423 lines
54 KiB
XML
423 lines
54 KiB
XML
<document id="C305468C560802E2F39A7424C58FB979" ID-DOI="10.5281/zenodo.6120089" ID-ISSN="0035-418X" ID-Zenodo-Dep="6120089" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645039399653" checkinUser="carolina" docAuthor="Makranczy, György" docDate="2014" docId="03E287D4FFACFF8B24BB5D12FE26FEE5" docLanguage="en" docName="RevSuisseZool.121.3.319-347.pdf.imf" docOrigin="Revue suisse de Zoologie 121 (3)" docStyle="DocumentStyle:0846B485C0C87F6EB7648DEEBF34B77F.2:RevSuisseZool.2011-2014.journal_article" docStyleId="0846B485C0C87F6EB7648DEEBF34B77F" docStyleName="RevSuisseZool.2011-2014.journal_article" docStyleVersion="2" docTitle="Thinobius injae Makranczy & H- 2014, sp. nov." docType="treatment" docUuid="96FA66C4-399A-4E1E-AB12-B03CC3BC2B91" docUuidSource="ZooBank" docVersion="4" lastPageNumber="330" masterDocId="FFDBFFACFFAEFF8024335818FFFBFFA0" masterDocTitle="The Far Eastern species of Thinobius Kiesenwetter, 1844 (Coleoptera: Staphylinidae, Oxytelinae) lacking female modified genital appendage" masterLastPageNumber="347" masterPageNumber="319" pageNumber="321" updateTime="1699297514242" updateUser="plazi">
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<mods:mods id="BCA01229C818E4B397C651AB19C16947" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="7116A7E66CCAFF8C849453455B8F7078">
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<mods:title id="42C31FF419AB46D59732AFB7AC77B15B">The Far Eastern species of Thinobius Kiesenwetter, 1844 (Coleoptera: Staphylinidae, Oxytelinae) lacking female modified genital appendage</mods:title>
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</mods:titleInfo>
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<mods:name id="ACA41E9D082CFDE10A66A4694664107E" type="personal">
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<mods:roleTerm id="6C793EA685D655D22B17A6ED781825DF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="78D560A0848EA574D0D28BA6069886DC">Makranczy, György</mods:namePart>
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<mods:typeOfResource id="EEF819E5E521D751E7751A419AEA76B9">text</mods:typeOfResource>
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<mods:title id="897BB581F8A248E29892A9828CD72409">Revue suisse de Zoologie</mods:title>
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</mods:titleInfo>
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<mods:part id="525AE6718787EA96DD4C0ED7D48BE4D0">
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<mods:date id="5515E78847D24CF7393A6A17E719A61D">2014</mods:date>
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<mods:detail id="A31222E2DB512E50D8632AAB7BA4AFDD" type="pubDate">
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<mods:number id="2C1B0C54D4C630DF5FF8D04F63ECCE0C">2014-09-30</mods:number>
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<mods:detail id="8EBFE9BD6D0AA13860E962999D0B1483" type="volume">
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<mods:number id="240BFDC049C7CBD4A9A1044A1FC72E51">121</mods:number>
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</mods:detail>
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<mods:detail id="52B4990AF0CF1471C5370E3A6AA8ABF6" type="issue">
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<mods:number id="BC643F71136CED24A0CD1AB7144522F9">3</mods:number>
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</mods:detail>
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<mods:extent id="E5BD9F8C3537DE4901F0CFB05076F8B6" unit="page">
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<mods:start id="903E94A652410BCE3320030C3762615E">319</mods:start>
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<mods:end id="1B2ACC1610B607610DE20A00C99D737F">347</mods:end>
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</mods:extent>
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<mods:classification id="E70CF062BA2B4A8C53FCA14620BA2F97">journal article</mods:classification>
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<mods:identifier id="E5866DC5B14966D61BC4E181F818027A" type="DOI">10.5281/zenodo.6120089</mods:identifier>
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<mods:identifier id="3580FCE85892D553ED972040BDCB989F" type="ISSN">0035-418X</mods:identifier>
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<mods:identifier id="E9E821FE675C525C12E5735AA7D8CF95" type="Zenodo-Dep">6120089</mods:identifier>
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<mods:identifier id="DF93B7837A50DD458173C9A1DE3D322B" type="ZooBank">AC5D93F0-A61E-4550-8973-C4BF90E30A14</mods:identifier>
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</mods:mods>
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<treatment id="03E287D4FFACFF8B24BB5D12FE26FEE5" LSID="urn:lsid:zoobank.org:act:96FA66C4-399A-4E1E-AB12-B03CC3BC2B91" httpUri="http://treatment.plazi.org/id/03E287D4FFACFF8B24BB5D12FE26FEE5" lastPageId="11" lastPageNumber="330" pageId="2" pageNumber="321">
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<subSubSection id="C3516549FFACFF8224BB5D12FBD8FA82" box="[136,1059,1290,1314]" pageId="2" pageNumber="321" type="nomenclature">
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<paragraph id="8BF436C2FFACFF8224BB5D12FBD8FA82" blockId="2.[136,1059,1290,1314]" box="[136,1059,1290,1314]" pageId="2" pageNumber="321">
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<heading id="D0BC81AEFFACFF8224BB5D12FBD8FA82" box="[136,1059,1290,1314]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="321" reason="2">
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<taxonomicName id="4C4B4D41FFACFF8224BB5D12FECCFA82" ID-CoL="7BWPD" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[136,311,1290,1314]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="species" species="injae" status="sp. nov.">
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<emphasis id="B93FEAD0FFACFF8224BB5D12FECCFA82" bold="true" box="[136,311,1290,1314]" italics="true" pageId="2" pageNumber="321">Thinobius injae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57ABFFACFF82250D5D13FE6AFA82" box="[318,401,1291,1314]" pageId="2" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
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Figs 1, 9-10, 23, 31, 43, 51-52, 73-77
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3516549FFACFF8224E75D36FCDBFA34" pageId="2" pageNumber="321" type="materials_examined">
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<paragraph id="8BF436C2FFACFF8224E75D36FCDBFA34" blockId="2.[136,1058,1325,1428]" pageId="2" pageNumber="321">
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<materialsCitation id="3B233C9FFFACFF8224E75D36FCE1FA34" collectingDate="2010-09-08" collectionCode="NIBR" county="Naerincheon river" elevation="200" latitude="38.066387" location="Gangwon-do" longLatPrecision="19" longitude="128.19084" municipality="Injae-gun district" pageId="2" pageNumber="321" specimenCode="MHNG 1" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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TYPE MATERIAL: HNHM,
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<typeStatus id="54F08860FFACFF8225E75D35FDD3FAE3" box="[468,552,1325,1347]" pageId="2" pageNumber="321" type="holotype">holotype</typeStatus>
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(3); “S-KOREA,
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<location id="8E946019FFACFF8226DB5D35FC9EFAE3" LSID="urn:lsid:plazi:treatment:03E287D4FFACFF8B24BB5D12FE26FEE5:8E946019FFACFF8226DB5D35FC9EFAE3" box="[744,869,1325,1347]" county="Naerincheon river" latitude="38.066387" longLatPrecision="19" longitude="128.19084" municipality="Injae-gun district" name="Gangwon-do" pageId="2" pageNumber="321">Gangwon-do</location>
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,
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<collectingMunicipality id="6B90ACB8FFACFF8227475D35FBE6FAE3" box="[884,1053,1325,1347]" pageId="2" pageNumber="321">Injae-gun district</collectingMunicipality>
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,
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<locationDeviation id="9A462FE0FFACFF8224BB5D5EFF05FAFC" box="[136,254,1350,1372]" location="Inje" pageId="2" pageNumber="321">
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<quantity id="4CB39B27FFACFF8224BB5D5EFF40FAFC" box="[136,187,1350,1372]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="321" unit="km" value="2.0">2 km</quantity>
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E Inje
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</locationDeviation>
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,
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<collectingCounty id="62954E4EFFACFF82253F5D5EFE40FAFC" box="[268,443,1350,1372]" pageId="2" pageNumber="321">Naerincheon river</collectingCounty>
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, sand at W end of bridge,
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<quantity id="4CB39B27FFACFF8226F65D5EFCFFFAFC" box="[709,772,1350,1372]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="321" unit="m" value="200.0">
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<elevation id="0066D1F1FFACFF8226F65D5EFCFFFAFC" box="[709,772,1350,1372]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="321" unit="m" value="200.0">200 m</elevation>
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</quantity>
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, gravel/sand border at water, flotation (2B),
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<geoCoordinate id="EE7F5005FFACFF82252A5D78FE72FAD5" box="[281,393,1376,1398]" degrees="38" direction="north" minutes="03" orientation="latitude" pageId="2" pageNumber="321" precision="15" seconds="59" value="38.066387">38°03'59"N</geoCoordinate>
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,
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<geoCoordinate id="EE7F5005FFACFF8225A55D78FDEBFAD5" box="[406,528,1376,1398]" degrees="128" direction="east" minutes="11" orientation="longitude" pageId="2" pageNumber="321" precision="15" seconds="27" value="128.19084">128°11'27"E</geoCoordinate>
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;
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<date id="FFF51002FFACFF82262E5D78FD71FAD6" box="[541,650,1376,1398]" pageId="2" pageNumber="321" value="2010-09-08">
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<collectingDate id="EFB1E9EAFFACFF82262E5D78FD71FAD6" box="[541,650,1376,1398]" pageId="2" pageNumber="321" value="2010-09-08">08.IX.2010</collectingDate>
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</date>
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; [leg.] Makranczy & al”.
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<typeStatus id="54F08860FFACFF8224E75D67FEC4FA33" box="[212,319,1407,1428]" pageId="2" pageNumber="321" type="paratype">PARATYPES</typeStatus>
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(2):
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<specimenCode id="DBED9EB9FFACFF82255C5D66FE30FA34" box="[367,459,1406,1428]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="321" type="Museum">MHNG 1</specimenCode>
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,
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<collectionCode id="ED5AAE07FFACFF8225EB5D67FDEAFA34" box="[472,529,1407,1428]" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="2" pageNumber="321">NIBR</collectionCode>
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<specimenCount id="9D4DFD4BFFACFF82262B5D66FDC3FA34" box="[536,568,1406,1428]" pageId="2" pageNumber="321" type="female">1♀</specimenCount>
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; same data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3516549FFACFF8624E75DBAFCD4FEF9" lastPageId="6" lastPageNumber="325" pageId="2" pageNumber="321" type="description">
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<paragraph id="8BF436C2FFACFF8324E75DBAFD76F9E6" blockId="2.[136,1059,1441,1655]" lastBlockId="3.[151,1074,1584,1662]" lastPageId="3" lastPageNumber="322" pageId="2" pageNumber="321">DESCRIPTION: Habitus as in Fig. 1. Measurements (n=3): HW = 0.23 (0.22- 0.25); TW = 0.22 (0.21-0.24); PW = 0.23 (0.22-0.26); SW = 0.25 (0.23-0.28); AW = 0.28 (0.25-0.31); HL = 0.21 (0.20-0.22); EL = 0.08 (0.075-0.08); TL = 0.07 (0.065- 0.07); PL = 0.19 (0.18-0.21); SL = 0.36 (0.335-0.39); SC = 0.32 (0.30-0.36); FB = 0.76 (0.71-0.85); BL = 1.43 (1.30-1.53) mm. BODY rather conspicuously bicoloured. Head and pronotum slightly reddish dark brown, abdomen blackish dark brown, elytra medium to light brown but scutellar area broadly dark brown. Legs, mouthparts and FIGS 1-4</paragraph>
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<paragraph id="8BF436C2FFADFF8324A45E57FC43F9DE" blockId="3.[151,1074,1584,1662]" pageId="3" pageNumber="322">
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(1-2) Habitusofnew
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<taxonomicName id="4C4B4D41FFADFF83255F5E57FDE4F9C4" box="[364,543,1615,1637]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B93FEAD0FFADFF83255F5E57FE37F9C5" box="[364,460,1615,1637]" italics="true" pageId="3" pageNumber="322">Thinobius</emphasis>
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species.
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</taxonomicName>
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<taxonomicName id="4C4B4D41FFADFF8326145E48FD8BF9C5" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[551,624,1616,1637]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="injae" status="sp. nov.">
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<emphasis id="B93FEAD0FFADFF8326145E48FD8BF9C5" box="[551,624,1616,1637]" italics="true" pageId="3" pageNumber="322">T. injae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57ABFFADFF8326445E57FD57F9C4" box="[631,684,1615,1636]" pageId="3" pageNumber="322" rank="species">sp. n.</taxonomicNameLabel>
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(1),
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<taxonomicName id="4C4B4D41FFADFF8326EE5E48FCA6F9C5" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[733,861,1616,1637]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="paraminor" status="sp. nov.">
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<emphasis id="B93FEAD0FFADFF8326EE5E48FCA6F9C5" box="[733,861,1616,1637]" italics="true" pageId="3" pageNumber="322">T. paraminor</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57ABFFADFF8327565E57FC61F9C4" box="[869,922,1615,1636]" pageId="3" pageNumber="322" rank="species">sp. n.</taxonomicNameLabel>
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(2). (3-4)
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<typeStatus id="54F08860FFADFF8320315E57FBC9F9C4" box="[1026,1074,1615,1636]" pageId="3" pageNumber="322">Type</typeStatus>
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locality of both
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<taxonomicName id="4C4B4D41FFADFF8325015E71FE81F9DE" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[306,378,1641,1662]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="injae" status="sp. nov.">
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<emphasis id="B93FEAD0FFADFF8325015E71FE81F9DE" box="[306,378,1641,1662]" italics="true" pageId="3" pageNumber="322">P. injae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57ABFFADFF8325B35E71FE4FF9DE" box="[384,436,1641,1662]" pageId="3" pageNumber="322" rank="species">sp. n.</taxonomicNameLabel>
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and
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<taxonomicName id="4C4B4D41FFADFF8325D75E71FD98F9DE" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[484,611,1641,1662]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="paraminor" status="sp. nov.">
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<emphasis id="B93FEAD0FFADFF8325D75E71FD98F9DE" box="[484,611,1641,1662]" italics="true" pageId="3" pageNumber="322">T. paraminor</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57ABFFADFF8326595E71FD65F9DE" box="[618,670,1641,1662]" pageId="3" pageNumber="322" rank="species">sp. n.</taxonomicNameLabel>
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; wide view (3), close-up (4).
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</paragraph>
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<paragraph id="8BF436C2FFAAFF84261F5E0EFD85F98B" blockId="4.[136,1058,1557,1661]" box="[556,638,1557,1579]" pageId="4" pageNumber="323">FIGS 5-8</paragraph>
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<paragraph id="8BF436C2FFAAFF8424BB5E2CFF51F9DD" blockId="4.[136,1058,1557,1661]" pageId="4" pageNumber="323">
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(5-6) Habitusofnew
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<taxonomicName id="4C4B4D41FFAAFF8425695E2CFDF6F9EA" box="[346,525,1588,1610]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B93FEAD0FFAAFF8425695E2CFE41F9EA" box="[346,442,1588,1610]" italics="true" pageId="4" pageNumber="323">Thinobius</emphasis>
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species.
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</taxonomicName>
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<taxonomicName id="4C4B4D41FFAAFF8426205E2DFD4EF9EA" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[531,693,1588,1610]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="schillhammeri" status="sp. nov.">
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<emphasis id="B93FEAD0FFAAFF8426205E2DFD4EF9EA" box="[531,693,1588,1610]" italics="true" pageId="4" pageNumber="323">T. schillhammeri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57ABFFAAFF84268F5E2DFD0BF9EA" box="[700,752,1589,1610]" pageId="4" pageNumber="323" rank="species">sp. n.</taxonomicNameLabel>
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(5),
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<taxonomicName id="4C4B4D41FFAAFF8427135E2DFC73F9EA" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[800,904,1588,1610]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="shavrini" status="sp. nov.">
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<emphasis id="B93FEAD0FFAAFF8427135E2DFC73F9EA" box="[800,904,1588,1610]" italics="true" pageId="4" pageNumber="323">T. shavrini</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57ABFFAAFF8427BD5E2DFC39F9EA" box="[910,962,1589,1610]" pageId="4" pageNumber="323" rank="species">sp. n.</taxonomicNameLabel>
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(6). (7-8) Habitats.
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<taxonomicName id="4C4B4D41FFAAFF8424D95E56FE71F9C3" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[234,394,1613,1635]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="schillhammeri" status="sp. nov.">
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<emphasis id="B93FEAD0FFAAFF8424D95E56FE71F9C3" box="[234,394,1613,1635]" italics="true" pageId="4" pageNumber="323">T. schillhammeri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57ABFFAAFF8425A35E56FE3FF9C3" box="[400,452,1614,1635]" pageId="4" pageNumber="323" rank="species">sp. n.</taxonomicNameLabel>
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, type locality (7),
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<taxonomicName id="4C4B4D41FFAAFF8426475E56FD20F9C3" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[628,731,1613,1635]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="shavrini" status="sp. nov.">
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<emphasis id="B93FEAD0FFAAFF8426475E56FD20F9C3" box="[628,731,1613,1635]" italics="true" pageId="4" pageNumber="323">T. shavrini</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57ABFFAAFF8426D35E56FCEFF9C3" box="[736,788,1614,1635]" pageId="4" pageNumber="323" rank="species">sp. n.</taxonomicNameLabel>
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,
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<typeStatus id="54F08860FFAAFF84272C5E56FC89F9C3" box="[799,882,1614,1635]" pageId="4" pageNumber="323" type="paratype">paratype</typeStatus>
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locality in Siberia (8).
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</paragraph>
|
||
<paragraph id="8BF436C2FFABFF8524A458FBFDCCFCD0" blockId="5.[151,1074,227,1660]" pageId="5" pageNumber="324">first two antennomeres medium to light brown, antennal articles 3-11 dark brown. All main body parts with coriaceous (or rather imbricate on abdomen and some other parts, like sides of head) microsculpture intermixed with fine and dense punctation, making body moderately lustrous; in contrast, clypeus almost unsculptured. Punctation strongest on head, on pronotum very fine, as also on elytra, indistinct on abdomen, latter being strongly microsculptured. Tergite VII not more lustrous than preceding tergites, but on the mostly unexposed tergite VIII microsculpture is faded, so shinier. Pubescence on forebody depressed, very fine, short; on abdomen even finer but somewhat longer setae plus a row of stronger and longer setae on apices of tergites. Abdominal sides and apex with a few darker, much longer and stronger, erect bristles. Strong dark bristle on outer side of supraantennal prominence at anterior border of eye and on posterior edge of vertex adjacent to temples, similar ones right behind anterior pronotal corner and another before posterior corner. Direction of setation on sides of head anterior, on disc mostly medial, but posterior in midline and near hind edge; on pronotal sides medial, most of disc antero-medial, midline anterior, posterior edge lateral. Posterior edge of vertex as well as middle of anterior pronotal edge with conspicuously long setae. Setation on abdomen posteriorly directed, but slightly postero-lateral on elytra. Antennae, legs and mouthparts with very short, not conspicuous setation, except for the stiff, darker bristle slightly distad from middle of each tibia and ones near apices of femora. Elytra with three long, stiff and erect lateral setae, posterior one slightly closer to middle.</paragraph>
|
||
<paragraph id="8BF436C2FFABFF8524D05B6FFF02FB29" blockId="5.[151,1074,227,1660]" pageId="5" pageNumber="324">Head (Fig. 23) 1.15x wider than long, temples almost straight, with only a gentle curve anteriorly, but narrowly rounded on posterior part. Vertex rather flat, supraantennal prominences strongly developed, anterior edges rather shiny. Anterior border of neck not marked with groove, only by change of microsculpture. Frontoclypeal suture appearing as fine, often shinier groove and dark line connecting supraantennal tubercles, similarly darker spots situated at both sides on middle of vertex. ANTENNAE (Fig. 51) with antennomeres 4-8 moderately to strongly transverse, as opposed to terminal antennomeres (Fig. 52) with article 9 as long as broad and 10 elongate.</paragraph>
|
||
<paragraph id="8BF436C2FFABFF8524D05C89FC1AF9BE" blockId="5.[151,1074,227,1660]" pageId="5" pageNumber="324">Pronotum (Fig. 9) slightly transverse, 1.30x wider than long, not appearing wider than head, with narrowly rounded anterior and rather narrowly rounded posterior corners. Anterior margin almost straight, as also sides (parallel) with slight concavity behind middle; posterior margin strongly but not evenly arched, almost straight in outer thirds, giving pronotum a faint pentagonal feel. No consistent and characteristic impressions, but both sides of anterior half of disc gently impressed as well as sides behind middle. Pronotal marginal bead visible on posterior margin. Scutellum large, setose and very strongly sculptured (Fig. 31). ELYTRA (Fig. 10) long, one elytron more than twice as long as broad, gently curved in cross-section, parallel-sided (very little dilation posteriorly). Shoulders moderately developed. Along suture with thin marginal bead, but broader right behind scutellum. Sutural corners broadly rounded. LEGS of medium length, tarsal lobes thin but relatively long (but not reaching apex of last tarsomere), tibiae strongly fusiform, appear a lot wider in middle than at ends.</paragraph>
|
||
<paragraph id="8BF436C2FFABFF8524D05E3EFC7EF9DC" blockId="5.[151,1074,227,1660]" pageId="5" pageNumber="324">Abdomen (Fig. 43) very weakly fusiform, sides more or less straight and parallel; widest at posterior 1/3, very slightly less wide than elytra at broadest point. Posterior margin of tergite VII with moderately broad palisade fringe.</paragraph>
|
||
<paragraph id="8BF436C2FFA8FF8624E758FCFCD4FEF9" blockId="6.[136,1059,227,345]" pageId="6" pageNumber="325">PRIMARY AND SECONDARY SEXUAL FEATURES: Sexes not appearing different in regular dorsal view. Posterior corners of tergite VIII (similar in both sexes) gently protruding, posterior edge otherwise straight. Male sternite VIII as in Fig. 73, tergite IX as in Fig. 74, sternite IX as in Fig. 75. Aedeagus as in Figs 76-77.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3516549FFA8FF8624E75974FD5FFE03" pageId="6" pageNumber="325" type="etymology">
|
||
<paragraph id="8BF436C2FFA8FF8624E75974FD5FFE03" blockId="6.[136,1058,363,419]" pageId="6" pageNumber="325">
|
||
ETYMOLOGY: The species is named after the city near the
|
||
<typeStatus id="54F08860FFA8FF86276F5974FC71FE24" box="[860,906,364,388]" pageId="6" pageNumber="325">type</typeStatus>
|
||
locality, deliberately in the older spelling (noun in apposition).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3516549FFA8FF8624E759AEFBECFC25" pageId="6" pageNumber="325" type="discussion">
|
||
<paragraph id="8BF436C2FFA8FF8624E759AEFBECFC25" blockId="6.[136,1059,437,902]" pageId="6" pageNumber="325">
|
||
REMARKS: The male
|
||
<typeStatus id="54F08860FFA8FF86258F59ADFDE2FE6D" box="[444,537,437,461]" pageId="6" pageNumber="325" type="holotype">holotype</typeStatus>
|
||
has slightly larger body size than the
|
||
<specimenCount id="9D4DFD4BFFA8FF86278859AEFF3BFE4D" pageId="6" pageNumber="325" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
. However, with only
|
||
<specimenCount id="9D4DFD4BFFA8FF86259959CCFDAEFE4C" box="[426,597,468,492]" pageId="6" pageNumber="325" type="generic">three specimens</specimenCount>
|
||
known this does not allow for conclusions on possible sexual differences. This taxon belongs in the
|
||
<taxonomicName id="4C4B4D41FFA8FF86272459EDFC50FDAC" authorityName="Kraatz" authorityYear="1857" box="[791,939,500,524]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="species" species="delicatulus">
|
||
<emphasis id="B93FEAD0FFA8FF86272459EDFC50FDAC" box="[791,939,500,524]" italics="true" pageId="6" pageNumber="325">T. delicatulus</emphasis>
|
||
</taxonomicName>
|
||
group. Its aedeagal structures are very similar to the sibling species
|
||
<taxonomicName id="4C4B4D41FFA8FF8626CE5A0CFBD9FD8B" authority="Kraatz, 1857" authorityName="Kraatz" authorityYear="1857" box="[765,1058,531,555]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="species" species="delicatulus">
|
||
<emphasis id="B93FEAD0FFA8FF8626CE5A0CFC75FD8B" box="[765,910,531,555]" italics="true" pageId="6" pageNumber="325">T. delicatulus</emphasis>
|
||
Kraatz, 1857
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4B4D41FFA8FF8624895A2CFE04FDEB" authority="Bernhauer, 1940" authorityName="Bernhauer" authorityYear="1940" box="[186,511,563,587]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="species" species="hummleri">
|
||
<emphasis id="B93FEAD0FFA8FF8624895A2CFEC4FDEB" box="[186,319,563,587]" italics="true" pageId="6" pageNumber="325">T. hummleri</emphasis>
|
||
Bernhauer, 1940
|
||
</taxonomicName>
|
||
. During the checking of
|
||
<taxonomicName id="4C4B4D41FFA8FF8627115A2BFC76FDEB" authorityName="Kiesenwetter" authorityYear="1844" box="[802,909,563,587]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FEAD0FFA8FF8627115A2BFC76FDEB" box="[802,909,563,587]" italics="true" pageId="6" pageNumber="325">Thinobius</emphasis>
|
||
</taxonomicName>
|
||
taxon names described from the East Palaearctic, it was discovered that the specimens Smetana identified as
|
||
<taxonomicName id="4C4B4D41FFA8FF86252B5A6BFE51FD2A" authorityName="Kraatz" authorityYear="1857" box="[280,426,626,650]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="species" species="delicatulus">
|
||
<emphasis id="B93FEAD0FFA8FF86252B5A6BFE51FD2A" box="[280,426,626,650]" italics="true" pageId="6" pageNumber="325">T. delicatulus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F35C7652FFA8FF8625C25A6AFDA2FD2A" box="[497,601,626,650]" name="Mongolia" pageId="6" pageNumber="325">Mongolia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFDA4B33FFA8FF8626585A6AFCF7FD2A" author="SMETANA, A." box="[619,780,626,650]" pageId="6" pageNumber="325" pagination="153 - 179" refId="ref12679" refString="SMETANA, A. 1975. Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei. 340. Staphylinidae IV. Unterfamilien Omaliinae bis Staphylininae (Coleoptera). Acta Zoologica Academiae Scientiarum Hungaricae 21 (1 - 2): 153 - 179." type="journal article" year="1975">Smetana, 1975</bibRefCitation>
|
||
) are in fact
|
||
<taxonomicName id="4C4B4D41FFA8FF8627AB5A6BFBE0FD2A" authorityName="Bernhauer" authorityYear="1940" box="[920,1051,626,650]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="species" species="hummleri">
|
||
<emphasis id="B93FEAD0FFA8FF8627AB5A6BFBE0FD2A" box="[920,1051,626,650]" italics="true" pageId="6" pageNumber="325">T. hummleri</emphasis>
|
||
</taxonomicName>
|
||
. The proper distinguishing features were not known at the time. Comparison of the new species to the European relatives also revealed that a pair of specimens with conspicuously transversal antennomeres (data:
|
||
<collectingCountry id="F35C7652FFA8FF86267F5AC8FD57FD48" box="[588,684,720,744]" name="Romania" pageId="6" pageNumber="325">Romania</collectingCountry>
|
||
, jud.
|
||
<collectingRegion id="498FF820FFA8FF8626C15AC9FC90FD48" box="[754,875,721,744]" country="Romania" name="Maramures" pageId="6" pageNumber="325">Maramureş</collectingRegion>
|
||
, Valea Vişeului, [river] Vişeu,
|
||
<quantity id="4CB39B27FFA8FF8625145AE8FE8DFCA8" box="[295,374,752,776]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="325" unit="km" value="1.5">1.5 km</quantity>
|
||
S Tisa confluence, W bank, sandy shoal N hanging bridge,
|
||
<quantity id="4CB39B27FFA8FF8624BB5B17FF37FC87" box="[136,204,783,807]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="325" unit="m" value="360.0">360 m</quantity>
|
||
, from gravelly spots in sandbank, after high water, flotation (305),
|
||
<geoCoordinate id="EE7F5005FFA8FF8627AD5B17FBE0FC87" box="[926,1051,783,807]" degrees="47" direction="north" minutes="54" orientation="latitude" pageId="6" pageNumber="325" precision="15" seconds="04" value="47.90111">47°54'04"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7F5005FFA8FF8624BB5B36FEF9FCE6" box="[136,258,814,838]" degrees="24" direction="east" minutes="09" orientation="longitude" pageId="6" pageNumber="325" precision="15" seconds="33" value="24.159166">24°09'33"E</geoCoordinate>
|
||
,
|
||
<date id="FFF51002FFA8FF8625245B36FE74FCE6" box="[279,399,814,838]" pageId="6" pageNumber="325" value="2007-06-11">11.VI.2007</date>
|
||
, [leg.] Gy. Makranczy) actually belong to
|
||
<taxonomicName id="4C4B4D41FFA8FF8627B55B28FBE7FCE6" authorityName="Kraatz" authorityYear="1857" box="[902,1052,814,839]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="species" species="delicatulus">
|
||
<emphasis id="B93FEAD0FFA8FF8627B55B28FBE7FCE6" box="[902,1052,814,839]" italics="true" pageId="6" pageNumber="325">T. delicatulus</emphasis>
|
||
</taxonomicName>
|
||
, which confirms its occurance in
|
||
<collectingCountry id="F35C7652FFA8FF8625D75B56FDBFFCC6" box="[484,580,846,870]" name="Romania" pageId="6" pageNumber="325">Romania</collectingCountry>
|
||
. Shapes of individual antennomeres turn out to be rather variable in this genus, making this character difficult to use in diagnosis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3516549FFA8FF8624E75B81FEEBFC5B" pageId="6" pageNumber="325" type="distribution">
|
||
<paragraph id="8BF436C2FFA8FF8624E75B81FBFAFC10" blockId="6.[212,1025,920,945]" box="[212,1025,920,945]" pageId="6" pageNumber="325">
|
||
DISTRIBUTION: The species is known only from the
|
||
<typeStatus id="54F08860FFA8FF8627305B81FCCAFC11" box="[771,817,921,945]" pageId="6" pageNumber="325">type</typeStatus>
|
||
locality (Figs 3-4).
|
||
</paragraph>
|
||
<paragraph id="8BF436C2FFA8FF8624E75BDCFEEBFC5B" blockId="6.[136,1059,963,1019]" pageId="6" pageNumber="325">BIONOMICS: Seems to be associated with lower rivers, more open, sunny gravelbanks.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3516549FFA8FF8624BB5C01FC4CFBF1" pageId="6" pageNumber="325" type="reference_group">
|
||
<paragraph id="8BF436C2FFA8FF8624BB5C01FC4CFBF1" blockId="6.[136,1059,1048,1212]" pageId="6" pageNumber="325">
|
||
<treatmentCitationGroup id="AB5B11ECFFA8FF8624BB5C01FC4CFBF1" pageId="6" pageNumber="325">
|
||
<taxonomicName id="4C4B4D41FFA8FF8624BB5C01FE0BFB90" ID-CoL="56GQF" authority="Naomi, 1995" authorityName="Naomi" authorityYear="1995" box="[136,496,1048,1073]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="species" species="ootsukai">
|
||
<emphasis id="B93FEAD0FFA8FF8624BB5C01FEA6FB91" bold="true" box="[136,349,1049,1073]" italics="true" pageId="6" pageNumber="325">Thinobius ootsukai</emphasis>
|
||
<bibRefCitation id="EFDA4B33FFA8FF8625565C00FE0BFB90" author="NAOMI, S. - I." box="[357,496,1048,1072]" pageId="6" pageNumber="325" pagination="17 - 25" refId="ref12642" refString="NAOMI, S. - I. 1995. Taxonomic study on the genus Thinobius Kiesenwetter from Japan (Coleoptera, Staphylinidae). Entomological Review of Japan 50 (1): 17 - 25." type="journal article" year="1995">Naomi, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
Figs 11-12, 24, 37-39, 64-66, 81
|
||
<treatmentCitation id="0AEA10D3FFA8FF8624BB5C23FD31FBF1" author="NAOMI, S. - I." box="[136,714,1083,1105]" page="22" pageId="6" pageNumber="325" year="1995">
|
||
<taxonomicName id="4C4B4D41FFA8FF8624BB5C23FD31FBF1" ID-CoL="56GQF" authority="Naomi, 1995: 22" authorityName="Naomi" authorityPageNumber="22" authorityYear="1995" box="[136,714,1083,1105]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="species" species="ootsukai" subGenus="Platyderothinophilus">
|
||
<emphasis id="B93FEAD0FFA8FF8624BB5C23FDDBFBF1" box="[136,544,1083,1105]" italics="true" pageId="6" pageNumber="325">Thinobius (Platyderothinophilus) ootsukai</emphasis>
|
||
<bibRefCitation id="EFDA4B33FFA8FF8626145C23FD31FBF1" author="NAOMI, S. - I." box="[551,714,1083,1105]" pageId="6" pageNumber="325" pagination="17 - 25" refId="ref12642" refString="NAOMI, S. - I. 1995. Taxonomic study on the genus Thinobius Kiesenwetter from Japan (Coleoptera, Staphylinidae). Entomological Review of Japan 50 (1): 17 - 25." type="journal article" year="1995">Naomi, 1995: 22</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
. –
|
||
<treatmentCitation id="0AEA10D3FFA8FF8626DA5C24FC4AFBF1" author="HERMAN, L. H." box="[745,945,1083,1105]" page="1754" pageId="6" pageNumber="325" year="2001">
|
||
<bibRefCitation id="EFDA4B33FFA8FF8626DA5C24FC4AFBF1" author="HERMAN, L. H." box="[745,945,1083,1105]" pageId="6" pageNumber="325" pagination="1 - 4218" refId="ref12443" refString="HERMAN, L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1785 to the end of the second Millennium. Bulletin of the American Museum of Natural History 265: 1 - 4218." type="journal article" year="2001">Herman, 2001: 1754</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3516549FFA8FF8624E75C43FE4CFB1C" pageId="6" pageNumber="325" type="materials_examined">
|
||
<paragraph id="8BF436C2FFA8FF8624E75C43FE4CFB1C" blockId="6.[136,1059,1048,1212]" pageId="6" pageNumber="325">
|
||
<materialsCitation id="3B233C9FFFA8FF8624E75C43FE49FB1C" collectingDate="1990-05-27" collectionCode="CMIC" collectorName="I. Oostsuka" country="Japan" county="Honshu" determinerName="Makranczy" location="Mt. Kiyosumi" municipality="Pref" pageId="6" pageNumber="325" specimenCount="3" stateProvince="Chiba" typeStatus="holotype">
|
||
TYPE MATERIAL EXAMINED:
|
||
<collectionCode id="ED5AAE07FFA8FF8625D15C42FDE4FBD0" box="[482,543,1114,1136]" pageId="6" pageNumber="325">CMIC</collectionCode>
|
||
,
|
||
<typeStatus id="54F08860FFA8FF86261D5C42FD79FBD0" box="[558,642,1114,1136]" pageId="6" pageNumber="325" type="holotype">holotype</typeStatus>
|
||
(3); “[
|
||
<collectingCountry id="F35C7652FFA8FF8626E35C42FCE3FBD0" box="[720,792,1114,1136]" name="Japan" pageId="6" pageNumber="325">JAPAN</collectingCountry>
|
||
:
|
||
<collectingCounty id="62954E4EFFA8FF8627145C42FC89FBD0" box="[807,882,1114,1136]" pageId="6" pageNumber="325">Honshu</collectingCounty>
|
||
,]
|
||
<collectingRegion id="498FF820FFA8FF8627BB5C42FC3AFBD0" box="[904,961,1114,1136]" country="Japan" name="Chiba" pageId="6" pageNumber="325">Chiba</collectingRegion>
|
||
<collectingMunicipality id="6B90ACB8FFA8FF8627FA5C42FC0AFBD0" box="[969,1009,1114,1136]" pageId="6" pageNumber="325">Pref</collectingMunicipality>
|
||
;
|
||
<location id="8E946019FFA8FF8620335C43FF1CFB29" LSID="urn:lsid:plazi:treatment:03E287D4FFACFF8B24BB5D12FE26FEE5:8E946019FFA8FF8620335C43FF1CFB29" country="Japan" county="Honshu" municipality="Pref" name="Mt. Kiyosumi" pageId="6" pageNumber="325" stateProvince="Chiba">Mt. Kiyosumi</location>
|
||
;
|
||
<date id="FFF51002FFA8FF8624C65C6BFEA3FB29" box="[245,344,1139,1161]" pageId="6" pageNumber="325" value="1990-05-27">
|
||
<collectingDate id="EFB1E9EAFFA8FF8624C65C6BFEA3FB29" box="[245,344,1139,1161]" pageId="6" pageNumber="325" value="1990-05-27">27.V.1990</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26BE5314FFA8FF8625555C6CFE2EFB29" box="[358,469,1139,1161]" pageId="6" pageNumber="325">I. Oostsuka</collectorName>
|
||
col. \
|
||
<typeStatus id="54F08860FFA8FF8626245C6BFD8BFB29" box="[535,624,1139,1161]" pageId="6" pageNumber="325" type="holotype">Holotype</typeStatus>
|
||
;
|
||
<taxonomicName id="4C4B4D41FFA8FF86264D5C6BFCC2FB29" authorityName="Naomi" authorityYear="1995" box="[638,825,1139,1161]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="species" species="ootsukai">Thinobius ootsukai</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFDA4B33FFA8FF8627745C6BFC3AFB29" author="NAOMI, S. - I." box="[839,961,1139,1161]" pageId="6" pageNumber="325" pagination="17 - 25" refId="ref12642" refString="NAOMI, S. - I. 1995. Taxonomic study on the genus Thinobius Kiesenwetter from Japan (Coleoptera, Staphylinidae). Entomological Review of Japan 50 (1): 17 - 25." type="journal article" year="1995">Naomi 1995</bibRefCitation>
|
||
. \ CBM - ZI; 33876 \
|
||
<typeStatus id="54F08860FFA8FF8624C85C95FEA4FB03" box="[251,351,1165,1187]" pageId="6" pageNumber="325" type="holotype">Holotypus</typeStatus>
|
||
;
|
||
<taxonomicName id="4C4B4D41FFA8FF86255F5C95FE36FB03" authorityName="Kiesenwetter" authorityYear="1844" box="[364,461,1165,1187]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="genus">Thinobius</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C4B4D41FFA8FF8625E95C95FD88FB03" authority="Naomi" authorityName="Naomi" authorityYear="1995" box="[474,627,1165,1187]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="species" species="ootsukai">ootsukai Naomi</taxonomicName>
|
||
; ver. Makranczy, 2011 \
|
||
<taxonomicName id="4C4B4D41FFA8FF8627505C95FC3FFB03" authorityName="Kiesenwetter" authorityYear="1844" box="[867,964,1165,1187]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="genus">Thinobius</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C4B4D41FFA8FF8627E25C95FF32FB1C" authority="Naomi" authorityName="Naomi" authorityYear="1995" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="species" species="ootsukai">ootsukai Naomi</taxonomicName>
|
||
; det.
|
||
<determinerName id="7E28DA06FFA8FF8625325CBEFE90FB1C" box="[257,363,1190,1212]" pageId="6" pageNumber="325">Makranczy</determinerName>
|
||
, 2011”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3516549FFA8FF8924E75CD4FE6EFA33" lastPageId="9" lastPageNumber="328" pageId="6" pageNumber="325" type="description">
|
||
<paragraph id="8BF436C2FFA8FF8724E75CD4FD68F98A" blockId="6.[136,1059,1228,1661]" lastBlockId="7.[151,1074,1556,1659]" lastPageId="7" lastPageNumber="326" pageId="6" pageNumber="325">
|
||
REDESCRIPTION: Measurements (n=1): HW = 0.33; TW = 0.31; PW = 0.38; SW = 0.44; AW = 0.46; HL = 0.27; EL = 0.11; TL = 0.075; PL = 0.295; SL = 0.54; SC = 0.50; FB = 1.08; BL =
|
||
<quantity id="4CB39B27FFA8FF8625AA5D12FE06FA82" box="[409,509,1290,1314]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" pageId="6" pageNumber="325" unit="mm" value="1.93">1.93 mm</quantity>
|
||
. BODY unicoloured. Head dark brown with only supraantennal tubercles appearing lighter. Pronotum dark brown, elytra somewhat lighter (but unicolorous) dark brown. Abdomen dark brown to blackish dark brown. Mouthparts and antennae slightly reddish dark brown, legs medium to dark brown, tarsi somewhat lighter. All main body parts with coriaceous microsculpture intermixed with fine and dense punctation, making body moderately lustrous. Punctation on head (especially anterior half of vertex) stronger, moderately deep, pronotum slightly less strong and more dense, elytra even finer and denser, on abdomen (Fig. 37) more indistinct; on tergite VII both coriaceous microsculpture and punctation fading posteriorly (as also on mostly unexposed tergite VIII), making tip of abdomen much shinier than rest. Pubescence on forebody rather depressed, short, fine and dense (on elytra the most dense), on abdomen somewhat less dense but short and extremely fine setae, FIGS 9-13
|
||
</paragraph>
|
||
<paragraph id="8BF436C2FFA9FF8724A45E2BFDD3F9DB" blockId="7.[151,1074,1556,1659]" pageId="7" pageNumber="326">
|
||
(9-10)
|
||
<emphasis id="B93FEAD0FFA9FF8724E85E2BFE88F9E9" box="[219,371,1587,1609]" italics="true" pageId="7" pageNumber="326">Thinobiusinjae</emphasis>
|
||
<taxonomicNameLabel id="A20C57ABFFA9FF87254F5E2BFE49F9E8" box="[380,434,1587,1608]" pageId="7" pageNumber="326" rank="species">sp. n.</taxonomicNameLabel>
|
||
; headandpronotum (9), elytra (10). (11-12)
|
||
<taxonomicName id="4C4B4D41FFA9FF8727455E2CFBD1F9E9" authority="Naomi" authorityName="Naomi" authorityYear="1995" box="[886,1066,1587,1609]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="ootsukai">
|
||
<emphasis id="B93FEAD0FFA9FF8727455E2CFC1AF9E9" box="[886,993,1587,1609]" italics="true" pageId="7" pageNumber="326">T. ootsukai</emphasis>
|
||
Naomi
|
||
</taxonomicName>
|
||
; head and pronotum (11), eye (12). (13)
|
||
<taxonomicName id="4C4B4D41FFA9FF8726255E55FD86F9C2" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[534,637,1612,1634]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="shavrini" status="sp. nov.">
|
||
<emphasis id="B93FEAD0FFA9FF8726255E55FD86F9C2" box="[534,637,1612,1634]" italics="true" pageId="7" pageNumber="326">T. shavrini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57ABFFA9FF8726B05E55FD4CF9C2" box="[643,695,1613,1634]" pageId="7" pageNumber="326" rank="species">sp. n.</taxonomicNameLabel>
|
||
; side of head. Scale bar =
|
||
<quantity id="4CB39B27FFA9FF87278A5E54FBF4F9C2" box="[953,1039,1612,1634]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="326" unit="mm" value="0.05">0.05 mm</quantity>
|
||
for 12, 0.1 mmfor 9-10, 13, 0.14 mmfor 11.
|
||
</paragraph>
|
||
<paragraph id="8BF436C2FFA6FF8826135E00FD71F98D" blockId="8.[136,1059,1559,1662]" box="[544,650,1559,1581]" pageId="8" pageNumber="327">FIGS 14-18</paragraph>
|
||
<footnote id="E8502ACCFFA6FF8824BB5E2EFC6FF9DE" pageId="8" pageNumber="327">
|
||
<paragraph id="8BF436C2FFA6FF8824BB5E2EFC6FF9DE" blockId="8.[136,1059,1559,1662]" pageId="8" pageNumber="327">
|
||
(14)
|
||
<taxonomicName id="4C4B4D41FFA6FF8824845E2EFE96F9EC" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[183,365,1590,1612]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="327" phylum="Arthropoda" rank="species" species="shavrini" status="sp. nov.">
|
||
<emphasis id="B93FEAD0FFA6FF8824845E2EFE96F9EC" box="[183,365,1590,1612]" italics="true" pageId="8" pageNumber="327">Thinobius shavrini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57ABFFA6FF8825405E2EFE53F9EB" box="[371,424,1590,1611]" pageId="8" pageNumber="327" rank="species">sp. n.</taxonomicNameLabel>
|
||
; elytra. (15-16)
|
||
<taxonomicName id="4C4B4D41FFA6FF88267B5E2FFD12F9EC" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[584,745,1590,1612]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="327" phylum="Arthropoda" rank="species" species="schillhammeri" status="sp. nov.">
|
||
<emphasis id="B93FEAD0FFA6FF88267B5E2FFD12F9EC" box="[584,745,1590,1612]" italics="true" pageId="8" pageNumber="327">T. schillhammeri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57ABFFA6FF8826C35E2EFCDEF9EB" box="[752,805,1590,1611]" pageId="8" pageNumber="327" rank="species">sp. n.</taxonomicNameLabel>
|
||
; head and pronotum (15), side of head (16). (17)
|
||
<taxonomicName id="4C4B4D41FFA6FF88255B5E48FDD4F9C5" authority="Naomi" authorityName="Naomi" authorityYear="1995" box="[360,559,1615,1637]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="327" phylum="Arthropoda" rank="species" species="yabakeinis">
|
||
<emphasis id="B93FEAD0FFA6FF88255B5E48FE13F9C5" box="[360,488,1615,1637]" italics="true" pageId="8" pageNumber="327">T. yabakeinis</emphasis>
|
||
Naomi
|
||
</taxonomicName>
|
||
; head and pronotum. (18)
|
||
<taxonomicName id="4C4B4D41FFA6FF8827075E48FC48F9C5" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[820,947,1616,1637]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="327" phylum="Arthropoda" rank="species" species="paraminor" status="sp. nov.">
|
||
<emphasis id="B93FEAD0FFA6FF8827075E48FC48F9C5" box="[820,947,1616,1637]" italics="true" pageId="8" pageNumber="327">T. paraminor</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57ABFFA6FF8827895E48FC15F9C5" box="[954,1006,1616,1637]" pageId="8" pageNumber="327" rank="species">sp. n.</taxonomicNameLabel>
|
||
; side of head (11). Scale bar = 0.1 mm for 16, 18, 0.2 mm for 15, 0.25 mm for 14, 17.
|
||
</paragraph>
|
||
</footnote>
|
||
<paragraph id="8BF436C2FFA7FF8924A458FBFF13FDD4" blockId="9.[151,1074,227,1291]" pageId="9" pageNumber="328">apices of tergites with a row of conspicuously long setae, abdominal sides and apex with a few darker and stronger bristles. Strong dark bristle on outer side of supraantennal prominence at anterior border of eye and on posterior edge of vertex adjacent to temples, similar ones right behind anterior pronotal corner and on side at 3/5 length. Setation on head medially directed except distinctly posterior on anterior part of vertex (a bristle on each side). Pronotal setation mostly antero-medially directed, but near posterior and side edges and at midline direction more anterior. Posterior edge of vertex as well as middle of anterior pronotal edge with rather large setae. Elytra and abdomen with posteriorly directed setae. Antennae, legs and mouthparts with very short, not conspicuous setation, except for the stiff, darker bristle around middle of each tibia and ones near apices of femora. Sides of elytra with three such stiff but inconspicuous setae distributed on side, the middle one shifted towards apex from middle.</paragraph>
|
||
<paragraph id="8BF436C2FFA7FF8924D05A64FCB7FCF0" blockId="9.[151,1074,227,1291]" pageId="9" pageNumber="328">Head (Fig. 11) 1.40x wider than long, temples shorter than eyes (Fig. 12), gently curved anteriorly, narrowly rounded posteriorly. Middle of vertex with slight impression, supraantennal tubercles moderately developed. Anterior border of neck not marked with groove, only by change of microsculpture. Frontoclypeal suture appearing as fine, often shinier groove and darker line connecting supraantennal tubercles. ANTENNAE moderately elongate (Fig. 39), antennomeres 4 and 6 smaller than adjacent ones and as long as broad, other antennomeres slightly elongate.</paragraph>
|
||
<paragraph id="8BF436C2FFA7FF8924D05B41FDE8FB0B" blockId="9.[151,1074,227,1291]" pageId="9" pageNumber="328">Pronotum transverse, 1.40x wider than long, significantly wider than head, anterior margin almost straight, anterior corners noticeable but obtuse-angled, sides strongly arched, forming a rather unbroken arch with posterior margin, posterior angles barely marked. Pronotal midline slightly elevated, rest of middle of disc gently impressed. Pronotal marginal bead visible on posterior margin. Scutellum large, setose and dull. ELYTRA (Fig. 38) long, one elytron slightly more than twice as long as broad, gently curved in cross-section and parallel-sided (very little dilation posteriorly). Shoulders moderately developed. Along suture only with traces of marginal bead, mostly at broadly rounded sutural corners. LEGS of medium length, tarsal lobes thin but relatively long (not reaching apex of last tarsomere), tibiae slightly fusiform, appear much wider in middle than at ends.</paragraph>
|
||
<paragraph id="8BF436C2FFA7FF8924D05CABFC94FAAB" blockId="9.[151,1074,227,1291]" pageId="9" pageNumber="328">Abdomen very weakly fusiform, sides more or less straight and parallel; widest in the middle, with just about the same width as elytra at broadest point. Posterior margin of tergite VII with palisade fringe, strongly widening in middle.</paragraph>
|
||
<paragraph id="8BF436C2FFA7FF8924D05D04FE6EFA33" blockId="9.[151,1074,1307,1427]" pageId="9" pageNumber="328">PRIMARY AND SECONDARY SEXUAL FEATURES: Posterior corners of tergite VIII (similar in both sexes) gently protruding (Fig. 24), posterior edge otherwise straight. Male sternite VIII as in Fig. 64, tergite IX as in Fig. 65, sternite IX as in Fig. 66. Aedeagus as in Fig. 81.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3516549FFA7FF8A24D05DBCFD71F989" lastPageId="10" lastPageNumber="329" pageId="9" pageNumber="328" type="discussion">
|
||
<paragraph id="8BF436C2FFA7FF8924D05DBCFF1EF9DB" blockId="9.[151,1074,1443,1659]" pageId="9" pageNumber="328">
|
||
<materialsCitation id="3B233C9FFFA7FF8924D05DBCFF25F9DB" collectionCode="CMIC, KUEC" pageId="9" pageNumber="328" specimenCount="6" specimenCount-male="1" typeStatus="paratype">
|
||
REMARKS: The description also mentions
|
||
<specimenCount id="9D4DFD4BFFA7FF89269D5DBCFCEAFA1B" box="[686,785,1443,1467]" pageId="9" pageNumber="328" type="male">one male</specimenCount>
|
||
<typeStatus id="54F08860FFA7FF8927295DBCFC8DFA1C" box="[794,886,1444,1468]" pageId="9" pageNumber="328" type="paratype">paratype</typeStatus>
|
||
. As the
|
||
<typeStatus id="54F08860FFA7FF8927E75DBBFBCAFA1B" box="[980,1073,1443,1467]" pageId="9" pageNumber="328" type="holotype">holotype</typeStatus>
|
||
was received undissected, obviously the aedeagus drawing in the original description must be based on the
|
||
<typeStatus id="54F08860FFA7FF8925B35DFCFE27FA5C" box="[384,476,1508,1532]" pageId="9" pageNumber="328" type="paratype">paratype</typeStatus>
|
||
; its whereabouts are unknown. It was neither deposited with the
|
||
<typeStatus id="54F08860FFA7FF8925325E1BFEA5F9BB" box="[257,350,1539,1563]" pageId="9" pageNumber="328" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED5AAE07FFA7FF8925475E1BFE43F9BB" box="[372,440,1539,1563]" pageId="9" pageNumber="328">CMIC</collectionCode>
|
||
) nor with the
|
||
<typeStatus id="54F08860FFA7FF89265A5E1CFD34F9BC" box="[617,719,1540,1564]" pageId="9" pageNumber="328" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4B4D41FFA7FF8926CC5E1CFC69F9BB" authorityName="Naomi" authorityYear="1995" box="[767,914,1539,1563]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="328" phylum="Arthropoda" rank="species" species="yabakeinis">
|
||
<emphasis id="B93FEAD0FFA7FF8926CC5E1CFC69F9BB" box="[767,914,1539,1563]" italics="true" pageId="9" pageNumber="328">T. yabakeinis</emphasis>
|
||
</taxonomicName>
|
||
, of which no previously dissected male specimen was included in the available set of
|
||
<typeStatus id="54F08860FFA7FF8927F85E3CFBCAF99C" box="[971,1073,1572,1596]" pageId="9" pageNumber="328" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<collectionCode id="ED5AAE07FFA7FF8924935E5BFF12F9FB" box="[160,233,1603,1627]" country="Japan" httpUri="http://grbio.org/cool/txbq-e9hf" name="Kyushu University Entomology Collection" pageId="9" pageNumber="328">KUEC</collectionCode>
|
||
) although an aedeagus drawing appears among the illustrations in
|
||
<bibRefCitation id="EFDA4B33FFA7FF8927DA5E5BFF25F9DB" author="NAOMI, S. - I." pageId="9" pageNumber="328" pagination="17 - 25" refId="ref12642" refString="NAOMI, S. - I. 1995. Taxonomic study on the genus Thinobius Kiesenwetter from Japan (Coleoptera, Staphylinidae). Entomological Review of Japan 50 (1): 17 - 25." type="journal article" year="1995">Naomi (1995)</bibRefCitation>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF436C2FFA4FF8A26135E0BFD71F989" blockId="10.[136,1058,1555,1658]" box="[544,650,1555,1577]" pageId="10" pageNumber="329">FIGS 19-24</paragraph>
|
||
</subSubSection>
|
||
<footnote id="E8502ACCFFA4FF8A24BB5E29FC24F9DA" pageId="10" pageNumber="329">
|
||
<paragraph id="8BF436C2FFA4FF8A24BB5E29FC24F9DA" blockId="10.[136,1058,1555,1658]" pageId="10" pageNumber="329">
|
||
(19)
|
||
<emphasis id="B93FEAD0FFA4FF8A24855E29FE78F9E7" box="[182,387,1585,1607]" italics="true" pageId="10" pageNumber="329">Thinobiusparaminor</emphasis>
|
||
<taxonomicNameLabel id="A20C57ABFFA4FF8A25BB5E2AFE40F9E7" box="[392,443,1586,1607]" pageId="10" pageNumber="329" rank="species">sp. n.</taxonomicNameLabel>
|
||
; pronotum. (20-22)
|
||
<taxonomicName id="4C4B4D41FFA4FF8A264C5E2AFD1EF9E7" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[639,741,1585,1607]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="329" phylum="Arthropoda" rank="species" species="shavrini" status="sp. nov.">
|
||
<emphasis id="B93FEAD0FFA4FF8A264C5E2AFD1EF9E7" box="[639,741,1585,1607]" italics="true" pageId="10" pageNumber="329">T. shavrini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57ABFFA4FF8A26D85E2AFCE5F9E7" box="[747,798,1586,1607]" pageId="10" pageNumber="329" rank="species">sp. n.</taxonomicNameLabel>
|
||
; head (20), pronotum (21), abdominal tip, dorsal view (22). (23)
|
||
<taxonomicName id="4C4B4D41FFA4FF8A25C85E54FDBFF9C1" authority="Makranczy & H-, 2014" authorityName="Makranczy & H-" authorityYear="2014" box="[507,580,1612,1633]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="329" phylum="Arthropoda" rank="species" species="injae" status="sp. nov.">
|
||
<emphasis id="B93FEAD0FFA4FF8A25C85E54FDBFF9C1" box="[507,580,1612,1633]" italics="true" pageId="10" pageNumber="329">T. injae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57ABFFA4FF8A267F5E53FD7AF9C0" box="[588,641,1611,1632]" pageId="10" pageNumber="329" rank="species">sp. n.</taxonomicNameLabel>
|
||
; head. (24)
|
||
<taxonomicName id="4C4B4D41FFA4FF8A26C85E54FC55F9C1" authority="Naomi" authorityName="Naomi" authorityYear="1995" box="[763,942,1611,1633]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="329" phylum="Arthropoda" rank="species" species="ootsukai">
|
||
<emphasis id="B93FEAD0FFA4FF8A26C85E54FC9EF9C1" box="[763,869,1611,1633]" italics="true" pageId="10" pageNumber="329">T. ootsukai</emphasis>
|
||
Naomi
|
||
</taxonomicName>
|
||
; abdominal tip, dorsal view. Scale bar = 0.1 mm for 19, 23, 0.18 mm for 20-21, 24, 0.22 mm for 22.
|
||
</paragraph>
|
||
</footnote>
|
||
<subSubSection id="C3516549FFA5FF8B24D058FCFE26FEE5" pageId="11" pageNumber="330" type="distribution">
|
||
<paragraph id="8BF436C2FFA5FF8B24D058FCFE66FEBB" blockId="11.[151,1073,227,283]" pageId="11" pageNumber="330">
|
||
DISTRIBUTION: The species is known only from Mt. Kiyosumi (
|
||
<collectingRegion id="498FF820FFA5FF8B27AE58FBFF3BFEBB" country="Japan" name="Chiba" pageId="11" pageNumber="330">Chiba prefecture</collectingRegion>
|
||
) in
|
||
<collectingCountry id="F35C7652FFA5FF8B24DE591CFED1FEBB" box="[237,298,260,283]" name="Japan" pageId="11" pageNumber="330">Japan</collectingCountry>
|
||
(Honshu).
|
||
</paragraph>
|
||
<paragraph id="8BF436C2FFA5FF8B24D05936FE26FEE5" blockId="11.[227,477,301,325]" box="[227,477,301,325]" pageId="11" pageNumber="330">BIONOMICS: Unknown.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |