId="121" pageNumber="122">
<taxonomicName id="A1964D04FFC14C62FF573BC6D4EFE5E8" authority="Gibbs, 2023" authorityName="Gibbs" authorityYear="2023" box="[189,404,1758,1783]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="cyrenaica" status="gen. et sp. nov.">
<emphasis id="54E2EA95FFC14C62FF573BC6D4EFE5E8" box="[189,404,1758,1783]" italics="true" pageId="121" pageNumber="122">Parusia cyrenaica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFC14C62FE713BC6D730E5E7" box="[411,587,1758,1784]" pageId="121" pageNumber="122" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="66293687FFC14C62FF573A1AD75FE404" blockId="121.[189,911,1652,1925]" box="[189,548,1793,1819]" pageId="121" pageNumber="122">
<taxonomicName id="A1964D04FFC14C62FF573A1AD417E404" authority="Gibbs, 2023" authorityName="Gibbs" authorityYear="2023" box="[189,364,1793,1819]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="faesae" status="gen. et sp. nov.">
<emphasis id="54E2EA95FFC14C62FF573A1AD417E404" box="[189,364,1793,1819]" italics="true" pageId="121" pageNumber="122">Parusia faesae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFC14C62FE993A19D75FE404" box="[371,548,1793,1819]" pageId="121" pageNumber="122" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="66293687FFC14C62FF573A3DD78EE420" blockId="121.[189,911,1652,1925]" box="[189,757,1828,1855]" pageId="121" pageNumber="122">
<taxonomicName id="A1964D04FFC14C62FF573A3DD76CE420" authority="(Becker, 1906)" authorityName="Gibbs" authorityYear="2023" baseAuthorityName="Becker" baseAuthorityYear="1906" box="[189,535,1828,1855]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="loewi" status="gen. et comb. nov.">
<emphasis id="54E2EA95FFC14C62FF573A3DD41AE421" box="[189,353,1828,1854]" italics="true" pageId="121" pageNumber="122">Parusia loewi</emphasis>
(
<bibRefCitation id="02074B76FFC14C62FE9A3A3CD775E420" author="Becker T." box="[368,526,1828,1855]" pageId="121" pageNumber="122" pagination="193 - 228" refId="ref73899" refString="Becker T. 1906. Usia Latr. Berliner Entomologische Zeitschrift 50 [1905]: 193 - 228." type="journal article" year="1906">Becker, 1906</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFC14C62FDF43A3DD78EE420" box="[542,757,1828,1855]" pageId="121" pageNumber="122" rank="species">gen. et comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="66293687FFC14C62FF573A50D729E47D" blockId="121.[189,911,1652,1925]" box="[189,594,1864,1890]" pageId="121" pageNumber="122">
<taxonomicName id="A1964D04FFC14C62FF573A50D4E1E47E" authority="Gibbs, 2023" authorityName="Gibbs" authorityYear="2023" box="[189,410,1864,1889]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="propinqua" status="gen. et sp. nov.">
<emphasis id="54E2EA95FFC14C62FF573A50D4E1E47E" box="[189,410,1864,1889]" italics="true" pageId="121" pageNumber="122">Parusia propinqua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFC14C62FE483A50D729E47D" box="[418,594,1864,1890]" pageId="121" pageNumber="122" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="66293687FFC14C62FF573A73D6F4E49A" blockId="121.[189,911,1652,1925]" box="[189,911,1898,1925]" pageId="121" pageNumber="122">
<taxonomicName id="A1964D04FFC14C62FF573A73D744E49A" authority="(Costa, 1883)" authorityName="Gibbs" authorityYear="2023" baseAuthorityName="Costa" baseAuthorityYear="1883" box="[189,575,1898,1925]" class="Insecta" family="Bombyliidae" genus="Parusia" kingdom="Animalia" order="Diptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="taeniolata" status="stat. nov.">
<emphasis id="54E2EA95FFC14C62FF573A73D4E3E49B" box="[189,408,1898,1924]" italics="true" pageId="121" pageNumber="122">Parusia taeniolata</emphasis>
(Costa, 1883)
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFC14C62FDAD3A73D7D7E49A" box="[583,684,1899,1925]" pageId="121" pageNumber="122">stat. rev.</taxonomicNameLabel>
, gen. et comb. nov.
</paragraph>
</subSubSection>
</treatment>
</document>
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