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<document id="D3F4E5BF231BDFF88FADA2E3F9CFA262" ID-CLB-Dataset="5890" ID-DOI="10.11646/zootaxa.3773.1.1" ID-GBIF-Dataset="6da85b06-8386-41bc-9b34-b16eba941eab" ID-ISSN="1175-5326" ID-Zenodo-Dep="4909796" ID-ZooBank="19F28753-B2D0-4D1F-9D47-88886F7333FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623114397150" checkinUser="felipe" docAuthor="Ng, Peter K. L. &amp; Rahayu, Dwi Listyo" docDate="2014" docId="03E287AE545BE2258A9D4616FB150A42" docLanguage="en" docName="zt03773p063.pdf" docOrigin="Zootaxa 3773 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Raoulia galea Ng &amp; Rahayu 2014, sp. nov." docType="treatment" docVersion="6" lastPageNumber="31" masterDocId="FFDBFFD65445E23B8A0A4416FF8A0F11" masterDocTitle="Revision of the family Acidopsidae Števčić, 2005, and the systematic position of Typhlocarcinodes Alcock, 1900, Caecopilumnus Borradaile, 1902, and Raoulia Ng, 1987, with descriptions of two new genera and five new species (Crustacea: Brachyura: Goneplacoidea)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="31" updateTime="1699003055515" updateUser="ExternalLinkService">
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<mods:title id="884D3826FF854F47A2CFDA60EDD490BD">Revision of the family Acidopsidae Števčić, 2005, and the systematic position of Typhlocarcinodes Alcock, 1900, Caecopilumnus Borradaile, 1902, and Raoulia Ng, 1987, with descriptions of two new genera and five new species (Crustacea: Brachyura: Goneplacoidea)</mods:title>
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<mods:namePart id="01826B14CB791B7169DDE8A154FDA32C">Ng, Peter K. L.</mods:namePart>
<mods:nameIdentifier id="634306BC33CE7083E668BEF365CDCD42" type="email">dbsngkl@nus.edu.sg</mods:nameIdentifier>
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<mods:namePart id="1F6661340D4F6AD76E60741DA08DB469">Rahayu, Dwi Listyo</mods:namePart>
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<mods:title id="3E317021D2305393A780CEA79B00F8AB">Zootaxa</mods:title>
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<mods:date id="52806EDF1ED850C199399DE7D31333F4">2014</mods:date>
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<mods:number id="D1E30458B2F135254D56B20F1BA4FEE2">3773</mods:number>
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<treatment id="03E287AE545BE2258A9D4616FB150A42" ID-DOI="http://doi.org/10.5281/zenodo.4909774" ID-GBIF-Taxon="183228357" ID-Zenodo-Dep="4909774" LSID="urn:lsid:plazi:treatment:03E287AE545BE2258A9D4616FB150A42" httpUri="http://treatment.plazi.org/id/03E287AE545BE2258A9D4616FB150A42" lastPageNumber="31" pageId="30" pageNumber="31">
<subSubSection id="C3516533545BE2258A9D4616FE2E0D0B" box="[151,420,512,538]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BF436B8545BE2258A9D4616FE2E0D0B" blockId="30.[151,420,512,571]" box="[151,420,512,538]" pageId="30" pageNumber="31">
<heading id="D0BC81D4545BE2258A9D4616FE2E0D0B" bold="true" box="[151,420,512,538]" fontSize="11" level="2" pageId="30" pageNumber="31" reason="4">
<emphasis id="B93FEAAA545BE2258A9D4616FE2E0D0B" bold="true" box="[151,420,512,538]" pageId="30" pageNumber="31">
<taxonomicName id="4C4B4D3B545BE2258A9D4616FEB50D0B" authority="Ng &amp; Rahayu, 2014" authorityName="Ng &amp; Rahayu" authorityYear="2014" box="[151,319,512,538]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="galea" status="sp. nov.">
<emphasis id="B93FEAAA545BE2258A9D4616FEB50D0B" bold="true" box="[151,319,512,538]" italics="true" pageId="30" pageNumber="31">Raoulia galea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C57D1545BE2258B4D4616FE2E0D0B" box="[327,420,512,538]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3516533545BE2258A9D4634FE030D2A" box="[151,393,546,571]" pageId="30" pageNumber="31" type="description">
<paragraph id="8BF436B8545BE2258A9D4634FE030D2A" blockId="30.[151,420,512,571]" box="[151,393,546,571]" pageId="30" pageNumber="31">
(
<figureCitation id="13702A3D545BE2258A954634FE8A0D2A" box="[159,256,546,571]" captionStart="FIGURE 5" captionStartId="11.[151,249,1549,1571]" captionTargetBox="[151,1436,193,1527]" captionTargetId="figure-230@11.[151,1436,193,1527]" captionTargetPageId="11" captionText="FIGURE 5. Right third maxillipeds. A, Acidops cessacii (A. Milne-Edwards, 1878), syntype female (14.1 × 10.0 mm) (NHM 99.3.23.66) Cape Verde (laterally inverted); B, Parapilumnus cristimanus (A. Milne-Edwards, 1873), male (16.3 × 12.5 mm) (ZRC 2003.90), Nansha Is. (Spratly Is.), South China Sea (laterally inverted); C, Raoulia limosa Ng, 1987, holotype male (8.3 × 6.3 mm) (ZMUC M58), off Marongas, Jolo, Philippines; D, Raoulia piroculata (Rathbun, 1911), lectotype female (7.3 × 5.5 mm) (USNM 41359a), Amirante Is., Indian Ocean; E, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand; F, Raoulia fortis sp. nov., holotype male (12.0 × 9.2 mm) (MNHN), Santo, Vanuatu (laterally inverted); G, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia; H, Caecopilumnus hirsutus (Borradaile, 1902), male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia (laterally inverted); I, Caecopilumnus crassipes (Tesch, 1918), male (11.8 × 10.1 mm) (ZRC 2013.1710) Lombok, Indonesia; J, Caecopilumnus loculatus sp. nov., holotype male (8.0 × 6.7 mm) (ZRC 2013.1723), Moorea, French Polynesia (laterally inverted); K, Thecaplax capillosa gen. et. sp. nov., holotype male (11.6 × 9.5 mm) (MZB), Lombok, Indonesia; L, Typhlocarcinodes integrifrons (Miers, 1881), lectotype male (8.5 × 6.9 mm) (NHM 81.24a), Goree I., Senegal." figureDoi="http://doi.org/10.5281/zenodo.4909806" httpUri="https://zenodo.org/record/4909806/files/figure.png" pageId="30" pageNumber="31">Figs. 5E</figureCitation>
,
<figureCitation id="13702A3D545BE2258B064634FEA00D2A" box="[268,298,546,571]" captionStart="FIGURE 18" captionStartId="28.[151,249,1877,1899]" captionTargetBox="[174,1413,193,1856]" captionTargetId="figure-65@28.[174,1413,193,1856]" captionTargetPageId="28" captionText="FIGURE 18. Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909832" httpUri="https://zenodo.org/record/4909832/files/figure.png" pageId="30" pageNumber="31">18</figureCitation>
,
<figureCitation id="13702A3D545BE2258B3F4634FE0B0D2A" box="[309,385,546,571]" captionStart="FIGURE 19" captionStartId="31.[151,249,1936,1958]" captionTargetBox="[204,1363,213,1901]" captionTargetId="figure-93@31.[179,1399,193,1917]" captionTargetPageId="31" captionText="FIGURE 19. AE, Raoulia piroculata (Rathbun, 1911), male (11.0 × 8.5 mm), South Africa; FL, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, male abdomen; B, left third maxilliped; C, D, left G1; E, F, distal part of left G1; G, left G2. Scales: A, B = 1.0 mm; C, D, G = 0.5 mm; E, F = 0.1 mm. AE after Barnard (1955: fig. 16)." figureDoi="http://doi.org/10.5281/zenodo.4909834" httpUri="https://zenodo.org/record/4909834/files/figure.png" pageId="30" pageNumber="31">19FL</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3516533545BE2258A9D467CFE4A0DB6" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BF436B8545BE2258A9D467CFE4A0DB6" blockId="30.[151,1436,618,1363]" pageId="30" pageNumber="31">
<emphasis id="B93FEAAA545BE2258A9D467CFEF20D92" bold="true" box="[151,376,618,643]" pageId="30" pageNumber="31">Material examined</emphasis>
.
<materialsCitation id="3B233CE5545BE2258B8C467CFE4A0DB6" ID-GBIF-Occurrence="3312327333" collectingDate="2000-05-03" collectingDateMax="2000-05-06" collectingDateMin="2000-05-03" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Thailand" location="Phuket" pageId="30" pageNumber="31" specimenCode="ZRC 2000.0980" specimenCount="1" stateProvince="Phuket" typeStatus="holotype">
<typeStatus id="54F0881A545BE2258B8C467CFE780D92" box="[390,498,618,643]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
: male (9.4 ×
<quantity id="4CB39B5D545BE2258889467CFD500D92" box="[643,730,618,643]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="30" pageNumber="31" unit="mm" value="7.4">7.4 mm</quantity>
) (
<specimenCode id="DBED9EC3545BE22588FA467CFC200D92" box="[752,938,618,643]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="30" pageNumber="31">ZRC 2000.0980</specimenCode>
), intertidal mud flats,
<collectingRegion id="498FF85A545BE2258E94467CFB650D92" box="[1182,1263,618,643]" country="Thailand" name="Phuket" pageId="30" pageNumber="31">Phuket</collectingRegion>
,
<collectingCountry id="F35C7628545BE2258EF0467CFAEA0D92" box="[1274,1376,618,643]" name="Thailand" pageId="30" pageNumber="31">Thailand</collectingCountry>
, coll.
<collectorName id="26BE536E545BE2258A9D4699FE800DB6" box="[151,266,655,679]" pageId="30" pageNumber="31">P.K.L. Ng</collectorName>
,
<date id="FFF51078545BE2258B1F4698FE360DB6" box="[277,444,654,679]" pageId="30" pageNumber="31" value="2000-05-03" valueMax="2000-05-06" valueMin="2000-05-03">
<collectingDate id="EFB1E990545BE2258B1F4698FE360DB6" box="[277,444,654,679]" pageId="30" pageNumber="31" value="2000-05-03" valueMax="2000-05-06" valueMin="2000-05-03">36 May 2000</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3516533545BE2258ACD46A4FADC0C6E" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="8BF436B8545BE2258ACD46A4FF700C4A" blockId="30.[151,1436,618,1363]" pageId="30" pageNumber="31">
<emphasis id="B93FEAAA545BE2258ACD46A4FEB00DDA" bold="true" box="[199,314,690,715]" pageId="30" pageNumber="31">Diagnosis</emphasis>
. Carapace width 1.3 times length (
<figureCitation id="13702A3D545BE22588B446A4FCAC0DDA" box="[702,806,690,715]" captionStart="FIGURE 18" captionStartId="28.[151,249,1877,1899]" captionTargetBox="[174,1413,193,1856]" captionTargetId="figure-65@28.[174,1413,193,1856]" captionTargetPageId="28" captionText="FIGURE 18. Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909832" httpUri="https://zenodo.org/record/4909832/files/figure.png" pageId="30" pageNumber="31">Fig. 18A</figureCitation>
). Junction between frontal, supraorbital margins appears gently curved in frontal view (
<figureCitation id="13702A3D545BE2258BFB46C0FDD20DFE" box="[497,600,726,751]" captionStart="FIGURE 18" captionStartId="28.[151,249,1877,1899]" captionTargetBox="[174,1413,193,1856]" captionTargetId="figure-65@28.[174,1413,193,1856]" captionTargetPageId="28" captionText="FIGURE 18. Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909832" httpUri="https://zenodo.org/record/4909832/files/figure.png" pageId="30" pageNumber="31">Fig. 18C</figureCitation>
). Third maxilliped merus shorter than ischium; ischium quadrangular, 1.1 times as long as broad (
<figureCitation id="13702A3D545BE2258BAC46ECFD820C02" box="[422,520,762,787]" captionStart="FIGURE 5" captionStartId="11.[151,249,1549,1571]" captionTargetBox="[151,1436,193,1527]" captionTargetId="figure-230@11.[151,1436,193,1527]" captionTargetPageId="11" captionText="FIGURE 5. Right third maxillipeds. A, Acidops cessacii (A. Milne-Edwards, 1878), syntype female (14.1 × 10.0 mm) (NHM 99.3.23.66) Cape Verde (laterally inverted); B, Parapilumnus cristimanus (A. Milne-Edwards, 1873), male (16.3 × 12.5 mm) (ZRC 2003.90), Nansha Is. (Spratly Is.), South China Sea (laterally inverted); C, Raoulia limosa Ng, 1987, holotype male (8.3 × 6.3 mm) (ZMUC M58), off Marongas, Jolo, Philippines; D, Raoulia piroculata (Rathbun, 1911), lectotype female (7.3 × 5.5 mm) (USNM 41359a), Amirante Is., Indian Ocean; E, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand; F, Raoulia fortis sp. nov., holotype male (12.0 × 9.2 mm) (MNHN), Santo, Vanuatu (laterally inverted); G, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia; H, Caecopilumnus hirsutus (Borradaile, 1902), male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia (laterally inverted); I, Caecopilumnus crassipes (Tesch, 1918), male (11.8 × 10.1 mm) (ZRC 2013.1710) Lombok, Indonesia; J, Caecopilumnus loculatus sp. nov., holotype male (8.0 × 6.7 mm) (ZRC 2013.1723), Moorea, French Polynesia (laterally inverted); K, Thecaplax capillosa gen. et. sp. nov., holotype male (11.6 × 9.5 mm) (MZB), Lombok, Indonesia; L, Typhlocarcinodes integrifrons (Miers, 1881), lectotype male (8.5 × 6.9 mm) (NHM 81.24a), Goree I., Senegal." figureDoi="http://doi.org/10.5281/zenodo.4909806" httpUri="https://zenodo.org/record/4909806/files/figure.png" pageId="30" pageNumber="31">Figs. 5E</figureCitation>
,
<figureCitation id="13702A3D545BE225881F46ECFDC30C02" box="[533,585,762,787]" captionStart="FIGURE 19" captionStartId="31.[151,249,1936,1958]" captionTargetBox="[204,1363,213,1901]" captionTargetId="figure-93@31.[179,1399,193,1917]" captionTargetPageId="31" captionText="FIGURE 19. AE, Raoulia piroculata (Rathbun, 1911), male (11.0 × 8.5 mm), South Africa; FL, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, male abdomen; B, left third maxilliped; C, D, left G1; E, F, distal part of left G1; G, left G2. Scales: A, B = 1.0 mm; C, D, G = 0.5 mm; E, F = 0.1 mm. AE after Barnard (1955: fig. 16)." figureDoi="http://doi.org/10.5281/zenodo.4909834" httpUri="https://zenodo.org/record/4909834/files/figure.png" pageId="30" pageNumber="31">19G</figureCitation>
). Ambulatory legs short; merus of last ambulatory leg 2.8 times as long as broad (
<figureCitation id="13702A3D545BE2258AEF4708FEC70C26" box="[229,333,798,823]" captionStart="FIGURE 18" captionStartId="28.[151,249,1877,1899]" captionTargetBox="[174,1413,193,1856]" captionTargetId="figure-65@28.[174,1413,193,1856]" captionTargetPageId="28" captionText="FIGURE 18. Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909832" httpUri="https://zenodo.org/record/4909832/files/figure.png" pageId="30" pageNumber="31">Fig. 18A</figureCitation>
). G1 with distal half distinctly curving towards abdomen in situ, distal part gradually tapering (
<figureCitation id="13702A3D545BE2258F654708FF640C4A" captionStart="FIGURE 19" captionStartId="31.[151,249,1936,1958]" captionTargetBox="[204,1363,213,1901]" captionTargetId="figure-93@31.[179,1399,193,1917]" captionTargetPageId="31" captionText="FIGURE 19. AE, Raoulia piroculata (Rathbun, 1911), male (11.0 × 8.5 mm), South Africa; FL, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, male abdomen; B, left third maxilliped; C, D, left G1; E, F, distal part of left G1; G, left G2. Scales: A, B = 1.0 mm; C, D, G = 0.5 mm; E, F = 0.1 mm. AE after Barnard (1955: fig. 16)." figureDoi="http://doi.org/10.5281/zenodo.4909834" httpUri="https://zenodo.org/record/4909834/files/figure.png" pageId="30" pageNumber="31">Fig. 19HK</figureCitation>
).
</paragraph>
<paragraph id="8BF436B8545BE2258ACD4770FADC0C6E" blockId="30.[151,1436,618,1363]" box="[199,1366,870,895]" pageId="30" pageNumber="31">
<emphasis id="B93FEAAA545BE2258ACD4770FE900C6E" bold="true" box="[199,282,870,895]" pageId="30" pageNumber="31">Colour</emphasis>
. The freshly collected specimen has a dirty white carapace and pereopods, with the setae brown.
</paragraph>
</subSubSection>
<subSubSection id="C3516533545BE2258ACD479CFD790CD6" pageId="30" pageNumber="31" type="etymology">
<paragraph id="8BF436B8545BE2258ACD479CFD790CD6" blockId="30.[151,1436,618,1363]" pageId="30" pageNumber="31">
<emphasis id="B93FEAAA545BE2258ACD479CFECC0CB2" bold="true" box="[199,326,906,931]" pageId="30" pageNumber="31">Etymology</emphasis>
. The name is derived from Latin
<taxonomicName id="4C4B4D3B545BE22588CB479CFD750CB2" authorityName="Ng &amp; Rahayu" authorityYear="2014" box="[705,767,906,931]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="galea">
<emphasis id="B93FEAAA545BE22588CB479CFD750CB2" box="[705,767,906,931]" italics="true" pageId="30" pageNumber="31">galea</emphasis>
</taxonomicName>
for helmet, alluding to the general shape of the carapace of the species. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3516533545BE2258ACD47C4FC2C0A3E" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BF436B8545BE2258ACD47C4FD9F0BF6" blockId="30.[151,1436,618,1363]" pageId="30" pageNumber="31">
<emphasis id="B93FEAAA545BE2258ACD47C4FEBE0CFA" bold="true" box="[199,308,978,1003]" pageId="30" pageNumber="31">Remarks</emphasis>
.
<taxonomicName id="4C4B4D3B545BE2258B4D47C4FE600CFA" authority="Ng &amp; Rahayu, 2014" authorityName="Ng &amp; Rahayu" authorityYear="2014" box="[327,490,978,1003]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="galea" status="sp. nov.">
<emphasis id="B93FEAAA545BE2258B4D47C4FE600CFA" box="[327,490,978,1003]" italics="true" pageId="30" pageNumber="31">Raoulia galea</emphasis>
</taxonomicName>
<emphasis id="B93FEAAA545BE2258BFD47C4FDDD0CFA" bold="true" box="[503,599,978,1003]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="A20C57D1545BE2258BFD47C4FDDD0CFA" box="[503,599,978,1003]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="4C4B4D3B545BE225894847C5FC3B0CFA" authorityName="Ng" authorityYear="1987" box="[834,945,978,1003]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="limosa">
<emphasis id="B93FEAAA545BE225894847C5FC3B0CFA" box="[834,945,978,1003]" italics="true" pageId="30" pageNumber="31">R. limosa</emphasis>
</taxonomicName>
, but it can be separated by possessing a somewhat a more quadrate carapace (
<figureCitation id="13702A3D545BE225883647E0FD050B1E" box="[572,655,1014,1039]" captionStart="FIGURE 18" captionStartId="28.[151,249,1877,1899]" captionTargetBox="[174,1413,193,1856]" captionTargetId="figure-65@28.[174,1413,193,1856]" captionTargetPageId="28" captionText="FIGURE 18. Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909832" httpUri="https://zenodo.org/record/4909832/files/figure.png" pageId="30" pageNumber="31">Fig. 18</figureCitation>
;
<figureCitation id="13702A3D545BE225889147E0FC860B1E" box="[667,780,1014,1039]" captionStart="FIGURE 13" captionStartId="23.[151,249,1877,1899]" captionTargetBox="[192,1395,193,1856]" captionTargetId="figure-66@23.[192,1395,193,1856]" captionTargetPageId="23" captionText="FIGURE 13. Raoulia limosa Ng, 1987, holotype male (8.3 × 6.3 mm) (ZMUC M58), Jolo, Philippines. A, dorsal view; B, male abdomen; C, anterior thoracic sternum and sterno-abdominal cavity; D, frontal view of carapace; E, posterior thoracic sternum; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909822" httpUri="https://zenodo.org/record/4909822/files/figure.png" pageId="30" pageNumber="31">Figs. 13A</figureCitation>
,
<figureCitation id="13702A3D545BE225891247E0FCC10B1E" box="[792,843,1014,1039]" captionStart="FIGURE 14" captionStartId="24.[151,249,1708,1730]" captionTargetBox="[151,1436,193,1687]" captionTargetId="figure-139@24.[151,1436,193,1687]" captionTargetPageId="24" captionText="FIGURE 14. Raoulia limosa Ng, 1987, male (8.3 × 6.5 mm) (ZRC 2013.1703), Lombok, Indonesia. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909824" httpUri="https://zenodo.org/record/4909824/files/figure.png" pageId="30" pageNumber="31">14A</figureCitation>
,
<figureCitation id="13702A3D545BE225895C47E0FC0D0B1E" box="[854,903,1014,1039]" captionStart="FIGURE 15" captionStartId="25.[151,249,1978,2000]" captionTargetBox="[151,1436,545,1956]" captionTargetId="figure-190@25.[151,1436,545,1956]" captionTargetPageId="25" captionText="FIGURE 15. Raoulia limosa Ng, 1987, male (4.8 × 3.5 mm) (ZMA 240126), Talaut Is., Philippines. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity." figureDoi="http://doi.org/10.5281/zenodo.4909826" httpUri="https://zenodo.org/record/4909826/files/figure.png" pageId="30" pageNumber="31">15A</figureCitation>
for
<taxonomicName id="4C4B4D3B545BE22589BF47E1FB950B1E" authorityName="Ng" authorityYear="1987" box="[949,1055,1014,1039]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="limosa">
<emphasis id="B93FEAAA545BE22589BF47E1FB950B1E" box="[949,1055,1014,1039]" italics="true" pageId="30" pageNumber="31">R. limosa</emphasis>
</taxonomicName>
); the junction between the frontal and supraorbital margins of
<taxonomicName id="4C4B4D3B545BE2258BD8400DFDA50B22" authorityName="Ng &amp; Rahayu" authorityYear="2014" box="[466,559,1050,1075]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="galea">
<emphasis id="B93FEAAA545BE2258BD8400DFDA50B22" box="[466,559,1050,1075]" italics="true" pageId="30" pageNumber="31">R. galea</emphasis>
</taxonomicName>
is gently curved (in frontal view) (
<figureCitation id="13702A3D545BE22589BD400CFB910B22" box="[951,1051,1050,1075]" captionStart="FIGURE 18" captionStartId="28.[151,249,1877,1899]" captionTargetBox="[174,1413,193,1856]" captionTargetId="figure-65@28.[174,1413,193,1856]" captionTargetPageId="28" captionText="FIGURE 18. Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909832" httpUri="https://zenodo.org/record/4909832/files/figure.png" pageId="30" pageNumber="31">Fig. 18C</figureCitation>
) (at right angles in
<taxonomicName id="4C4B4D3B545BE2258EFD400DFAEB0B22" authorityName="Ng" authorityYear="1987" box="[1271,1377,1050,1075]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="limosa">
<emphasis id="B93FEAAA545BE2258EFD400DFAEB0B22" box="[1271,1377,1050,1075]" italics="true" pageId="30" pageNumber="31">R. limosa</emphasis>
</taxonomicName>
;
<figureCitation id="13702A3D545BE2258F7A400CFF400B46" captionStart="FIGURE 13" captionStartId="23.[151,249,1877,1899]" captionTargetBox="[192,1395,193,1856]" captionTargetId="figure-66@23.[192,1395,193,1856]" captionTargetPageId="23" captionText="FIGURE 13. Raoulia limosa Ng, 1987, holotype male (8.3 × 6.3 mm) (ZMUC M58), Jolo, Philippines. A, dorsal view; B, male abdomen; C, anterior thoracic sternum and sterno-abdominal cavity; D, frontal view of carapace; E, posterior thoracic sternum; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909822" httpUri="https://zenodo.org/record/4909822/files/figure.png" pageId="30" pageNumber="31">Fig. 13D</figureCitation>
,
<figureCitation id="13702A3D545BE2258ADC4028FE820B46" box="[214,264,1086,1111]" captionStart="FIGURE 14" captionStartId="24.[151,249,1708,1730]" captionTargetBox="[151,1436,193,1687]" captionTargetId="figure-139@24.[151,1436,193,1687]" captionTargetPageId="24" captionText="FIGURE 14. Raoulia limosa Ng, 1987, male (8.3 × 6.5 mm) (ZRC 2013.1703), Lombok, Indonesia. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909824" httpUri="https://zenodo.org/record/4909824/files/figure.png" pageId="30" pageNumber="31">14C</figureCitation>
); the merus of the third maxilliped is more subovate, with the anterolateral margin gently convex (
<figureCitation id="13702A3D545BE2258F6E4028FF320B6A" captionStart="FIGURE 5" captionStartId="11.[151,249,1549,1571]" captionTargetBox="[151,1436,193,1527]" captionTargetId="figure-230@11.[151,1436,193,1527]" captionTargetPageId="11" captionText="FIGURE 5. Right third maxillipeds. A, Acidops cessacii (A. Milne-Edwards, 1878), syntype female (14.1 × 10.0 mm) (NHM 99.3.23.66) Cape Verde (laterally inverted); B, Parapilumnus cristimanus (A. Milne-Edwards, 1873), male (16.3 × 12.5 mm) (ZRC 2003.90), Nansha Is. (Spratly Is.), South China Sea (laterally inverted); C, Raoulia limosa Ng, 1987, holotype male (8.3 × 6.3 mm) (ZMUC M58), off Marongas, Jolo, Philippines; D, Raoulia piroculata (Rathbun, 1911), lectotype female (7.3 × 5.5 mm) (USNM 41359a), Amirante Is., Indian Ocean; E, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand; F, Raoulia fortis sp. nov., holotype male (12.0 × 9.2 mm) (MNHN), Santo, Vanuatu (laterally inverted); G, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia; H, Caecopilumnus hirsutus (Borradaile, 1902), male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia (laterally inverted); I, Caecopilumnus crassipes (Tesch, 1918), male (11.8 × 10.1 mm) (ZRC 2013.1710) Lombok, Indonesia; J, Caecopilumnus loculatus sp. nov., holotype male (8.0 × 6.7 mm) (ZRC 2013.1723), Moorea, French Polynesia (laterally inverted); K, Thecaplax capillosa gen. et. sp. nov., holotype male (11.6 × 9.5 mm) (MZB), Lombok, Indonesia; L, Typhlocarcinodes integrifrons (Miers, 1881), lectotype male (8.5 × 6.9 mm) (NHM 81.24a), Goree I., Senegal." figureDoi="http://doi.org/10.5281/zenodo.4909806" httpUri="https://zenodo.org/record/4909806/files/figure.png" pageId="30" pageNumber="31">Figs. 5E</figureCitation>
,
<figureCitation id="13702A3D545BE2258ACE4074FF7C0B6A" box="[196,246,1122,1147]" captionStart="FIGURE 19" captionStartId="31.[151,249,1936,1958]" captionTargetBox="[204,1363,213,1901]" captionTargetId="figure-93@31.[179,1399,193,1917]" captionTargetPageId="31" captionText="FIGURE 19. AE, Raoulia piroculata (Rathbun, 1911), male (11.0 × 8.5 mm), South Africa; FL, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, male abdomen; B, left third maxilliped; C, D, left G1; E, F, distal part of left G1; G, left G2. Scales: A, B = 1.0 mm; C, D, G = 0.5 mm; E, F = 0.1 mm. AE after Barnard (1955: fig. 16)." figureDoi="http://doi.org/10.5281/zenodo.4909834" httpUri="https://zenodo.org/record/4909834/files/figure.png" pageId="30" pageNumber="31">19G</figureCitation>
) (more rounded, with the anterolateral margin markedly convex in
<taxonomicName id="4C4B4D3B545BE22589E44075FBD30B6A" authorityName="Ng" authorityYear="1987" box="[1006,1113,1122,1147]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="limosa">
<emphasis id="B93FEAAA545BE22589E44075FBD30B6A" box="[1006,1113,1122,1147]" italics="true" pageId="30" pageNumber="31">R. limosa</emphasis>
</taxonomicName>
;
<figureCitation id="13702A3D545BE2258E624074FB400B6A" box="[1128,1226,1122,1147]" captionStart="FIGURE 5" captionStartId="11.[151,249,1549,1571]" captionTargetBox="[151,1436,193,1527]" captionTargetId="figure-230@11.[151,1436,193,1527]" captionTargetPageId="11" captionText="FIGURE 5. Right third maxillipeds. A, Acidops cessacii (A. Milne-Edwards, 1878), syntype female (14.1 × 10.0 mm) (NHM 99.3.23.66) Cape Verde (laterally inverted); B, Parapilumnus cristimanus (A. Milne-Edwards, 1873), male (16.3 × 12.5 mm) (ZRC 2003.90), Nansha Is. (Spratly Is.), South China Sea (laterally inverted); C, Raoulia limosa Ng, 1987, holotype male (8.3 × 6.3 mm) (ZMUC M58), off Marongas, Jolo, Philippines; D, Raoulia piroculata (Rathbun, 1911), lectotype female (7.3 × 5.5 mm) (USNM 41359a), Amirante Is., Indian Ocean; E, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand; F, Raoulia fortis sp. nov., holotype male (12.0 × 9.2 mm) (MNHN), Santo, Vanuatu (laterally inverted); G, Raoulia calva sp. nov., holotype male (6.6 × 5.2 mm) (ZRC 2013.1721), Society Is., French Polynesia; H, Caecopilumnus hirsutus (Borradaile, 1902), male (7.1 × 5.6 mm) (ZMA 261027), Flores, Indonesia (laterally inverted); I, Caecopilumnus crassipes (Tesch, 1918), male (11.8 × 10.1 mm) (ZRC 2013.1710) Lombok, Indonesia; J, Caecopilumnus loculatus sp. nov., holotype male (8.0 × 6.7 mm) (ZRC 2013.1723), Moorea, French Polynesia (laterally inverted); K, Thecaplax capillosa gen. et. sp. nov., holotype male (11.6 × 9.5 mm) (MZB), Lombok, Indonesia; L, Typhlocarcinodes integrifrons (Miers, 1881), lectotype male (8.5 × 6.9 mm) (NHM 81.24a), Goree I., Senegal." figureDoi="http://doi.org/10.5281/zenodo.4909806" httpUri="https://zenodo.org/record/4909806/files/figure.png" pageId="30" pageNumber="31">Figs. 5C</figureCitation>
,
<figureCitation id="13702A3D545BE2258EDC4074FA820B6A" box="[1238,1288,1122,1147]" captionStart="FIGURE 16" captionStartId="26.[151,249,1634,1656]" captionTargetBox="[168,1435,209,1606]" captionTargetId="figure-197@26.[151,1436,193,1613]" captionTargetPageId="26" captionText="FIGURE 16. Raoulia limosa Ng, 1987. AH, holotype male (8.3 × 6.3 mm) (ZMUC M58), Jolo, Philippines; I, female (9.5 × 7.3 mm) (ZRC 2013.1705), Lombok, Indonesia. A, male abdomen; B, left third maxilliped; C, frontal view of carapace showing antennules, antennae, epistome and orbit; D, E, left G1; F, G, distal part of left G1; H, left G2; I, female abdomen. Scales: AC, I = 1.0 mm; D, E, H = 0.5 mm; F, G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4909828" httpUri="https://zenodo.org/record/4909828/files/figure.png" pageId="30" pageNumber="31">16B</figureCitation>
). The G1s of the two species are superficially similar, but in
<taxonomicName id="4C4B4D3B545BE22588BB4091FC850B8E" authority="Ng &amp; Rahayu, 2014" authorityName="Ng &amp; Rahayu" authorityYear="2014" box="[689,783,1158,1183]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="galea" status="sp. nov.">
<emphasis id="B93FEAAA545BE22588BB4091FC850B8E" box="[689,783,1158,1183]" italics="true" pageId="30" pageNumber="31">R. galea</emphasis>
</taxonomicName>
<emphasis id="B93FEAAA545BE22589124090FCF90B8E" bold="true" box="[792,883,1158,1183]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="A20C57D1545BE22589124090FCF90B8E" box="[792,883,1158,1183]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the distal half curves towards the abdomen (i.e. directed ventrally) in situ (
<figureCitation id="13702A3D545BE2258BCE40BCFDDA0BD2" box="[452,592,1194,1219]" captionStart="FIGURE 19" captionStartId="31.[151,249,1936,1958]" captionTargetBox="[204,1363,213,1901]" captionTargetId="figure-93@31.[179,1399,193,1917]" captionTargetPageId="31" captionText="FIGURE 19. AE, Raoulia piroculata (Rathbun, 1911), male (11.0 × 8.5 mm), South Africa; FL, Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, male abdomen; B, left third maxilliped; C, D, left G1; E, F, distal part of left G1; G, left G2. Scales: A, B = 1.0 mm; C, D, G = 0.5 mm; E, F = 0.1 mm. AE after Barnard (1955: fig. 16)." figureDoi="http://doi.org/10.5281/zenodo.4909834" httpUri="https://zenodo.org/record/4909834/files/figure.png" pageId="30" pageNumber="31">Fig. 19HK</figureCitation>
), while in
<taxonomicName id="4C4B4D3B545BE22588C240BDFCB90BD2" authorityName="Ng" authorityYear="1987" box="[712,819,1194,1219]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="limosa">
<emphasis id="B93FEAAA545BE22588C240BDFCB90BD2" box="[712,819,1194,1219]" italics="true" pageId="30" pageNumber="31">R. limosa</emphasis>
</taxonomicName>
, it is bent outwards (
<figureCitation id="13702A3D545BE2258E2940BCFB250BD2" box="[1059,1199,1194,1219]" captionStart="FIGURE 16" captionStartId="26.[151,249,1634,1656]" captionTargetBox="[168,1435,209,1606]" captionTargetId="figure-197@26.[151,1436,193,1613]" captionTargetPageId="26" captionText="FIGURE 16. Raoulia limosa Ng, 1987. AH, holotype male (8.3 × 6.3 mm) (ZMUC M58), Jolo, Philippines; I, female (9.5 × 7.3 mm) (ZRC 2013.1705), Lombok, Indonesia. A, male abdomen; B, left third maxilliped; C, frontal view of carapace showing antennules, antennae, epistome and orbit; D, E, left G1; F, G, distal part of left G1; H, left G2; I, female abdomen. Scales: AC, I = 1.0 mm; D, E, H = 0.5 mm; F, G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4909828" httpUri="https://zenodo.org/record/4909828/files/figure.png" pageId="30" pageNumber="31">Fig. 16DG</figureCitation>
). This argues against the two species being conspecific.
</paragraph>
<paragraph id="8BF436B8545BE2258ACD40E4FC2C0A3E" blockId="30.[151,1436,618,1363]" pageId="30" pageNumber="31">
Although
<taxonomicName id="4C4B4D3B545BE2258B3140E5FE100A1A" authority="Ng &amp; Rahayu, 2014" authorityName="Ng &amp; Rahayu" authorityYear="2014" box="[315,410,1266,1291]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="galea" status="sp. nov.">
<emphasis id="B93FEAAA545BE2258B3140E5FE100A1A" box="[315,410,1266,1291]" italics="true" pageId="30" pageNumber="31">R. galea</emphasis>
</taxonomicName>
<emphasis id="B93FEAAA545BE2258BAE40E4FD8B0A1A" bold="true" box="[420,513,1266,1291]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="A20C57D1545BE2258BAE40E4FD8B0A1A" box="[420,513,1266,1291]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is also found in the Indian Ocean like
<taxonomicName id="4C4B4D3B545BE22589C640E5FBE80A1A" baseAuthorityName="Rathbun" baseAuthorityYear="1911" box="[972,1122,1266,1291]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="piroculata">
<emphasis id="B93FEAAA545BE22589C640E5FBE80A1A" box="[972,1122,1266,1291]" italics="true" pageId="30" pageNumber="31">R. piroculata</emphasis>
</taxonomicName>
, its carapace appears more rounded in general appearance (
<figureCitation id="13702A3D545BE2258BF74100FDEE0A3E" box="[509,612,1302,1327]" captionStart="FIGURE 18" captionStartId="28.[151,249,1877,1899]" captionTargetBox="[174,1413,193,1856]" captionTargetId="figure-65@28.[174,1413,193,1856]" captionTargetPageId="28" captionText="FIGURE 18. Raoulia galea sp. nov., holotype male (9.4 × 7.4 mm) (ZRC 2000.0980), Phuket, Thailand. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909832" httpUri="https://zenodo.org/record/4909832/files/figure.png" pageId="30" pageNumber="31">Fig. 18A</figureCitation>
;
<figureCitation id="13702A3D545BE225887A4100FD5E0A3E" box="[624,724,1302,1327]" captionStart="FIGURE 17" captionStartId="27.[151,249,1905,1927]" captionTargetBox="[151,1436,373,1884]" captionTargetId="figure-144@27.[151,1436,373,1884]" captionTargetPageId="27" captionText="FIGURE 17. Raoulia piroculata (Rathbun, 1911), lectotype female (7.3 × 5.5 mm) (USNM 41359a), Amirante Is., Indian Ocean. A, dorsal view; B, anterior thoracic sternum and sterno-abdominal cavity; C, frontal view of carapace; D, posterior thoracic sternum; E, outer view of right chela; F, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.4909830" httpUri="https://zenodo.org/record/4909830/files/figure.png" pageId="30" pageNumber="31">Fig. 17A</figureCitation>
for
<taxonomicName id="4C4B4D3B545BE22589084101FC1F0A3E" baseAuthorityName="Rathbun" baseAuthorityYear="1911" box="[770,917,1302,1327]" class="Malacostraca" family="Chasmocarcinidae" genus="Raoulia" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="piroculata">
<emphasis id="B93FEAAA545BE22589084101FC1F0A3E" box="[770,917,1302,1327]" italics="true" pageId="30" pageNumber="31">R. piroculata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3516533545BE2258ACD412CFB150A42" box="[199,1183,1338,1363]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BF436B8545BE2258ACD412CFB150A42" blockId="30.[151,1436,618,1363]" box="[199,1183,1338,1363]" pageId="30" pageNumber="31">
<emphasis id="B93FEAAA545BE2258ACD412CFED30A42" bold="true" box="[199,345,1338,1363]" pageId="30" pageNumber="31">Distribution</emphasis>
.
<collectingRegion id="498FF85A545BE2258B6D412CFE320A42" box="[359,440,1338,1363]" country="Thailand" name="Phuket" pageId="30" pageNumber="31">Phuket</collectingRegion>
, Andaman Sea coast of
<collectingCountry id="F35C7628545BE22588CF412CFCA10A42" box="[709,811,1338,1363]" name="Thailand" pageId="30" pageNumber="31">Thailand</collectingCountry>
, eastern Indian Ocean; intertidal.
</paragraph>
</subSubSection>
</treatment>
</document>