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<document id="AFF710295091A4D6623AF54C12E09D05" ID-DOI="10.5252/geodiversitas2023v45a13" ID-ISSN="1638-9395" ID-Zenodo-Dep="8157175" ID-ZooBank="urn:lsid:zoobank.org:pub:DAE211A9-C735-42E8-A396-26110863CD96" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1689623470392" checkinUser="felipe" docAuthor="Alfonso-Rojas, Andrés, Vanegas, Rubén D., Mariño-Morejón, Elizabeth &amp; Cadena, Edwin-Alberto" docDate="2023" docId="527587BD3655DD2AFF4E3ACD6ACFFA4F" docLanguage="en" docName="Geodiversitas.45.13.377-399.pdf" docOrigin="Geodiversitas 45 (13)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Colombophis portai Hoffstetter &amp; Rage 1977" docType="treatment" docVersion="1" lastPageNumber="382" masterDocId="AE4CFFC53652DD2DFF8A3B756836FF9A" masterDocTitle="Newly discovered fossils provide novel insights on the biology of the South American Miocene snake Colombophis Hoffstetter &amp; Rage, 1977" masterLastPageNumber="399" masterPageNumber="377" pageNumber="382" updateTime="1689703536584" updateUser="valdenar" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="C9C13CE64534E521F1D268BF2BD0FD1C">Newly discovered fossils provide novel insights on the biology of the South American Miocene snake Colombophis Hoffstetter &amp; Rage, 1977</mods:title>
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<mods:namePart id="F97AA22CB0CD3E403690C4404E74EF58">Alfonso-Rojas, Andrés</mods:namePart>
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<mods:namePart id="5F9105F929AD9F2D024B65A176C0D41A">Vanegas, Rubén D.</mods:namePart>
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<mods:namePart id="BCB54EFDF42524370B95DB6672561022">Mariño-Morejón, Elizabeth</mods:namePart>
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<mods:namePart id="BF83B1B480065104203EFF85979BF738">Cadena, Edwin-Alberto</mods:namePart>
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<mods:title id="CAE56788EE0F8185A861BEA7E8D01A3C">Geodiversitas</mods:title>
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<mods:date id="B3FDDB45351D2264394897BE20488B58">2023</mods:date>
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<treatment id="527587BD3655DD2AFF4E3ACD6ACFFA4F" LSID="urn:lsid:plazi:treatment:527587BD3655DD2AFF4E3ACD6ACFFA4F" httpUri="http://treatment.plazi.org/id/527587BD3655DD2AFF4E3ACD6ACFFA4F" lastPageNumber="382" pageId="7" pageNumber="382">
<subSubSection id="92C665203655DD2AFF4E3ACD6AF1FE4B" box="[196,711,438,466]" pageId="7" pageNumber="382" type="nomenclature">
<paragraph id="DA6336AB3655DD2AFF4E3ACD6AF1FE4B" blockId="7.[196,711,438,497]" box="[196,711,438,466]" pageId="7" pageNumber="382">
<taxonomicName id="1DDC4D283655DD2AFF4E3ACD6AF1FE4B" authority="Hoffstetter &amp; Rage, 1977" authorityName="Hoffstetter &amp; Rage" authorityYear="1977" box="[196,711,438,466]" class="Squamata" family="Aniliidae" genus="Colombophis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="382" phylum="Chordata" rank="species" species="portai">
<emphasis id="E8A8EAB93655DD2AFF4E3ACD6996FE4A" bold="true" box="[196,416,438,466]" italics="true" pageId="7" pageNumber="382">Colombophis portai</emphasis>
<bibRefCitation id="BE4D4B5A3655DD2AFE233AC36AF1FE4B" author="HOFFSTETTER R. &amp; RAGE J. C." box="[425,711,438,465]" pageId="7" pageNumber="382" pagination="161 - 190" refId="ref8705" refString="HOFFSTETTER R. &amp; RAGE J. C. 1977. - Le gisement de vertebres miocenes de la Venta (Colombie) et sa faune de serpents. Annales de Paleontologie (Vertebres) 63 (2): 161 - 190." type="journal article" year="1977">Hoffstetter &amp; Rage, 1977</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="92C665203655DD2AFE053AA369CAFE6A" box="[399,508,470,497]" pageId="7" pageNumber="382" type="description">
<paragraph id="DA6336AB3655DD2AFE053AA369CAFE6A" blockId="7.[196,711,438,497]" box="[399,508,470,497]" pageId="7" pageNumber="382">
(
<figureCitation id="42E72A2E3655DD2AFE133AA369C4FE6B" box="[409,498,470,497]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="8.[132,143,1650,1667]" captionStartId-1="9.[132,143,1511,1528]" captionStartId-2="10.[132,143,1793,1810]" captionTargetBox-0="[153,1431,238,1609]" captionTargetBox-1="[152,1430,234,1483]" captionTargetBox-2="[163,1428,215,1759]" captionTargetId-0="figure-159@8.[723,1267,767,1330]" captionTargetId-1="figure-268@9.[620,990,582,1509]" captionTargetId-2="figure-109@10.[701,1255,748,1312]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIG. 3. — Colombophis portai Hoffstetter &amp; Rage, 1977 specimens! A, B, VPPLT-1006; C, VPPLT-0067; D, VPPLT-0068; E, VPPLT-1739; F, VPPLT-1735. Views: 1, anterior;2, dorsal; 3, lateral;4, ventral, 5, posterior views. Scale bars: 10 mm." captionText-1="FIG. 4. — Colombophis portai Hoffstetter &amp; Rage,1977 VPPLT-1740 specimen,which consists of seven associated vertebrae:A, B, mid-trunk vertebra;C, D, midtrunk vertebra; E, F, posterior-trunk vertebra. Views: 1, anterior; 2, dorsal; 3, lateral; 4, ventral; 5, posterior views. Scale bar: 10 mm." captionText-2="FIG. 5. — Colombophis portai Hoffstetter &amp; Rage, 1977 VPPLT-1564 specimen: A, B, nine articulated precloacal vertebrae; C, D, several fragments of precloacal vertebrae and ribs embedded in the rock matrix; E-I, three articulated precloacal vertebrae in dorsal (E, F), anterior (G) and ventral (H, I) views; J-M, isolated vertebra in anterior (J, K) and posterior (L, M) views. Scale bars: A, B, 50 mm; C-N, 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8157181" figureDoi-1="http://doi.org/10.5281/zenodo.8157183" figureDoi-2="http://doi.org/10.5281/zenodo.8157185" httpUri-0="https://zenodo.org/record/8157181/files/figure.png" httpUri-1="https://zenodo.org/record/8157183/files/figure.png" httpUri-2="https://zenodo.org/record/8157185/files/figure.png" pageId="7" pageNumber="382">Figs 3-5</figureCitation>
)
</paragraph>
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<subSubSection id="92C665203655DD2AFF0E397A6ABEFDBD" box="[132,648,527,552]" pageId="7" pageNumber="382" type="reference_group">
<paragraph id="DA6336AB3655DD2AFF0E397A6ABEFDBD" blockId="7.[132,648,527,552]" box="[132,648,527,552]" pageId="7" pageNumber="382">
<taxonomicName id="1DDC4D283655DD2AFF0E397A6AB2FDBD" authority="Hoffstetter &amp; Rage, 1977: 174" authorityName="Hoffstetter &amp; Rage" authorityPageNumber="174" authorityYear="1977" box="[132,644,527,552]" class="Squamata" family="Aniliidae" genus="Colombophis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="382" phylum="Chordata" rank="species" species="portai">
<emphasis id="E8A8EAB93655DD2AFF0E397A6974FDBD" box="[132,322,527,551]" italics="true" pageId="7" pageNumber="382">Colombophis portai</emphasis>
<bibRefCitation id="BE4D4B5A3655DD2AFEC3397A6AB2FDBD" author="HOFFSTETTER R. &amp; RAGE J. C." box="[329,644,527,552]" pageId="7" pageNumber="382" pagination="161 - 190" refId="ref8705" refString="HOFFSTETTER R. &amp; RAGE J. C. 1977. - Le gisement de vertebres miocenes de la Venta (Colombie) et sa faune de serpents. Annales de Paleontologie (Vertebres) 63 (2): 161 - 190." type="journal article" year="1977">Hoffstetter &amp; Rage, 1977: 174</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="92C665203655DD2AFF0E394A6B31FC53" pageId="7" pageNumber="382" type="diagnosis">
<paragraph id="DA6336AB3655DD2AFF0E394A6B31FC53" blockId="7.[131,777,575,970]" pageId="7" pageNumber="382">
<smallCapsWord id="DC85A0773655DD2AFF0E394A68EFFDCF" baselines="592,593" box="[132,217,575,598]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Revised" pageId="7" pageNumber="382">REVISED</smallCapsWord>
<smallCapsWord id="DC85A0773655DD2AFF683931697AFDCF" baselines="593" box="[226,332,580,597]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="diagnosis" pageId="7" pageNumber="382">DIAGNOSIS</smallCapsWord>
. — Fossil snake with pre-cloacal vertebrae medium to large in size (cl =
<emphasis id="E8A8EAB93655DD2AFE23392F698FFDEB" box="[425,441,602,625]" italics="true" pageId="7" pageNumber="382">c.</emphasis>
<quantity id="1D249B4E3655DD2AFE4A392C6A14FDE8" box="[448,546,601,626]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="11.0" metricValueMin="5.0" pageId="7" pageNumber="382" unit="mm" value="8.0" valueMax="11.0" valueMin="5.0">5-11 mm</quantity>
); the neural arch is depressed, with a shallow median notch in the posterior border; neural spine is reduced to a small tubercle with a circular or triangular outline from dorsal view, and is restricted to the posterior end of the neural arch; prezygapophyses are anterolaterally oriented and highly inclined above the horizontal plane, reaching the level of the zygosphene roof; prezygapophyzeal process is short; presence of paracotylar and parazygantral foramina is variable; synapophyses are undivided; haemal keel is broad with a single or divergent apophysis in the posterior end; subcentral foramina are usually placed close to the sagittal plane, usually small but can be enlarged or absent; the centrum length is longer than the neural arch width (cl&gt; naw); the neural arch is longer than width (pr-po&gt; naw) and the length is proportional to its high (pr-po =
<emphasis id="E8A8EAB93655DD2AFD5F38C76AD3FC53" box="[725,741,946,969]" italics="true" pageId="7" pageNumber="382">c.</emphasis>
h).
</paragraph>
</subSubSection>
<subSubSection id="92C665203655DD2AFF0E38946ACFFA4F" pageId="7" pageNumber="382" type="description">
<paragraph id="DA6336AB3655DD2AFF0E38946ACFFA4F" blockId="7.[131,777,993,1493]" pageId="7" pageNumber="382">
<smallCapsWord id="DC85A0773655DD2AFF0E389468DEFC6D" baselines="1010,1011" box="[132,232,993,1016]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Referred" pageId="7" pageNumber="382">REFERRED</smallCapsWord>
<smallCapsWord id="DC85A0773655DD2AFF673892697AFC62" baselines="1012" box="[237,332,999,1016]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="7" pageNumber="382">MATERIAL</smallCapsWord>
. — VPPLT-0067, an anterior-trunk vertebra (
<figureCitation id="42E72A2E3655DD2AFF07388E68E1FB8E" box="[141,215,1019,1044]" captionStart="FIG" captionStartId="8.[132,143,1650,1667]" captionTargetBox="[153,1431,238,1609]" captionTargetId="figure-159@8.[723,1267,767,1330]" captionTargetPageId="8" captionText="FIG. 3. — Colombophis portai Hoffstetter &amp; Rage, 1977 specimens! A, B, VPPLT-1006; C, VPPLT-0067; D, VPPLT-0068; E, VPPLT-1739; F, VPPLT-1735. Views: 1, anterior;2, dorsal; 3, lateral;4, ventral, 5, posterior views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8157181" httpUri="https://zenodo.org/record/8157181/files/figure.png" pageId="7" pageNumber="382">Fig. 3C</figureCitation>
); VPPLT-0068 (
<figureCitation id="42E72A2E3655DD2AFEF2388E69F3FB8E" box="[376,453,1019,1044]" captionStart="FIG" captionStartId="8.[132,143,1650,1667]" captionTargetBox="[153,1431,238,1609]" captionTargetId="figure-159@8.[723,1267,767,1330]" captionTargetPageId="8" captionText="FIG. 3. — Colombophis portai Hoffstetter &amp; Rage, 1977 specimens! A, B, VPPLT-1006; C, VPPLT-0067; D, VPPLT-0068; E, VPPLT-1739; F, VPPLT-1735. Views: 1, anterior;2, dorsal; 3, lateral;4, ventral, 5, posterior views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8157181" httpUri="https://zenodo.org/record/8157181/files/figure.png" pageId="7" pageNumber="382">Fig. 3D</figureCitation>
), an anterior-trunk vertebra; VPPLT-0070, a posterior-trunk vertebra; VPPLT-0430, a mid-trunk vertebra; VPPLT-0799, a mid-trunk vertebra; VPPLT-0845, a fragmentary mid-trunk vertebra; VPPLT-0869, a posterior-trunk vertebra; VPPLT-0871, a fragmentary mid-trunk vertebra; VPPLT-1006, a mid-trunk vertebra (
<figureCitation id="42E72A2E3655DD2AFE6E3FF56A64FB02" box="[484,594,1152,1176]" captionStart="FIG" captionStartId="8.[132,143,1650,1667]" captionTargetBox="[153,1431,238,1609]" captionTargetId="figure-159@8.[723,1267,767,1330]" captionTargetPageId="8" captionText="FIG. 3. — Colombophis portai Hoffstetter &amp; Rage, 1977 specimens! A, B, VPPLT-1006; C, VPPLT-0067; D, VPPLT-0068; E, VPPLT-1739; F, VPPLT-1735. Views: 1, anterior;2, dorsal; 3, lateral;4, ventral, 5, posterior views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8157181" httpUri="https://zenodo.org/record/8157181/files/figure.png" pageId="7" pageNumber="382">Fig. 3A, B</figureCitation>
); VPPLT-1160, a fragmentary posterior-trunk vertebra; VPPLT-1166, a posterior-trunk vertebra; VPPLT-1253, a posterior-trunk vertebra; VPPLT-1551, two associated mid-trunk vertebrae; VPPLT-1564, nine pre-cloacal articulated vertebrae together with seven associated and badly preserved fragments of rock matrix with ribs and vertebral fragments embedded (
<figureCitation id="42E72A2E3655DD2AFE2B3E6B69EFFAAC" box="[417,473,1310,1334]" captionStart="FIG" captionStartId="10.[132,143,1793,1810]" captionTargetBox="[163,1428,215,1759]" captionTargetId="figure-109@10.[701,1255,748,1312]" captionTargetPageId="10" captionText="FIG. 5. — Colombophis portai Hoffstetter &amp; Rage, 1977 VPPLT-1564 specimen: A, B, nine articulated precloacal vertebrae; C, D, several fragments of precloacal vertebrae and ribs embedded in the rock matrix; E-I, three articulated precloacal vertebrae in dorsal (E, F), anterior (G) and ventral (H, I) views; J-M, isolated vertebra in anterior (J, K) and posterior (L, M) views. Scale bars: A, B, 50 mm; C-N, 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8157185" httpUri="https://zenodo.org/record/8157185/files/figure.png" pageId="7" pageNumber="382">Fig. 5</figureCitation>
); VPPLT-1731, a fragmentary mid-trunk vertebra; VPPLT-1734, a mid-trunk vertebra; VPPLT-1735, four mid-trunk vertebrae fragments associated with a posterior-trunk vertebra (
<figureCitation id="42E72A2E3655DD2AFE103E1B69D0FA1C" box="[410,486,1390,1414]" captionStart="FIG" captionStartId="8.[132,143,1650,1667]" captionTargetBox="[153,1431,238,1609]" captionTargetId="figure-159@8.[723,1267,767,1330]" captionTargetPageId="8" captionText="FIG. 3. — Colombophis portai Hoffstetter &amp; Rage, 1977 specimens! A, B, VPPLT-1006; C, VPPLT-0067; D, VPPLT-0068; E, VPPLT-1739; F, VPPLT-1735. Views: 1, anterior;2, dorsal; 3, lateral;4, ventral, 5, posterior views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8157181" httpUri="https://zenodo.org/record/8157181/files/figure.png" pageId="7" pageNumber="382">Fig. 3F</figureCitation>
); VPPLT-1738, a mid-trunk vertebra; VPPLT-1739, an anterior-trunk vertebra (
<figureCitation id="42E72A2E3655DD2AFD0E3EFD6AF8FA3A" box="[644,718,1416,1440]" captionStart="FIG" captionStartId="8.[132,143,1650,1667]" captionTargetBox="[153,1431,238,1609]" captionTargetId="figure-159@8.[723,1267,767,1330]" captionTargetPageId="8" captionText="FIG. 3. — Colombophis portai Hoffstetter &amp; Rage, 1977 specimens! A, B, VPPLT-1006; C, VPPLT-0067; D, VPPLT-0068; E, VPPLT-1739; F, VPPLT-1735. Views: 1, anterior;2, dorsal; 3, lateral;4, ventral, 5, posterior views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8157181" httpUri="https://zenodo.org/record/8157181/files/figure.png" pageId="7" pageNumber="382">Fig. 3E</figureCitation>
); VPPLT-1740, five mid-trunk vertebrae fragments associated with a posterior-trunk vertebrae and a post-cloacal vertebrae (
<figureCitation id="42E72A2E3655DD2AFD3A3EC86ADCFA4F" box="[688,746,1469,1493]" captionStart="FIG" captionStartId="9.[132,143,1511,1528]" captionTargetBox="[152,1430,234,1483]" captionTargetId="figure-268@9.[620,990,582,1509]" captionTargetPageId="9" captionText="FIG. 4. — Colombophis portai Hoffstetter &amp; Rage,1977 VPPLT-1740 specimen,which consists of seven associated vertebrae:A, B, mid-trunk vertebra;C, D, midtrunk vertebra; E, F, posterior-trunk vertebra. Views: 1, anterior; 2, dorsal; 3, lateral; 4, ventral; 5, posterior views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8157183" httpUri="https://zenodo.org/record/8157183/files/figure.png" pageId="7" pageNumber="382">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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