treatments-xml/data/03/EA/E2/03EAE245435E9B1CCC631BF68A70FBAD.xml
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<document id="01B7555D25819B6D18075CB4E4A255C5" ID-CLB-Dataset="286006" ID-DOI="10.38201/btha.abc.v52.i1.11" ID-GBIF-Dataset="c2dbed47-91bf-4b32-ba19-6aac4b574387" ID-ISSN="2311-9284" ID-Zenodo-Dep="10515159" IM.bibliography_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1705345755780" checkinUser="felipe" docAuthor="Manning, John &amp; Klak, Cornelia" docDate="2022" docId="03EAE245435E9B1CCC631BF68A70FBAD" docLanguage="en" docName="Bothalia.52.1.a11.pdf" docOrigin="Bothalia (a 11) 52 (1)" docSource="http://dx.doi.org/10.38201/btha.abc.v52.i1.11" docStyle="DocumentStyle:C8C375161A5A5A90DC5116AFA74E0A4D.1:Bothalia.2020-.journal_article" docStyleId="C8C375161A5A5A90DC5116AFA74E0A4D" docStyleName="Bothalia.2020-.journal_article" docStyleVersion="1" docTitle="Tetragonia L." docType="treatment" docVersion="3" lastPageNumber="3" masterDocId="FFD39A3D435F9B1ECF5B1F3B8E10FFBF" masterDocTitle="New infrageneric taxa in Leucospermum (Proteaceae) and Tetragonia L. (Aizoaceae) and a new name in Aizoon subgenus Galenia (Aizoaceae) in southern Africa" masterLastPageNumber="5" masterPageNumber="1" pageNumber="2" updateTime="1705940359142" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="826408ECBE63827E512A78C2EC12901F">New infrageneric taxa in Leucospermum (Proteaceae) and Tetragonia L. (Aizoaceae) and a new name in Aizoon subgenus Galenia (Aizoaceae) in southern Africa</mods:title>
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<paragraph id="8BFC5353435E9B1FCC631BF68B6EFB54" blockId="1.[824,1406,1229,1259]" box="[824,1406,1229,1259]" pageId="1" pageNumber="2">
<heading id="D0B4E43F435E9B1FCC631BF68B6EFB54" box="[824,1406,1229,1259]" fontSize="12" level="2" pageId="1" pageNumber="2" reason="4">
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2.
<taxonomicName id="4C4328D0435E9B1FCC2F1BF68A19FB54" ID-CoL="64743" box="[884,1033,1229,1259]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="genus">Tetragonia</taxonomicName>
</emphasis>
L. (
<taxonomicName id="4C4328D0435E9B1FCB631BF68ADEFB54" box="[1080,1230,1229,1259]" class="Magnoliopsida" family="Aizoaceae" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="family">Aizoaceae</taxonomicName>
: Aizooideae)
</heading>
</paragraph>
<paragraph id="8BFC5353435E9B1FCC631A2B8A67F82C" blockId="1.[824,1437,1296,1941]" pageId="1" pageNumber="2">
<taxonomicName id="4C4328D0435E9B1FCC631A2B8DB9FA96" ID-CoL="64743" box="[824,937,1296,1321]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435E9B1FCC631A2B8DB9FA96" box="[824,937,1296,1321]" italics="true" pageId="1" pageNumber="2">Tetragonia</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C4328D0435E9B1FCC8A1A2B8A5BFA96" ID-CoL="65P" box="[977,1099,1296,1321]" class="Magnoliopsida" family="Aizoaceae" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="family">Aizoaceae</taxonomicName>
: Azooideae) as recently re­circumscribed, is a genus of ± 50 species from temperate and subtropical regions, mainly in the southern Hemisphere (
<bibRefCitation id="EFD22EA2435E9B1FCCCF1A4A8A49FA35" author="Klak, C. &amp; Hanacek, P. &amp; Bruyns, P. V." box="[916,1113,1393,1418]" pageId="1" pageNumber="2" pagination="1147 - 1170" refId="ref3359" refString="Klak, C., Hanacek, P. &amp; Bruyns, P. V., 2017, ' Disentangling the Aizooideae: New generic concepts and a new subfamily in Aizoaceae', Taxon 66, 1147 - 1170." type="journal article" year="2017">Klak et al. 2017</bibRefCitation>
). The southern African taxa were last revised by Adamson (1955), and this remains the current treatment for the species in the region. In his classification, Adamson (1955) segregated the species among the four subgenera
<taxonomicName id="4C4328D0435E9B1FCBFA1AC88D88F990" ID-CoL="64743" authority=", Tetragonoides DC." authorityName="Tetragonoides DC." class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435E9B1FCBFA1AC88B00F9B3" box="[1185,1296,1523,1548]" italics="true" pageId="1" pageNumber="2">Tetragonia</emphasis>
,
<emphasis id="B9378F41435E9B1FCA7A1AC88D4DF992" italics="true" pageId="1" pageNumber="2">Tetragonoides</emphasis>
DC.
</taxonomicName>
,
<taxonomicName id="4C4328D0435E9B1FCCFD192F8ADBF992" ID-CoL="WGW" authority="(Schinz) Engl." authorityName="Engl." baseAuthorityName="Schinz" box="[934,1227,1556,1583]" class="Magnoliopsida" family="Aizoaceae" genus="Anisostigma" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435E9B1FCCFD192F8A39F992" box="[934,1065,1556,1581]" italics="true" pageId="1" pageNumber="2">Anisostigma</emphasis>
(Schinz) Engl.
</taxonomicName>
and
<taxonomicName id="4C4328D0435E9B1FCA52192F8A0EF9F1" ID-CoL="7YZP" authority="(S. Moore) Adamson" authorityName="Adamson" baseAuthorityName="S. Moore" class="Magnoliopsida" family="Aizoaceae" genus="Tribulocarpus" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435E9B1FCA52192F8B8DF992" box="[1289,1437,1556,1581]" italics="true" pageId="1" pageNumber="2">Tribulocarpus</emphasis>
(S.Moore) Adamson
</taxonomicName>
, defined by several characters of the fruits and flowers. The recent molecular phylogenetic analysis of the subfamily by
<bibRefCitation id="EFD22EA2435E9B1FCB92194D8B8AF930" author="Klak, C. &amp; Hanacek, P. &amp; Bruyns, P. V." box="[1225,1434,1654,1679]" pageId="1" pageNumber="2" pagination="1147 - 1170" refId="ref3359" refString="Klak, C., Hanacek, P. &amp; Bruyns, P. V., 2017, ' Disentangling the Aizooideae: New generic concepts and a new subfamily in Aizoaceae', Taxon 66, 1147 - 1170." type="journal article" year="2017">Klak et al. (2017)</bibRefCitation>
, however, uncovered some unexpected relationships among the species that necessitated realignments among several taxa in order to render them monophyletic. Among these was the removal of
<taxonomicName id="4C4328D0435E9B1FCA7019C38B8CF8AE" ID-CoL="64743" box="[1323,1436,1784,1809]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435E9B1FCA7019C38B8CF8AE" box="[1323,1436,1784,1809]" italics="true" pageId="1" pageNumber="2">Tetragonia</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="4C4328D0435E9B1FCCF718238A3FF88E" ID-CoL="WGW" box="[940,1071,1816,1841]" class="Magnoliopsida" family="Aizoaceae" genus="Anisostigma" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435E9B1FCCF718238A3FF88E" box="[940,1071,1816,1841]" italics="true" pageId="1" pageNumber="2">Anisostigma</emphasis>
</taxonomicName>
and subgenus
<taxonomicName id="4C4328D0435E9B1FCBBD18238B6AF88E" ID-CoL="7YZP" box="[1254,1402,1816,1841]" class="Magnoliopsida" family="Aizoaceae" genus="Tribulocarpus" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435E9B1FCBBD18238B6AF88E" box="[1254,1402,1816,1841]" italics="true" pageId="1" pageNumber="2">Tribulocarpus</emphasis>
</taxonomicName>
to the new tribe
<taxonomicName id="4C4328D0435E9B1FCC8118028AD6F8ED" authority="Klak" authorityName="Klak" box="[986,1222,1849,1874]" class="Magnoliopsida" family="Aizoaceae" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="tribe" tribe="Anisostigmateae">Anisostigmateae Klak</taxonomicName>
in subfamily
<taxonomicName id="4C4328D0435E9B1FCA3A18018D87F8CD" ID-CoL="KBT" class="Magnoliopsida" family="Aizoaceae" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="subFamily" subFamily="Sesuvioideae">Sesuvioideae</taxonomicName>
as the genera
<taxonomicName id="4C4328D0435E9B1FCB6018628B1EF8CA" ID-CoL="WGW" authority="Schinz" authorityName="Schinz" box="[1083,1294,1881,1909]" class="Magnoliopsida" family="Aizoaceae" genus="Anisostigma" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435E9B1FCB6018628AAEF8CD" box="[1083,1214,1881,1906]" italics="true" pageId="1" pageNumber="2">Anisostigma</emphasis>
Schinz
</taxonomicName>
and
<taxonomicName id="4C4328D0435E9B1FCA1218628DF7F82A" ID-CoL="7YZP" authority="S. Moore" authorityName="S. Moore" class="Magnoliopsida" family="Aizoaceae" genus="Tribulocarpus" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435E9B1FCA1218628D90F82C" italics="true" pageId="1" pageNumber="2">Tribulocarpus</emphasis>
S.Moore
</taxonomicName>
respectively.
</paragraph>
<paragraph id="8BFC5353435E9B1CCC6318828C94FE70" blockId="1.[824,1436,1977,2099]" lastBlockId="2.[150,765,212,463]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
In addition to these changes, the circumscription of
<taxonomicName id="4C4328D0435E9B1FCC6318E28DB9F84D" ID-CoL="64743" box="[824,937,2009,2034]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435E9B1FCC6318E28DB9F84D" box="[824,937,2009,2034]" italics="true" pageId="1" pageNumber="2">Tetragonia</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="4C4328D0435E9B1FCB6318E28AC0F84D" box="[1080,1232,2009,2034]" pageId="1" pageNumber="2" rank="subGenus" subGenus="Tetragonoides">
<emphasis id="B9378F41435E9B1FCB6318E28AC0F84D" box="[1080,1232,2009,2034]" italics="true" pageId="1" pageNumber="2">Tetragonoides</emphasis>
</taxonomicName>
was greatly altered by the removal from it of most of the southern African taxa previously included there by Adamson (1955).
<taxonomicName id="4C4328D0435D9B1CCFA91FEF8F73FF52" ID-CoL="64743" box="[242,355,212,237]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435D9B1CCFA91FEF8F73FF52" box="[242,355,212,237]" italics="true" pageId="2" pageNumber="3">Tetragonia</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="4C4328D0435D9B1CCE851FEF8C66FF52" box="[478,630,212,237]" pageId="2" pageNumber="3" rank="subGenus" subGenus="Tetragonoides">
<emphasis id="B9378F41435D9B1CCE851FEF8C66FF52" box="[478,630,212,237]" italics="true" pageId="2" pageNumber="3">Tetragonoides</emphasis>
</taxonomicName>
is now understood to be essentially Australasian in distribution, with the exception of the cosmopolitan littoral species
<taxonomicName id="4C4328D0435D9B1CCFC31E0E8FCDFEF1" ID-CoL="7C2YJ" authority="(Pall.) Kuntze" authorityName="Kuntze" baseAuthorityName="Pall." box="[152,477,309,334]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="tetragonoides">
<emphasis id="B9378F41435D9B1CCFC31E0E8F55FEF1" box="[152,325,309,334]" italics="true" pageId="2" pageNumber="3">T. tetragonoides</emphasis>
(Pall.) Kuntze
</taxonomicName>
, the
<typeStatus id="54F8EDF1435D9B1CCD4B1E0E8C51FEF1" box="[528,577,309,334]" pageId="2" pageNumber="3">type</typeStatus>
of the subgenus. Two additional subgenera, subgenus
<taxonomicName id="4C4328D0435D9B1CCD761E6E8ED6FE30" authority="(Miq.) Klak" authorityName="Klak" baseAuthorityName="Miq." class="Magnoliopsida" family="Aizoaceae" genus="Tetragonella" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435D9B1CCD761E6E8CA2FED1" box="[557,690,341,366]" italics="true" pageId="2" pageNumber="3">Tetragonella</emphasis>
(Miq.) Klak
</taxonomicName>
and subgenus
<taxonomicName id="4C4328D0435D9B1CCE231E4D8C23FE30" authority="Klak" authorityName="Klak" box="[376,563,374,399]" pageId="2" pageNumber="3" rank="subGenus" subGenus="Americanae">
<emphasis id="B9378F41435D9B1CCE231E4D8FEAFE30" box="[376,506,374,399]" italics="true" pageId="2" pageNumber="3">Americanae</emphasis>
Klak
</taxonomicName>
were established to accommodate respectively the other Australasian and the American species (
<bibRefCitation id="EFD22EA2435D9B1CCE981E8D8C6AFE70" author="Klak, C. &amp; Hanacek, P. &amp; Bruyns, P. V." box="[451,634,438,463]" pageId="2" pageNumber="3" pagination="1147 - 1170" refId="ref3359" refString="Klak, C., Hanacek, P. &amp; Bruyns, P. V., 2017, ' Disentangling the Aizooideae: New generic concepts and a new subfamily in Aizoaceae', Taxon 66, 1147 - 1170." type="journal article" year="2017">Klak et al. 2017</bibRefCitation>
).
</paragraph>
<paragraph id="8BFC5353435D9B1CCC411FB88D0FFF25" box="[794,799,131,154]" pageId="2" pageNumber="3">|</paragraph>
<paragraph id="8BFC5353435D9B1CCFCC1ECE8CF6FCEF" blockId="2.[151,764,501,848]" pageId="2" pageNumber="3">
The remaining four southern African species that were included by Adamson (1955) in subgenus
<taxonomicName id="4C4328D0435D9B1CCDDB1D2E8EACFDF1" pageId="2" pageNumber="3" rank="subGenus" subGenus="Tetragonoides">
<emphasis id="B9378F41435D9B1CCDDB1D2E8EACFDF1" italics="true" pageId="2" pageNumber="3">Tetragonoides</emphasis>
</taxonomicName>
were retrieved by
<bibRefCitation id="EFD22EA2435D9B1CCED61D0E8C5AFDF1" author="Klak, C. &amp; Hanacek, P. &amp; Bruyns, P. V." box="[397,586,565,590]" pageId="2" pageNumber="3" pagination="1147 - 1170" refId="ref3359" refString="Klak, C., Hanacek, P. &amp; Bruyns, P. V., 2017, ' Disentangling the Aizooideae: New generic concepts and a new subfamily in Aizoaceae', Taxon 66, 1147 - 1170." type="journal article" year="2017">Klak et al. (2017)</bibRefCitation>
as a clade sister to subgenus
<taxonomicName id="4C4328D0435D9B1CCE7D1D6E8F87FDD1" box="[294,407,597,622]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435D9B1CCE7D1D6E8F87FDD1" box="[294,407,597,622]" italics="true" pageId="2" pageNumber="3">Tetragonia</emphasis>
</taxonomicName>
and were accordingly included in it, with the observation that the series erected by Adamson (1955) should be maintained. This alludes to the seven sections recognised by Adamson (1955) in subgenus
<taxonomicName id="4C4328D0435D9B1CCE5D1DED8F67FD50" box="[262,375,726,751]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435D9B1CCE5D1DED8F67FD50" box="[262,375,726,751]" italics="true" pageId="2" pageNumber="3">Tetragonia</emphasis>
</taxonomicName>
. Although they included these additional four former members of subgenus
<taxonomicName id="4C4328D0435D9B1CCDD91DCC8EACFC8F" pageId="2" pageNumber="3" rank="subGenus" subGenus="Tetragonoides">
<emphasis id="B9378F41435D9B1CCDD91DCC8EACFC8F" italics="true" pageId="2" pageNumber="3">Tetragonoides</emphasis>
</taxonomicName>
in subgenus
<taxonomicName id="4C4328D0435D9B1CCE0D1C2C8C8EFC8F" authority=", Klak et al. (2017)" authorityName="Klak" authorityYear="2017" box="[342,670,791,816]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435D9B1CCE0D1C2C8FDFFC8F" box="[342,463,791,816]" italics="true" pageId="2" pageNumber="3">Tetragonia,</emphasis>
<bibRefCitation id="EFD22EA2435D9B1CCE821C2C8C8EFC8F" author="Klak, C. &amp; Hanacek, P. &amp; Bruyns, P. V." box="[473,670,791,816]" pageId="2" pageNumber="3" pagination="1147 - 1170" refId="ref3359" refString="Klak, C., Hanacek, P. &amp; Bruyns, P. V., 2017, ' Disentangling the Aizooideae: New generic concepts and a new subfamily in Aizoaceae', Taxon 66, 1147 - 1170." type="journal article" year="2017">Klak et al. (2017)</bibRefCitation>
</taxonomicName>
did not accommodate them within the existing classification.
</paragraph>
<paragraph id="8BFC5353435D9B1CCFCC1C4C8CF7FB2F" blockId="2.[151,765,885,1168]" pageId="2" pageNumber="3">
Significantly, the removal of
<taxonomicName id="4C4328D0435D9B1CCE871C4D8C9CFC30" box="[476,652,886,911]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="tetragonoides">
<emphasis id="B9378F41435D9B1CCE871C4D8C9CFC30" box="[476,652,886,911]" italics="true" pageId="2" pageNumber="3">T. tetragonoides</emphasis>
</taxonomicName>
from this group renders it morphologically more homogenous. The four species share a prostrate, ± annual habit and half­inferior ovary, and differ from all other members of subgenus
<taxonomicName id="4C4328D0435D9B1CCE7D1CCD8F87FBB0" box="[294,407,1014,1039]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435D9B1CCE7D1CCD8F87FBB0" box="[294,407,1014,1039]" italics="true" pageId="2" pageNumber="3">Tetragonia</emphasis>
</taxonomicName>
in having the stamens less than twice as many as the perianth segments vs. more than twice as many as the perianth segments (Adamson 1955). The new series
<taxonomicName id="4C4328D0435D9B1CCEFE1B6C8CEBFBCF" authority="J. C. Manning &amp; Klak" authorityName="J. C. Manning &amp; Klak" box="[421,763,1111,1138]" pageId="2" pageNumber="3" rank="series" series="Echinatae">
<emphasis id="B9378F41435D9B1CCEFE1B6C8C1FFBCF" box="[421,527,1111,1136]" italics="true" pageId="2" pageNumber="3">Echinatae</emphasis>
J.C.Manning &amp; Klak
</taxonomicName>
is accordingly proposed here to accommodate them.
</paragraph>
<paragraph id="8BFC5353435D9B1CCFCC1B8D8FA7F8CB" blockId="2.[150,764,1206,1908]" pageId="2" pageNumber="3">
There are two other nomenclatural corrections required in subgenus
<taxonomicName id="4C4328D0435D9B1CCE261BED8FFEFB50" box="[381,494,1238,1263]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435D9B1CCE261BED8FFEFB50" box="[381,494,1238,1263]" italics="true" pageId="2" pageNumber="3">Tetragonia</emphasis>
</taxonomicName>
. The first concerns the authorities for the infrageneric names that were published by Fenzl (1862). Adamson (1955) treats these as series, with Fenzl (1862) as author. However, examination of Fenzl (1862) reveals that they were published as third tier names two ranks below sectional level but without a definite rank. Although they are to be accepted as validly published by Fenzl (1862) they are to be treated as unranked (Turland et al. 2018: Art.
<emphasis id="B9378F41435D9B1CCFCC1AC18C73F94C" italics="true" pageId="2" pageNumber="3">
<emphasis id="B9378F41435D9B1CCFCC1AC18EC2F9AD" bold="true" box="[151,210,1530,1554]" italics="true" pageId="2" pageNumber="3">37.3.</emphasis>
<taxonomicName id="4C4328D0435D9B1CCF821AC38FB0F9AE" box="[217,416,1528,1553]" class="Magnoliopsida" family="Aizoaceae" genus="Anisostigma" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="subSpecies" species="name" subSpecies="published">A name published</taxonomicName>
before
<date id="FFFD7593435D9B1CCEAE1AC38CB0F9AE" box="[501,672,1528,1553]" pageId="2" pageNumber="3" value="1953-01-01">1 January 1953</date>
without a clear indication of its rank is validly published provided that all other requirements for valid publication are fulfilled; it is, however, inoperative in questions of priority except for homonymy. If it is the
<taxonomicName id="4C4328D0435D9B1CCD0C19428C85F92D" box="[599,661,1657,1682]" class="Magnoliopsida" family="Aizoaceae" genus="Anisostigma" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="name">name</taxonomicName>
of a new taxon, it may serve as a basionym or replaced synonym for subsequent new combinations, names at new ranks, or replacement names at definite ranks.
</emphasis>
). Their later publication by Adamson (1955) with a clear indication of the rank of series serves as the first publication at that rank and they are therefore to be treated as combinations by Adamson (1955).
</paragraph>
<paragraph id="8BFC5353435D9B1CCFCC18A28DB6FC8B" blockId="2.[151,766,1945,2099]" lastBlockId="2.[823,1437,212,820]" pageId="2" pageNumber="3">
The second correction concerns the status of the series that includes the
<typeStatus id="54F8EDF1435D9B1CCED518818FAFF86C" box="[398,447,1978,2003]" pageId="2" pageNumber="3">type</typeStatus>
of the genus,
<taxonomicName id="4C4328D0435D9B1CCD3C18818EECF84C" authority="L. Although" authorityName="L. Although" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="fruticosa">
<emphasis id="B9378F41435D9B1CCD3C18818CCFF86C" box="[615,735,1978,2003]" italics="true" pageId="2" pageNumber="3">T. fruticosa</emphasis>
L. Although
</taxonomicName>
the genus
<taxonomicName id="4C4328D0435D9B1CCED818E18FE4F84C" box="[387,500,2010,2035]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435D9B1CCED818E18FE4F84C" box="[387,500,2010,2035]" italics="true" pageId="2" pageNumber="3">Tetragonia</emphasis>
</taxonomicName>
was only lectotypified by Britton (1918) after the treatment by Fenzl (1862), the name
<emphasis id="B9378F41435D9B1CCE4817218F38F78C" box="[275,296,2074,2099]" italics="true" pageId="2" pageNumber="3">T.</emphasis>
[unranked]
<taxonomicName id="4C4328D0435D9B1CCEEC17218C2AF78C" box="[439,570,2074,2099]" pageId="2" pageNumber="3" rank="series" series="Fruticulosae">
<emphasis id="B9378F41435D9B1CCEEC17218C2AF78C" box="[439,570,2074,2099]" italics="true" pageId="2" pageNumber="3">Fruticulosae</emphasis>
</taxonomicName>
Fenzl. (1862) is to be treated as not validly published [Turland et al. 2018: Art.
<emphasis id="B9378F41435D9B1CCCE81FCD8D4BFD8E" italics="true" pageId="2" pageNumber="3">
<emphasis id="B9378F41435D9B1CCCE81FCD8DFDFEB1" bold="true" box="[947,1005,246,270]" italics="true" pageId="2" pageNumber="3">22.2.</emphasis>
<taxonomicName id="4C4328D0435D9B1CCCAF1FCE8A58FEB1" box="[1012,1096,245,270]" class="Magnoliopsida" family="Aizoaceae" genus="Anisostigma" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="name">A name</taxonomicName>
of a subdivision of a genus that includes the
<typeStatus id="54F8EDF1435D9B1CCC931E2E8DE8FE91" box="[968,1016,277,302]" pageId="2" pageNumber="3">type</typeStatus>
(i.e. the original
<typeStatus id="54F8EDF1435D9B1CCBE01E2E8AFBFE91" box="[1211,1259,277,302]" pageId="2" pageNumber="3">type</typeStatus>
or all elements eligible as
<typeStatus id="54F8EDF1435D9B1CCCF11E0D8DCAFEF0" box="[938,986,310,335]" pageId="2" pageNumber="3">type</typeStatus>
or the previously designated
<typeStatus id="54F8EDF1435D9B1CCA451E0C8B5CFEEE" box="[1310,1356,311,337]" pageId="2" pageNumber="3">type</typeStatus>
) of the adopted, legitimate
<taxonomicName id="4C4328D0435D9B1CCB7B1E6D8A4EFED0" box="[1056,1118,342,367]" class="Magnoliopsida" family="Aizoaceae" genus="Anisostigma" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="name">name</taxonomicName>
of the genus is not validly published unless its epithet repeats the generic
<taxonomicName id="4C4328D0435D9B1CCA041E4D8B8DFE30" box="[1375,1437,374,399]" class="Magnoliopsida" family="Aizoaceae" genus="Anisostigma" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="name">name</taxonomicName>
unaltered. For the purposes of this provision, explicit indication that the nomenclaturally typical element is included is considered as equivalent to inclusion of the
<typeStatus id="54F8EDF1435D9B1CCC621EC38D7CFDAE" box="[825,876,504,529]" pageId="2" pageNumber="3">type</typeStatus>
, whether or not it has been previously designated.
</emphasis>
]. Similarly, the publication of this taxon at series rank (spelled
<emphasis id="B9378F41435D9B1CCC8E1D038A53FDEE" box="[981,1091,568,593]" italics="true" pageId="2" pageNumber="3">Fruticosae</emphasis>
) by Adamson (1955) is to be treated as not validly published. The correct name for this taxon is series
<taxonomicName id="4C4328D0435D9B1CCB511D428A6BFD2D" box="[1034,1147,633,658]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435D9B1CCB511D428A6BFD2D" box="[1034,1147,633,658]" italics="true" pageId="2" pageNumber="3">Tetragonia</emphasis>
</taxonomicName>
(Turland et al. 2018: Art.
<emphasis id="B9378F41435D9B1CCC621DA08D80FC8B" italics="true" pageId="2" pageNumber="3">
<emphasis id="B9378F41435D9B1CCC621DA08D63FD0C" bold="true" box="[825,883,667,691]" italics="true" pageId="2" pageNumber="3">22.1.</emphasis>
The
<taxonomicName id="4C4328D0435D9B1CCCF41DA28DFDFD0D" box="[943,1005,665,690]" class="Magnoliopsida" family="Aizoaceae" genus="Anisostigma" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="name">name</taxonomicName>
of any subdivision of a genus that includes the
<typeStatus id="54F8EDF1435D9B1CCCE81D818DF3FD6C" box="[947,995,698,723]" pageId="2" pageNumber="3">type</typeStatus>
of the adopted, legitimate
<taxonomicName id="4C4328D0435D9B1CCA4E1D818B43FD6C" box="[1301,1363,698,723]" class="Magnoliopsida" family="Aizoaceae" genus="Anisostigma" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="name">name</taxonomicName>
of the genus to which it is assigned is to repeat that generic
<taxonomicName id="4C4328D0435D9B1CCC631DC08D66FCAB" box="[824,886,763,788]" class="Magnoliopsida" family="Aizoaceae" genus="Anisostigma" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="name">name</taxonomicName>
unaltered as its epithet, not followed by an author citation.
</emphasis>
).
</paragraph>
<paragraph id="8BFC5353435D9B1CCC631C618B13FC2C" blockId="2.[824,1436,857,915]" pageId="2" pageNumber="3">
The species of
<taxonomicName id="4C4328D0435D9B1CCCBA1C618A42FCCC" box="[993,1106,858,883]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435D9B1CCCBA1C618A42FCCC" box="[993,1106,858,883]" italics="true" pageId="2" pageNumber="3">Tetragonia</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="4C4328D0435D9B1CCB911C618B2BFCCC" box="[1226,1339,858,883]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435D9B1CCB911C618B2BFCCC" box="[1226,1339,858,883]" italics="true" pageId="2" pageNumber="3">Tetragonia</emphasis>
</taxonomicName>
are thus distributed among eight series as follows:
</paragraph>
</subSubSection>
<heading id="D0B4E43F435D9B1CCC631C838A70FBAD" centered="true" fontSize="10" level="2" pageId="2" pageNumber="3" reason="7">
<treatmentCitationGroup id="AB53747D435D9B1CCC631C838B48FC6E" box="[824,1368,952,980]" pageId="2" pageNumber="3">
<subSubSection id="C35900D8435D9B1CCC631C838DC5FC6E" box="[824,981,952,977]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BFC5353435D9B1CCC631C838DC5FC6E" blockId="2.[824,1437,952,1042]" box="[824,981,952,977]" pageId="2" pageNumber="3">
<taxonomicName id="4C4328D0435D9B1CCC631C838DDEFC6E" ID-CoL="64743" authority="L." authorityName="L." box="[824,974,952,977]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B9378F41435D9B1CCC631C838DA3FC6E" bold="true" box="[824,947,952,977]" pageId="2" pageNumber="3">Tetragonia</emphasis>
<emphasis id="B9378F41435D9B1CCCE01C838DDEFC6E" box="[955,974,952,977]" italics="true" pageId="2" pageNumber="3">
<collectionCode id="ED52CB96435D9B1CCCE01C838DDAFC6E" box="[955,970,952,977]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="2" pageNumber="3" type="Herbarium">L</collectionCode>
.
</emphasis>
</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="C35900D8435D9B1CCC861C818B48FC6E" box="[989,1368,952,980]" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="8BFC5353435D9B1CCC861C818B48FC6E" blockId="2.[824,1437,952,1042]" box="[989,1368,952,980]" pageId="2" pageNumber="3">
<treatmentCitation id="0AE27542435D9B1CCC861C818B44FC6E" author="" box="[989,1364,952,980]" journal="Species plantarum" page="480" pageId="2" pageNumber="3" score="1.6765355566371578" volume="1" year="1753">
<bibCitation id="5E695B3F435D9B1CCC861C818B44FC6E" box="[989,1364,952,980]" journal="Species plantarum" page="480" pageId="2" pageNumber="3" score="1.6765355566371578" volume="1" year="1753">Species plantarum 1: 480 (1753)</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
</treatmentCitationGroup>
<subSubSection id="C35900D8435D9B1CCA041C838A70FBAD" pageId="2" pageNumber="3" type="type_taxon">
<paragraph id="8BFC5353435D9B1CCA041C838A70FBAD" blockId="2.[824,1437,952,1042]" pageId="2" pageNumber="3">
<typeStatus id="54F8EDF1435D9B1CCA041C838B8AFC6E" box="[1375,1434,952,977]" pageId="2" pageNumber="3">Type</typeStatus>
:
<taxonomicName id="4C4328D0435D9B1CCC3D1CE28DCBFC4D" box="[870,987,985,1010]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="fruticosa">
<emphasis id="B9378F41435D9B1CCC3D1CE28DCBFC4D" box="[870,987,985,1010]" italics="true" pageId="2" pageNumber="3">T. fruticosa</emphasis>
</taxonomicName>
<collectionCode id="ED52CB96435D9B1CCCBB1CE28DE3FC4D" box="[992,1011,985,1010]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="2" pageNumber="3" type="Herbarium">L</collectionCode>
., lecto., designated by
<bibCitation id="5E695B3F435D9B1CCBB51CE28A4CFBAD" author="Britton" page="125" pageId="2" pageNumber="3" score="0.08437413277588224" volumeTitle="Flora of Bermuda" year="1918">
Britton,
<bibCitation id="5E695B3F435D9B1CCA131CE28A4CFBAD" page="125" pageId="2" pageNumber="3" score="0.08437413277588224" volumeTitle="Flora of Bermuda" year="1918">
Flora of
<collectingCountry id="F35413C3435D9B1CCC3F1CC28DC0FBAD" box="[868,976,1017,1042]" name="Bermuda" pageId="2" pageNumber="3">Bermuda</collectingCountry>
: 125 (1918)
</bibCitation>
</bibCitation>
.
</paragraph>
</subSubSection>
</heading>
</treatment>
</document>