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<document id="936937C8098D01A2C65E93FD29D98D4C" ID-DOI="10.3897/zoologia.37.e36514" ID-ISSN="1984-4689" ID-Zenodo-Dep="13175881" ID-ZooBank="E03C0430-68C6-449B-A0AF-9FB0968FB38C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722590035522" checkinUser="felipe" docAuthor="Cláudio, Vinícius C., Barbosa, Gedimar P., Rocha, Vlamir J. &amp; Rassy, Ricardo Moratelli Fabrício B." docDate="2020" docId="03FEB34E8906FFF982C9ADF18779FADA" docLanguage="en" docName="Zoologia.37.e36514.pdf" docOrigin="Zoologia (e 36514) 37" docSource="http://dx.doi.org/10.3897/zoologia.37.e36514" docStyle="DocumentStyle:9D87517C92C8E430512793D922E147D4.1:Zoologia.2017-2020.journal_article" docStyleId="9D87517C92C8E430512793D922E147D4" docStyleName="Zoologia.2017-2020.journal_article" docStyleVersion="1" docTitle="Artibeus lituratus" docType="treatment" docVersion="2" lastPageNumber="14" masterDocId="FFC7CB36890BFFF48168AD378369FFE4" masterDocTitle="The bat fauna (Mammalia: Chiroptera) of Carlos Botelho State Park, Atlantic Forest of Southeastern Brazil, including new distribution records for the state of São Paulo" masterLastPageNumber="32" masterPageNumber="1" pageNumber="14" updateTime="1722627286571" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="924CB6D26CD3598D1C9B15890092744C">The bat fauna (Mammalia: Chiroptera) of Carlos Botelho State Park, Atlantic Forest of Southeastern Brazil, including new distribution records for the state of São Paulo</mods:title>
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<mods:namePart id="66ACBFCA37B779A39B7932561DDCA9F6">Cláudio, Vinícius C.</mods:namePart>
<mods:affiliation id="B2D9F3ADCD16FE3E50A2DBBE412F566F">Centro de Ciências Biológicas e da Saúde, Universidade Federal de São Carlos. 13565 - 905 São Carlos, SP, Brazil. &amp; Fundação Parque Zoológico de São Paulo. 04301 - 905 São Paulo, SP, Brazil. &amp; Instituto de Biologia, Programa de Pós-Graduação em Biodiversidade e Biologia Evolutiva, Universidade Federal do</mods:affiliation>
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<mods:namePart id="6D6011C83E2203B32AF4ADC1016E9A57">Barbosa, Gedimar P.</mods:namePart>
<mods:affiliation id="3F01C082AEF1E2536F4562134D352F79">Centro de Ciências Biológicas e da Saúde, Universidade Federal de São Carlos. 13565 - 905 São Carlos, SP, Brazil. &amp; Fundação Parque Zoológico de São Paulo. 04301 - 905 São Paulo, SP, Brazil.</mods:affiliation>
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<mods:namePart id="A2B6266F72950DC07A534D882132EE92">Rocha, Vlamir J.</mods:namePart>
<mods:affiliation id="9A942656F57D99969FA2E66E6C5B811F">Centro de Ciências Agrárias, Universidade Federal de São Carlos. 13600 - 970 Araras, SP, Brazil.</mods:affiliation>
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<mods:date id="064A6AC459F6076A01AB6FE3871192D2">2020</mods:date>
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<subSubSection id="C34D51D38906FFF982C9ADF18674FF05" box="[929,1309,198,225]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BE802588906FFF982C9ADF18674FF05" blockId="13.[929,1309,198,225]" box="[929,1309,198,225]" pageId="13" pageNumber="14">
<heading id="D0A0B5348906FFF982C9ADF18674FF05" box="[929,1309,198,225]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="5">
<taxonomicName id="4C5779DB8906FFF982C9ADF18674FF05" ID-CoL="7R5ZG" authority="(Olfers, 1818)" baseAuthorityName="Olfers" baseAuthorityYear="1818" box="[929,1309,198,225]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="lituratus">
<emphasis id="B923DE4A8906FFF982C9ADF18700FF05" box="[929,1129,198,225]" italics="true" pageId="13" pageNumber="14">Artibeus lituratus</emphasis>
(Olfers, 1818)
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<subSubSection id="C34D51D38906FFF98556ADDB8779FADA" pageId="13" pageNumber="14" type="description">
<paragraph id="8BE802588906FFF98556ADDB8716FEE4" blockId="13.[1086,1151,236,256]" box="[1086,1151,236,256]" pageId="13" pageNumber="14">
<figureCitation id="136C1EDD8906FFF98556ADDB8716FEE4" box="[1086,1151,236,256]" captionStart="Figures 1829" captionStartId="14.[159,226,1708,1728]" captionTargetBox="[236,1359,197,1694]" captionTargetId="figure-62@14.[236,1359,197,1694]" captionTargetPageId="14" captionText="Figures 1829. Bat species captured in Carlos Botelho State Park, São Paulo state, Brazil: (18) Artibeus fimbriatus; (19) Artibeus lituratus; (20) Artibeus obscurus; (21)Dermanura cinerea; (22) Platyrrhinus lineatus; (23)Platyrrhinus recifinus; (24) Pygoderma bilabiatum; (25)Sturnira lilium; (26) Sturnira tildae; (27) Vampyressa pusilla; (28) Cynomops abrasus; (29) Molossops neglectus." pageId="13" pageNumber="14">Fig. 19</figureCitation>
</paragraph>
<paragraph id="8BE802588906FFF9820DAC21874FFC33" blockId="13.[807,1430,277,1342]" pageId="13" pageNumber="14">
Taxonomy. The distinction between
<taxonomicName id="4C5779DB8906FFF985B4AC218621FECC" baseAuthorityName="Olfers" baseAuthorityYear="1818" box="[1244,1352,277,297]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="lituratus">
<emphasis id="B923DE4A8906FFF985B4AC218621FECC" box="[1244,1352,277,297]" italics="true" pageId="13" pageNumber="14">A. lituratus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5779DB8906FFF984E8AC2180E2FEA2" authorityName="Gray" authorityYear="1838" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="fimbriatus">
<emphasis id="B923DE4A8906FFF984E8AC2180E2FEA2" italics="true" pageId="13" pageNumber="14">A. fimbriatus</emphasis>
</taxonomicName>
is discussed above.
<taxonomicName id="4C5779DB8906FFF9850CAC0387BCFEA2" baseAuthorityName="Olfers" baseAuthorityYear="1818" box="[1124,1237,307,327]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="lituratus">
<emphasis id="B923DE4A8906FFF9850CAC0387BCFEA2" box="[1124,1237,307,327]" italics="true" pageId="13" pageNumber="14">A. lituratus</emphasis>
</taxonomicName>
can be externally distinguished from
<taxonomicName id="4C5779DB8906FFF98287AC66871DFE80" authorityName="Nogueira et al." authorityYear="2014" baseAuthorityName="Spix" baseAuthorityYear="1832" box="[1007,1140,337,356]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="planirostris">
<emphasis id="B923DE4A8906FFF98287AC66871DFE80" box="[1007,1140,337,356]" italics="true" pageId="13" pageNumber="14">A. planirostris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5779DB8906FFF985C1AC66867FFE80" baseAuthorityName="Schinz" baseAuthorityYear="1821" box="[1193,1302,337,356]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="obscurus">
<emphasis id="B923DE4A8906FFF985C1AC66867FFE80" box="[1193,1302,337,356]" italics="true" pageId="13" pageNumber="14">A. obscurus</emphasis>
</taxonomicName>
by its larger size; well-marked facial stripes (poorly marked to absent on the other two); lack of ventral frosting (present on the other two species); base of the noseleaf attached to the upper lip, while is always separated in
<taxonomicName id="4C5779DB8906FFF98287ACFE871BFE38" authorityName="Nogueira et al." authorityYear="2014" baseAuthorityName="Spix" baseAuthorityYear="1832" box="[1007,1138,457,476]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="planirostris">
<emphasis id="B923DE4A8906FFF98287ACFE871BFE38" box="[1007,1138,457,476]" italics="true" pageId="13" pageNumber="14">A. planirostris</emphasis>
</taxonomicName>
and in some
<taxonomicName id="4C5779DB8906FFF98590ACFE860AFE38" baseAuthorityName="Schinz" baseAuthorityYear="1821" box="[1272,1379,457,476]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="obscurus">
<emphasis id="B923DE4A8906FFF98590ACFE860AFE38" box="[1272,1379,457,476]" italics="true" pageId="13" pageNumber="14">A. obscurus</emphasis>
</taxonomicName>
; and dorsal side of the uropatagium densely furred, which is almost naked in
<taxonomicName id="4C5779DB8906FFF982E3AF328778FDF3" authorityName="Nogueira et al." authorityYear="2014" baseAuthorityName="Spix" baseAuthorityYear="1832" box="[907,1041,516,536]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="planirostris">
<emphasis id="B923DE4A8906FFF982E3AF328778FDF3" box="[907,1041,516,536]" italics="true" pageId="13" pageNumber="14">A. planirostris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5779DB8906FFF98520AF3287DFFDF3" baseAuthorityName="Schinz" baseAuthorityYear="1821" box="[1096,1206,516,536]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="obscurus">
<emphasis id="B923DE4A8906FFF98520AF3287DFFDF3" box="[1096,1206,516,536]" italics="true" pageId="13" pageNumber="14">A. obscurus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C5779DB8906FFF985ADAF338601FDF3" baseAuthorityName="Olfers" baseAuthorityYear="1818" box="[1221,1384,516,535]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="lituratus">
<emphasis id="B923DE4A8906FFF985ADAF338601FDF3" box="[1221,1384,516,535]" italics="true" pageId="13" pageNumber="14">Artibeus lituratus</emphasis>
</taxonomicName>
can also be distinguished from
<taxonomicName id="4C5779DB8906FFF9852CAF1587A5FDD1" authorityName="Nogueira et al." authorityYear="2014" baseAuthorityName="Spix" baseAuthorityYear="1832" box="[1092,1228,546,565]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="planirostris">
<emphasis id="B923DE4A8906FFF9852CAF1587A5FDD1" box="[1092,1228,546,565]" italics="true" pageId="13" pageNumber="14">A. planirostris</emphasis>
</taxonomicName>
by the presence of dense fur on the dorsal side of the forearm (almost naked in
<taxonomicName id="4C5779DB8906FFF984E8AF7780F8FD95" authorityName="Nogueira et al." authorityYear="2014" baseAuthorityName="Spix" baseAuthorityYear="1832" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="planirostris">
<emphasis id="B923DE4A8906FFF984E8AF7780F8FD95" italics="true" pageId="13" pageNumber="14">A. planirostris</emphasis>
</taxonomicName>
). Additionally,
<taxonomicName id="4C5779DB8906FFF98547AF6987F0FD95" baseAuthorityName="Olfers" baseAuthorityYear="1818" box="[1071,1177,606,625]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="lituratus">
<emphasis id="B923DE4A8906FFF98547AF6987F0FD95" box="[1071,1177,606,625]" italics="true" pageId="13" pageNumber="14">A. lituratus</emphasis>
</taxonomicName>
has short dorsal fur (
<quantity id="4CAFAFBD8906FFF9841AAF69803CFD6B" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="13" pageNumber="14" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 mm</quantity>
), which is longer in
<taxonomicName id="4C5779DB8906FFF9854CAF4B87F9FD6B" baseAuthorityName="Schinz" baseAuthorityYear="1821" box="[1060,1168,636,655]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="obscurus">
<emphasis id="B923DE4A8906FFF9854CAF4B87F9FD6B" box="[1060,1168,636,655]" italics="true" pageId="13" pageNumber="14">A. obscurus</emphasis>
</taxonomicName>
(
<quantity id="4CAFAFBD8906FFF985F6AF4B8794FD6B" box="[1182,1277,636,655]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="13" pageNumber="14" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
<bibRefCitation id="EFC67FA98906FFF9847EAF4B80F8FD49" author="Koepcke J &amp; Kraft R" pageId="13" pageNumber="14" pagination="75 - 84" refId="ref25283" refString="Koepcke J, Kraft R (1984) Cranial and external characters of the larger fruit bats of the genus Artibeus from Amazonian Peru (Chiroptera: Phyllostomidae). Spixiana 7: 75 - 84." type="journal article" year="1984">Koepcke and Kraft 1984</bibRefCitation>
,
<bibRefCitation id="EFC67FA98906FFF982F1AFAE8705FD49" author="Marques-Aguiar SA" box="[921,1132,665,685]" pageId="13" pageNumber="14" pagination="1 - 83" refId="ref26013" refString="Marques-Aguiar SA (1994) Systematic review of the large species of Artibeus Leach, 1821 (Mammalia: Chiroptera) with some phylogenetic inferences. Boletim do Museu Paraense Emilio Goeldi 10: 1 - 83." type="journal article" year="1994">Marques-Aguiar 1994</bibRefCitation>
,
<bibRefCitation id="EFC67FA98906FFF9851CAFAD87CEFD49" author="Marques-Aguiar SA" box="[1140,1191,666,685]" pageId="13" pageNumber="14" pagination="301 - 321" refId="ref26051" refString="Marques-Aguiar SA (2008) Genus Artibeus Leach, 1821. In: Gardner AL (Ed.) Mammals of South America. The University of Chicago Press, Chicago, vol. 1, 301 - 321." type="journal article" year="2008">2008</bibRefCitation>
,
<bibRefCitation id="EFC67FA98906FFF985C7AFAD86FBFD49" author="Haynes MA &amp; Lee TE Jr" box="[1199,1426,666,685]" pageId="13" pageNumber="14" pagination="1 - 5" refId="ref25050" refString="Haynes MA, Lee TE Jr (2004) Artibeus obscurus. Mammalian Species 752: 1 - 5. https: // doi. org / 10.1644 / 752" type="journal article" year="2004">Haynes and Lee Jr 2004</bibRefCitation>
,
<bibRefCitation id="EFC67FA98906FFF9824FAF8080F5FD2E" author="Hollis L" box="[807,924,695,714]" pageId="13" pageNumber="14" pagination="1 - 6" refId="ref25080" refString="Hollis L (2005) Artibeus planirostris. Mammalian Species 775: 1 - 6. https: // doi. org / 10.1644 / 1545 - 1410 (2005) 775 [0001: AP] 2.0. CO; 2" type="journal article" year="2005">Hollis 2005</bibRefCitation>
,
<bibRefCitation id="EFC67FA98906FFF982CFAF8F8749FD2F" author="Zortea M" box="[935,1056,696,715]" pageId="13" pageNumber="14" pagination="107 - 128" refId="ref29250" refString="Zortea M (2007) Subfamilia Stenodermatinae. In: Reis NR, Peracchi AL, Pedro WA, Lima IP (Eds) Morcegos do Brasil. Editora UEL, Londrina, 107 - 128." type="book chapter" year="2007">Zortéa 2007</bibRefCitation>
,
<bibRefCitation id="EFC67FA98906FFF98544AF808670FD2F" author="Dias D &amp; Peracchi AL" box="[1068,1305,695,715]" pageId="13" pageNumber="14" pagination="333 - 369" refId="ref23859" refString="Dias D, Peracchi AL (2008) Quiropteros da Reserva Biologica do Tingua, estado do Rio de Janeiro, sudeste do Brasil (Mammalia: Chiroptera). Revista Brasileira de Zoologia 25: 333 - 369. https: // doi. org / 10.1590 / S 0101 - 81752008000200023" type="journal article" year="2008">Dias and Peracchi 2008</bibRefCitation>
,
<bibRefCitation id="EFC67FA98906FFF9844CAF8080AAFD0C" author="Araujo P &amp; Langguth A" pageId="13" pageNumber="14" pagination="715 - 722" refId="ref22522" refString="Araujo P, Langguth A (2010) Caracteres distintivos das quatro especies de grandes Artibeus (Phyllostomidae) de Paraiba e Pernambuco, Brasil. Chiroptera Neotropical 16: 715 - 722." type="journal article" year="2010">Araújo and Langguth 2010</bibRefCitation>
). The PECB specimens (ZSP 015, 056; see
<tableCitation id="C6D537E38906FFF98439AFE286FFFD0C" box="[1361,1430,725,744]" captionStart="Table 6" captionStartId="10.[159,209,905,925]" captionText="Table 6. Selected measurements (mm) and weigth (g) for specimens of Carolliinae and Stenodermatinae from PECB, São Paulo state, Brazil. See Material and Methods for description of measurements." pageId="13" pageNumber="14">Table 6</tableCitation>
for measurements) identified as
<taxonomicName id="4C5779DB8906FFF98533AFC387A8FCE2" baseAuthorityName="Olfers" baseAuthorityYear="1818" box="[1115,1217,755,775]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="lituratus">
<emphasis id="B923DE4A8906FFF98533AFC387A8FCE2" box="[1115,1217,755,775]" italics="true" pageId="13" pageNumber="14">A. lituratus</emphasis>
</taxonomicName>
have bicolored dorsal fur, with grayish brown basal band and chocolate brown tips; ventral fur weakly bicolored with pale brown basis and grayish tips, frosting absent. Uropatagium and forearm are densely furred, the facial stripes are bright and well markedwell-marked, the edge of ears and tragus are yellowish, the dorsal fur ranges from
<quantity id="4CAFAFBD8906FFF9820AAE90809DFC5E" box="[866,1012,935,954]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="8.9" metricValueMin="7.7" pageId="13" pageNumber="14" unit="mm" value="8.3" valueMax="8.9" valueMin="7.7">7.7 to 8.9 mm</quantity>
in lenghtlength and the noseleaf base is attached to the upper lip.
</paragraph>
<paragraph id="8BE802588906FFF9820DAED58750FBAB" blockId="13.[807,1430,277,1342]" pageId="13" pageNumber="14">
Distribution. In
<collectingCountry id="F34042C88906FFF98568AED58750FC11" box="[1024,1081,994,1013]" name="Brazil" pageId="13" pageNumber="14">Brazil</collectingCountry>
, the species is recorded in all biomes and states (
<bibRefCitation id="EFC67FA98906FFF982FAA937872FFBF7" author="Tavares VC &amp; Gregorin R &amp; Peracchi AL" box="[914,1094,1024,1043]" pageId="13" pageNumber="14" pagination="25 - 58" refId="ref28526" refString="Tavares VC; Gregorin R, Peracchi AL (2008) Diversidade de morcegos no Brasil: lista atualizada com comentarios sobre distribuicao e taxonomia. In: Pacheco SM, Marques RV, Esberard CEL (Eds) Morcegos no Brasil: Biologia, Sistematica, Ecologia e Conservacao. Armazem Digital Comunicacao Ltda, Porto Alegre, 25 - 58." type="book chapter" year="2008">Tavares et al. 2008</bibRefCitation>
,
<bibRefCitation id="EFC67FA98906FFF98527A9378788FBF7" author="Reis NR &amp; Peracchi AL &amp; Batista CB &amp; Lima IP &amp; Pereira AD" box="[1103,1249,1024,1043]" pageId="13" pageNumber="14" refId="ref27752" refString="Reis NR, Peracchi AL, Batista CB, Lima IP, Pereira AD (2017) Historia Natural dos Morcegos Brasileiros: Chave de Identificacao de Especies. Technical Books, Rio de Janeiro, 416 pp." type="book" year="2017">Reis et al. 2017</bibRefCitation>
). In
<collectingRegion id="4993CCBA8906FFF98462A9378602FBF7" box="[1290,1387,1024,1043]" country="Brazil" name="Sao Paulo" pageId="13" pageNumber="14">São Paulo</collectingRegion>
, the species is also widely distributed, occurring in all the vegetational formations (
<bibRefCitation id="EFC67FA98906FFF982C9A90B8746FBAB" author="Garbino GST" box="[929,1071,1084,1103]" pageId="13" pageNumber="14" pagination="43 - 128" refId="ref24390" refString="Garbino GST (2016) Research on bats (Chiroptera) from the state of Sao Paulo, southeastern Brazil: annotated species list and bibliographic review. Arquivos de Zoologia (Sao Paulo) 47: 43 - 128." type="journal article" year="2016">Garbino 2016</bibRefCitation>
).
</paragraph>
<paragraph id="8BE802588906FFF9820DA96D8779FADA" blockId="13.[807,1430,277,1342]" pageId="13" pageNumber="14">
Field observations. We captured
<specimenCount id="9D51C9D18906FFF985C5A96D865FFB89" box="[1197,1334,1114,1133]" count="10" pageId="13" pageNumber="14" type="generic">10 specimens</specimenCount>
, seven of which (
<specimenCount id="9D51C9D18906FFF9821CA94F80D6FB6E" box="[884,959,1143,1163]" count="4" pageId="13" pageNumber="14" type="male">4 males</specimenCount>
and
<specimenCount id="9D51C9D18906FFF98287A94F8725FB6E" box="[1007,1100,1143,1163]" count="3" pageId="13" pageNumber="14" type="female">3 females</specimenCount>
) were taken in ground-level mist-nets in sampling sites M6, M18, M19, M21, M22 and M32; and three (
<specimenCount id="9D51C9D18906FFF98203A98480C5FB22" box="[875,940,1203,1222]" count="1" pageId="13" pageNumber="14" type="male">1 male</specimenCount>
and
<specimenCount id="9D51C9D18906FFF982B6A9848755FB22" box="[990,1084,1203,1222]" count="2" pageId="13" pageNumber="14" type="female">2 females</specimenCount>
) in mist-nets elevated
<quantity id="4CAFAFBD8906FFF98476A984862DFB23" box="[1310,1348,1203,1223]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="14" unit="m" value="8.0">8 m</quantity>
in sampling sites M15, M20 and M24 (Appendix 1). Captures occurred in November, January, February, March, May and September. We netted lactating females in January, March and November; and juveniles in September.
</paragraph>
</subSubSection>
</treatment>
</document>