397 lines
62 KiB
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397 lines
62 KiB
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<mods:title id="D3E4071A144C6D9B3E6F44FD9B90D4FF">Revision of the family Acidopsidae Števčić, 2005, and the systematic position of Typhlocarcinodes Alcock, 1900, Caecopilumnus Borradaile, 1902, and Raoulia Ng, 1987, with descriptions of two new genera and five new species (Crustacea: Brachyura: Goneplacoidea)</mods:title>
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<mods:namePart id="55301709ECE7AF4593555336F5A9CDAB">Ng, Peter K. L.</mods:namePart>
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<treatment id="03E287AE5460E2138A9D448EFC610CD6" ID-DOI="http://doi.org/10.5281/zenodo.4909782" ID-GBIF-Taxon="183228350" ID-Zenodo-Dep="4909782" LSID="urn:lsid:plazi:treatment:03E287AE5460E2138A9D448EFC610CD6" httpUri="http://treatment.plazi.org/id/03E287AE5460E2138A9D448EFC610CD6" lastPageId="40" lastPageNumber="41" pageId="37" pageNumber="38">
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<taxonomicName id="4C4B4D3B5460E21E8A9D448EFD1C0FA3" authority="Borradaile, 1902" authorityName="Borradaile" authorityYear="1902" box="[151,662,152,178]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="hirsutus">
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<emphasis id="B93FEAAA5460E21E8A9D448EFD1C0FA3" bold="true" box="[151,662,152,178]" pageId="37" pageNumber="38">
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<emphasis id="B93FEAAA5460E21E8A9D448EFE340FA3" bold="true" box="[151,446,152,178]" italics="true" pageId="37" pageNumber="38">Caecopilumnus hirsutus</emphasis>
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<bibRefCitation id="EFDA4B495460E21E8BCC448EFD1C0FA3" author="Borradaile, L. A." box="[454,662,152,178]" pageId="37" pageNumber="38" pagination="237 - 271" refId="ref30082" refString="Borradaile, L. A. (1902) Marine Crustaceans. III. The Xanthidae and Other Crabs. In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, 1 (3), 237 - 271." type="journal article" year="1902">Borradaile, 1902</bibRefCitation>
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<paragraph id="8BF436B85460E21E8A9D44ACFED40FC2" blockId="37.[151,662,152,211]" box="[151,350,186,211]" pageId="37" pageNumber="38">
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(
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<figureCitation id="13702A3D5460E21E8A9544ACFE890FC2" box="[159,259,186,211]" captionStart="FIGURE 5" captionStartId="11.[151,249,1549,1571]" captionTargetBox="[151,1436,193,1527]" captionTargetId="figure-230@11.[151,1436,193,1527]" captionTargetPageId="11" captionText="FIGURE 5. Right third maxillipeds. A, Acidops cessacii (A. Milne-Edwards, 1878), syntype female (14.1 × 10.0 mm) (NHM 99.3.23.66) Cape Verde (laterally inverted); B, Parapilumnus cristimanus (A. Milne-Edwards, 1873), male (16.3 × 12.5 mm) (ZRC 2003.90), Nansha Is. (Spratly Is.), South China Sea (laterally inverted); C, Raoulia limosa Ng, 1987, holotype male (8.3 × 6.3 mm) (ZMUC M58), off Marongas, Jolo, Philippines; D, Raoulia piroculata (Rathbun, 1911), lectotype female (7.3 × 5.5 mm) (USNM 41359a), Amirante Is., Indian Ocean; E, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand; F, Raoulia fortis sp. nov., holotype male (12.0 × 9.2 mm) (MNHN), Santo, Vanuatu (laterally inverted); G, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia; H, Caecopilumnus hirsutus (Borradaile, 1902), male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia (laterally inverted); I, Caecopilumnus crassipes (Tesch, 1918), male (11.8 × 10.1 mm) (ZRC 2013.1710) Lombok, Indonesia; J, Caecopilumnus loculatus sp. nov., holotype male (8.0 × 6.7 mm) (ZRC 2013.1723), Moorea, French Polynesia (laterally inverted); K, Thecaplax capillosa gen. et. sp. nov., holotype male (11.6 × 9.5 mm) (MZB), Lombok, Indonesia; L, Typhlocarcinodes integrifrons (Miers, 1881), lectotype male (8.5 × 6.9 mm) (NHM 81.24a), Goree I., Senegal." figureDoi="http://doi.org/10.5281/zenodo.4909806" httpUri="https://zenodo.org/record/4909806/files/figure.png" pageId="37" pageNumber="38">Figs. 5H</figureCitation>
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,
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<figureCitation id="13702A3D5460E21E8B0444ACFEDC0FC2" box="[270,342,186,211]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStart-2="FIGURE 26" captionStart-3="FIGURE 27" captionStartId-0="37.[151,249,1907,1929]" captionStartId-1="38.[151,249,1877,1899]" captionStartId-2="39.[151,249,1493,1515]" captionStartId-3="40.[151,249,1929,1951]" captionTargetBox-0="[173,1411,541,1861]" captionTargetBox-1="[152,1434,193,1856]" captionTargetBox-2="[151,1436,181,1472]" captionTargetBox-3="[193,1393,1043,1890]" captionTargetId-0="figure-179@37.[151,1436,512,1885]" captionTargetId-1="figure-57@38.[152,1434,193,1856]" captionTargetId-2="figure-223@39.[151,1436,181,1472]" captionTargetId-3="figure-425@40.[185,1401,1023,1905]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionTargetPageId-2="39" captionTargetPageId-3="40" captionText-0="FIGURE 24. Caecopilumnus hirsutus Borradaile, 1902, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, frontal view of carapace showing buccal cavity and third maxilliped; C, frontal view of carapace showing front and orbits; D, right eye; E, outer view of left chela. After Borradaile (1902: fig. 59)." captionText-1="FIGURE 25. Caecopilumnus hirsutus Borradaile, 1902, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, right third maxilliped; C, left third and fourth ambulatory legs; D, frontal view of carapace; E, outer view of left chela." captionText-2="FIGURE 26. Caecopilumnus hirsutus Borradaile, 1902, male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, right fourth ambulatory leg." captionText-3="FIGURE 27. Caecopilumnus hirsutus Borradaile, 1902, male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia. A, male abdomen; B, C, left G1. Scales: A = 0.5 mm; B, C = 0.25 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4909844" figureDoi-1="http://doi.org/10.5281/zenodo.4909846" figureDoi-2="http://doi.org/10.5281/zenodo.4909848" figureDoi-3="http://doi.org/10.5281/zenodo.4909850" httpUri-0="https://zenodo.org/record/4909844/files/figure.png" httpUri-1="https://zenodo.org/record/4909846/files/figure.png" httpUri-2="https://zenodo.org/record/4909848/files/figure.png" httpUri-3="https://zenodo.org/record/4909850/files/figure.png" pageId="37" pageNumber="38">24–27</figureCitation>
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)
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</paragraph>
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<paragraph id="8BF436B85460E21E8A9D4514FC720E09" blockId="37.[151,1435,258,342]" box="[151,1016,258,280]" pageId="37" pageNumber="38">
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<treatmentCitation id="0AEA10A95460E21E8A9D4514FDFB0E09" author="Borradaile, L. A." box="[151,625,258,280]" page="237" pageId="37" pageNumber="38" year="1902">
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<taxonomicName id="4C4B4D3B5460E21E8A9D4514FDFB0E09" authority="Borradaile, 1902: 237" authorityName="Borradaile" authorityPageNumber="237" authorityYear="1902" box="[151,625,258,280]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="hirsutus">
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<emphasis id="B93FEAAA5460E21E8A9D4514FE070E09" box="[151,397,258,280]" italics="true" pageId="37" pageNumber="38">Caecopilumnus hirsutus</emphasis>
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<bibRefCitation id="EFDA4B495460E21E8B994514FDFB0E09" author="Borradaile, L. A." box="[403,625,258,280]" pageId="37" pageNumber="38" pagination="237 - 271" refId="ref30082" refString="Borradaile, L. A. (1902) Marine Crustaceans. III. The Xanthidae and Other Crabs. In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, 1 (3), 237 - 271." type="journal article" year="1902">Borradaile, 1902: 237</bibRefCitation>
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.—
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<treatmentCitation id="0AEA10A95460E21E88854514FC870E09" author="Ng, P. K. L." box="[655,781,258,280]" page="91" pageId="37" pageNumber="38" year="1987">
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<bibRefCitation id="EFDA4B495460E21E88854514FC870E09" author="Ng, P. K. L." box="[655,781,258,280]" pageId="37" pageNumber="38" pagination="69 - 111" refId="ref31014" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858, and the status of the Rhizopinae Stimpson, 1858 (Crustacea, Decapoda, Brachyura). Indo-Malayan Zoology, 4 (1), 69 - 111, pl. 1, table 1." type="journal article" year="1987">Ng 1987: 91</bibRefCitation>
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</treatmentCitation>
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.—
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<treatmentCitation id="0AEA10A95460E21E89264514FC7E0E09" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[812,1012,258,280]" page="143" pageId="37" pageNumber="38" year="2008">
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<bibRefCitation id="EFDA4B495460E21E89264514FC7E0E09" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[812,1012,258,280]" pageId="37" pageNumber="38" pagination="1 - 286" refId="ref31276" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
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Ng
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<emphasis id="B93FEAAA5460E21E89594515FC0C0E09" box="[851,902,258,280]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
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2008: 143
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</bibRefCitation>
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</treatmentCitation>
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.
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</paragraph>
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<paragraph id="8BF436B85460E21E8A9D4537FA110E47" blockId="37.[151,1435,258,342]" pageId="37" pageNumber="38">
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<treatmentCitationGroup id="AB5B11965460E21E8A9D4537FA110E47" pageId="37" pageNumber="38">
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<taxonomicName id="4C4B4D3B5460E21E8A9D4537FE290E26" baseAuthorityName="Borradaile" baseAuthorityYear="1902" box="[151,419,289,311]" class="Malacostraca" family="Chasmocarcinidae" genus="Typhlocarcinodes" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="hirsutus">
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<emphasis id="B93FEAAA5460E21E8A9D4537FE290E26" box="[151,419,289,311]" italics="true" pageId="37" pageNumber="38">Typhlocarcinodes hirsutus</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="0AEA10A95460E21E8BB64537FDED0E26" author="Tesch, J. J." box="[444,615,289,311]" page="228" pageId="37" pageNumber="38" year="1918">
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<bibRefCitation id="EFDA4B495460E21E8BB64537FDED0E26" author="Tesch, J. J." box="[444,615,289,311]" pageId="37" pageNumber="38" pagination="149 - 295" refId="ref32198" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch 1918: 228</bibRefCitation>
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</treatmentCitation>
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, pl. 15 fig. 3.—
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<treatmentCitation id="0AEA10A95460E21E89014537FC4D0E26" author="Yokoya, Y." box="[779,967,289,311]" page="200" pageId="37" pageNumber="38" year="1933">
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<bibRefCitation id="EFDA4B495460E21E89014537FC4D0E26" author="Yokoya, Y." box="[779,967,289,311]" pageId="37" pageNumber="38" pagination="1 - 226" refId="ref32233" refString="Yokoya, Y. (1933) On the distribution of Decapod Crustaceans inhabiting the continental shelf around Japan, chiefly based upon the materials collected by the S. S. Soyo-Maru, during the year 1923 - 1930. Journal of the College of Agriculture, Imperial University of Tokyo, 12 (1), 1 - 226." type="journal article" year="1933">Yokoya 1933: 200</bibRefCitation>
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</treatmentCitation>
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||
.—
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||
<treatmentCitation id="0AEA10A95460E21E89EC4537FB040E26" author="Serene, R." box="[998,1166,289,311]" page="92" pageId="37" pageNumber="38" year="1968">
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<bibRefCitation id="EFDA4B495460E21E89EC4537FB040E26" author="Serene, R." box="[998,1166,289,311]" pageId="37" pageNumber="38" pagination="33 - 112" refId="ref31824" refString="Serene, R. (1968) The Brachyura of the Indo-West Pacific Region. In: Prodromus for a Check List of the (non-planctonic) Marine Fauna of South East Asia. Singapore National Academy of Sciences, Special Publication No. 1 (Fauna IIICc 3), 33 - 112." type="journal article" year="1968">Serène 1968: 92</bibRefCitation>
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</treatmentCitation>
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||
.—
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||
<treatmentCitation id="0AEA10A95460E21E8EA74537FADF0E26" author="Sakai, T." box="[1197,1365,289,311]" page="192" pageId="37" pageNumber="38" year="1936">
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<bibRefCitation id="EFDA4B495460E21E8EA74537FADF0E26" author="Sakai, T." box="[1197,1365,289,311]" pageId="37" pageNumber="38" refId="ref31554" refString="Sakai, T. (1936) Crabs of Japan, 66 Plates in Life Colours with Descriptions. Sanseido, Tokyo, x + 239 + 27 pp. [bibliography and index], 66 pls, frontispiece. [in Japanese, dated as 1935]" type="book" year="1936">Sakai 1936: 192</bibRefCitation>
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</treatmentCitation>
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||
, pl. 55 fig. 1.—
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||
<treatmentCitation id="0AEA10A95460E21E8B114556FE480E47" author="Sakai, T." box="[283,450,320,342]" page="574" pageId="37" pageNumber="38" year="1939">
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<bibRefCitation id="EFDA4B495460E21E8B114556FE480E47" author="Sakai, T." box="[283,450,320,342]" pageId="37" pageNumber="38" pagination="365 - 741" refId="ref31603" refString="Sakai, T. (1939) Studies on the crabs of Japan. IV. Brachygnatha, Brachyrhyncha. Yokendo, Tokyo, 365 - 741, pls. 42 - 111." type="book chapter" year="1939">Sakai 1939: 574</bibRefCitation>
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</treatmentCitation>
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, pl. 68 fig. 1.—
|
||
<treatmentCitation id="0AEA10A95460E21E88554556FC8E0E47" author="Sakai, T." box="[607,772,320,342]" page="170" pageId="37" pageNumber="38" year="1965">
|
||
<bibRefCitation id="EFDA4B495460E21E88554556FC8E0E47" author="Sakai, T." box="[607,772,320,342]" pageId="37" pageNumber="38" pagination="1 - 32" refId="ref31669" refString="Sakai, T. (1965) The Crabs of Sagami Bay. Tokyo, i - xvi, 1 - 206, 1 - 92, 1 - 32, pls. 1 - 100." type="journal article" year="1965">Sakai 1965: 170</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 84 fig. 4.—
|
||
<treatmentCitation id="0AEA10A95460E21E89A84556FBCC0E47" author="Serene, R." box="[930,1094,320,342]" page="92" pageId="37" pageNumber="38" year="1968">
|
||
<bibRefCitation id="EFDA4B495460E21E89A84556FBCC0E47" author="Serene, R." box="[930,1094,320,342]" pageId="37" pageNumber="38" pagination="33 - 112" refId="ref31824" refString="Serene, R. (1968) The Brachyura of the Indo-West Pacific Region. In: Prodromus for a Check List of the (non-planctonic) Marine Fauna of South East Asia. Singapore National Academy of Sciences, Special Publication No. 1 (Fauna IIICc 3), 33 - 112." type="journal article" year="1968">Serène 1968: 92</bibRefCitation>
|
||
</treatmentCitation>
|
||
.—
|
||
<treatmentCitation id="0AEA10A95460E21E8E6F4556FA810E47" author="Sakai, T." box="[1125,1291,320,342]" page="550" pageId="37" pageNumber="38" year="1976">
|
||
<bibRefCitation id="EFDA4B495460E21E8E6F4556FA810E47" author="Sakai, T." box="[1125,1291,320,342]" pageId="37" pageNumber="38" pagination="1 - 251" refId="ref31706" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo, vol. 1 [English text], xxxix + 773 pp.; Vol. 2 [Japanese text], 461 pp.; Vol. 3 [plates], 16 pp., pls. 1 - 251." type="book chapter" year="1976">Sakai 1976: 550</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 194 fig. 3.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165335460E21E8A9D456BFD270ECF" pageId="37" pageNumber="38" type="materials_examined">
|
||
<paragraph id="8BF436B85460E21E8A9D456BFD270ECF" blockId="37.[151,1436,381,478]" pageId="37" pageNumber="38">
|
||
<emphasis id="B93FEAAA5460E21E8A9D456BFEF20E87" bold="true" box="[151,376,381,406]" pageId="37" pageNumber="38">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="3B233CE55460E21E8B8C456BFD270ECF" ID-GBIF-Occurrence="3312327303" collectingDate="1899-07" collectionCode="CUMZ" country="Indonesia" county="Expedition" elevation="72" location="Madura Bay" municipality="Flores" pageId="37" pageNumber="38" specimenCode="ZMA 261027" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<typeStatus id="54F0881A5460E21E8B8C456BFE790E87" box="[390,499,381,406]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
|
||
: female (8.4 ×
|
||
<quantity id="4CB39B5D5460E21E8893456BFD650E87" box="[665,751,381,406]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="37" pageNumber="38" unit="mm" value="7.2">7.2 mm</quantity>
|
||
) (
|
||
<collectionCode id="ED5AAE7D5460E21E890C456BFCD70E87" box="[774,861,381,406]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="37" pageNumber="38">CUMZ</collectionCode>
|
||
),
|
||
<collectorName id="26BE536E5460E21E8965456BFB890E87" box="[879,1027,381,406]" pageId="37" pageNumber="38">Haifaro Reef</collectorName>
|
||
,
|
||
<collectorName id="26BE536E5460E21E8E07456BFBF70E87" box="[1037,1149,381,406]" pageId="37" pageNumber="38">Fadiffoha</collectorName>
|
||
,
|
||
<collectorName id="26BE536E5460E21E8E8D456BFAAD0E87" box="[1159,1319,381,406]" pageId="37" pageNumber="38">Maldives, J. S</collectorName>
|
||
. Gardiner, 1900.
|
||
<collectorName id="26BE536E5460E21E8AD545B7FEA30EAB" box="[223,297,417,442]" pageId="37" pageNumber="38">Others</collectorName>
|
||
—
|
||
<emphasis id="B93FEAAA5460E21E8B4F45B7FE300EAB" bold="true" box="[325,442,417,442]" pageId="37" pageNumber="38">
|
||
<collectingCountry id="F35C76285460E21E8B4F45B7FE300EAB" box="[325,442,417,442]" name="Indonesia" pageId="37" pageNumber="38">Indonesia</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D4DFD315460E21E8BC145B7FD920EAB" box="[459,536,417,442]" pageId="37" pageNumber="38" type="male">1 male</specimenCount>
|
||
(7.1 ×
|
||
<quantity id="4CB39B5D5460E21E886445B7FD420EAB" box="[622,712,417,442]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="37" pageNumber="38" unit="mm" value="5.6">5.6 mm</quantity>
|
||
) (
|
||
<specimenCode id="DBED9EC35460E21E88EA45B7FC0B0EAB" box="[736,897,417,442]" collectionCode="ZMA" country="Netherlands" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="37" pageNumber="38">ZMA 261027</specimenCode>
|
||
), station 51,
|
||
<location id="8E9460635460E21E8E1F45B7FB290EAB" LSID="urn:lsid:plazi:treatment:03E287AE5460E2138A9D448EFC610CD6:8E9460635460E21E8E1F45B7FB290EAB" box="[1045,1187,417,442]" country="Indonesia" county="Expedition" municipality="Flores" name="Madura Bay" pageId="37" pageNumber="38">Madura Bay</location>
|
||
, west coast of
|
||
<collectingMunicipality id="6B90ACC25460E21E8F4545B7FA120EAB" box="[1359,1432,417,442]" pageId="37" pageNumber="38">Flores</collectingMunicipality>
|
||
,
|
||
<quantity id="4CB39B5D5460E21E8A9D45D3FF760ECC" box="[151,252,453,478]" metricMagnitude="1" metricUnit="m" metricValue="7.2" metricValueMax="9.0" metricValueMin="5.4" pageId="37" pageNumber="38" unit="m" value="72.0" valueMax="90.0" valueMin="54.0">
|
||
<elevation id="0066D18B5460E21E8A9D45D3FF760ECC" box="[151,252,453,478]" metricMagnitude="1" metricUnit="m" metricValue="7.2" metricValueMax="9.0" metricValueMin="5.4" pageId="37" pageNumber="38" unit="m" value="72.0" valueMax="90.0" valueMin="54.0">54–90 m</elevation>
|
||
</quantity>
|
||
, coll. SIBOGA
|
||
<collectingCounty id="62954E345460E21E8BA545D3FDA70ECF" box="[431,557,453,478]" pageId="37" pageNumber="38">Expedition</collectingCounty>
|
||
,
|
||
<date id="FFF510785460E21E883245D3FD220ECF" box="[568,680,453,478]" pageId="37" pageNumber="38" value="1899-07">
|
||
<collectingDate id="EFB1E9905460E21E883245D3FD220ECF" box="[568,680,453,478]" pageId="37" pageNumber="38" value="1899-07">July 1899</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3466305460E21E8A9D4365FC6208D7" ID-DOI="http://doi.org/10.5281/zenodo.4909844" ID-Zenodo-Dep="4909844" httpUri="https://zenodo.org/record/4909844/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,249,1907,1929]" targetBox="[173,1411,541,1861]" targetPageId="37">
|
||
<paragraph id="8BF436B85460E21E8A9D4365FC6208D7" blockId="37.[151,1435,1907,1990]" pageId="37" pageNumber="38">
|
||
<emphasis id="B93FEAAA5460E21E8A9D4365FE910899" bold="true" box="[151,283,1907,1929]" pageId="37" pageNumber="38">FIGURE 24</emphasis>
|
||
.
|
||
<taxonomicName id="4C4B4D3B5460E21E8B204365FD530898" authority="Borradaile, 1902" authorityName="Borradaile" authorityYear="1902" box="[298,729,1907,1929]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="B93FEAAA5460E21E8B204365FDA80898" box="[298,546,1907,1929]" italics="true" pageId="37" pageNumber="38">Caecopilumnus hirsutus</emphasis>
|
||
<bibRefCitation id="EFDA4B495460E21E88204365FD530898" author="Borradaile, L. A." box="[554,729,1907,1929]" pageId="37" pageNumber="38" pagination="237 - 271" refId="ref30082" refString="Borradaile, L. A. (1902) Marine Crustaceans. III. The Xanthidae and Other Crabs. In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, 1 (3), 237 - 271." type="journal article" year="1902">Borradaile, 1902</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, frontal view of carapace showing buccal cavity and third maxilliped; C, frontal view of carapace showing front and orbits; D, right eye; E, outer view of left chela. After
|
||
<bibRefCitation id="EFDA4B495460E21E88D743A6FC0708D7" author="Borradaile, L. A." box="[733,909,1968,1990]" pageId="37" pageNumber="38" pagination="237 - 271" refId="ref30082" refString="Borradaile, L. A. (1902) Marine Crustaceans. III. The Xanthidae and Other Crabs. In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, 1 (3), 237 - 271." type="journal article" year="1902">Borradaile (1902</bibRefCitation>
|
||
: fig. 59).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3466305463E21D8A9D4343FF5908B9" ID-DOI="http://doi.org/10.5281/zenodo.4909846" ID-Zenodo-Dep="4909846" httpUri="https://zenodo.org/record/4909846/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,249,1877,1899]" targetBox="[152,1434,193,1856]" targetPageId="38">
|
||
<paragraph id="8BF436B85463E21D8A9D4343FF5908B9" blockId="38.[151,1436,1877,1960]" pageId="38" pageNumber="39">
|
||
<emphasis id="B93FEAAA5463E21D8A9D4343FE91087B" bold="true" box="[151,283,1877,1899]" pageId="38" pageNumber="39">FIGURE 25</emphasis>
|
||
.
|
||
<taxonomicName id="4C4B4D3B5463E21D8B204343FD53087A" authority="Borradaile, 1902" authorityName="Borradaile" authorityYear="1902" box="[298,729,1877,1899]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="B93FEAAA5463E21D8B204343FDA8087A" box="[298,546,1877,1899]" italics="true" pageId="38" pageNumber="39">Caecopilumnus hirsutus</emphasis>
|
||
<bibRefCitation id="EFDA4B495463E21D88204343FD53087A" author="Borradaile, L. A." box="[554,729,1877,1899]" pageId="38" pageNumber="39" pagination="237 - 271" refId="ref30082" refString="Borradaile, L. A. (1902) Marine Crustaceans. III. The Xanthidae and Other Crabs. In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, 1 (3), 237 - 271." type="journal article" year="1902">Borradaile, 1902</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, right third maxilliped; C, left third and fourth ambulatory legs; D, frontal view of carapace; E, outer view of left chela.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3466305462E21C8A9D41C3FBD6091B" ID-DOI="http://doi.org/10.5281/zenodo.4909848" ID-Zenodo-Dep="4909848" httpUri="https://zenodo.org/record/4909848/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,249,1493,1515]" targetBox="[151,1436,181,1472]" targetPageId="39">
|
||
<paragraph id="8BF436B85462E21C8A9D41C3FBD6091B" blockId="39.[151,1436,1493,1546]" pageId="39" pageNumber="40">
|
||
<emphasis id="B93FEAAA5462E21C8A9D41C3FE910AFB" bold="true" box="[151,283,1493,1515]" pageId="39" pageNumber="40">FIGURE 26</emphasis>
|
||
.
|
||
<taxonomicName id="4C4B4D3B5462E21C8B2141C3FD560AFA" authority="Borradaile, 1902" authorityName="Borradaile" authorityYear="1902" box="[299,732,1493,1515]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="B93FEAAA5462E21C8B2141C3FDAE0AFA" box="[299,548,1493,1515]" italics="true" pageId="39" pageNumber="40">Caecopilumnus hirsutus</emphasis>
|
||
<bibRefCitation id="EFDA4B495462E21C882641C3FD560AFA" author="Borradaile, L. A." box="[556,732,1493,1515]" pageId="39" pageNumber="40" pagination="237 - 271" refId="ref30082" refString="Borradaile, L. A. (1902) Marine Crustaceans. III. The Xanthidae and Other Crabs. In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, 1 (3), 237 - 271." type="journal article" year="1902">Borradaile, 1902</bibRefCitation>
|
||
</taxonomicName>
|
||
, male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, right fourth ambulatory leg.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35165335462E21C8ACD4220FE5E0879" pageId="39" pageNumber="40" type="diagnosis">
|
||
<paragraph id="8BF436B85462E21C8ACD4220FED00832" blockId="39.[151,1436,1590,2037]" pageId="39" pageNumber="40">
|
||
<emphasis id="B93FEAAA5462E21C8ACD4220FEB0095E" bold="true" box="[199,314,1590,1615]" pageId="39" pageNumber="40">Diagnosis</emphasis>
|
||
. Carapace subquadrate; dorsal surface prominently granular; regions separated by distinct deep grooves (
|
||
<figureCitation id="13702A3D5462E21C8B0E424CFEF70962" box="[260,381,1626,1651]" captionStart="FIGURE 24" captionStartId="37.[151,249,1907,1929]" captionTargetBox="[173,1411,541,1861]" captionTargetId="figure-179@37.[151,1436,512,1885]" captionTargetPageId="37" captionText="FIGURE 24. Caecopilumnus hirsutus Borradaile, 1902, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, frontal view of carapace showing buccal cavity and third maxilliped; C, frontal view of carapace showing front and orbits; D, right eye; E, outer view of left chela. After Borradaile (1902: fig. 59)." figureDoi="http://doi.org/10.5281/zenodo.4909844" httpUri="https://zenodo.org/record/4909844/files/figure.png" pageId="39" pageNumber="40">Figs. 24A</figureCitation>
|
||
,
|
||
<figureCitation id="13702A3D5462E21C8B87424CFE4A0962" box="[397,448,1626,1651]" captionStart="FIGURE 25" captionStartId="38.[151,249,1877,1899]" captionTargetBox="[152,1434,193,1856]" captionTargetId="figure-57@38.[152,1434,193,1856]" captionTargetPageId="38" captionText="FIGURE 25. Caecopilumnus hirsutus Borradaile, 1902, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, right third maxilliped; C, left third and fourth ambulatory legs; D, frontal view of carapace; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909846" httpUri="https://zenodo.org/record/4909846/files/figure.png" pageId="39" pageNumber="40">25A</figureCitation>
|
||
,
|
||
<figureCitation id="13702A3D5462E21C8BDB424CFD8F0962" box="[465,517,1626,1651]" captionStart="FIGURE 26" captionStartId="39.[151,249,1493,1515]" captionTargetBox="[151,1436,181,1472]" captionTargetId="figure-223@39.[151,1436,181,1472]" captionTargetPageId="39" captionText="FIGURE 26. Caecopilumnus hirsutus Borradaile, 1902, male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4909848" httpUri="https://zenodo.org/record/4909848/files/figure.png" pageId="39" pageNumber="40">26A</figureCitation>
|
||
). Anterolateral margin arcuate, granular, with low granuliform lobes, may be indistinct (
|
||
<figureCitation id="13702A3D5462E21C8B1A426BFE0E0987" box="[272,388,1661,1686]" captionStart="FIGURE 24" captionStartId="37.[151,249,1907,1929]" captionTargetBox="[173,1411,541,1861]" captionTargetId="figure-179@37.[151,1436,512,1885]" captionTargetPageId="37" captionText="FIGURE 24. Caecopilumnus hirsutus Borradaile, 1902, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, frontal view of carapace showing buccal cavity and third maxilliped; C, frontal view of carapace showing front and orbits; D, right eye; E, outer view of left chela. After Borradaile (1902: fig. 59)." figureDoi="http://doi.org/10.5281/zenodo.4909844" httpUri="https://zenodo.org/record/4909844/files/figure.png" pageId="39" pageNumber="40">Figs. 24A</figureCitation>
|
||
,
|
||
<figureCitation id="13702A3D5462E21C8B9A426BFE490987" box="[400,451,1661,1686]" captionStart="FIGURE 25" captionStartId="38.[151,249,1877,1899]" captionTargetBox="[152,1434,193,1856]" captionTargetId="figure-57@38.[152,1434,193,1856]" captionTargetPageId="38" captionText="FIGURE 25. Caecopilumnus hirsutus Borradaile, 1902, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, right third maxilliped; C, left third and fourth ambulatory legs; D, frontal view of carapace; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909846" httpUri="https://zenodo.org/record/4909846/files/figure.png" pageId="39" pageNumber="40">25A</figureCitation>
|
||
,
|
||
<figureCitation id="13702A3D5462E21C8BDA426BFD8E0987" box="[464,516,1661,1686]" captionStart="FIGURE 26" captionStartId="39.[151,249,1493,1515]" captionTargetBox="[151,1436,181,1472]" captionTargetId="figure-223@39.[151,1436,181,1472]" captionTargetPageId="39" captionText="FIGURE 26. Caecopilumnus hirsutus Borradaile, 1902, male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4909848" httpUri="https://zenodo.org/record/4909848/files/figure.png" pageId="39" pageNumber="40">26A</figureCitation>
|
||
). Merus of third maxilliped rounded to subovate (
|
||
<figureCitation id="13702A3D5462E21C8E37426BFB280987" box="[1085,1186,1661,1686]" captionStart="FIGURE 5" captionStartId="11.[151,249,1549,1571]" captionTargetBox="[151,1436,193,1527]" captionTargetId="figure-230@11.[151,1436,193,1527]" captionTargetPageId="11" captionText="FIGURE 5. Right third maxillipeds. A, Acidops cessacii (A. Milne-Edwards, 1878), syntype female (14.1 × 10.0 mm) (NHM 99.3.23.66) Cape Verde (laterally inverted); B, Parapilumnus cristimanus (A. Milne-Edwards, 1873), male (16.3 × 12.5 mm) (ZRC 2003.90), Nansha Is. (Spratly Is.), South China Sea (laterally inverted); C, Raoulia limosa Ng, 1987, holotype male (8.3 × 6.3 mm) (ZMUC M58), off Marongas, Jolo, Philippines; D, Raoulia piroculata (Rathbun, 1911), lectotype female (7.3 × 5.5 mm) (USNM 41359a), Amirante Is., Indian Ocean; E, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand; F, Raoulia fortis sp. nov., holotype male (12.0 × 9.2 mm) (MNHN), Santo, Vanuatu (laterally inverted); G, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia; H, Caecopilumnus hirsutus (Borradaile, 1902), male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia (laterally inverted); I, Caecopilumnus crassipes (Tesch, 1918), male (11.8 × 10.1 mm) (ZRC 2013.1710) Lombok, Indonesia; J, Caecopilumnus loculatus sp. nov., holotype male (8.0 × 6.7 mm) (ZRC 2013.1723), Moorea, French Polynesia (laterally inverted); K, Thecaplax capillosa gen. et. sp. nov., holotype male (11.6 × 9.5 mm) (MZB), Lombok, Indonesia; L, Typhlocarcinodes integrifrons (Miers, 1881), lectotype male (8.5 × 6.9 mm) (NHM 81.24a), Goree I., Senegal." figureDoi="http://doi.org/10.5281/zenodo.4909806" httpUri="https://zenodo.org/record/4909806/files/figure.png" pageId="39" pageNumber="40">Figs. 5H</figureCitation>
|
||
,
|
||
<figureCitation id="13702A3D5462E21C8EA5426BFB6B0987" box="[1199,1249,1661,1686]" captionStart="FIGURE 24" captionStartId="37.[151,249,1907,1929]" captionTargetBox="[173,1411,541,1861]" captionTargetId="figure-179@37.[151,1436,512,1885]" captionTargetPageId="37" captionText="FIGURE 24. Caecopilumnus hirsutus Borradaile, 1902, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, frontal view of carapace showing buccal cavity and third maxilliped; C, frontal view of carapace showing front and orbits; D, right eye; E, outer view of left chela. After Borradaile (1902: fig. 59)." figureDoi="http://doi.org/10.5281/zenodo.4909844" httpUri="https://zenodo.org/record/4909844/files/figure.png" pageId="39" pageNumber="40">24B</figureCitation>
|
||
,
|
||
<figureCitation id="13702A3D5462E21C8EE7426BFAAA0987" box="[1261,1312,1661,1686]" captionStart="FIGURE 25" captionStartId="38.[151,249,1877,1899]" captionTargetBox="[152,1434,193,1856]" captionTargetId="figure-57@38.[152,1434,193,1856]" captionTargetPageId="38" captionText="FIGURE 25. Caecopilumnus hirsutus Borradaile, 1902, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, right third maxilliped; C, left third and fourth ambulatory legs; D, frontal view of carapace; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909846" httpUri="https://zenodo.org/record/4909846/files/figure.png" pageId="39" pageNumber="40">25B</figureCitation>
|
||
). Merus of last ambulatory leg 2.1 times as long as broad; propodus subovate, longer than broad (
|
||
<figureCitation id="13702A3D5462E21C8E9E42B6FA8309A8" box="[1172,1289,1696,1721]" captionStart="FIGURE 25" captionStartId="38.[151,249,1877,1899]" captionTargetBox="[152,1434,193,1856]" captionTargetId="figure-57@38.[152,1434,193,1856]" captionTargetPageId="38" captionText="FIGURE 25. Caecopilumnus hirsutus Borradaile, 1902, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, right third maxilliped; C, left third and fourth ambulatory legs; D, frontal view of carapace; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909846" httpUri="https://zenodo.org/record/4909846/files/figure.png" pageId="39" pageNumber="40">Figs. 25C</figureCitation>
|
||
,
|
||
<figureCitation id="13702A3D5462E21C8F1242B6FAC109A8" box="[1304,1355,1696,1721]" captionStart="FIGURE 26" captionStartId="39.[151,249,1493,1515]" captionTargetBox="[151,1436,181,1472]" captionTargetId="figure-223@39.[151,1436,181,1472]" captionTargetPageId="39" captionText="FIGURE 26. Caecopilumnus hirsutus Borradaile, 1902, male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4909848" httpUri="https://zenodo.org/record/4909848/files/figure.png" pageId="39" pageNumber="40">26C</figureCitation>
|
||
).
|
||
<collectingRegion id="498FF85A5462E21C8F6842B6FA1609A8" box="[1378,1436,1696,1721]" country="Maldives" name="Maale" pageId="39" pageNumber="40">Male</collectingRegion>
|
||
abdomen relatively wide; a1 with lateral margins straight; a3–6 proportionately wide (
|
||
<figureCitation id="13702A3D5462E21C8E6742D5FB5D09CD" box="[1133,1239,1731,1756]" captionStart="FIGURE 27" captionStartId="40.[151,249,1929,1951]" captionTargetBox="[193,1393,1043,1890]" captionTargetId="figure-425@40.[185,1401,1023,1905]" captionTargetPageId="40" captionText="FIGURE 27. Caecopilumnus hirsutus Borradaile, 1902, male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia. A, male abdomen; B, C, left G1. Scales: A = 0.5 mm; B, C = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4909850" httpUri="https://zenodo.org/record/4909850/files/figure.png" pageId="39" pageNumber="40">Fig. 27A</figureCitation>
|
||
). Gl stout, gently curved, proximally dilated, distal half more slender than proximal, tip tapered, with subdistal spinules (
|
||
<figureCitation id="13702A3D5462E21C8F3A42F0FF270832" captionStart="FIGURE 27" captionStartId="40.[151,249,1929,1951]" captionTargetBox="[193,1393,1043,1890]" captionTargetId="figure-425@40.[185,1401,1023,1905]" captionTargetPageId="40" captionText="FIGURE 27. Caecopilumnus hirsutus Borradaile, 1902, male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia. A, male abdomen; B, C, left G1. Scales: A = 0.5 mm; B, C = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4909850" httpUri="https://zenodo.org/record/4909850/files/figure.png" pageId="39" pageNumber="40">Fig. 27B, C</figureCitation>
|
||
). G2 unknown.
|
||
</paragraph>
|
||
<paragraph id="8BF436B85462E21C8ACD433BFE5E0879" blockId="39.[151,1436,1590,2037]" pageId="39" pageNumber="40">
|
||
<emphasis id="B93FEAAA5462E21C8ACD433BFE900857" bold="true" box="[199,282,1837,1862]" pageId="39" pageNumber="40">Colour</emphasis>
|
||
.
|
||
<bibRefCitation id="EFDA4B495462E21C8B21433BFE330857" author="Sakai, T." box="[299,441,1837,1862]" pageId="39" pageNumber="40" pagination="1 - 251" refId="ref31706" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo, vol. 1 [English text], xxxix + 773 pp.; Vol. 2 [Japanese text], 461 pp.; Vol. 3 [plates], 16 pp., pls. 1 - 251." type="book chapter" year="1976">Sakai (1976</bibRefCitation>
|
||
: pl. 194 fig. 3) figured the species as cream-coloured with several regularly arranged brown spots on its carapace.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165335462E2138ACD4365FE660CB4" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="discussion">
|
||
<paragraph id="8BF436B85462E21C8ACD4365FEAD08E4" blockId="39.[151,1436,1590,2037]" pageId="39" pageNumber="40">
|
||
<emphasis id="B93FEAAA5462E21C8ACD4365FEBE089D" bold="true" box="[199,308,1907,1932]" pageId="39" pageNumber="40">Remarks</emphasis>
|
||
. This rare species remains somewhat enigmatic. The
|
||
<typeStatus id="54F0881A5462E21C89AE4365FB8C089D" box="[932,1030,1907,1932]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
|
||
female is in poor condition but its characters essentially agree with the detailed figures and descriptions by
|
||
<bibRefCitation id="EFDA4B495462E21C89F04380FB4D08BE" author="Borradaile, L. A." box="[1018,1223,1942,1967]" pageId="39" pageNumber="40" pagination="237 - 271" refId="ref30082" refString="Borradaile, L. A. (1902) Marine Crustaceans. III. The Xanthidae and Other Crabs. In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, 1 (3), 237 - 271." type="journal article" year="1902">Borradaile (1902)</bibRefCitation>
|
||
(see
|
||
<figureCitation id="13702A3D5462E21C8F064380FAEE08BE" box="[1292,1380,1942,1967]" captionStart="FIGURE 24" captionStartId="37.[151,249,1907,1929]" captionTargetBox="[173,1411,541,1861]" captionTargetId="figure-179@37.[151,1436,512,1885]" captionTargetPageId="37" captionText="FIGURE 24. Caecopilumnus hirsutus Borradaile, 1902, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, frontal view of carapace showing buccal cavity and third maxilliped; C, frontal view of carapace showing front and orbits; D, right eye; E, outer view of left chela. After Borradaile (1902: fig. 59)." figureDoi="http://doi.org/10.5281/zenodo.4909844" httpUri="https://zenodo.org/record/4909844/files/figure.png" pageId="39" pageNumber="40">Fig. 24</figureCitation>
|
||
). As discussed under
|
||
<taxonomicName id="4C4B4D3B5462E21C8B5A43ACFE5308C3" authorityName="Tesch" authorityYear="1918" box="[336,473,1978,2002]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="crassipes">
|
||
<emphasis id="B93FEAAA5462E21C8B5A43ACFE5308C3" box="[336,473,1978,2002]" italics="true" pageId="39" pageNumber="40">C. crassipes</emphasis>
|
||
</taxonomicName>
|
||
, the differences in the carapace and ambulatory legs easily separates
|
||
<taxonomicName id="4C4B4D3B5462E21C8EE943ACFAD408C2" authorityName="Borradaile" authorityYear="1902" box="[1251,1374,1978,2003]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="B93FEAAA5462E21C8EE943ACFAD408C2" box="[1251,1374,1978,2003]" italics="true" pageId="39" pageNumber="40">C. hirsutus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C4B4D3B5462E21C8A9D43C8FE9508E4" authorityName="Tesch" authorityYear="1918" box="[151,287,2014,2037]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="crassipes">
|
||
<emphasis id="B93FEAAA5462E21C8A9D43C8FE9508E4" box="[151,287,2014,2037]" italics="true" pageId="39" pageNumber="40">C. crassipes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF436B8546DE2138ACD448EFE220EC1" blockId="40.[151,1436,151,968]" pageId="40" pageNumber="41">
|
||
We have doubts if the
|
||
<typeStatus id="54F0881A546DE2138BCA448EFE7A0FA1" box="[448,496,152,176]" pageId="40" pageNumber="41">type</typeStatus>
|
||
female (from the
|
||
<collectingCountry id="F35C7628546DE21388B04481FCAE0FA1" box="[698,804,151,176]" name="Maldives" pageId="40" pageNumber="41">Maldives</collectingCountry>
|
||
) is conspecific with the present specimen from
|
||
<collectingCountry id="F35C7628546DE2138F334481FAF50FA1" box="[1337,1407,151,176]" name="Indonesia" pageId="40" pageNumber="41">Flores</collectingCountry>
|
||
in the
|
||
<collectingRegion id="498FF85A546DE2138AC844AAFE3F0FC4" box="[194,437,188,213]" country="Indonesia" name="Nusa Tenggara Timur" pageId="40" pageNumber="41">Lesser Sunda Islands</collectingRegion>
|
||
. reported by
|
||
<bibRefCitation id="EFDA4B49546DE213884144AAFD690FC4" author="Tesch, J. J." box="[587,739,188,213]" pageId="40" pageNumber="41" pagination="149 - 295" refId="ref32198" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch (1918)</bibRefCitation>
|
||
. Although their carapaces, chelipeds and ambulatory legs are very similar, their third maxillipeds differ. In the
|
||
<typeStatus id="54F0881A546DE21388CA44F6FD7A0FE9" box="[704,752,224,248]" pageId="40" pageNumber="41">type</typeStatus>
|
||
female, the ischium is prominently rounded and the ischium relatively short (
|
||
<figureCitation id="13702A3D546DE2138B554512FE530E0C" box="[351,473,260,285]" captionStart="FIGURE 24" captionStartId="37.[151,249,1907,1929]" captionTargetBox="[173,1411,541,1861]" captionTargetId="figure-179@37.[151,1436,512,1885]" captionTargetPageId="37" captionText="FIGURE 24. Caecopilumnus hirsutus Borradaile, 1902, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, frontal view of carapace showing buccal cavity and third maxilliped; C, frontal view of carapace showing front and orbits; D, right eye; E, outer view of left chela. After Borradaile (1902: fig. 59)." figureDoi="http://doi.org/10.5281/zenodo.4909844" httpUri="https://zenodo.org/record/4909844/files/figure.png" pageId="40" pageNumber="41">Figs. 24B</figureCitation>
|
||
,
|
||
<figureCitation id="13702A3D546DE2138BE64512FD950E0C" box="[492,543,260,285]" captionStart="FIGURE 25" captionStartId="38.[151,249,1877,1899]" captionTargetBox="[152,1434,193,1856]" captionTargetId="figure-57@38.[152,1434,193,1856]" captionTargetPageId="38" captionText="FIGURE 25. Caecopilumnus hirsutus Borradaile, 1902, holotype female (8.4 × 7.2 mm) (CUMZ), Fadiffoha, Maldives. A, dorsal view; B, right third maxilliped; C, left third and fourth ambulatory legs; D, frontal view of carapace; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909846" httpUri="https://zenodo.org/record/4909846/files/figure.png" pageId="40" pageNumber="41">25B</figureCitation>
|
||
). In the male from
|
||
<collectingCountry id="F35C7628546DE21389144512FCED0E0C" box="[798,871,260,285]" name="Indonesia" pageId="40" pageNumber="41">Flores</collectingCountry>
|
||
, the merus is more quadrate and the ischium proportionately longer (
|
||
<figureCitation id="13702A3D546DE2138BA84531FE770E51" box="[418,509,295,320]" captionStart="FIGURE 5" captionStartId="11.[151,249,1549,1571]" captionTargetBox="[151,1436,193,1527]" captionTargetId="figure-230@11.[151,1436,193,1527]" captionTargetPageId="11" captionText="FIGURE 5. Right third maxillipeds. A, Acidops cessacii (A. Milne-Edwards, 1878), syntype female (14.1 × 10.0 mm) (NHM 99.3.23.66) Cape Verde (laterally inverted); B, Parapilumnus cristimanus (A. Milne-Edwards, 1873), male (16.3 × 12.5 mm) (ZRC 2003.90), Nansha Is. (Spratly Is.), South China Sea (laterally inverted); C, Raoulia limosa Ng, 1987, holotype male (8.3 × 6.3 mm) (ZMUC M58), off Marongas, Jolo, Philippines; D, Raoulia piroculata (Rathbun, 1911), lectotype female (7.3 × 5.5 mm) (USNM 41359a), Amirante Is., Indian Ocean; E, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand; F, Raoulia fortis sp. nov., holotype male (12.0 × 9.2 mm) (MNHN), Santo, Vanuatu (laterally inverted); G, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia; H, Caecopilumnus hirsutus (Borradaile, 1902), male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia (laterally inverted); I, Caecopilumnus crassipes (Tesch, 1918), male (11.8 × 10.1 mm) (ZRC 2013.1710) Lombok, Indonesia; J, Caecopilumnus loculatus sp. nov., holotype male (8.0 × 6.7 mm) (ZRC 2013.1723), Moorea, French Polynesia (laterally inverted); K, Thecaplax capillosa gen. et. sp. nov., holotype male (11.6 × 9.5 mm) (MZB), Lombok, Indonesia; L, Typhlocarcinodes integrifrons (Miers, 1881), lectotype male (8.5 × 6.9 mm) (NHM 81.24a), Goree I., Senegal." figureDoi="http://doi.org/10.5281/zenodo.4909806" httpUri="https://zenodo.org/record/4909806/files/figure.png" pageId="40" pageNumber="41">Fig. 5H</figureCitation>
|
||
). While we have observed some variation in the form of the third maxilliped in
|
||
<taxonomicName id="4C4B4D3B546DE2138F88453EFF750E75" authorityName="Tesch" authorityYear="1918" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="crassipes">
|
||
<emphasis id="B93FEAAA546DE2138F88453EFF750E75" italics="true" pageId="40" pageNumber="41">C. crassipes</emphasis>
|
||
</taxonomicName>
|
||
, which is represented by a good series of specimens, it is not to the degree observed here. Nevertheless, with only
|
||
<specimenCount id="9D4DFD31546DE2138B024566FE3A0E99" box="[264,432,367,392]" pageId="40" pageNumber="41" type="generic">two specimens</specimenCount>
|
||
, we prefer to err on the side of caution and recognise the
|
||
<collectingCountry id="F35C7628546DE2138E394579FBF30E99" box="[1075,1145,367,392]" name="Indonesia" pageId="40" pageNumber="41">Flores</collectingCountry>
|
||
specimen as belonging to
|
||
<taxonomicName id="4C4B4D3B546DE2138A9D4582FE990EBC" authorityName="Borradaile" authorityYear="1902" box="[151,275,404,429]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="B93FEAAA546DE2138A9D4582FE990EBC" box="[151,275,404,429]" italics="true" pageId="40" pageNumber="41">C. hirsutus</emphasis>
|
||
</taxonomicName>
|
||
for the time being. More material may show otherwise. For more comparisons with
|
||
<taxonomicName id="4C4B4D3B546DE2138ED44582FAE20EBD" authorityName="Tesch" authorityYear="1918" box="[1246,1384,404,428]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="crassipes">
|
||
<emphasis id="B93FEAAA546DE2138ED44582FAE20EBD" box="[1246,1384,404,428]" italics="true" pageId="40" pageNumber="41">C. crassipes</emphasis>
|
||
</taxonomicName>
|
||
, see remarks for this species.
|
||
</paragraph>
|
||
<paragraph id="8BF436B8546DE2138ACD45CAFDC40DB9" blockId="40.[151,1436,151,968]" pageId="40" pageNumber="41">
|
||
We are also uncertain of the identity of the Japanese material identified as “
|
||
<taxonomicName id="4C4B4D3B546DE2138E3545CAFAE30EE4" baseAuthorityName="Borradaile" baseAuthorityYear="1902" box="[1087,1385,476,501]" class="Malacostraca" family="Chasmocarcinidae" genus="Typhlocarcinodes" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="B93FEAAA546DE2138E3545CAFAE30EE4" box="[1087,1385,476,501]" italics="true" pageId="40" pageNumber="41">Typhlocarcinodes hirsutus</emphasis>
|
||
</taxonomicName>
|
||
” by
|
||
<bibRefCitation id="EFDA4B49546DE2138A9D45E9FEB50D09" author="Yokoya, Y." box="[151,319,511,536]" pageId="40" pageNumber="41" pagination="1 - 226" refId="ref32233" refString="Yokoya, Y. (1933) On the distribution of Decapod Crustaceans inhabiting the continental shelf around Japan, chiefly based upon the materials collected by the S. S. Soyo-Maru, during the year 1923 - 1930. Journal of the College of Agriculture, Imperial University of Tokyo, 12 (1), 1 - 226." type="journal article" year="1933">Yokoya (1933)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFDA4B49546DE2138B7245E9FD890D09" author="Sakai, T." box="[376,515,511,536]" pageId="40" pageNumber="41" refId="ref31554" refString="Sakai, T. (1936) Crabs of Japan, 66 Plates in Life Colours with Descriptions. Sanseido, Tokyo, x + 239 + 27 pp. [bibliography and index], 66 pls, frontispiece. [in Japanese, dated as 1935]" type="book" year="1936">Sakai (1936</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDA4B49546DE213880545E9FDC00D09" author="Sakai, T." box="[527,586,511,536]" pageId="40" pageNumber="41" pagination="365 - 741" refId="ref31603" refString="Sakai, T. (1939) Studies on the crabs of Japan. IV. Brachygnatha, Brachyrhyncha. Yokendo, Tokyo, 365 - 741, pls. 42 - 111." type="book chapter" year="1939">1939</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDA4B49546DE213885D45E9FD180D09" author="Sakai, T." box="[599,658,511,536]" pageId="40" pageNumber="41" pagination="1 - 32" refId="ref31669" refString="Sakai, T. (1965) The Crabs of Sagami Bay. Tokyo, i - xvi, 1 - 206, 1 - 92, 1 - 32, pls. 1 - 100." type="journal article" year="1965">1965</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDA4B49546DE213889445E9FD5D0D09" author="Sakai, T." box="[670,727,511,536]" pageId="40" pageNumber="41" pagination="1 - 251" refId="ref31706" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo, vol. 1 [English text], xxxix + 773 pp.; Vol. 2 [Japanese text], 461 pp.; Vol. 3 [plates], 16 pp., pls. 1 - 251." type="book chapter" year="1976">1976</bibRefCitation>
|
||
) from Sagami Bay, Izu Peninsula, Tosa Bay, Bungo Strait and the southern part of the Sea of
|
||
<collectingCountry id="F35C7628546DE2138BF14632FDB40D2A" box="[507,574,548,571]" name="Japan" pageId="40" pageNumber="41">Japan</collectingCountry>
|
||
. The figures are rather schematic but if accurate, on the basis of the deeper carapace grooves (highlighted by Sakai in his key and description) and relatively longer ambulatory propodus, their specimens can be referred to
|
||
<taxonomicName id="4C4B4D3B546DE2138BED467AFD720D94" authorityName="Borradaile" authorityYear="1902" box="[487,760,620,645]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="B93FEAAA546DE2138BED467AFD720D94" box="[487,760,620,645]" italics="true" pageId="40" pageNumber="41">Caecopilumnus hirsutus</emphasis>
|
||
</taxonomicName>
|
||
as defined here. These specimens should be examined at a later date to confirm this identification.
|
||
</paragraph>
|
||
<paragraph id="8BF436B8546DE2138ACD46A2FE660CB4" blockId="40.[151,1436,151,968]" pageId="40" pageNumber="41">
|
||
We collected a good series of
|
||
<taxonomicName id="4C4B4D3B546DE213881E46A2FD160DDD" authorityName="Tesch" authorityYear="1918" box="[532,668,692,716]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="crassipes">
|
||
<emphasis id="B93FEAAA546DE213881E46A2FD160DDD" box="[532,668,692,716]" italics="true" pageId="40" pageNumber="41">C. crassipes</emphasis>
|
||
</taxonomicName>
|
||
from intertidal areas of Lombok, which is adjacent to the Banda Sea area where the
|
||
<typeStatus id="54F0881A546DE2138B4F46C1FE2D0DE1" box="[325,423,727,752]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
|
||
was collected, presumably also from intertidal areas (stated as “reef collection”). We did not, however, collect any
|
||
<taxonomicName id="4C4B4D3B546DE2138BC746EAFDC70C04" authorityName="Borradaile" authorityYear="1902" box="[461,589,764,789]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="B93FEAAA546DE2138BC746EAFDC70C04" box="[461,589,764,789]" italics="true" pageId="40" pageNumber="41">C. hirsutus</emphasis>
|
||
</taxonomicName>
|
||
specimens. The specimen of
|
||
<taxonomicName id="4C4B4D3B546DE21389BE46EAFBBE0C04" authorityName="Borradaile" authorityYear="1902" box="[948,1076,764,789]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="B93FEAAA546DE21389BE46EAFBBE0C04" box="[948,1076,764,789]" italics="true" pageId="40" pageNumber="41">C. hirsutus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F35C7628546DE2138E8E46EAFB400C04" box="[1156,1226,764,789]" name="Indonesia" pageId="40" pageNumber="41">Flores</collectingCountry>
|
||
was collected by dredging from depths of
|
||
<quantity id="4CB39B5D546DE2138BA74709FD990C26" box="[429,531,799,824]" metricMagnitude="1" metricUnit="m" metricValue="7.2" metricValueMax="9.0" metricValueMin="5.4" pageId="40" pageNumber="41" unit="m" value="72.0" valueMax="90.0" valueMin="54.0">
|
||
54–
|
||
<specimenCount id="9D4DFD31546DE2138BDD4709FD990C26" box="[471,531,799,824]" pageId="40" pageNumber="41" type="generic">90 m</specimenCount>
|
||
</quantity>
|
||
. The habitat of the type female of
|
||
<taxonomicName id="4C4B4D3B546DE21389994736FB870C29" authorityName="Borradaile" authorityYear="1902" box="[915,1037,799,824]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="B93FEAAA546DE21389994736FB870C29" box="[915,1037,799,824]" italics="true" pageId="40" pageNumber="41">C. hirsutus</emphasis>
|
||
</taxonomicName>
|
||
was not stated;
|
||
<bibRefCitation id="EFDA4B49546DE2138EC84709FA160C29" author="Borradaile, L. A." box="[1218,1436,799,824]" pageId="40" pageNumber="41" pagination="237 - 271" refId="ref30082" refString="Borradaile, L. A. (1902) Marine Crustaceans. III. The Xanthidae and Other Crabs. In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes: Being the Account of the Work carried on and of the Collections made by an Expedition during the years 1899 and 1900, 1 (3), 237 - 271." type="journal article" year="1902">Borradaile’s (1902)</bibRefCitation>
|
||
brachyuran material including both intertidal as well as dredged material. Interestingly,
|
||
<bibRefCitation id="EFDA4B49546DE2138E9E4752FAEF0C4C" author="Sakai, T." box="[1172,1381,836,861]" pageId="40" pageNumber="41" pagination="1 - 251" refId="ref31706" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo, vol. 1 [English text], xxxix + 773 pp.; Vol. 2 [Japanese text], 461 pp.; Vol. 3 [plates], 16 pp., pls. 1 - 251." type="book chapter" year="1976">Sakai (1976: 550)</bibRefCitation>
|
||
also indicates that his Japanese specimens of
|
||
<taxonomicName id="4C4B4D3B546DE213887B477EFD650C91" authorityName="Borradaile" authorityYear="1902" box="[625,751,871,896]" class="Malacostraca" family="Pilumnidae" genus="Caecopilumnus" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="B93FEAAA546DE213887B477EFD650C91" box="[625,751,871,896]" italics="true" pageId="40" pageNumber="41">C. hirsutus</emphasis>
|
||
</taxonomicName>
|
||
were collected from the “Bottoms of sand, mud or broken shells; depth 35 to 95 metres”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3516533546DE2138ACD47B9FC610CD6" box="[199,1003,943,968]" pageId="40" pageNumber="41" type="distribution">
|
||
<paragraph id="8BF436B8546DE2138ACD47B9FC610CD6" blockId="40.[151,1436,151,968]" box="[199,1003,943,968]" pageId="40" pageNumber="41">
|
||
<emphasis id="B93FEAAA546DE2138ACD47B9FED30CD9" bold="true" box="[199,345,943,968]" pageId="40" pageNumber="41">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F35C7628546DE2138B6D47B9FE580CD9" box="[359,466,943,968]" name="Maldives" pageId="40" pageNumber="41">Maldives</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35C7628546DE2138BD747B9FD200CD9" box="[477,682,943,968]" name="Indonesia" pageId="40" pageNumber="41">Flores (Indonesia)</collectingCountry>
|
||
and
|
||
<collectingCountry id="F35C7628546DE21388EA47A6FCA90CD6" box="[736,803,944,967]" name="Japan" pageId="40" pageNumber="41">Japan</collectingCountry>
|
||
; subtidal to
|
||
<specimenCount id="9D4DFD31546DE21389A147B9FC610CD6" box="[939,1003,943,968]" pageId="40" pageNumber="41" type="generic">
|
||
<quantity id="4CB39B5D546DE21389A147B9FC6D0CD6" box="[939,999,943,968]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="40" pageNumber="41" unit="m" value="95.0">95 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |