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<mods:title id="9FF5CA4A87E995BD4EFA15C950F0E54A">A new species of Athyreus MacLeay, 1819 (Coleoptera: Geotrupidae: Bolboceratinae: Athyreini) from northeastern Brazil</mods:title>
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<mods:namePart id="D3D638FF2FBC464AA28B2F3E72C06805">Celante, Gabriel Luiz</mods:namePart>
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<mods:namePart id="F7067C6084D5D1E05D44A01BCE97E448">Jahyny, Benoit Jean Bernard</mods:namePart>
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<taxonomicName id="4C4322ED351CFFA7C6FCFD13FD84198C" ID-CoL="JFC9" authority="Felsche, 1909" authorityName="Felsche" authorityYear="1909" box="[151,548,740,767]" class="Insecta" family="Geotrupidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="396" phylum="Arthropoda" rank="species" species="tuberifer">
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<emphasis id="B937857C351CFFA7C6FCFD13FED6198D" bold="true" box="[151,374,740,767]" italics="true" pageId="7" pageNumber="396">Athyreus tuberifer</emphasis>
Felsche, 1909
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<emphasis id="B937857C351CFFA7C6FCFCDFFE201830" bold="true" box="[151,384,808,834]" pageId="7" pageNumber="396">Material examined.</emphasis>
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<specimenCount id="9D4592E7351CFFA7C7EEFCDEFE6F1830" box="[389,463,808,835]" count="1" pageId="7" pageNumber="396" type="male">1 male</specimenCount>
at
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labeled “
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(
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?),
<taxonomicName id="4C4322ED351CFFA7C26BFCDFFAC91830" authority=", P. Grossi" authorityName="P. Grossi" box="[1024,1385,808,835]" class="Insecta" family="Geotrupidae" genus="Athyreus" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="396" phylum="Arthropoda" rank="species" species="tuberifer">
<emphasis id="B937857C351CFFA7C26BFCDFFBC31830" box="[1024,1123,808,834]" italics="true" pageId="7" pageNumber="396">Athyreus</emphasis>
aff.
<emphasis id="B937857C351CFFA7C2FEFCDFFB571830" box="[1173,1271,808,834]" italics="true" pageId="7" pageNumber="396">tuberifer</emphasis>
, P. Grossi
</taxonomicName>
, det. 2014, Campus Univ., SE,
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<collectingDate id="EFB98646351CFFA7C7D1FCBBFD951814" box="[442,565,844,870]" pageId="7" pageNumber="396" value="1984-06-11">11/vi/1984</collectingDate>
</date>
, Valverde, G.” (
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)
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.
</paragraph>
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<emphasis id="B937857C351CFFA7C6ACFC87FEC118F8" bold="true" box="[199,353,880,906]" pageId="7" pageNumber="396">Distribution.</emphasis>
<collectingCountry id="F35419FE351CFFA7C70DFC86FE7018F9" box="[358,464,881,907]" name="Brazil" pageId="7" pageNumber="396">BRAZIL</collectingCountry>
:
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,
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,
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,
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,
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,
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,
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,
<collectingRegion id="4987978C351CFFA7C289FC86FA8E18F9" box="[1250,1326,881,907]" country="Brazil" name="Parana" pageId="7" pageNumber="396">Paraná</collectingRegion>
(
<bibRefCitation id="EFD2249F351CFFA7C350FC87FEFC18DD" author="Howden, H. F. &amp; Martinez, A." pageId="7" pageNumber="396" pagination="1 - 70" refId="ref5829" refString="Howden, H. F. &amp; Martinez, A. (1978) A review of the New World genus Athyreus MacLeay (Scarabaeidae, Geotrupinae, Athyreini). Contributions of the American Entomological Institute, 15, 1 - 70." type="journal article" year="1978">Howden &amp; Martínez 1978</bibRefCitation>
),
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(
<emphasis id="B937857C351CFFA7C7BEFC62FDFB18DD" bold="true" box="[469,603,917,943]" pageId="7" pageNumber="396">new record</emphasis>
);
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(
<bibRefCitation id="EFD2249F351CFFA7C549FC63FBEE18DD" author="Howden, H. F. &amp; Martinez, A." box="[802,1102,916,943]" pageId="7" pageNumber="396" pagination="1 - 70" refId="ref5829" refString="Howden, H. F. &amp; Martinez, A. (1978) A review of the New World genus Athyreus MacLeay (Scarabaeidae, Geotrupinae, Athyreini). Contributions of the American Entomological Institute, 15, 1 - 70." type="journal article" year="1978">Howden &amp; Martínez 1978</bibRefCitation>
). (
<figureCitation id="137845EB351CFFA7C201FC63FB6418DD" box="[1130,1220,916,943]" captionStart="FIGURE 5" captionStartId="6.[152,255,1716,1740]" captionTargetBox="[151,1435,181,1692]" captionTargetId="figure-24@6.[151,1435,181,1692]" captionTargetPageId="6" captionText="FIGURE 5AD. Distribution records of some Bolboceratinae species. A, Athyreus arretado new species, Athyreus vavini Boucomont, 1902 and Athyreus tuberifer Felsche, 1909 distribution records; B, Neoathyreus anthracinus (Klug, 1843) and Neoathyreus braziliensis Howden, 1985 distribution records; C, Neoathyreus centromaculatus (Felsche, 1909), Neoathyreus lanuginosus (Klug, 1843), and Parathyreus trituberculatus (Klug, 1843) distribution records; D, Bolbapium caesum (Klug, 1843), Bolbapium parcepunctatum (Arrow, 1913), and Bolbapium sulcifrons Ide &amp; Martínez, 1993 distribution records. The black symbols represent new records, the unfilled symbols are literature records." figureDoi="http://doi.org/10.5281/zenodo.13848925" httpUri="https://zenodo.org/record/13848925/files/figure.png" pageId="7" pageNumber="396">Fig. 5A</figureCitation>
).
</paragraph>
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