273 lines
31 KiB
XML
273 lines
31 KiB
XML
<document id="E1B2883E9D486B7710216270B7E600C3" ID-CLB-Dataset="8143" ID-DOI="10.2478/aemnp-2019-0020" ID-GBIF-Dataset="a03ecce2-b526-4f94-aa65-620016956099" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488918" ID-ZooBank="9F309FCC-A2ED-47B9-BC37-D0C4A3B482E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612219281920" checkinUser="felipe" docAuthor="Fikáček, Martin & Liu, Hsing-Che" docDate="2019" docId="03F387C57A3FFFAFFEA902163141B5FC" docLanguage="en" docName="ActaEntMusNatPra.59.1.229-252.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 59 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Thysanarthria persica Fikáček & Liu 2019, sp. nov." docType="treatment" docVersion="10" lastPageNumber="246" masterDocId="FFCAFFBD7A2EFFBEFF8901283431B709" masterDocTitle="A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)" masterLastPageNumber="252" masterPageNumber="229" pageNumber="246" updateTime="1698926077238" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="5255DE89019379D876BE1875633CFCAB">A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)</mods:title>
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<mods:name id="9DAB4A7AFEA3AEB7E214EAF55FE7D188" type="personal">
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<mods:roleTerm id="82F06A5C6B05BA269F5C2E9F82D30141">Author</mods:roleTerm>
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<mods:namePart id="EAE3C3EF07BA52F8E8ACE81CB6079087">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="E336C5E9BE9722DE85BE08E9029A49B7">Department of Entomology, National Museum, Cirkusová 1740, CZ- 19100 Praha 9 – Horní Počernice, Czech Republic & Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 12844 Praha 2, Czech Republic</mods:affiliation>
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<mods:nameIdentifier id="AF2632A0C96B8E82E9F9D6F648AD6A4F" type="email">mfikacek@gmail.com</mods:nameIdentifier>
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<mods:namePart id="31E2125FA248E900F6E862F2BD9D1EC8">Liu, Hsing-Che</mods:namePart>
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<mods:affiliation id="1AA4F5FD79FAC4B302EC48F9F1CB4A05">Department of Environmental Engineering and Management, Chaoyang University of Technology, No. 168, Jifeng E. Rd., Taichung City 413, Taiwan</mods:affiliation>
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<mods:nameIdentifier id="A96210B62569DB7AC47CBF7797C472F1" type="email">td965771@gmail.com</mods:nameIdentifier>
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<mods:title id="AAC2C14BD599FF50B11BE78354F632A5">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="08947861E5E5C83F2E8CE0292F07754C">2019</mods:date>
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<mods:number id="070B52B0BC1116B5CA6CB68FF9B8A5A3">2019-06-20</mods:number>
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<mods:number id="5693417003668D622BEB5DA1DFE64A38">59</mods:number>
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<treatment id="03F387C57A3FFFAFFEA902163141B5FC" ID-DOI="http://doi.org/10.5281/zenodo.4488719" ID-GBIF-Taxon="177423411" ID-Zenodo-Dep="4488719" LSID="urn:lsid:plazi:treatment:03F387C57A3FFFAFFEA902163141B5FC" httpUri="http://treatment.plazi.org/id/03F387C57A3FFFAFFEA902163141B5FC" lastPageNumber="246" pageId="17" pageNumber="246">
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<subSubSection id="C34065587A3FFFAFFEA90216365EB45C" box="[288,623,830,853]" pageId="17" pageNumber="246" type="nomenclature">
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<paragraph id="8BE536D37A3FFFAFFEA90216365EB45C" blockId="17.[288,623,830,853]" box="[288,623,830,853]" pageId="17" pageNumber="246">
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<heading id="D0AD81BF7A3FFFAFFEA90216365EB45C" box="[288,623,830,853]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="246" reason="3">
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<emphasis id="B92EEAC17A3FFFAFFEA90216365EB45C" bold="true" box="[288,623,830,853]" pageId="17" pageNumber="246">
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<taxonomicName id="4C5A4D507A3FFFAFFEA902163625B45C" ID-CoL="8YBVV" authority="Fikáček & Liu, 2019" authorityName="Fikáček & Liu" authorityYear="2019" box="[288,532,830,853]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="persica" status="sp. nov.">
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<emphasis id="B92EEAC17A3FFFAFFEA902163625B45C" bold="true" box="[288,532,830,853]" italics="true" pageId="17" pageNumber="246">Thysanarthria persica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21D57BA7A3FFFAFFD930216365EB45C" box="[538,623,830,853]" pageId="17" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C34065587A3FFFAFFEE502743612B466" box="[364,547,860,879]" pageId="17" pageNumber="246" type="description">
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<paragraph id="8BE536D37A3FFFAFFEE502743612B466" blockId="17.[151,761,860,1195]" box="[364,547,860,879]" pageId="17" pageNumber="246">
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(
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||
<figureCitation id="13612A567A3FFFAFFEFB02743587B466" box="[370,438,860,879]" captionStart="Fig" captionStartId="2.[151,182,2004,2023]" captionTargetBox="[165,1409,612,1979]" captionTargetId="figure-13@2.[151,1437,604,1996]" captionTargetPageId="2" captionText="Fig. 1. Habitus of the Thysanarthria species and related Old Word Chaetarthriini.A – Thysanarthria bengalensis Hebauer, 2001, holotype; B – T. persica sp. nov., paratype; C – T. siamensis Hebauer, 2001, holotype; D – T. hongsonensis Hebauer, 2001, holotype; E – European Chaetarthria: C. seminulum (Herbst, 1797); F – large Old World Chaetarthria: C. nigerrima (Blackburn, 1891)." figureDoi="http://doi.org/10.5281/zenodo.4488920" httpUri="https://zenodo.org/record/4488920/files/figure.png" pageId="17" pageNumber="246">Figs 1B</figureCitation>
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,
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<figureCitation id="13612A567A3FFFAFFE49027435E5B466" box="[448,468,860,879]" captionStart="Fig" captionStartId="19.[151,182,2009,2028]" captionTargetBox="[159,1399,787,1981]" captionTargetId="figure-14@19.[151,1436,777,1989]" captionTargetPageId="19" captionText="Fig. 10. Male genitalia of Thysanarthria species, holotypes.A–E – T. persica sp. nov. (A–C – aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. wadicola sp. nov. (F–H – aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488944" httpUri="https://zenodo.org/record/4488944/files/figure.png" pageId="17" pageNumber="246">10</figureCitation>
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A–E,
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<figureCitation id="13612A567A3FFFAFFD8E0274362DB466" box="[519,540,860,879]" captionStart="Fig" captionStartId="20.[151,182,1949,1968]" captionTargetBox="[151,1436,401,1931]" captionTargetId="figure-13@20.[151,1437,401,1931]" captionTargetPageId="20" captionText="Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu." figureDoi="http://doi.org/10.5281/zenodo.4488946" httpUri="https://zenodo.org/record/4488946/files/figure.png" pageId="17" pageNumber="246">11</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C34065587A3FFFAFFF1E02563532B4A5" pageId="17" pageNumber="246" type="reference_group">
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<paragraph id="8BE536D37A3FFFAFFF1E02563532B4A5" blockId="17.[151,761,860,1195]" pageId="17" pageNumber="246">
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<treatmentCitationGroup id="AB4A11FD7A3FFFAFFF1E02563532B4A5" pageId="17" pageNumber="246">
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<taxonomicName id="4C5A4D507A3FFFAFFF1E02563545B498" authorityName=": HANSEN" authorityYear="1991" box="[151,372,894,913]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="sulcata">
|
||
<emphasis id="B92EEAC17A3FFFAFFF1E02563520B498" box="[151,273,894,913]" italics="true" pageId="17" pageNumber="246">Thysanarthria</emphasis>
|
||
cf.
|
||
<emphasis id="B92EEAC17A3FFFAFFEBF02563545B498" box="[310,372,894,913]" italics="true" pageId="17" pageNumber="246">sulcata</emphasis>
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||
</taxonomicName>
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||
(part):
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||
<treatmentCitation id="0AFB10C27A3FFFAFFE3E025636A7B498" author="FIKACEK M. & GENTILI E. & SHORT A. E. Z." box="[439,662,894,913]" page="139" pageId="17" pageNumber="246" year="2010">
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<bibRefCitation id="EFCB4B227A3FFFAFFE3E025636A7B498" author="FIKACEK" box="[439,662,894,913]" etAl="et al." firstAuthor="FIKACEK" page="139" pageId="17" pageNumber="246" pagination="135 - 165" refId="ref17868" refString="FIKACEK M., GENTILI E. & SHORT A. E. Z. 2010: Order Coleoptera, family Hydrophilidae. Pp. 135 - 165. In: HARTEN A. VAN (ed.): Arthropod Fauna of the United Arab Emirates. Volume 3. Al Ummah Printing, Publishing, Distribution and Advertising, Abu Dhabi, 700 pp." type="book chapter" year="2010">
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<subScript id="17DE34967A3FFFAFFE3E025635CAB499" attach="right" box="[439,507,894,913]" fontSize="6" pageId="17" pageNumber="246">FIKÁČEK</subScript>
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||
et al. (2010: 139
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||
</bibRefCitation>
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||
</treatmentCitation>
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||
; misidentification).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
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<subSubSection id="C34065587A3FFFAFFF1E02EB354DB3A2" pageId="17" pageNumber="246" type="materials_examined">
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<paragraph id="8BE536D37A3FFFAFFF1E02EB354DB3A2" blockId="17.[151,761,860,1195]" pageId="17" pageNumber="246">
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<emphasis id="B92EEAC17A3FFFAFFF1E02EB3529B4DF" bold="true" box="[151,280,963,982]" pageId="17" pageNumber="246">Type material.</emphasis>
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<materialsCitation id="3B323C8E7A3FFFAFFE9502EB36ABB302" ID-GBIF-Occurrence="3756360323" collectingDate="1973-04-10" collectionCode="NMPC" country="Iran" location="Baluchistan" municipality="Iran" pageId="17" pageNumber="246" specimenCount="1" specimenCount-male="1" stateProvince="SISTAN AND BALUCHESTAN" typeStatus="holotype">
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||
<typeStatus id="54E188717A3FFFAFFE9502EB355FB4DD" box="[284,366,963,982]" pageId="17" pageNumber="246" type="holotype">HOLOTYPE</typeStatus>
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:
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<specimenCount id="9D5CFD5A7A3FFFAFFEFE029435B6B4DC" box="[375,391,956,981]" pageId="17" pageNumber="246" type="male">♂</specimenCount>
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(
|
||
<collectionCode id="ED4BAE167A3FFFAFFE1802EB35E0B4DF" box="[401,465,963,982]" country="Czech Republic" name="National Museum Prague" pageId="17" pageNumber="246">NMPC</collectionCode>
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),
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<emphasis id="B92EEAC17A3FFFAFFE5702EB3667B4DD" bold="true" box="[478,598,963,982]" pageId="17" pageNumber="246">
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<collectingCountry id="F34D76437A3FFFAFFE5702EB3626B4DF" box="[478,535,963,982]" name="Iran" pageId="17" pageNumber="246">IRAN</collectingCountry>
|
||
:
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<collectingRegion id="499EF8317A3FFFAFFD9702EB3667B4DD" box="[542,598,963,982]" pageId="17" pageNumber="246">SISTAN</collectingRegion>
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</emphasis>
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AND
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<emphasis id="B92EEAC17A3FFFAFFDF402EB36C9B4DF" bold="true" box="[637,760,963,982]" pageId="17" pageNumber="246">BALUCHESTAN:</emphasis>
|
||
<collectingMunicipality id="6B81ACA97A3FFFAFFF1E02F6348AB4F8" box="[151,187,990,1009]" pageId="17" pageNumber="246">Iran</collectingMunicipality>
|
||
,
|
||
<location id="8E8560087A3FFFAFFF4A02F6351BB4F8" LSID="urn:lsid:plazi:treatment:03F387C57A3FFFAFFEA902163141B5FC:8E8560087A3FFFAFFF4A02F6351BB4F8" box="[195,298,990,1009]" country="Iran" municipality="Iran" name="Baluchistan" pageId="17" pageNumber="246" stateProvince="SISTAN AND BALUCHESTAN">Baluchistan</location>
|
||
,
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<locationDeviation id="9A572FF17A3FFFAFFEB802F6363AB4F8" box="[305,523,990,1009]" pageId="17" pageNumber="246">
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<quantity id="4CA29B367A3FFFAFFEB802F63557B4F8" box="[305,358,990,1009]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="17" pageNumber="246" unit="km" value="16.0">16 km</quantity>
|
||
SE of Tange-Sarhe
|
||
</locationDeviation>
|
||
,
|
||
<locationDeviation id="9A572FF17A3FFFAFFD9B02F636C4B4F8" box="[530,757,990,1009]" pageId="17" pageNumber="246">
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<quantity id="4CA29B367A3FFFAFFD9B02F63678B4F8" box="[530,585,990,1009]" metricMagnitude="4" metricUnit="m" metricValue="6.1" pageId="17" pageNumber="246" unit="km" value="61.0">61 km</quantity>
|
||
, NNW of Nik-shahr
|
||
</locationDeviation>
|
||
,
|
||
<date id="FFE410137A3FFFAFFF1E02D034C0B302" box="[151,241,1016,1035]" pageId="17" pageNumber="246" value="1973-04-10">
|
||
<collectingDate id="EFA0E9FB7A3FFFAFFF1E02D034C0B302" box="[151,241,1016,1035]" pageId="17" pageNumber="246" value="1973-04-10">10.iv.1973</collectingDate>
|
||
</date>
|
||
, Expedition National Museum Prague,locality 154.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B323C8E7A3FFFAFFD1502D03624B32F" ID-GBIF-Occurrence="3756360349" pageId="17" pageNumber="246" typeStatus="paratype">
|
||
<typeStatus id="54E188717A3FFFAFFD1502D036C3B300" box="[668,754,1016,1035]" pageId="17" pageNumber="246" type="paratype">PARATYPES</typeStatus>
|
||
:
|
||
<specimenCount id="9D5CFD5A7A3FFFAFFF1E053B34E4B32F" box="[151,213,1043,1062]" pageId="17" pageNumber="246" type="generic">4 spec.</specimenCount>
|
||
(
|
||
<collectionCode id="ED4BAE167A3FFFAFFF56053B3511B32F" box="[223,288,1043,1062]" country="Czech Republic" name="National Museum Prague" pageId="17" pageNumber="246">NMPC</collectionCode>
|
||
), same data as the
|
||
<typeStatus id="54E188717A3FFFAFFE4D053B3620B32F" box="[452,529,1043,1062]" pageId="17" pageNumber="246" type="holotype">holotype</typeStatus>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B323C8E7A3FFFAFFD93053B3616B352" ID-GBIF-Occurrence="3756360312" pageId="17" pageNumber="246">
|
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<emphasis id="B92EEAC17A3FFFAFFD93053B367DB32F" bold="true" box="[538,588,1043,1062]" pageId="17" pageNumber="246">
|
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<collectingRegion id="499EF8317A3FFFAFFD93053B3674B32D" box="[538,581,1043,1062]" country="Iran" name="Fars" pageId="17" pageNumber="246">FARS</collectingRegion>
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||
:
|
||
</emphasis>
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||
1 ♂ (
|
||
<collectionCode id="ED4BAE167A3FFFAFFDF6053B36F0B32F" box="[639,705,1043,1062]" country="Czech Republic" name="National Museum Prague" pageId="17" pageNumber="246">NMPC</collectionCode>
|
||
): Iran, Fars,Ali-abad,
|
||
<quantity id="4CA29B367A3FFFAFFE9D05063579B348" box="[276,328,1070,1089]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="17" pageNumber="246" unit="km" value="75.0">75 km</quantity>
|
||
NW of Djahrom,wadi of the river Shur,
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||
<date id="FFE410137A3FFFAFFD1D050636C4B348" box="[660,757,1070,1089]" pageId="17" pageNumber="246" value="1970-07-10">10.vii.1970</date>
|
||
, Expedition National Museum Praha,locality 53.
|
||
</materialsCitation>
|
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<materialsCitation id="3B323C8E7A3FFFAFFDA30560354DB3A2" ID-GBIF-Occurrence="3756360320" pageId="17" pageNumber="246">
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<emphasis id="B92EEAC17A3FFFAFFDA305603648B352" bold="true" box="[554,633,1096,1115]" pageId="17" pageNumber="246">
|
||
<collectingRegion id="499EF8317A3FFFAFFDA305603643B350" box="[554,626,1096,1115]" country="Iran" name="Kerman" pageId="17" pageNumber="246">KERMAN</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
1 ♂ (
|
||
<collectionCode id="ED4BAE167A3FFFAFFD2D056036DEB352" box="[676,751,1096,1115]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="246" type="Museum">NHMW</collectionCode>
|
||
): Iran, Kerman, Manujan,
|
||
<quantity id="4CA29B367A3FFFAFFEE3054B359BB37F" box="[362,426,1123,1142]" metricMagnitude="5" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="246" unit="km" value="110.0">110 km</quantity>
|
||
E Bandarabaas, at light,
|
||
<date id="FFE410137A3FFFAFFD09054B36E2B37F" box="[640,723,1123,1142]" pageId="17" pageNumber="246" value="1974-06-02">2.vi.1974</date>
|
||
, lgt. Pretzmann, Exp. Nat. Hist. Mus. Vindob. [= expedition of the Natural History Museum, Vienna].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A3FFFAFFF1E05EC358EB072" pageId="17" pageNumber="246" type="description">
|
||
<paragraph id="8BE536D37A3FFFAFFF1E05EC358EB072" blockId="17.[151,761,1220,2075]" pageId="17" pageNumber="246">
|
||
<emphasis id="B92EEAC17A3FFFAFFF1E05EC356DB3D2" bold="true" box="[151,348,1220,1243]" pageId="17" pageNumber="246">
|
||
Description.
|
||
<emphasis id="B92EEAC17A3FFFAFFEAC05EC356DB3D2" bold="true" box="[293,348,1220,1243]" italics="true" pageId="17" pageNumber="246">Body</emphasis>
|
||
</emphasis>
|
||
length
|
||
<quantity id="4CA29B367A3FFFAFFE2305EC3619B3D2" box="[426,552,1220,1243]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="17" pageNumber="246" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.7–1.9 mm</quantity>
|
||
(
|
||
<typeStatus id="54E188717A3FFFAFFDBC05EC36A3B3D2" box="[565,658,1220,1243]" pageId="17" pageNumber="246" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CA29B367A3FFFAFFD1105EC36DDB3D2" box="[664,748,1220,1243]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="246" unit="mm" value="1.7">1.7 mm</quantity>
|
||
), maximum body width 1.0–
|
||
<quantity id="4CA29B367A3FFFAFFE2205CC35C7B3F2" box="[427,502,1252,1275]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="246" unit="mm" value="1.1">1.1 mm</quantity>
|
||
(
|
||
<typeStatus id="54E188717A3FFFAFFD8805CC366CB3F2" box="[513,605,1252,1275]" pageId="17" pageNumber="246" type="holotype">holotype</typeStatus>
|
||
1.0 mm). Head and labrum black; pronotum dark brown in centre, becoming slightly paler towards margins; elytra uniformly dark brown; legs brown.
|
||
<emphasis id="B92EEAC17A3FFFAFFE97046C3568B252" bold="true" box="[286,345,1348,1371]" italics="true" pageId="17" pageNumber="246">Head</emphasis>
|
||
without microsculpture on interstices; punctation sparse, each puncture bearing pointed seta. Eyes separated by 2.5× the width of one eye in dorsal view.
|
||
<emphasis id="B92EEAC17A3FFFAFFF1E048C3535B2B2" bold="true" box="[151,260,1444,1467]" italics="true" pageId="17" pageNumber="246">Pronotum</emphasis>
|
||
with sparse setiferous punctation similar to that on head; interstices without microsculpture.
|
||
<emphasis id="B92EEAC17A3FFFAFFDE804EC3695B2D2" bold="true" box="[609,676,1476,1499]" italics="true" pageId="17" pageNumber="246">Elytra</emphasis>
|
||
with 10 striae sharply impressed except in basal fourth where neither striae nor serial punctures are visible; intervals weakly convex at midlength and near apex; interval punctation sparse, setiferous; interstices without microsculpture.
|
||
<emphasis id="B92EEAC17A3FFFAFFD07076C36C9B152" bold="true" box="[654,760,1604,1627]" italics="true" pageId="17" pageNumber="246">Aedeagus</emphasis>
|
||
(
|
||
<figureCitation id="13612A567A3FFFAFFF17074C34DEB172" box="[158,239,1636,1659]" captionStart="Fig" captionStartId="19.[151,182,2009,2028]" captionTargetBox="[159,1399,787,1981]" captionTargetId="figure-14@19.[151,1436,777,1989]" captionTargetPageId="19" captionText="Fig. 10. Male genitalia of Thysanarthria species, holotypes.A–E – T. persica sp. nov. (A–C – aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. wadicola sp. nov. (F–H – aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488944" httpUri="https://zenodo.org/record/4488944/files/figure.png" pageId="17" pageNumber="246">Figs 10</figureCitation>
|
||
A–E)
|
||
<quantity id="4CA29B367A3FFFAFFEBB074C35B4B172" box="[306,389,1636,1659]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="246" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long. Phallobase slightly widened at base of parameres, c. as wide as parameres combined, slightly narrowing towards base, arcuately bent in lateral view. Parameres narrowly elongate, gradually narrowing towards apex, outer face arcuate or indistinctly sinuate. Median lobe narrow with widely rounded membranous apex nearly reaching level of parameral apices, paired subapical projections absent; gonopore transversely oval, situated in distal fourth of median lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A3FFFAFFF1E06AC31B7B6BC" pageId="17" pageNumber="246" type="diagnosis">
|
||
<paragraph id="8BE536D37A3FFFAFFF1E06AC31B7B6BC" blockId="17.[151,761,1220,2075]" lastBlockId="17.[828,1437,190,757]" pageId="17" pageNumber="246">
|
||
<emphasis id="B92EEAC17A3FFFAFFF1E06AC35A0B092" bold="true" box="[151,401,1924,1947]" pageId="17" pageNumber="246">Differential diagnosis.</emphasis>
|
||
Unlike most other
|
||
<taxonomicName id="4C5A4D507A3FFFAFFDD606AC36C9B092" authorityName="Fikáček & Liu" authorityYear="2019" box="[607,760,1924,1947]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEAC17A3FFFAFFDD606AC36C9B092" box="[607,760,1924,1947]" italics="true" pageId="17" pageNumber="246">Thysanarthria</emphasis>
|
||
</taxonomicName>
|
||
except
|
||
<taxonomicName id="4C5A4D507A3FFFAFFF6B068C356FB0B2" authority="Fikáček & Liu, 2019" authorityName="Fikáček & Liu" authorityYear="2019" box="[226,350,1956,1979]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="wadicola" status="sp. nov.">
|
||
<emphasis id="B92EEAC17A3FFFAFFF6B068C356FB0B2" box="[226,350,1956,1979]" italics="true" pageId="17" pageNumber="246">T. wadicola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21D57BA7A3FFFAFFEEA068C3586B0B2" box="[355,439,1956,1979]" pageId="17" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C5A4D507A3FFFAFFE48068C3618B0B2" authorityName="Fikáček & Liu" authorityYear="2019" box="[449,553,1956,1979]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="persica">
|
||
<emphasis id="B92EEAC17A3FFFAFFE48068C3618B0B2" box="[449,553,1956,1979]" italics="true" pageId="17" pageNumber="246">T. persica</emphasis>
|
||
</taxonomicName>
|
||
has uniformly dark pronotum and elytra. Its genitalia are characteristic by rather elongate parameres with arcuate lateral face and widely rounded apex (in contrast to
|
||
<taxonomicName id="4C5A4D507A3FFFAFFDB0092C368DBF12" authorityName="Fikáček & Liu" authorityYear="2019" box="[569,700,2052,2075]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="wadicola">
|
||
<emphasis id="B92EEAC17A3FFFAFFDB0092C368DBF12" box="[569,700,2052,2075]" italics="true" pageId="17" pageNumber="246">T. wadicola</emphasis>
|
||
</taxonomicName>
|
||
with prolonged acuminate apices of parameres). It also differs from
|
||
<taxonomicName id="4C5A4D507A3FFFAFFCFE01F637C4B7FC" authorityName="Fikáček & Liu" authorityYear="2019" box="[887,1013,222,245]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="wadicola">
|
||
<emphasis id="B92EEAC17A3FFFAFFCFE01F637C4B7FC" box="[887,1013,222,245]" italics="true" pageId="17" pageNumber="246">T. wadicola</emphasis>
|
||
</taxonomicName>
|
||
in the median lobe nearly reaching the level of apices of parameres (in contrast to very short median lobe in
|
||
<taxonomicName id="4C5A4D507A3FFFAFFC6B0036306EB63C" authorityName="Fikáček & Liu" authorityYear="2019" box="[994,1119,286,309]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="wadicola">
|
||
<emphasis id="B92EEAC17A3FFFAFFC6B0036306EB63C" box="[994,1119,286,309]" italics="true" pageId="17" pageNumber="246">T. wadicola</emphasis>
|
||
</taxonomicName>
|
||
), in parameres c. one third as long as phallobase when seen in lateral view (c. as long as phallobase in
|
||
<taxonomicName id="4C5A4D507A3FFFAFFC580076307FB67C" authorityName="Fikáček & Liu" authorityYear="2019" box="[977,1102,350,373]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="wadicola">
|
||
<emphasis id="B92EEAC17A3FFFAFFC580076307FB67C" box="[977,1102,350,373]" italics="true" pageId="17" pageNumber="246">T. wadicola</emphasis>
|
||
</taxonomicName>
|
||
) and in moderately wide bases of parameres in lateral view (very wide base narrowing into a very narrow apex in lateral view in
|
||
<taxonomicName id="4C5A4D507A3FFFAFFB7300B63147B6BC" authorityName="Fikáček & Liu" authorityYear="2019" box="[1274,1398,414,437]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="wadicola">
|
||
<emphasis id="B92EEAC17A3FFFAFFB7300B63147B6BC" box="[1274,1398,414,437]" italics="true" pageId="17" pageNumber="246">T. wadicola</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A3FFFAFFCB500963143B6FC" pageId="17" pageNumber="246" type="etymology">
|
||
<paragraph id="8BE536D37A3FFFAFFCB500963143B6FC" blockId="17.[828,1437,190,757]" pageId="17" pageNumber="246">
|
||
<emphasis id="B92EEAC17A3FFFAFFCB50096378DB6DC" bold="true" box="[828,956,446,469]" pageId="17" pageNumber="246">Etymology.</emphasis>
|
||
The name refers to
|
||
<collectingCountry id="F34D76437A3FFFAFFB00009630FCB6DC" box="[1161,1229,446,469]" name="Iran" pageId="17" pageNumber="246">Persia</collectingCountry>
|
||
, the historical name of
|
||
<collectingCountry id="F34D76437A3FFFAFFCD000F637B5B6FC" box="[857,900,478,501]" name="Iran" pageId="17" pageNumber="246">Iran</collectingCountry>
|
||
this new species is described from. Adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A3FFFAFFCB500D63024B5DC" pageId="17" pageNumber="246" type="biology_ecology">
|
||
<paragraph id="8BE536D37A3FFFAFFCB500D63024B5DC" blockId="17.[828,1437,190,757]" pageId="17" pageNumber="246">
|
||
<emphasis id="B92EEAC17A3FFFAFFCB500D637A6B51C" bold="true" box="[828,919,510,533]" pageId="17" pageNumber="246">Biology.</emphasis>
|
||
Based on
|
||
<bibRefCitation id="EFCB4B227A3FFFAFFB8200D630D0B51C" author="HOBERLANDT" box="[1035,1249,510,533]" firstAuthor="HOBERLANDT" pageId="17" pageNumber="246" pagination="5 - 32" refId="ref18257" refString="HOBERLANDT L. 1981: Results of the Czechoslovak-Iranian entomological expeditions to Iran. Introduction to the Second expedition 1973. Acta Entomologica Musei Nationalis Pragae 40: 5 - 32 + photos 1 - 42." type="journal article" year="1981">HOBERLANDT (1981)</bibRefCitation>
|
||
, the type locality was a rocky bank of a mountain torrent without vegetation at c.
|
||
<quantity id="4CA29B367A3FFFAFFC1B031637E7B55C" box="[914,982,574,597]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="246" unit="m" value="900.0">900 m</quantity>
|
||
of altitude on stony mountains slopes; the specimens were collected from small gravel strands on the bank of the brook. The
|
||
<typeStatus id="54E188717A3FFFAFFBB1035630A5B59C" box="[1080,1172,638,661]" pageId="17" pageNumber="246" type="paratype">paratype</typeStatus>
|
||
from
|
||
<collectingRegion id="499EF8317A3FFFAFFB5F03563135B59C" box="[1238,1284,638,661]" country="Iran" name="Fars" pageId="17" pageNumber="246">Fars</collectingRegion>
|
||
was collected in a wadi with a stream with muddy and sandy banks (
|
||
<bibRefCitation id="EFCB4B227A3FFFAFFCCA03963038B5DC" author="HOBERLANDT" box="[835,1033,702,725]" firstAuthor="HOBERLANDT" pageId="17" pageNumber="246" pagination="9 - 20" refId="ref18218" refString="HOBERLANDT L. 1974: Results of the Czechoslovak-Iranian entomological expedition to Iran 1970. Np. 1: Introduction. Acta Entomologica Musei Nationalis Pragae, Supplementum 6: 9 - 20 + Figs 1 - 22." type="journal article" year="1974">HOBERLANDT 1974</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34065587A3FFFAFFCB503F63141B5FC" box="[828,1392,734,757]" pageId="17" pageNumber="246" type="distribution">
|
||
<paragraph id="8BE536D37A3FFFAFFCB503F63141B5FC" blockId="17.[828,1437,190,757]" box="[828,1392,734,757]" pageId="17" pageNumber="246">
|
||
<emphasis id="B92EEAC17A3FFFAFFCB503F637FFB5FC" bold="true" box="[828,974,734,757]" pageId="17" pageNumber="246">Distribution.</emphasis>
|
||
So far only known from southern
|
||
<collectingCountry id="F34D76437A3FFFAFFAB703F6315DB5FC" box="[1342,1388,734,757]" name="Iran" pageId="17" pageNumber="246">Iran</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |