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<document id="643E9F2FCBCFCBF393E88EAB7820DAED" ID-CLB-Dataset="22328" ID-DOI="10.11646/zootaxa.4772.1.9" ID-GBIF-Dataset="7f1c7500-472a-4211-beaf-3277b400f99f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3813997" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588834923358" checkinUser="plazi" docAuthor="Boubakri, Abir, Bosmans, Robert, Pétillon, Julien &amp; Achouri, Mohamed Sghaier" docDate="2020" docId="03E2878E6918FFA11BE6F9DDA842F3F3" docLanguage="en" docName="zootaxa.4772.1.9.pdf" docOrigin="Zootaxa 4772 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Agraecina salsicola Bosmans &amp; Boubakri 2020, sp. n." docType="treatment" docVersion="8" lastPageNumber="196" masterDocId="FFDBFFF66918FFA01B71FF9BA94FF221" masterDocTitle="Description of a new Agraecina species from Tunisia (Araneae: Liocranidae), with a review of all species of the genus" masterLastPageNumber="200" masterPageNumber="195" pageNumber="195" updateTime="1698838880390" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8A7D99557F8B2038754455B880495479">Description of a new Agraecina species from Tunisia (Araneae: Liocranidae), with a review of all species of the genus</mods:title>
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<mods:namePart id="10471B835EAB4A453301B4DFE4B89979">Boubakri, Abir</mods:namePart>
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<mods:affiliation id="DD528BFAC9B4363A215A4FE387108C1E">Terrestrial Ecology Unit, Gent University, Gent, Belgium. https: // orcid. org / 0000 - 0003 - 4705 - 8587</mods:affiliation>
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<mods:namePart id="4719DB7C970618B85C27974D253AF3C4">Pétillon, Julien</mods:namePart>
<mods:affiliation id="FAF4AE59C76E342C505A5D5D0DEE5FFE">UMR CNRS 6553 Ecobio, Universit de Rennes, Rennes, France. https: // orcid. org / 0000 - 0002 - 7611 - 5133</mods:affiliation>
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<treatment id="03E2878E6918FFA11BE6F9DDA842F3F3" ID-DOI="http://doi.org/10.5281/zenodo.3815556" ID-GBIF-Taxon="163975602" ID-Zenodo-Dep="3815556" LSID="urn:lsid:plazi:treatment:03E2878E6918FFA11BE6F9DDA842F3F3" httpUri="http://treatment.plazi.org/id/03E2878E6918FFA11BE6F9DDA842F3F3" lastPageId="1" lastPageNumber="196" pageId="0" pageNumber="195">
<subSubSection id="C35165136918FFA01BE6F9DDAA29F47E" box="[151,870,1605,1632]" pageId="0" pageNumber="195" type="nomenclature">
<paragraph id="8BF436986918FFA01BE6F9DDAA29F47E" blockId="0.[151,1437,1498,2028]" box="[151,870,1605,1632]" pageId="0" pageNumber="195">
<heading id="D0BC81F46918FFA01BE6F9DDAA29F47E" bold="true" box="[151,870,1605,1632]" fontSize="11" level="1" pageId="0" pageNumber="195" reason="1">
<emphasis id="B93FEA8A6918FFA01BE6F9DDAA29F47E" bold="true" box="[151,870,1605,1632]" pageId="0" pageNumber="195">
<taxonomicName id="4C4B4D1B6918FFA01BE6F9DDABD5F47E" ID-CoL="98ZQT" authority="Bosmans &amp; Boubakri" authorityName="Bosmans &amp; Boubakri" authorityYear="2020" box="[151,666,1605,1632]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="195" phylum="Arthropoda" rank="species" species="salsicola" status="sp. nov.">
<emphasis id="B93FEA8A6918FFA01BE6F9DDA8CBF441" bold="true" box="[151,388,1605,1632]" italics="true" pageId="0" pageNumber="195">Agraecina salsicola</emphasis>
Bosmans &amp; Boubakri
</taxonomicName>
<taxonomicNameLabel id="A20C57F16918FFA019D3F9DDABABF441" box="[674,740,1606,1632]" pageId="0" pageNumber="195" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="13702A1D6918FFA01982F9DEAA12F47E" box="[755,861,1605,1631]" captionStart="FIGURES 14" captionStartId="1.[151,258,1031,1053]" captionTargetBox="[151,1435,472,1002]" captionTargetId="figure@1.[151,1435,472,1002]" captionTargetPageId="1" captionText="FIGURES 14. Agraecina salsicola sp. n. 1. Male, dorsal view; 2. Male palp, retrolateral view; 3. Idem, ventral view; 4. Idem, prolateral view. Abbreviations: E—embolus; MA—median apophysis; RTA—retrolateral tibial apohysis." figureDoi="http://doi.org/10.5281/zenodo.3813999" httpUri="https://zenodo.org/record/3813999/files/figure.png" pageId="0" pageNumber="195">Figs 14</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C35165136918FFA01BE6F9F0AA10F486" pageId="0" pageNumber="195" type="materials_examined">
<paragraph id="8BF436986918FFA01BE6F9F0AA10F486" blockId="0.[151,1437,1498,2028]" pageId="0" pageNumber="195">
<emphasis id="B93FEA8A6918FFA01BE6F9F0A873F4A2" bold="true" box="[151,316,1643,1667]" inLineHeading="true" pageId="0" pageNumber="195" reason="1">Type material.</emphasis>
<materialsCitation id="3B233CC56918FFA01A30F9F0AA10F486" ID-GBIF-Occurrence="2610673310" collectionCode="MNHN" collectorName="A. Boubakri" country="Tunisia" location="Ariana" pageId="0" pageNumber="195" specimenCount="3" specimenCount-male="3" stateProvince="L'Ariana" typeStatus="holotype">
<typeStatus id="54F0883A6918FFA01A30F9F0A8EBF4A2" box="[321,420,1643,1667]" pageId="0" pageNumber="195" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4DFD116918FFA01ADBF9F7A8F1F4A2" box="[426,446,1644,1667]" pageId="0" pageNumber="195" type="male"></specimenCount>
and
<specimenCount id="9D4DFD116918FFA01A81F9F0AB5EF4A2" box="[496,529,1643,1667]" pageId="0" pageNumber="195" type="male">2♂</specimenCount>
<typeStatus id="54F0883A6918FFA01966F9F7AB32F4A5" box="[535,637,1644,1668]" pageId="0" pageNumber="195" type="paratype">paratypes</typeStatus>
from
<collectingCountry id="F35C76086918FFA019CDF9F0AA66F4A2" box="[700,809,1643,1667]" name="Tunisia" pageId="0" pageNumber="195">TUNISIA</collectingCountry>
,
<collectingRegion id="498FF87A6918FFA01840F9F0AA33F4A2" box="[817,892,1643,1667]" country="Tunisia" name="L'Ariana" pageId="0" pageNumber="195">Ariana</collectingRegion>
, Kalaât Landaluz (370251N, 100620E), spring 2018,
<collectingMethod id="520A4E8F6918FFA01BABF914A86BF486" box="[218,292,1679,1703]" pageId="0" pageNumber="195">pitfalls</collectingMethod>
in salt marsh, leg.
<collectorName id="26BE534E6918FFA01A9CF914AB3CF486" box="[493,627,1679,1703]" pageId="0" pageNumber="195">A. Boubakri</collectorName>
, deposited in
<collectionCode id="ED5AAE5D6918FFA01877F90BAA14F486" box="[774,859,1680,1703]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="0" pageNumber="195">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35165136918FFA01BB6F928AAA3F4CE" pageId="0" pageNumber="195" type="diagnosis">
<paragraph id="8BF436986918FFA01BB6F928AAA3F4CE" blockId="0.[151,1437,1498,2028]" pageId="0" pageNumber="195">
<emphasis id="B93FEA8A6918FFA01BB6F928A873F4EA" bold="true" box="[199,316,1715,1739]" inLineHeading="true" pageId="0" pageNumber="195" reason="1">Diagnosis.</emphasis>
<taxonomicName id="4C4B4D1B6918FFA01A32F92FAB57F4EA" authority="Boubakri &amp; Bosmans &amp; Pétillon &amp; Achouri, 2020" authorityName="Boubakri &amp; Bosmans &amp; Pétillon &amp; Achouri" authorityYear="2020" box="[323,536,1715,1739]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="195" phylum="Arthropoda" rank="species" species="salsicola" status="sp. nov.">
<emphasis id="B93FEA8A6918FFA01A32F92FAB57F4EA" box="[323,536,1715,1739]" italics="true" pageId="0" pageNumber="195">Agraecina salsicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C57F16918FFA0196EF92FAB16F4EA" box="[543,601,1716,1739]" pageId="0" pageNumber="195" rank="species">sp. n.</taxonomicNameLabel>
is closely related to
<taxonomicName id="4C4B4D1B6918FFA01846F92FAAD7F4EA" authorityName="Bosmans" authorityYear="1999" box="[823,920,1715,1739]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="195" phylum="Arthropoda" rank="species" species="hodna">
<emphasis id="B93FEA8A6918FFA01846F92FAAD7F4EA" box="[823,920,1715,1739]" italics="true" pageId="0" pageNumber="195">A. hodna</emphasis>
</taxonomicName>
from
<collectingCountry id="F35C76086918FFA018A9F928AD64F4EA" box="[984,1067,1715,1739]" name="Algeria" pageId="0" pageNumber="195">Algeria</collectingCountry>
, but clearly differs by the strongly elongated tibial apophysis and the relatively short median apophysis (
<figureCitation id="13702A1D6918FFA0180FF94CAAAFF4CE" box="[894,992,1751,1775]" captionStart="FIGURES 14" captionStartId="1.[151,258,1031,1053]" captionTargetBox="[151,1435,472,1002]" captionTargetId="figure@1.[151,1435,472,1002]" captionTargetPageId="1" captionText="FIGURES 14. Agraecina salsicola sp. n. 1. Male, dorsal view; 2. Male palp, retrolateral view; 3. Idem, ventral view; 4. Idem, prolateral view. Abbreviations: E—embolus; MA—median apophysis; RTA—retrolateral tibial apohysis." figureDoi="http://doi.org/10.5281/zenodo.3813999" httpUri="https://zenodo.org/record/3813999/files/figure.png" pageId="0" pageNumber="195">Figs 23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35165136918FFA01BB6F960AD6CF532" box="[199,1059,1787,1811]" pageId="0" pageNumber="195" type="etymology">
<paragraph id="8BF436986918FFA01BB6F960AD6CF532" blockId="0.[151,1437,1498,2028]" box="[199,1059,1787,1811]" pageId="0" pageNumber="195">
<emphasis id="B93FEA8A6918FFA01BB6F960A807F532" bold="true" box="[199,328,1787,1811]" inLineHeading="true" pageId="0" pageNumber="195" reason="1">Etymology.</emphasis>
The species was collected in a salt marsh, hence the name
<taxonomicName id="4C4B4D1B6918FFA018CCF960AD53F532" authorityName="Boubakri &amp; Bosmans &amp; Pétillon &amp; Achouri" authorityYear="2020" box="[957,1052,1787,1811]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="195" phylum="Arthropoda" rank="species" species="salsicola">
<emphasis id="B93FEA8A6918FFA018CCF960AD53F532" box="[957,1052,1787,1811]" italics="true" pageId="0" pageNumber="195">salsicola</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35165136918FFA11BB6F884A8DBF347" lastPageId="1" lastPageNumber="196" pageId="0" pageNumber="195" type="description">
<paragraph id="8BF436986918FFA01BB6F884AA41F57A" blockId="0.[151,1437,1498,2028]" pageId="0" pageNumber="195">
<emphasis id="B93FEA8A6918FFA01BB6F884A81DF516" bold="true" box="[199,338,1823,1847]" inLineHeading="true" pageId="0" pageNumber="195" reason="1">Description.</emphasis>
Measurements (n=3;
<typeStatus id="54F0883A6918FFA01933F884ABD0F516" box="[578,671,1823,1847]" pageId="0" pageNumber="195" type="holotype">holotype</typeStatus>
in parentheses): Total length 4.25.0 (5.0); prosoma 1.602.15 (1.81) long, 1.161.60 (1.56) wide. Leg measurements in
<tableCitation id="C6C903236918FFA019C9F8D8AA48F57A" box="[696,775,1859,1883]" captionStart="TABLE 1" captionStartId="2.[151,239,1251,1275]" captionText="TABLE 1. Agraecina salsicola sp. n., leg measurements of male holotype." pageId="0" pageNumber="195">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BF436986918FFA01BB6F8FCAB3EF582" blockId="0.[151,1437,1498,2028]" pageId="0" pageNumber="195">
Colour: Carapace yellowish-brown, with anastomosing greyish stripes; legs yellowish-brown; abdomen pale grey, somewhat darkened antero-dorsally (
<figureCitation id="13702A1D6918FFA01953F810AB2EF582" box="[546,609,1931,1955]" captionStart="FIGURES 14" captionStartId="1.[151,258,1031,1053]" captionTargetBox="[151,1435,472,1002]" captionTargetId="figure@1.[151,1435,472,1002]" captionTargetPageId="1" captionText="FIGURES 14. Agraecina salsicola sp. n. 1. Male, dorsal view; 2. Male palp, retrolateral view; 3. Idem, ventral view; 4. Idem, prolateral view. Abbreviations: E—embolus; MA—median apophysis; RTA—retrolateral tibial apohysis." figureDoi="http://doi.org/10.5281/zenodo.3813999" httpUri="https://zenodo.org/record/3813999/files/figure.png" pageId="0" pageNumber="195">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BF436986918FFA01BB6F82BACD3F5CA" blockId="0.[151,1437,1498,2028]" pageId="0" pageNumber="195">Eyes: Anterior median eyes slightly smaller than other eyes, separated by half their diameter, from the anterior lateral eyes by their diameter; posterior eyes separated by their diameter. Chelicerae with 3 pro- and 2 retromarginal small</paragraph>
<paragraph id="8BF436986919FFA11BE6FF01A851F2F7" blockId="1.[151,1437,154,466]" pageId="1" pageNumber="196">teeth. Legs spinous, as in all other species of the genus, typically tibiae and metatarsi with 3 and 2 pairs of ventral spines, respectively.</paragraph>
<paragraph id="8BF436986919FFA11BB6FF79ADC6F363" blockId="1.[151,1437,154,466]" pageId="1" pageNumber="196">
Male palp (
<figureCitation id="13702A1D6919FFA11A39FF79A8E1F2DB" box="[328,430,226,250]" captionStart="FIGURES 14" captionStartId="1.[151,258,1031,1053]" captionTargetBox="[151,1435,472,1002]" captionTargetId="figure@1.[151,1435,472,1002]" captionTargetPageId="1" captionText="FIGURES 14. Agraecina salsicola sp. n. 1. Male, dorsal view; 2. Male palp, retrolateral view; 3. Idem, ventral view; 4. Idem, prolateral view. Abbreviations: E—embolus; MA—median apophysis; RTA—retrolateral tibial apohysis." figureDoi="http://doi.org/10.5281/zenodo.3813999" httpUri="https://zenodo.org/record/3813999/files/figure.png" pageId="1" pageNumber="196">Figs 24</figureCitation>
): Tibial apophysis elongated, nearly linear, 1.6 times longer than the tibias length without apophysis, in lateral view with 2 bends; median apophysis S-shaped, relatively short compared to other
<taxonomicName id="4C4B4D1B6919FFA11F88FE9DAC27F33C" authorityName="Simon" authorityYear="1932" box="[1273,1384,262,285]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEA8A6919FFA11F88FE9DAC27F33C" box="[1273,1384,262,285]" italics="true" pageId="1" pageNumber="196">Agraecina</emphasis>
</taxonomicName>
species; embolus threadlike, representing one-fifth of a circle, as in the other species of the genus.
</paragraph>
<paragraph id="8BF436986919FFA11BB6FED5A8DBF347" blockId="1.[151,1437,154,466]" box="[199,404,334,358]" pageId="1" pageNumber="196">Female: Unknown.</paragraph>
</subSubSection>
<subSubSection id="C35165136919FFA11BB6FEEAA842F3F3" pageId="1" pageNumber="196" type="distribution">
<paragraph id="8BF436986919FFA11BB6FEEAAA60F3A8" blockId="1.[151,1437,154,466]" box="[199,815,369,394]" pageId="1" pageNumber="196">
<emphasis id="B93FEA8A6919FFA11BB6FEEAA815F3A8" bold="true" box="[199,346,369,393]" inLineHeading="true" pageId="1" pageNumber="196" reason="1">Distribution.</emphasis>
Only known from the
<typeStatus id="54F0883A6919FFA1193CFEE9AB34F3AB" box="[589,635,370,394]" pageId="1" pageNumber="196">type</typeStatus>
locality (
<figureCitation id="13702A1D6919FFA11991FEE9AA6EF3A8" box="[736,801,369,394]" captionStart="FIGURE 5" captionStartId="1.[151,244,2013,2035]" captionText="FIGURE 5. Situation of the salt marsh of Kalaât Landalouz in Tunisia." pageId="1" pageNumber="196">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BF436986919FFA11BB6FE0EA842F3F3" blockId="1.[151,1437,154,466]" pageId="1" pageNumber="196">
<emphasis id="B93FEA8A6919FFA11BB6FE0EA869F38C" bold="true" box="[199,294,405,429]" inLineHeading="true" pageId="1" pageNumber="196" reason="1">Habitat.</emphasis>
A coastal salt marsh dominated by
<taxonomicName id="4C4B4D1B6919FFA119D1FE0DAA36F38C" box="[672,889,406,430]" class="Magnoliopsida" family="Amaranthaceae" genus="Salicornia" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="196" phylum="Tracheophyta" rank="species" species="europaea">
<emphasis id="B93FEA8A6919FFA119D1FE0DAA36F38C" box="[672,889,406,430]" italics="true" pageId="1" pageNumber="196">Salicornia europaea</emphasis>
</taxonomicName>
L.,
<taxonomicName id="4C4B4D1B6919FFA118D5FE0DAD1BF38F" box="[932,1108,406,430]" class="Magnoliopsida" family="Amaranthaceae" genus="Atriplex" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="196" phylum="Tracheophyta" rank="species" species="halimus">
<emphasis id="B93FEA8A6919FFA118D5FE0DAD1BF38F" box="[932,1108,406,430]" italics="true" pageId="1" pageNumber="196">Atriplex halimus</emphasis>
</taxonomicName>
L. and
<taxonomicName id="4C4B4D1B6919FFA11FD4FE0DAC30F38C" authorityName="Willd" authorityYear="1816" box="[1189,1407,406,429]" class="Magnoliopsida" family="Tamaricaceae" genus="Tamarix" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="196" phylum="Tracheophyta" rank="species" species="canariensis">
<emphasis id="B93FEA8A6919FFA11FD4FE0DAC30F38C" box="[1189,1407,406,429]" italics="true" pageId="1" pageNumber="196">Tamarix canariensis</emphasis>
</taxonomicName>
L. (
<figureCitation id="13702A1D6919FFA11BEEFE21A84EF3F3" box="[159,257,442,466]" captionStart-0="FIGURES 6" captionStart-1="FIGURE 7" captionStartId-0="2.[151,258,1181,1203]" captionStartId-1="3.[151,244,1436,1458]" captionTargetBox-0="[429,1159,182,1154]" captionTargetId-0="figure@2.[429,1159,182,1154]" captionTargetPageId-0="2" captionText-0="FIGURES 6. General view of the salt marsh of Kalaât Landalouz where Agroecina salsicola sp. n. was collected" captionText-1="FIGURE 7. Detail of halophytic vegetation (Atriplex and Salicornia spp.)." figureDoi-0="http://doi.org/10.5281/zenodo.3814001" httpUri-0="https://zenodo.org/record/3814001/files/figure.png" pageId="1" pageNumber="196">Figs 67</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>