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<mods:title id="5A7094EC6367D65A007BB566E94109AE">Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar</mods:title>
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<mods:affiliation id="DC88F854C424F7D7FD8F4290468AA940">Institut de Systématique, Evolution, Biodiversité (ISYEB), MNHN-CNRS-Sorbonne Université- EPHE-Université des Antilles, Muséum national d’Histoire naturelle, CP 50, 45 rue Buffon, 75005 Paris, France.</mods:affiliation>
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<mods:namePart id="3C6CF062ADA936E63021C2E79E483B0B">Bouteille, Emilien</mods:namePart>
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<mods:affiliation id="CFAF67C88047106553EA1E9B8055AE69">Mécanismes adaptatifs et évolution (MECADEV), Muséum national d’Histoire naturelle, CNRS, 57 rue Cuvier, CP 50, 75005 Paris, France.</mods:affiliation>
<mods:nameIdentifier id="364C72D1F5E46F029CCA3708B00E91BB" type="email">emilien.bouteille@gmail.com</mods:nameIdentifier>
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<mods:namePart id="31F503E7ED9D23AEDE207F836285BC08">Cesne, Maxime Le</mods:namePart>
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<mods:affiliation id="EEBCD5596DB1032A3F8D56CBC3848B19">Institut de Systématique, Evolution, Biodiversité (ISYEB), MNHN-CNRS-Sorbonne Université- EPHE-Université des Antilles, Muséum national d’Histoire naturelle, CP 50, 45 rue Buffon, 75005 Paris, France.</mods:affiliation>
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<treatment id="E8713702D105FFB5925EB3E0FB45F9CB" ID-DOI="http://doi.org/10.5281/zenodo.11400834" ID-GBIF-Taxon="232142718" ID-Zenodo-Dep="11400834" LSID="urn:lsid:plazi:treatment:E8713702D105FFB5925EB3E0FB45F9CB" httpUri="http://treatment.plazi.org/id/E8713702D105FFB5925EB3E0FB45F9CB" lastPageId="23" lastPageNumber="189" pageId="21" pageNumber="187">
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<heading id="3B2F3178D105FFB7925EB3E0FBC9F94C" box="[512,1076,1674,1701]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="187" reason="2">
<taxonomicName id="A7D8FD97D105FFB7925EB3E0FC56F94D" authority="(Stal, 1866)" authorityName="Bucher &amp; Bouteille &amp; Cesne" authorityYear="2024" baseAuthorityName="Stal" baseAuthorityYear="1866" box="[512,939,1674,1701]" class="Insecta" family="Cercopidae" genus="Soulierana" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="187" phylum="Arthropoda" rank="species" species="laeviuscula" status="comb. nov.">
<emphasis id="52AC5A06D105FFB7925EB3E0FCE5F94C" bold="true" box="[512,792,1674,1701]" italics="true" pageId="21" pageNumber="187">Soulierana laeviuscula</emphasis>
(
<bibRefCitation id="0449FBE5D105FFB79377B3E0FC5EF94D" author="Stal C." box="[809,931,1674,1701]" firstAuthor="Stal" journalOrPublisher="Hemiptera Africana" pageId="21" pageNumber="187" pagination="1 - 276" part="4" refId="ref15853" refString="Stal C. 1866. Hemiptera Homoptera Latr. Hemiptera Africana 4: 1 - 276." title="Hemiptera Homoptera Latr" type="journal article" year="1866">Stål, 1866</bibRefCitation>
)
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<taxonomicNameLabel id="499FE77DD105FFB793ECB3E0FBC9F94C" box="[946,1076,1674,1701]" pageId="21" pageNumber="187" rank="species">comb. nov.</taxonomicNameLabel>
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<paragraph id="60678614D105FFB792A9B3C4FCC1F921" blockId="21.[512,1076,1674,1736]" box="[759,828,1710,1736]" pageId="21" pageNumber="187">
<figureCitation id="F8E39A91D105FFB792A9B3C4FCC1F921" box="[759,828,1710,1736]" captionStart="Fig" captionStartId="22.[189,232,1892,1918]" captionTargetBox="[265,1278,290,1817]" captionTargetId="figure-81@22.[240,1320,265,1825]" captionTargetPageId="22" captionText="Fig. 9. Soulierana laeviuscula (Stål, 1866), ♂ (CASENT 3001283). A. Dorsal view. B. Lateral view. C. Frontal view.D. Labels.E. Male terminalia in lateral view, pygofer, anal tube, aedeagus, left paramere and left subgenital plate. F. Aedeagus. G. Left paramere, inside view on the left and outside view on the rigth. H. Distribution map. Scale bars: A–C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11400846" httpUri="https://zenodo.org/record/11400846/files/figure.png" pageId="21" pageNumber="187">Fig. 9</figureCitation>
</paragraph>
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<paragraph id="60678614D105FFB790E3B39AFDABF8E0" blockId="21.[189,598,1775,1802]" box="[189,598,1775,1802]" pageId="21" pageNumber="187">
<treatmentCitationGroup id="40C8A13AD105FFB790E3B39AFDABF8E0" box="[189,598,1775,1802]" pageId="21" pageNumber="187">
<treatmentCitation id="E179A005D105FFB790E3B39AFDAFF8E0" author="Stal C." box="[189,594,1775,1802]" page="64" pageId="21" pageNumber="187" year="1866">
<taxonomicName id="A7D8FD97D105FFB790E3B39AFDAFF8E0" ID-CoL="3VHDF" authority="Stal, 1866: 64" authorityName="Stal" authorityPageNumber="64" authorityYear="1866" box="[189,594,1775,1802]" class="Insecta" family="Cercopidae" genus="Literna" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="187" phylum="Arthropoda" rank="species" species="laeviuscula">
<emphasis id="52AC5A06D105FFB790E3B39AFE5EF8E0" box="[189,419,1775,1801]" italics="true" pageId="21" pageNumber="187">Literna laeviuscula</emphasis>
<bibRefCitation id="0449FBE5D105FFB791F5B385FDAFF8E0" author="Stal C." box="[427,594,1775,1802]" firstAuthor="Stal" journalOrPublisher="Hemiptera Africana" page="64" pageId="21" pageNumber="187" pagination="1 - 276" part="4" refId="ref15853" refString="Stal C. 1866. Hemiptera Homoptera Latr. Hemiptera Africana 4: 1 - 276." title="Hemiptera Homoptera Latr" type="journal article" year="1866">Stål, 1866: 64</bibRefCitation>
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.
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</paragraph>
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<subSubSection id="28C2D59FD105FFB590E3B25AFAD4FDD4" lastPageId="23" lastPageNumber="189" pageId="21" pageNumber="187" type="materials_examined">
<paragraph id="60678614D105FFB790E3B25AFE57F8A3" blockId="21.[189,426,1840,1866]" box="[189,426,1840,1866]" pageId="21" pageNumber="187">
<heading id="3B2F3178D105FFB790E3B25AFE57F8A3" bold="true" box="[189,426,1840,1866]" fontSize="11" level="3" pageId="21" pageNumber="187" reason="3">
<emphasis id="52AC5A06D105FFB790E3B25AFE57F8A3" bold="true" box="[189,426,1840,1866]" pageId="21" pageNumber="187">Material examined</emphasis>
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•
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a.s.l.
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leg.;
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•
<materialsCitation id="D0B08C49D105FFB79409B2E9FE61F80D" ID-GBIF-Occurrence="4873642321" collectedFrom="foret seche" collectingDate="2011-01-12" collectionCode="MNHN" collectorName="D. Ouvrard" country="Madagascar" elevation="243" latitude="-21.6655556" location="massif du Makay" longitude="45.22888888888889" pageId="21" pageNumber="187" specimenCode="MNHN EH 24541" specimenCount="1" specimenCount-male="1" stateProvince="Toliara">
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; Province de
<collectingRegion id="A21C48F6D105FFB79541B2E8FA8EF875" box="[1311,1395,1922,1948]" pageId="21" pageNumber="187">Toliara</collectingRegion>
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<collectedFrom id="D3C4C5E9D105FFB791CDB2CFFDE8F856" box="[403,533,1957,1983]" pageId="21" pageNumber="187">forêt sèche</collectedFrom>
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a.s.l.
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<collectingDate id="0422593CD105FFB7944DB2CCFB56F829" box="[1043,1195,1958,1984]" pageId="21" pageNumber="187" value="2011-01-12">12 Jan. 2011</collectingDate>
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leg.;
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<specimenCode id="307E2E6FD105FFB79146B2A3FE61F80D" box="[280,412,1993,2020]" pageId="21" pageNumber="187">(EH) 24541</specimenCode>
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•
<materialsCitation id="D0B08C49D105FFB591ECB2A3FC4CFECA" ID-GBIF-Occurrence="4873642313" collectedFrom="foret seche" collectingDate="2011-01-12" collectionCode="MNHN" collectorName="D. Ouvrard" country="Madagascar" elevation="243" lastPageId="23" lastPageNumber="189" latitude="-21.6655556" location="massif du Makay" longitude="45.22888888888889" pageId="21" pageNumber="187" specimenCode="MNHN EH 24700" specimenCount="1" specimenCount-male="1" stateProvince="Toliara">
<specimenCount id="76DE4D9DD105FFB791ECB2A3FE23F80A" box="[434,478,1993,2019]" count="1" pageId="21" pageNumber="187" type="male">1 ♂</specimenCount>
; Province de
<collectingRegion id="A21C48F6D105FFB79228B2A3FD37F80A" box="[630,714,1993,2019]" pageId="21" pageNumber="187">Toliara</collectingRegion>
,
<location id="6507D0CFD105FFB7928DB2A3FC6AF80A" LSID="urn:lsid:plazi:treatment:E8713702D105FFB5925EB3E0FB45F9CB:6507D0CFD105FFB7928DB2A3FC6AF80A" box="[723,919,1993,2019]" country="Madagascar" name="massif du Makay" pageId="21" pageNumber="187" stateProvince="Toliara">massif du Makay</location>
,
<collectedFrom id="D3C4C5E9D105FFB793FFB2A3FBDDF80A" box="[929,1056,1993,2019]" pageId="21" pageNumber="187">forêt sèche</collectedFrom>
;
<geoCoordinate id="05ECE0D3D105FFB79474B2ADFB39F80A" box="[1066,1220,1991,2019]" degrees="21" direction="south" minutes="36" orientation="latitude" pageId="21" pageNumber="187" seconds="236" value="-21.6655556">21°36′236″ S</geoCoordinate>
,
<geoCoordinate id="05ECE0D3D105FFB79491B2ADFA8FF80A" box="[1231,1394,1991,2019]" degrees="45" direction="east" minutes="06" orientation="longitude" pageId="21" pageNumber="187" seconds="464" value="45.22888888888889">45°06′464″ E</geoCoordinate>
;
<elevation id="EBF56127D107FFB590E3B463FEB7FECA" box="[189,330,265,291]" metricMagnitude="2" metricUnit="m" metricValue="2.43" pageId="23" pageNumber="189" unit="m" value="243.0">
<quantity id="A7202BF1D107FFB590E3B463FEF4FECA" box="[189,265,265,291]" metricMagnitude="2" metricUnit="m" metricValue="2.43" pageId="23" pageNumber="189" unit="m" value="243.0">243 m</quantity>
a.s.l.
</elevation>
;
<date id="1466A0D4D107FFB59107B463FE0EFECA" box="[345,499,265,291]" pageId="23" pageNumber="189" value="2011-01-12">
<collectingDate id="0422593CD107FFB59107B463FE0EFECA" box="[345,499,265,291]" pageId="23" pageNumber="189" value="2011-01-12">12 Jan. 2011</collectingDate>
</date>
;
<collectorName id="CD2DE3C2D107FFB5925FB463FD7AFECA" box="[513,647,265,291]" pageId="23" pageNumber="189">D. Ouvrard</collectorName>
leg.;
<collectionCode id="06C91ED1D107FFB59293B463FCD5FECA" box="[717,808,265,291]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="189">MNHN</collectionCode>
<specimenCode id="307E2E6FD107FFB59376B463FC4CFECA" box="[808,945,265,291]" pageId="23" pageNumber="189">(EH) 24700</specimenCode>
</materialsCitation>
•
<materialsCitation id="D0B08C49D107FFB59391B463FEF8FE80" ID-GBIF-Occurrence="4873642304" collectedFrom="foret seche" collectingDate="2011-01-12" collectionCode="MNHN" collectorName="D. Ouvrard" country="Madagascar" elevation="243" latitude="-21.6655556" location="massif du Makay" longitude="45.22888888888889" pageId="23" pageNumber="189" specimenCode="MNHN EH 24696" specimenCount="1" specimenCount-female="1" stateProvince="Toliara">
<specimenCount id="76DE4D9DD107FFB59391B463FC03FECA" box="[975,1022,265,291]" count="1" pageId="23" pageNumber="189" type="female">1 ♀</specimenCount>
; Province de
<collectingRegion id="A21C48F6D107FFB594FDB463FB0AFECA" box="[1187,1271,265,291]" pageId="23" pageNumber="189">Toliara</collectingRegion>
,
<location id="6507D0CFD107FFB5955BB463FEF0FEAF" LSID="urn:lsid:plazi:treatment:E8713702D105FFB5925EB3E0FB45F9CB:6507D0CFD107FFB5955BB463FEF0FEAF" country="Madagascar" name="massif du Makay" pageId="23" pageNumber="189" stateProvince="Toliara">massif du Makay</location>
<collectedFrom id="D3C4C5E9D107FFB5914AB446FE6EFEAF" box="[276,403,300,326]" pageId="23" pageNumber="189">forêt sèche</collectedFrom>
;
<geoCoordinate id="05ECE0D3D107FFB591C1B441FDC7FEAF" box="[415,570,299,327]" degrees="21" direction="south" minutes="36" orientation="latitude" pageId="23" pageNumber="189" seconds="236" value="-21.6655556">21°36′236″ S</geoCoordinate>
,
<geoCoordinate id="05ECE0D3D107FFB59218B441FD19FEAE" box="[582,740,299,327]" degrees="45" direction="east" minutes="06" orientation="longitude" pageId="23" pageNumber="189" seconds="464" value="45.22888888888889">45°06′464″ E</geoCoordinate>
;
<elevation id="EBF56127D107FFB592B1B446FC8BFEAF" box="[751,886,300,327]" metricMagnitude="2" metricUnit="m" metricValue="2.43" pageId="23" pageNumber="189" unit="m" value="243.0">
<quantity id="A7202BF1D107FFB592B1B446FCC4FEAE" box="[751,825,300,327]" metricMagnitude="2" metricUnit="m" metricValue="2.43" pageId="23" pageNumber="189" unit="m" value="243.0">243 m</quantity>
a.s.l.
</elevation>
;
<date id="1466A0D4D107FFB593DDB446FBEAFEAF" box="[899,1047,300,327]" pageId="23" pageNumber="189" value="2011-01-12">
<collectingDate id="0422593CD107FFB593DDB446FBEAFEAF" box="[899,1047,300,327]" pageId="23" pageNumber="189" value="2011-01-12">12 Jan. 2011</collectingDate>
</date>
;
<collectorName id="CD2DE3C2D107FFB5947CB447FB58FEAF" box="[1058,1189,300,327]" pageId="23" pageNumber="189">D. Ouvrard</collectorName>
leg.;
<collectionCode id="06C91ED1D107FFB594BAB446FABDFEAF" box="[1252,1344,300,326]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="189">MNHN</collectionCode>
<specimenCode id="307E2E6FD107FFB5951EB446FEF8FE80" pageId="23" pageNumber="189">(EH) 24696</specimenCode>
</materialsCitation>
•
<materialsCitation id="D0B08C49D107FFB59142B425FCEDFE65" ID-GBIF-Occurrence="4873642337" collectedFrom="foret seche" collectingDate="2011-01-12" collectionCode="MNHN" collectorName="D. Ouvrard" country="Madagascar" elevation="243" latitude="-21.6655556" location="massif du Makay" longitude="45.22888888888889" pageId="23" pageNumber="189" specimenCode="MNHN EH 24697" specimenCount="1" specimenCount-female="1" stateProvince="Toliara">
<specimenCount id="76DE4D9DD107FFB59142B425FEB4FE83" box="[284,329,335,362]" count="1" pageId="23" pageNumber="189" type="female">1 ♀</specimenCount>
; Province de
<collectingRegion id="A21C48F6D107FFB591BAB425FDC5FE80" box="[484,568,335,361]" pageId="23" pageNumber="189">Toliara</collectingRegion>
,
<location id="6507D0CFD107FFB5921DB425FCF5FE80" LSID="urn:lsid:plazi:treatment:E8713702D105FFB5925EB3E0FB45F9CB:6507D0CFD107FFB5921DB425FCF5FE80" box="[579,776,335,361]" country="Madagascar" name="massif du Makay" pageId="23" pageNumber="189" stateProvince="Toliara">massif du Makay</location>
<collectedFrom id="D3C4C5E9D107FFB59351B425FC72FE80" box="[783,911,335,361]" pageId="23" pageNumber="189">forêt sèche</collectedFrom>
;
<geoCoordinate id="05ECE0D3D107FFB593C4B424FBC8FE80" box="[922,1077,334,362]" degrees="21" direction="south" minutes="36" orientation="latitude" pageId="23" pageNumber="189" seconds="236" value="-21.6655556">21°36′236″ S</geoCoordinate>
,
<geoCoordinate id="05ECE0D3D107FFB5941FB424FB22FE83" box="[1089,1247,334,362]" degrees="45" direction="east" minutes="06" orientation="longitude" pageId="23" pageNumber="189" seconds="464" value="45.22888888888889">45°06′464″ E</geoCoordinate>
;
<elevation id="EBF56127D107FFB594B4B425FA8CFE80" box="[1258,1393,335,362]" metricMagnitude="2" metricUnit="m" metricValue="2.43" pageId="23" pageNumber="189" unit="m" value="243.0">
<quantity id="A7202BF1D107FFB594B4B425FACEFE83" box="[1258,1331,335,362]" metricMagnitude="2" metricUnit="m" metricValue="2.43" pageId="23" pageNumber="189" unit="m" value="243.0">243 m</quantity>
a.s.l.
</elevation>
;
<date id="1466A0D4D107FFB590E3B419FEABFE64" box="[189,342,371,397]" pageId="23" pageNumber="189" value="2011-01-12">
<collectingDate id="0422593CD107FFB590E3B419FEABFE64" box="[189,342,371,397]" pageId="23" pageNumber="189" value="2011-01-12">12 Jan. 2011</collectingDate>
</date>
;
<collectorName id="CD2DE3C2D107FFB5913DB419FE14FE65" box="[355,489,370,397]" pageId="23" pageNumber="189">D. Ouvrard</collectorName>
leg.;
<collectionCode id="06C91ED1D107FFB59273B419FD75FE64" box="[557,648,371,397]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="189">MNHN</collectionCode>
<specimenCode id="307E2E6FD107FFB592D7B419FCEDFE65" box="[649,784,370,397]" pageId="23" pageNumber="189">(EH) 24697</specimenCode>
</materialsCitation>
•
<materialsCitation id="D0B08C49D107FFB59373B419FDF0FE1E" ID-GBIF-Occurrence="4873642301" collectedFrom="tropical dry forest" collectingDate="2001-03-11" collectingDateMax="2001-03-16" collectingDateMin="2001-03-11" collectingMethod="malaise trap" collectionCode="BLF, CASENT" collectorName="Fisher &amp; Griswold" country="Madagascar" elevation="120" latitude="-14.308889" location="Foret d'Anabohazo" longLatPrecision="21" longitude="47.914448" pageId="23" pageNumber="189" specimenCode="BLF336, CASENT 3001856" specimenCount="1" specimenCount-male="1" stateProvince="Province d'Antsiranana">
<specimenCount id="76DE4D9DD107FFB59373B419FCA0FE64" box="[813,861,371,397]" count="1" pageId="23" pageNumber="189" type="male">1 ♂</specimenCount>
;
<collectingCounty id="8906FE98D107FFB59332B418FB7CFE65" box="[876,1153,370,396]" pageId="23" pageNumber="189">
<collectingRegion id="A21C48F6D107FFB59332B418FB7CFE65" box="[876,1153,370,396]" pageId="23" pageNumber="189">Province d’Antsiranana</collectingRegion>
</collectingCounty>
,
<location id="6507D0CFD107FFB594D0B419FA8FFE65" LSID="urn:lsid:plazi:treatment:E8713702D105FFB5925EB3E0FB45F9CB:6507D0CFD107FFB594D0B419FA8FFE65" box="[1166,1394,370,397]" country="Madagascar" county="Province d'Antsiranana" latitude="-14.308889" longLatPrecision="21" longitude="47.914448" municipality="Foret d'Anabohazo" name="Foret d'Anabohazo" pageId="23" pageNumber="189">Forêt d’Anabohazo</location>
,
<locationDeviation id="71D59F36D107FFB590E3B4FCFD9DFE59" box="[189,608,406,432]" location="Macromandia" pageId="23" pageNumber="189">
<quantity id="A7202BF1D107FFB590E3B4FCFEE2FE59" box="[189,287,406,432]" metricMagnitude="4" metricUnit="m" metricValue="2.16" pageId="23" pageNumber="189" unit="km" value="21.6">21.6 km</quantity>
247° WSW Macromandia
</locationDeviation>
;
<geoCoordinate id="05ECE0D3D107FFB59230B4FEFD02FE59" box="[622,767,404,432]" degrees="14" direction="south" minutes="18" orientation="latitude" pageId="23" pageNumber="189" precision="15" seconds="32" value="-14.308889">14°18′32″ S</geoCoordinate>
,
<geoCoordinate id="05ECE0D3D107FFB59351B4FEFC5FFE59" box="[783,930,404,432]" degrees="47" direction="east" minutes="54" orientation="longitude" pageId="23" pageNumber="189" precision="15" seconds="52" value="47.914448">47°54′52″ E</geoCoordinate>
;
<elevation id="EBF56127D107FFB593ECB4FCFBBDFE59" box="[946,1088,406,432]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="189" unit="m" value="120.0">
<quantity id="A7202BF1D107FFB593ECB4FCFC02FE59" box="[946,1023,406,432]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="189" unit="m" value="120.0">120 m</quantity>
a.s.l.
</elevation>
;
<date id="1466A0D4D107FFB5940EB4FCFAE2FE59" box="[1104,1311,406,432]" pageId="23" pageNumber="189" value="2001-03-11" valueMax="2001-03-16" valueMin="2001-03-11">
<collectingDate id="0422593CD107FFB5940EB4FCFAE2FE59" box="[1104,1311,406,432]" pageId="23" pageNumber="189" value="2001-03-11" valueMax="2001-03-16" valueMin="2001-03-11">11–16 Mar. 2001</collectingDate>
</date>
;
<collectorName id="CD2DE3C2D107FFB59571B4FCFA8AFE59" box="[1327,1399,406,432]" pageId="23" pageNumber="189">Fisher</collectorName>
and
<collectorName id="CD2DE3C2D107FFB590B1B4D3FEA7FE3A" box="[239,346,441,467]" pageId="23" pageNumber="189">Griswold</collectorName>
leg.;
<collectingMethod id="B999FE03D107FFB591C0B4D3FDD1FE3A" box="[414,556,441,467]" pageId="23" pageNumber="189">malaise trap</collectingMethod>
in
<collectedFrom id="D3C4C5E9D107FFB5920DB4D3FCD5FE3A" box="[595,808,441,467]" pageId="23" pageNumber="189">tropical dry forest</collectedFrom>
; California Academy of Sciences; collecting code:
<specimenCode id="307E2E6FD107FFB590E1B4B6FEDFFE1F" box="[191,290,476,502]" pageId="23" pageNumber="189">BLF336</specimenCode>
;
<collectionCode id="06C91ED1D107FFB59171B4B6FE62FE1F" box="[303,415,476,502]" pageId="23" pageNumber="189">CASENT</collectionCode>
<specimenCode id="307E2E6FD107FFB591F6B4B6FDF0FE1E" box="[424,525,476,503]" pageId="23" pageNumber="189">3001856</specimenCode>
</materialsCitation>
•
<materialsCitation id="D0B08C49D107FFB59276B4B6FAD8FDD4" ID-GBIF-Occurrence="4873642346" collectedFrom="in rainforest" collectingDate="2001-03-04" collectingDateMax="2001-03-09" collectingDateMin="2001-03-04" collectingMethod="malaise trap" collectionCode="BLF, CASENT" collectorName="Fischer &amp; Griswold" country="Madagascar" elevation="600" latitude="-13.798612" location="Foret d'Ambilanivy" longLatPrecision="21" longitude="48.161667" municipality="Ampasindava" pageId="23" pageNumber="189" specimenCode="BLF3251, CASENT 3001283" specimenCount="1" specimenCount-male="1" stateProvince="Province d'Antsiranana">
<specimenCount id="76DE4D9DD107FFB59276B4B6FDA8FE1E" box="[552,597,476,503]" count="1" pageId="23" pageNumber="189" type="male">1 ♂</specimenCount>
;
<collectingCounty id="8906FE98D107FFB5923AB4B6FC8AFE1F" box="[612,887,476,502]" pageId="23" pageNumber="189">
<collectingRegion id="A21C48F6D107FFB5923AB4B6FC8AFE1F" box="[612,887,476,502]" pageId="23" pageNumber="189">Province d’Antsiranana</collectingRegion>
</collectingCounty>
,
<collectingMunicipality id="80031C6ED107FFB593DDB4B6FBD9FE1F" box="[899,1060,476,502]" pageId="23" pageNumber="189">Ampasindava</collectingMunicipality>
,
<location id="6507D0CFD107FFB5946FB4B6FAE5FE1F" LSID="urn:lsid:plazi:treatment:E8713702D105FFB5925EB3E0FB45F9CB:6507D0CFD107FFB5946FB4B6FAE5FE1F" box="[1073,1304,476,502]" country="Madagascar" county="Province d'Antsiranana" latitude="-13.798612" longLatPrecision="21" longitude="48.161667" municipality="Ampasindava" name="Foret d'Ambilanivy" pageId="23" pageNumber="189">Forêt d’Ambilanivy</location>
,
<locationDeviation id="71D59F36D107FFB5957BB4B6FE7FFDF0" location="Ambaliha" pageId="23" pageNumber="189">
<quantity id="A7202BF1D107FFB5957BB4B6FA8BFE1F" box="[1317,1398,476,502]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="23" pageNumber="189" unit="km" value="3.9">3.9 km</quantity>
181° S Ambaliha
</locationDeviation>
;
<geoCoordinate id="05ECE0D3D107FFB591D2B494FDEAFDF0" box="[396,535,510,538]" degrees="13" direction="south" minutes="47" orientation="latitude" pageId="23" pageNumber="189" precision="15" seconds="55" value="-13.798612">13°47′55″ S</geoCoordinate>
,
<geoCoordinate id="05ECE0D3D107FFB5927CB494FD5FFDF3" box="[546,674,510,538]" degrees="48" direction="east" minutes="9" orientation="longitude" pageId="23" pageNumber="189" precision="15" seconds="42" value="48.161667">48°9′42″ E</geoCoordinate>
;
<elevation id="EBF56127D107FFB592F2B495FCCDFDF0" box="[684,816,511,538]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="189" unit="m" value="600.0">
<quantity id="A7202BF1D107FFB592F2B495FD09FDF3" box="[684,756,511,538]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="189" unit="m" value="600.0">600 m</quantity>
a.s.l.
</elevation>
;
<date id="1466A0D4D107FFB59365B495FC19FDF0" box="[827,996,511,538]" pageId="23" pageNumber="189" value="2001-03-04" valueMax="2001-03-09" valueMin="2001-03-04">
<collectingDate id="0422593CD107FFB59365B495FC19FDF0" box="[827,996,511,538]" pageId="23" pageNumber="189" value="2001-03-04" valueMax="2001-03-09" valueMin="2001-03-04">4–9 Mar. 2001</collectingDate>
</date>
;
<collectorName id="CD2DE3C2D107FFB593B0B495FBBEFDF0" box="[1006,1091,511,537]" pageId="23" pageNumber="189">Fischer</collectorName>
and
<collectorName id="CD2DE3C2D107FFB59429B495FB1FFDF0" box="[1143,1250,511,537]" pageId="23" pageNumber="189">Griswold</collectorName>
leg.;
<collectingMethod id="B999FE03D107FFB59541B495FF17FDD4" pageId="23" pageNumber="189">malaise trap</collectingMethod>
<collectedFrom id="D3C4C5E9D107FFB590AEB748FE82FDD5" box="[240,383,546,572]" pageId="23" pageNumber="189">in rainforest</collectedFrom>
; California Academy of Sciences; collection code:
<specimenCode id="307E2E6FD107FFB59396B748FBC7FDD4" box="[968,1082,546,573]" pageId="23" pageNumber="189">BLF3251</specimenCode>
;
<collectionCode id="06C91ED1D107FFB5941BB749FB48FDD4" box="[1093,1205,547,573]" pageId="23" pageNumber="189">CASENT</collectionCode>
<specimenCode id="307E2E6FD107FFB594E2B749FAD8FDD4" box="[1212,1317,547,573]" pageId="23" pageNumber="189">3001283</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="34A7D69CD106FFB490E3B20EFCDBF800" ID-DOI="http://doi.org/10.5281/zenodo.11400846" ID-Zenodo-Dep="11400846" httpUri="https://zenodo.org/record/11400846/files/figure.png" pageId="22" pageNumber="188" startId="22.[189,232,1892,1918]" targetBox="[265,1278,290,1817]" targetPageId="22">
<paragraph id="60678614D106FFB490E3B20EFCDBF800" blockId="22.[187,1399,1892,2025]" pageId="22" pageNumber="188">
<emphasis id="52AC5A06D106FFB490E3B20EFEF0F896" bold="true" box="[189,269,1892,1919]" pageId="22" pageNumber="188">Fig. 9.</emphasis>
<taxonomicName id="A7D8FD97D106FFB49148B20FFD3CF896" authority="(Stal, 1866)" authorityName="Bucher &amp; Bouteille &amp; Cesne" authorityYear="2024" baseAuthorityName="Stal" baseAuthorityYear="1866" box="[278,705,1892,1919]" class="Insecta" family="Cercopidae" genus="Soulierana" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="188" phylum="Arthropoda" rank="species" species="laeviuscula">
<emphasis id="52AC5A06D106FFB49148B20FFDD4F896" box="[278,553,1893,1919]" italics="true" pageId="22" pageNumber="188">Soulierana laeviuscula</emphasis>
(
<bibRefCitation id="0449FBE5D106FFB49265B20EFD45F896" author="Stal C." box="[571,696,1892,1919]" firstAuthor="Stal" journalOrPublisher="Hemiptera Africana" pageId="22" pageNumber="188" pagination="1 - 276" part="4" refId="ref15853" refString="Stal C. 1866. Hemiptera Homoptera Latr. Hemiptera Africana 4: 1 - 276." title="Hemiptera Homoptera Latr" type="journal article" year="1866">Stål, 1866</bibRefCitation>
)
</taxonomicName>
, ♂ (CASENT 3001283).
<emphasis id="52AC5A06D106FFB493ADB20EFBF5F897" bold="true" box="[1011,1032,1892,1918]" pageId="22" pageNumber="188">A</emphasis>
. Dorsal view.
<emphasis id="52AC5A06D106FFB494E4B20FFB31F896" bold="true" box="[1210,1228,1893,1919]" pageId="22" pageNumber="188">B</emphasis>
. Lateral view.
<emphasis id="52AC5A06D106FFB490E1B2E2FF2FF84B" bold="true" box="[191,210,1928,1954]" pageId="22" pageNumber="188">C</emphasis>
. Frontal view.
<emphasis id="52AC5A06D106FFB49125B2E2FE6DF84B" bold="true" box="[379,400,1928,1954]" pageId="22" pageNumber="188">D</emphasis>
. Labels.
<emphasis id="52AC5A06D106FFB491A6B2E2FDF1F84B" bold="true" box="[504,524,1928,1954]" pageId="22" pageNumber="188">E</emphasis>
. Male terminalia in lateral view, pygofer, anal tube, aedeagus, left paramere and left subgenital plate.
<emphasis id="52AC5A06D106FFB491BFB2C1FE0EF82C" bold="true" box="[481,499,1963,1989]" pageId="22" pageNumber="188">F</emphasis>
. Aedeagus.
<emphasis id="52AC5A06D106FFB492DEB2C1FD6AF82C" bold="true" box="[640,663,1963,1989]" pageId="22" pageNumber="188">G</emphasis>
. Left paramere, inside view on the left and outside view on the rigth.
<emphasis id="52AC5A06D106FFB4915DB2A4FEE7F801" bold="true" box="[259,282,1998,2024]" pageId="22" pageNumber="188">H</emphasis>
. Distribution map. Scale bars: A–C = 1 mm.
</paragraph>
</caption>
<subSubSection id="28C2D59FD107FFB590E3B70EFB45F9CB" pageId="23" pageNumber="189" type="description">
<paragraph id="60678614D107FFB590E3B70EFEB3FD97" blockId="23.[189,334,612,638]" box="[189,334,612,638]" pageId="23" pageNumber="189">
<heading id="3B2F3178D107FFB590E3B70EFEB3FD97" bold="true" box="[189,334,612,638]" fontSize="11" level="3" pageId="23" pageNumber="189" reason="3">
<emphasis id="52AC5A06D107FFB590E3B70EFEB3FD97" bold="true" box="[189,334,612,638]" pageId="23" pageNumber="189">Description</emphasis>
</heading>
</paragraph>
<paragraph id="60678614D107FFB590E3B7F8FE12FD1A" blockId="23.[189,1399,658,755]" pageId="23" pageNumber="189">
This species was first described by
<bibRefCitation id="0449FBE5D107FFB59234B7F8FD0DFD45" author="Stal C." box="[618,752,658,685]" firstAuthor="Stal" journalOrPublisher="Hemiptera Africana" pageId="23" pageNumber="189" pagination="1 - 276" part="4" refId="ref15853" refString="Stal C. 1866. Hemiptera Homoptera Latr. Hemiptera Africana 4: 1 - 276." title="Hemiptera Homoptera Latr" type="journal article" year="1866">Stål (1866)</bibRefCitation>
from a single female specimen. Here, we propose its integration into the genus
<taxonomicName id="A7D8FD97D107FFB591A7B7DCFC8EFD39" authority="Bucher &amp; Bouteille" authorityName="Bucher &amp; Bouteille" authorityYear="2024" box="[505,883,694,720]" class="Insecta" family="Cercopidae" genus="Soulierana" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="189" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="52AC5A06D107FFB591A7B7DCFD86FD39" box="[505,635,694,720]" italics="true" pageId="23" pageNumber="189">Soulierana</emphasis>
Bucher &amp; Bouteille
</taxonomicName>
<taxonomicNameLabel id="499FE77DD107FFB59323B7DCFC11FD39" box="[893,1004,694,720]" pageId="23" pageNumber="189" rank="genus">gen. nov.</taxonomicNameLabel>
with the description of the male genitalia for the first time.
</paragraph>
<paragraph id="60678614D107FFB590E3B670FD54FCDC" blockId="23.[189,681,794,821]" box="[189,681,794,821]" pageId="23" pageNumber="189">
TOTAL LENGTH. Between 5.2 and
<quantity id="A7202BF1D107FFB59216B670FD58FCDC" box="[584,677,794,821]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="23" pageNumber="189" unit="mm" value="5.5">5.5 mm</quantity>
.
</paragraph>
<paragraph id="60678614D107FFB590E3B636FD1DFBEA" blockId="23.[187,1399,860,1027]" pageId="23" pageNumber="189">HEAD. Median part of vertex relatively flat, forming very obtuse angle with postclypeus. Median furrow on postclypeus wide, deep, beginning below front margin of head and straight at upper extremity. Lateral bulges very thin and comparable to two carinae, parallel and keeping same width over entire length; in dorsal view, postclypeus slightly exceeding anterior margin of head; in lateral view, postclypeus convex and forming obtuse angle slightly pronounced.</paragraph>
<paragraph id="60678614D107FFB590E3B140FDADFBAD" blockId="23.[189,592,1066,1092]" box="[189,592,1066,1092]" pageId="23" pageNumber="189">THORAX. Scutellum relatively flat.</paragraph>
<paragraph id="60678614D107FFB590E3B106FDB0FB40" blockId="23.[189,1399,1131,1193]" pageId="23" pageNumber="189">TEGMINA. Three bumps clearly distinguishable on each tegmen. Apical network not dense with fewer than 15 concave cells per tegmen.</paragraph>
<paragraph id="60678614D107FFB590E3B1BAFAD7FA08" blockId="23.[189,1401,1232,1505]" pageId="23" pageNumber="189">MALE GENITALIA. Subgenital plates longer than length of pygofer, margin wider on first third and curved dorsally on apical third; on apical margin, flattened internal part covered with setae forming rounded protuberance. Sterno-lateral plates greatly exceeding posterior margin of pygofer. Parameres inserted almost entirely in pygofer and not very visible in lateral view, their dorsal margin bearing dorsal extension, posterior tip forming spine hook-shaped clearly visible in dorsal view. Aedeagus developed anteriorly in one-pointed protrusions; ventral margin forming convex arc and going straight up linearly towards dorsal part. Aedeagus bifid on its apical part; posterior extension with thin serrated apex, on last third; anterior dorsal extension with rounded apex representing one third length of posterior one.</paragraph>
<paragraph id="60678614D107FFB590E3B362FB45F9CB" blockId="23.[189,1208,1544,1570]" box="[189,1208,1544,1570]" pageId="23" pageNumber="189">COLORATION. Specimens entirely black except for thorax ochre-yellow in ventral view.</paragraph>
</subSubSection>
</treatment>
</document>
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