292 lines
40 KiB
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292 lines
40 KiB
XML
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<mods:title id="A96DB96F751050B72953EE6C06039E75">Resolving taxonomic issues of cicadas (Hemiptera: Cicadidae) including new combinations, new synonymies, and revised status, with updates on the diversity of the Brazilian cicada fauna and new records for four South American countries</mods:title>
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<treatment id="03E2878BFFE45922FF5FFE1BFBA4FE4F" ID-DOI="http://doi.org/10.5281/zenodo.8239536" ID-GBIF-Taxon="212427655" ID-Zenodo-Dep="8239536" LSID="urn:lsid:plazi:treatment:03E2878BFFE45922FF5FFE1BFBA4FE4F" httpUri="http://treatment.plazi.org/id/03E2878BFFE45922FF5FFE1BFBA4FE4F" lastPageId="15" lastPageNumber="354" pageId="14" pageNumber="353">
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<subSubSection id="C3516516FFE45923FF5FFE1BFA7BFB90" box="[152,1423,1112,1138]" pageId="14" pageNumber="353" type="nomenclature">
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<paragraph id="8BF4369DFFE45923FF5FFE1BFA7BFB90" blockId="14.[152,1423,1112,1138]" box="[152,1423,1112,1138]" pageId="14" pageNumber="353">
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<heading id="D0BC81F1FFE45923FF5FFE1BFA7BFB90" bold="true" box="[152,1423,1112,1138]" centered="true" fontSize="11" level="1" pageId="14" pageNumber="353" reason="1">
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<emphasis id="B93FEA8FFFE45923FF5FFE1BFA7BFB90" bold="true" box="[152,1423,1112,1138]" pageId="14" pageNumber="353">
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NEW COMBINATIONS AND SYNONYMIES IN
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<taxonomicName id="4C4B4D1EFFE45923FCF7FE1BFC1BFB90" ID-CoL="8HC2Y" ID-ENA="2218939" authority="Distant, 1905" authorityName="Distant" authorityYear="1905" box="[816,1007,1112,1138]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="359" phylum="Arthropoda" rank="tribe" tribe="Zammarini">ZAMMARINI</taxonomicName>
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BASED ON
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<bibRefCitation id="EFDA4B6CFFE45923FB5CFE1BFA7BFB90" author="GOEMANS" box="[1179,1423,1112,1138]" firstAuthor="GOEMANS" pageId="14" pageNumber="353" refId="ref13149" refString="Goemans, G. (2016) The classification and phylogeny of the Neotropical cicada tribe Zammarini (Hemiptera, Cicadidae) and a revision of its type genus Zammara Amyot & Audinet-Serville, 1843 and its sister genus Zammaralna Boulard & Sueur, 1996. Ph. D. dissertation, University of Connecticut, Storrs, 206 pp." type="book" year="2016">GOEMANS (2016)</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3516516FFE45922FF50FEE3FBA4FE4F" lastPageId="15" lastPageNumber="354" pageId="14" pageNumber="353" type="discussion">
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<paragraph id="8BF4369DFFE45923FF50FEE3FDDCFA70" blockId="14.[151,1437,1184,2039]" pageId="14" pageNumber="353">
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<bibRefCitation id="EFDA4B6CFFE45923FF50FEE3FEA2FB58" author="Goemans" box="[151,342,1184,1210]" firstAuthor="Goemans" pageId="14" pageNumber="353" refId="ref13149" refString="Goemans, G. (2016) The classification and phylogeny of the Neotropical cicada tribe Zammarini (Hemiptera, Cicadidae) and a revision of its type genus Zammara Amyot & Audinet-Serville, 1843 and its sister genus Zammaralna Boulard & Sueur, 1996. Ph. D. dissertation, University of Connecticut, Storrs, 206 pp." type="book" year="2016">Goemans (2016)</bibRefCitation>
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studied the
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<taxonomicName id="4C4B4D1EFFE45923FE21FEE3FD96FB58" authorityName="Distant" authorityYear="1905" box="[486,610,1184,1210]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="359" phylum="Arthropoda" rank="tribe" tribe="Zammarini">Zammarini</taxonomicName>
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providing evidence for numerous taxonomic changes. The taxonomical acts within the dissertation were not considered to be valid for nomenclatural purposes following Article 8 of the
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<emphasis id="B93FEA8FFFE45923FF50FEABFD46FAE0" box="[151,690,1256,1282]" italics="true" pageId="14" pageNumber="353">International Code of Zoological Nomenclature</emphasis>
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(ICZN 1999) and were disclaimed for such purposes in that work (
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<bibRefCitation id="EFDA4B6CFFE45923FF58FF4FFEBAFAC4" author="Goemans" box="[159,334,1292,1318]" firstAuthor="Goemans" pageId="14" pageNumber="353" refId="ref13149" refString="Goemans, G. (2016) The classification and phylogeny of the Neotropical cicada tribe Zammarini (Hemiptera, Cicadidae) and a revision of its type genus Zammara Amyot & Audinet-Serville, 1843 and its sister genus Zammaralna Boulard & Sueur, 1996. Ph. D. dissertation, University of Connecticut, Storrs, 206 pp." type="book" year="2016">Goemans 2016</bibRefCitation>
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). The specifics and long history of confusion for the application of
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<taxonomicName id="4C4B4D1EFFE45923FB86FF4FFB49FAC4" authorityName="Distant" authorityYear="1905" box="[1089,1213,1292,1318]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="359" phylum="Arthropoda" rank="tribe" tribe="Zammarini">Zammarini</taxonomicName>
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taxa is described in
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<bibRefCitation id="EFDA4B6CFFE45923FF50FF73FEA1FAA8" author="Goemans" box="[151,341,1328,1354]" firstAuthor="Goemans" pageId="14" pageNumber="353" refId="ref13149" refString="Goemans, G. (2016) The classification and phylogeny of the Neotropical cicada tribe Zammarini (Hemiptera, Cicadidae) and a revision of its type genus Zammara Amyot & Audinet-Serville, 1843 and its sister genus Zammaralna Boulard & Sueur, 1996. Ph. D. dissertation, University of Connecticut, Storrs, 206 pp." type="book" year="2016">Goemans (2016)</bibRefCitation>
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. He was able to study almost all
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<typeStatus id="54F0883FFFE45923FD05FF72FD06FAA9" box="[706,754,1329,1355]" pageId="14" pageNumber="353">type</typeStatus>
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specimens and clarify the taxonomy of the group as a whole. The original work (
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<bibRefCitation id="EFDA4B6CFFE45923FEB6FF17FDEAFA8C" author="Goemans" box="[369,542,1364,1390]" firstAuthor="Goemans" pageId="14" pageNumber="353" refId="ref13149" refString="Goemans, G. (2016) The classification and phylogeny of the Neotropical cicada tribe Zammarini (Hemiptera, Cicadidae) and a revision of its type genus Zammara Amyot & Audinet-Serville, 1843 and its sister genus Zammaralna Boulard & Sueur, 1996. Ph. D. dissertation, University of Connecticut, Storrs, 206 pp." type="book" year="2016">Goemans 2016</bibRefCitation>
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) should be consulted for summary information on the morphology, history, and distribution of each species studied.
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</paragraph>
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<paragraph id="8BF4369DFFE45923FF00FFDFFEDCF950" blockId="14.[151,1437,1184,2039]" pageId="14" pageNumber="353">
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Unfortunately, he seems to have moved on to a different area of study and has yet to publish the results formally. Some of his findings have already been applied to update the taxonomy of numerous species (e.g.,
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<bibRefCitation id="EFDA4B6CFFE45923FB2BFF83FA6CFA38" author="Sanborn" box="[1260,1432,1472,1498]" firstAuthor="Sanborn" pageId="14" pageNumber="353" pagination="1 - 104" refId="ref15227" refString="Sanborn, A. F. (2019 a) The cicadas (Hemiptera: Cicadidae) of Bolivia including the descriptions of fifteen new species, the resurrection of one genus and two species, seven new combinations, six new synonymies, and twenty-eight new records. Zootaxa, 4655 (1), 1 - 104. https: // doi. org / 10.11646 / zootaxa. 4655.1.1" type="journal article" year="2019" yearSuffix="a">Sanborn 2019a</bibRefCitation>
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;
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<bibRefCitation id="EFDA4B6CFFE45923FF50FFA7FF2AFA1C" author="Sanborn" box="[151,222,1508,1534]" firstAuthor="Sanborn" pageId="14" pageNumber="353" pagination="117 - 126" refId="ref15522" refString="Sanborn, A. F. (2020 c) Dyticodopoea and Pygmaeodopoea, new genera for the Central American cicada species previously assigned to Odopoea Stal, 1861 (Hemiptera: Cicadoidea: Cicadidae: Cicadinae: Zammarini). Proceedings of the Entomological Society of Washington, 122, 117 - 126. https: // doi. org / 10.4289 / 0013 - 8797.122.1.117" type="journal article" year="2020" yearSuffix="c">2020c</bibRefCitation>
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) and other changes are being reported in the literature even though the changes are unofficial under the
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<emphasis id="B93FEA8FFFE45923FAA6FFA7FA68FA1C" box="[1377,1436,1508,1534]" italics="true" pageId="14" pageNumber="353">Code</emphasis>
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(e.g.,
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<bibRefCitation id="EFDA4B6CFFE45923FF1FFC4AFE50F9C0" author="Nunes" box="[216,420,1544,1571]" etAl="et al." firstAuthor="Nunes" pageId="14" pageNumber="353" refId="ref14077" refString="Nunes, V. L., Ruschel, T. P., Maccagnan, D. H. B., Simies, P. C. & Acosta, R. C. (2023) Cigarras do Brasil, Guia Fotographica. 1 a edicao. ebook. 46 pp. [https: // zenodo. org / record / 7712483]" type="book" year="2023">
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Nunes
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<emphasis id="B93FEA8FFFE45923FEE0FC4AFEADF9C0" box="[295,345,1544,1570]" italics="true" pageId="14" pageNumber="353">et al</emphasis>
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. 2023
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</bibRefCitation>
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). Most of his remaining nomenclatural data is presented here to reassign species to clarify the taxonomy of the
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<taxonomicName id="4C4B4D1EFFE45923FE41FC6FFDF6F9A4" authorityName="Distant" authorityYear="1905" box="[390,514,1580,1606]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="359" phylum="Arthropoda" rank="tribe" tribe="Zammarini">Zammarini</taxonomicName>
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as his work is important in correcting historic misconceptions and the unofficial results are being incorporated into the more recent literature. The new combinations are provided here to make the changes suggested by
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<bibRefCitation id="EFDA4B6CFFE45923FE49FC37FDBCF96C" author="Goemans" box="[398,584,1652,1678]" firstAuthor="Goemans" pageId="14" pageNumber="353" refId="ref13149" refString="Goemans, G. (2016) The classification and phylogeny of the Neotropical cicada tribe Zammarini (Hemiptera, Cicadidae) and a revision of its type genus Zammara Amyot & Audinet-Serville, 1843 and its sister genus Zammaralna Boulard & Sueur, 1996. Ph. D. dissertation, University of Connecticut, Storrs, 206 pp." type="book" year="2016">Goemans (2016)</bibRefCitation>
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official under the
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<emphasis id="B93FEA8FFFE45923FCD4FC37FCBAF96C" box="[787,846,1652,1678]" italics="true" pageId="14" pageNumber="353">Code</emphasis>
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so that correct taxonomy can be applied to these taxa in the future.
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</paragraph>
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<paragraph id="8BF4369DFFE45923FF00FCFFFE03F815" blockId="14.[151,1437,1184,2039]" pageId="14" pageNumber="353">
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With respect to Brazilian species,
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<taxonomicName id="4C4B4D1EFFE45923FD85FCFEFC44F934" authority="Erichson, 1848" authorityName="Erichson" authorityYear="1848" box="[578,944,1724,1750]" class="Insecta" family="Cicadidae" genus="Zammara" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="eximia">
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<emphasis id="B93FEA8FFFE45923FD85FCFEFCF5F934" box="[578,769,1725,1750]" italics="true" pageId="14" pageNumber="353">Zammara eximia</emphasis>
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<bibRefCitation id="EFDA4B6CFFE45923FCC0FCFFFC44F934" author="Erichson" box="[775,944,1724,1750]" firstAuthor="Erichson" pageId="14" pageNumber="353" pagination="533 - 1260" refId="ref12784" refString="Erichson, W. F. (1848) Homoptera. In: Schomburgk, R. (Ed.), Reisen in Britisch-Guiana in den Jahren 1840 - 1844 im auftrag Sr. Majestat des Konigs von Preussen ausgef ¸ hrt von Richard Schomburgk. Nebst einer Fauna und Flora Guiana's nach Vorlagen von Johannes M ¸ ller, Ehrenberg, Erichson, Klotzsch, Troschel, Cabanis und andern. Band 3. Weber, Leipzig, pp. 533 - 1260. https: // doi. org / 10.5962 / bhl. title. 109982" type="book chapter" year="1848">Erichson, 1848</bibRefCitation>
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</taxonomicName>
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was determined to possess three tarsomeres and is reassigned to
|
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<taxonomicName id="4C4B4D1EFFE45923FEBCFCA3FCD6F918" authority="(Erichson, 1848)" authorityName="Sanborn" authorityYear="2023" baseAuthorityName="Erichson" baseAuthorityYear="1848" box="[379,802,1760,1786]" class="Insecta" family="Cicadidae" genus="Zammaralna" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="eximia" status="comb. nov.">
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<emphasis id="B93FEA8FFFE45923FEBCFCA3FDAAF918" box="[379,606,1760,1786]" italics="true" pageId="14" pageNumber="353">Zammaralna eximia</emphasis>
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(
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<bibRefCitation id="EFDA4B6CFFE45923FDAAFCA3FCEDF918" author="Erichson" box="[621,793,1760,1786]" firstAuthor="Erichson" pageId="14" pageNumber="353" pagination="533 - 1260" refId="ref12784" refString="Erichson, W. F. (1848) Homoptera. In: Schomburgk, R. (Ed.), Reisen in Britisch-Guiana in den Jahren 1840 - 1844 im auftrag Sr. Majestat des Konigs von Preussen ausgef ¸ hrt von Richard Schomburgk. Nebst einer Fauna und Flora Guiana's nach Vorlagen von Johannes M ¸ ller, Ehrenberg, Erichson, Klotzsch, Troschel, Cabanis und andern. Band 3. Weber, Leipzig, pp. 533 - 1260. https: // doi. org / 10.5962 / bhl. title. 109982" type="book chapter" year="1848">Erichson, 1848</bibRefCitation>
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)
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</taxonomicName>
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<emphasis id="B93FEA8FFFE45923FCEEFCA3FC7BF918" bold="true" box="[809,911,1760,1786]" pageId="14" pageNumber="353">
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<taxonomicNameLabel id="A20C57F4FFE45923FCEEFCA3FC7BF918" box="[809,911,1760,1786]" pageId="14" pageNumber="353" rank="species">n. comb.</taxonomicNameLabel>
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</emphasis>
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, as presented in the species list of
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<bibRefCitation id="EFDA4B6CFFE45923FAD2FCA2FF11F8FC" author="Nunes" etAl="et al." firstAuthor="Nunes" pageId="14" pageNumber="353" refId="ref14077" refString="Nunes, V. L., Ruschel, T. P., Maccagnan, D. H. B., Simies, P. C. & Acosta, R. C. (2023) Cigarras do Brasil, Guia Fotographica. 1 a edicao. ebook. 46 pp. [https: // zenodo. org / record / 7712483]" type="book" year="2023">
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Nunes
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<emphasis id="B93FEA8FFFE45923FAA3FCA2FA61F918" box="[1380,1429,1760,1786]" italics="true" pageId="14" pageNumber="353">et al</emphasis>
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. (2023)
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||
</bibRefCitation>
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||
. Similarly,
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<taxonomicName id="4C4B4D1EFFE45923FEA5FD46FC57F8FC" authority="Amyot & Audinet-Serville, 1843" authorityName="Amyot & Audinet-Serville" authorityYear="1843" box="[354,931,1796,1822]" class="Insecta" family="Cicadidae" genus="Zammara" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="strepens">
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<emphasis id="B93FEA8FFFE45923FEA5FD46FDC5F8FC" box="[354,561,1797,1822]" italics="true" pageId="14" pageNumber="353">Zammara strepens</emphasis>
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<bibRefCitation id="EFDA4B6CFFE45923FDF1FD47FC57F8FC" author="Amyot & Audinet-Serville" box="[566,931,1796,1822]" firstAuthor="Amyot" pageId="14" pageNumber="353" refId="ref11480" refString="Amyot, C. J. B. & Audinet-Serville, A. (1843) Histoire Naturelle des Insectes. Hemipteres. Deuxime partie. Homopteres. Homoptera Latr. Librairie encyclopedique de Roret, Paris, 675 pp. https: // doi. org / 10.5962 / bhl. title. 8471" type="book" year="1843">Amyot & Audinet-Serville, 1843</bibRefCitation>
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</taxonomicName>
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was determined to have three tarsomeses and a narrow head so it is reassigned to
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||
<taxonomicName id="4C4B4D1EFFE45923FDD9FD6BFB9DF8A0" authority="(Amyot & Audinet-Serville, 1843)" authorityName="Sanborn" authorityYear="2023" baseAuthorityName="Amyot & Audinet-Serville" baseAuthorityYear="1843" box="[542,1129,1832,1858]" class="Insecta" family="Cicadidae" genus="Orellana" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="strepens" status="comb. nov.">
|
||
<emphasis id="B93FEA8FFFE45923FDD9FD6BFD17F8A0" box="[542,739,1832,1858]" italics="true" pageId="14" pageNumber="353">Orellana strepens</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDA4B6CFFE45923FD36FD6BFB95F8A0" author="Amyot & Audinet-Serville" box="[753,1121,1832,1858]" firstAuthor="Amyot" pageId="14" pageNumber="353" refId="ref11480" refString="Amyot, C. J. B. & Audinet-Serville, A. (1843) Histoire Naturelle des Insectes. Hemipteres. Deuxime partie. Homopteres. Homoptera Latr. Librairie encyclopedique de Roret, Paris, 675 pp. https: // doi. org / 10.5962 / bhl. title. 8471" type="book" year="1843">Amyot & Audinet-Serville, 1843</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA8FFFE45923FBA8FD6BFB20F8A0" bold="true" box="[1135,1236,1832,1858]" pageId="14" pageNumber="353">
|
||
<taxonomicNameLabel id="A20C57F4FFE45923FBA8FD6BFB20F8A0" box="[1135,1236,1832,1858]" pageId="14" pageNumber="353" rank="species">n. comb.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, also as presented in the species list of
|
||
<bibRefCitation id="EFDA4B6CFFE45923FEBCFD0EFDA1F884" author="Nunes" box="[379,597,1868,1895]" etAl="et al." firstAuthor="Nunes" pageId="14" pageNumber="353" refId="ref14077" refString="Nunes, V. L., Ruschel, T. P., Maccagnan, D. H. B., Simies, P. C. & Acosta, R. C. (2023) Cigarras do Brasil, Guia Fotographica. 1 a edicao. ebook. 46 pp. [https: // zenodo. org / record / 7712483]" type="book" year="2023">
|
||
Nunes
|
||
<emphasis id="B93FEA8FFFE45923FE0EFD0EFE0EF884" box="[457,506,1868,1894]" italics="true" pageId="14" pageNumber="353">et al</emphasis>
|
||
. (2023)
|
||
</bibRefCitation>
|
||
. In addition, it was determined that the images and distribution of
|
||
<taxonomicName id="4C4B4D1EFFE45923FAFFFD0FFE57F868" authority="Sanborn, 2011 a" authorityName="Sanborn" authorityYear="2011" class="Insecta" family="Cicadidae" genus="Orellana" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="pollyae">
|
||
<emphasis id="B93FEA8FFFE45923FAFFFD0FFF1DF868" italics="true" pageId="14" pageNumber="353">Orellana pollyae</emphasis>
|
||
<bibRefCitation id="EFDA4B6CFFE45923FF34FD33FE57F868" author="Sanborn" box="[243,419,1904,1930]" firstAuthor="Sanborn" pageId="14" pageNumber="353" pagination="377 - 384" refId="ref14616" refString="Sanborn, A. F. (2011 a) A new species of Orellana Distant from Brazil (Hemiptera: Cicadoidea: Cicadidae). Proceedings of the Entomological Society of Washington, 113, 377 - 384. https: // doi. org / 10.4289 / 0013 - 8797.113.3.377" type="journal article" year="2011" yearSuffix="a">Sanborn, 2011a</bibRefCitation>
|
||
</taxonomicName>
|
||
were the same as
|
||
<taxonomicName id="4C4B4D1EFFE45923FDBCFD33FD0FF868" authorityName="Sanborn" authorityYear="2023" baseAuthorityName="Amyot & Audinet-Serville" baseAuthorityYear="1843" box="[635,763,1904,1930]" class="Insecta" family="Cicadidae" genus="Orellana" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="strepens" status="comb. nov.">
|
||
<emphasis id="B93FEA8FFFE45923FDBCFD33FD0FF868" box="[635,763,1904,1930]" italics="true" pageId="14" pageNumber="353">O. strepens</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA8FFFE45923FCC3FD33FC98F868" bold="true" box="[772,876,1904,1930]" pageId="14" pageNumber="353">
|
||
<taxonomicNameLabel id="A20C57F4FFE45923FCC3FD33FC98F868" box="[772,876,1904,1930]" pageId="14" pageNumber="353" rank="species">n. comb.</taxonomicNameLabel>
|
||
</emphasis>
|
||
so that
|
||
<taxonomicName id="4C4B4D1EFFE45923FC0DFD33FAAAF868" authority="Sanborn, 2011a" authorityName="Sanborn" authorityYear="2011" box="[970,1374,1904,1930]" class="Insecta" family="Cicadidae" genus="Orellana" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="pollyae">
|
||
<emphasis id="B93FEA8FFFE45923FC0DFD33FB7EF868" box="[970,1162,1904,1930]" italics="true" pageId="14" pageNumber="353">Orellana pollyae</emphasis>
|
||
<bibRefCitation id="EFDA4B6CFFE45923FB54FD33FAAAF868" author="Sanborn" box="[1171,1374,1904,1930]" firstAuthor="Sanborn" pageId="14" pageNumber="353" pagination="377 - 384" refId="ref14616" refString="Sanborn, A. F. (2011 a) A new species of Orellana Distant from Brazil (Hemiptera: Cicadoidea: Cicadidae). Proceedings of the Entomological Society of Washington, 113, 377 - 384. https: // doi. org / 10.4289 / 0013 - 8797.113.3.377" type="journal article" year="2011" yearSuffix="a">
|
||
Sanborn, 2011a
|
||
<taxonomicNameLabel id="A20C57F4FFE45923FA8AFD33FAAAF868" box="[1357,1374,1904,1930]" pageId="14" pageNumber="353">
|
||
<emphasis id="B93FEA8FFFE45923FA8AFD33FAAAF868" bold="true" box="[1357,1374,1904,1930]" pageId="14" pageNumber="353">n</emphasis>
|
||
</taxonomicNameLabel>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA8FFFE45923FA99FD33FA68F868" bold="true" box="[1374,1436,1904,1930]" pageId="14" pageNumber="353">. syn.</emphasis>
|
||
becomes a junior synonym of
|
||
<taxonomicName id="4C4B4D1EFFE45923FE37FDD7FBBCF84C" authority="(Amyot & Audinet-Serville, 1843)" authorityName="Sanborn" authorityYear="2023" baseAuthorityName="Amyot & Audinet-Serville" baseAuthorityYear="1843" box="[496,1096,1940,1966]" class="Insecta" family="Cicadidae" genus="Orellana" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="strepens" status="comb. nov.">
|
||
<emphasis id="B93FEA8FFFE45923FE37FDD7FD4CF84C" box="[496,696,1940,1966]" italics="true" pageId="14" pageNumber="353">Orellana strepens</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDA4B6CFFE45923FD0FFDD7FBB4F84C" author="Amyot & Audinet-Serville" box="[712,1088,1940,1966]" firstAuthor="Amyot" pageId="14" pageNumber="353" refId="ref11480" refString="Amyot, C. J. B. & Audinet-Serville, A. (1843) Histoire Naturelle des Insectes. Hemipteres. Deuxime partie. Homopteres. Homoptera Latr. Librairie encyclopedique de Roret, Paris, 675 pp. https: // doi. org / 10.5962 / bhl. title. 8471" type="book" year="1843">Amyot & Audinet-Serville, 1843</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA8FFFE45923FB96FDD7FB4CF84C" bold="true" box="[1105,1208,1940,1966]" pageId="14" pageNumber="353">
|
||
<taxonomicNameLabel id="A20C57F4FFE45923FB96FDD7FB4CF84C" box="[1105,1208,1940,1966]" pageId="14" pageNumber="353" rank="species">n. comb.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<bibRefCitation id="EFDA4B6CFFE45923FB07FDD6FA68F84C" author="Nunes" box="[1216,1436,1940,1967]" etAl="et al." firstAuthor="Nunes" pageId="14" pageNumber="353" refId="ref14077" refString="Nunes, V. L., Ruschel, T. P., Maccagnan, D. H. B., Simies, P. C. & Acosta, R. C. (2023) Cigarras do Brasil, Guia Fotographica. 1 a edicao. ebook. 46 pp. [https: // zenodo. org / record / 7712483]" type="book" year="2023">
|
||
Nunes
|
||
<emphasis id="B93FEA8FFFE45923FAD7FDD6FAB6F84C" box="[1296,1346,1940,1966]" italics="true" pageId="14" pageNumber="353">et al</emphasis>
|
||
. (2023)
|
||
</bibRefCitation>
|
||
included
|
||
<taxonomicName id="4C4B4D1EFFE45923FF39FDFBFE86F830" authorityName="Sanborn" authorityYear="2011" box="[254,370,1976,2003]" class="Insecta" family="Cicadidae" genus="Orellana" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="pollyae">
|
||
<emphasis id="B93FEA8FFFE45923FF39FDFBFEE6F830" box="[254,274,1976,2002]" italics="true" pageId="14" pageNumber="353">O</emphasis>
|
||
.
|
||
<emphasis id="B93FEA8FFFE45923FEE7FDFBFE86F830" box="[288,370,1976,2002]" italics="true" pageId="14" pageNumber="353">pollyae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA8FFFE45923FEBEFDFBFE32F830" bold="true" box="[377,454,1976,2002]" pageId="14" pageNumber="353">
|
||
<taxonomicNameLabel id="A20C57F4FFE45923FEBEFDFBFE32F830" box="[377,454,1976,2002]" pageId="14" pageNumber="353">n. syn.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in their list of species as a junior synonym of
|
||
<taxonomicName id="4C4B4D1EFFE45923FC00FDFBFBB1F830" authorityName="Sanborn" authorityYear="2023" baseAuthorityName="Amyot & Audinet-Serville" baseAuthorityYear="1843" box="[967,1093,1976,2002]" class="Insecta" family="Cicadidae" genus="Orellana" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="353" phylum="Arthropoda" rank="species" species="strepens" status="comb. nov.">
|
||
<emphasis id="B93FEA8FFFE45923FC00FDFBFBB1F830" box="[967,1093,1976,2002]" italics="true" pageId="14" pageNumber="353">O. strepens</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA8FFFE45923FB8CFDFBFB45F830" bold="true" box="[1099,1201,1976,2002]" pageId="14" pageNumber="353">
|
||
<taxonomicNameLabel id="A20C57F4FFE45923FB8CFDFBFB45F830" box="[1099,1201,1976,2002]" pageId="14" pageNumber="353" rank="species">n. comb.</taxonomicNameLabel>
|
||
</emphasis>
|
||
but did not make the change official under the
|
||
<emphasis id="B93FEA8FFFE45923FE72FD9FFE04F814" box="[437,496,2012,2038]" italics="true" pageId="14" pageNumber="353">Code</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF4369DFFE55922FF00FAD4FBA4FE4F" blockId="15.[151,1437,151,429]" pageId="15" pageNumber="354">
|
||
There are additional taxonomic changes reported in
|
||
<bibRefCitation id="EFDA4B6CFFE55922FCDCFAD4FC2CFF53" author="Goemans" box="[795,984,151,177]" firstAuthor="Goemans" pageId="15" pageNumber="354" refId="ref13149" refString="Goemans, G. (2016) The classification and phylogeny of the Neotropical cicada tribe Zammarini (Hemiptera, Cicadidae) and a revision of its type genus Zammara Amyot & Audinet-Serville, 1843 and its sister genus Zammaralna Boulard & Sueur, 1996. Ph. D. dissertation, University of Connecticut, Storrs, 206 pp." type="book" year="2016">Goemans (2016)</bibRefCitation>
|
||
in species of
|
||
<taxonomicName id="4C4B4D1EFFE55922FBBEFAD4FB01FF53" authorityName="Distant" authorityYear="1905" box="[1145,1269,151,177]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="359" phylum="Arthropoda" rank="tribe" tribe="Zammarini">Zammarini</taxonomicName>
|
||
not inhabiting
|
||
<collectingCountry id="F35C760DFFE55922FF50FAF8FF2BFF37" box="[151,223,187,213]" name="Brazil" pageId="15" pageNumber="354">Brazil</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C4B4D1EFFE55922FF2BFAF8FD6AFF37" authority="Goding, 1925" authorityName="Goding" authorityYear="1925" box="[236,670,186,213]" class="Insecta" family="Cicadidae" genus="Orellana" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="354" phylum="Arthropoda" rank="species" species="brunneipennis">
|
||
<emphasis id="B93FEA8FFFE55922FF2BFAF8FE0CFF37" box="[236,504,187,213]" italics="true" pageId="15" pageNumber="354">Orellana brunneipennis</emphasis>
|
||
<bibRefCitation id="EFDA4B6CFFE55922FDC5FAF8FD6AFF37" author="Goding" box="[514,670,186,213]" firstAuthor="Goding" pageId="15" pageNumber="354" pagination="1 - 34" refId="ref13121" refString="Goding, F. W. (1925) Synopsis of the Cicadidae of Ecuador. Revista del Colegio Nacional Rocafuerte, 7, 1 - 34." type="journal article" year="1925">Goding, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4B4D1EFFE55922FD1FFAF8FBD7FF37" authority="Goding, 1925" authorityName="Goding" authorityYear="1925" box="[728,1059,186,213]" class="Insecta" family="Cicadidae" genus="Orellana" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="354" phylum="Arthropoda" rank="species" species="pulla">
|
||
<emphasis id="B93FEA8FFFE55922FD1FFAF8FC8BFF37" box="[728,895,187,213]" italics="true" pageId="15" pageNumber="354">Orellana pulla</emphasis>
|
||
<bibRefCitation id="EFDA4B6CFFE55922FC40FAF8FBD7FF37" author="Goding" box="[903,1059,186,213]" firstAuthor="Goding" pageId="15" pageNumber="354" pagination="1 - 34" refId="ref13121" refString="Goding, F. W. (1925) Synopsis of the Cicadidae of Ecuador. Revista del Colegio Nacional Rocafuerte, 7, 1 - 34." type="journal article" year="1925">Goding, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
were shown to possess two tarsi instead of three so they are reassigned to become
|
||
<taxonomicName id="4C4B4D1EFFE55922FD03FA9CFB78FF1B" authority="(Goding, 1925)" authorityName="Sanborn" authorityYear="2023" baseAuthorityName="Goding" baseAuthorityYear="1925" box="[708,1164,222,249]" class="Insecta" family="Cicadidae" genus="Zammara" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="354" phylum="Arthropoda" rank="species" species="brunneipennis" status="comb. nov.">
|
||
<emphasis id="B93FEA8FFFE55922FD03FA9CFC2CFF1B" box="[708,984,223,249]" italics="true" pageId="15" pageNumber="354">Zammara brunneipennis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDA4B6CFFE55922FC20FA9CFB77FF1B" author="Goding" box="[999,1155,222,249]" firstAuthor="Goding" pageId="15" pageNumber="354" pagination="1 - 34" refId="ref13121" refString="Goding, F. W. (1925) Synopsis of the Cicadidae of Ecuador. Revista del Colegio Nacional Rocafuerte, 7, 1 - 34." type="journal article" year="1925">Goding, 1925</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA8FFFE55922FB54FA9CFB0DFF1B" bold="true" box="[1171,1273,223,249]" pageId="15" pageNumber="354">
|
||
<taxonomicNameLabel id="A20C57F4FFE55922FB54FA9CFB0DFF1B" box="[1171,1273,223,249]" pageId="15" pageNumber="354" rank="species">n. comb.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4B4D1EFFE55922FAE8FA9CFE70FEFF" authority="(Goding, 1925)" authorityName="Sanborn" authorityYear="2023" baseAuthorityName="Goding" baseAuthorityYear="1925" class="Insecta" family="Cicadidae" genus="Zammara" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="354" phylum="Arthropoda" rank="species" species="pulla" status="comb. nov.">
|
||
<emphasis id="B93FEA8FFFE55922FAE8FA9CFF25FEFF" italics="true" pageId="15" pageNumber="354">Zammara pulla</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDA4B6CFFE55922FF18FB40FE88FEFF" author="Goding" box="[223,380,258,285]" firstAuthor="Goding" pageId="15" pageNumber="354" pagination="1 - 34" refId="ref13121" refString="Goding, F. W. (1925) Synopsis of the Cicadidae of Ecuador. Revista del Colegio Nacional Rocafuerte, 7, 1 - 34." type="journal article" year="1925">Goding, 1925</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA8FFFE55922FE4CFB40FE05FEFF" bold="true" box="[395,497,259,285]" pageId="15" pageNumber="354">
|
||
<taxonomicNameLabel id="A20C57F4FFE55922FE4CFB40FE05FEFF" box="[395,497,259,285]" pageId="15" pageNumber="354" rank="species">n. comb.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, respectively.
|
||
<taxonomicName id="4C4B4D1EFFE55922FD55FB40FBAEFEFF" authority="(Goding, 1925)" authorityName="Sanborn" authorityYear="2023" baseAuthorityName="Goding" baseAuthorityYear="1925" box="[658,1114,258,285]" class="Insecta" family="Cicadidae" genus="Zammara" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="354" phylum="Arthropoda" rank="species" species="brunneipennis" status="n. comb., n. syn.">
|
||
<emphasis id="B93FEA8FFFE55922FD55FB40FC52FEFF" box="[658,934,259,285]" italics="true" pageId="15" pageNumber="354">Zammara brunneipennis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDA4B6CFFE55922FC72FB40FBA5FEFF" author="Goding" box="[949,1105,258,285]" firstAuthor="Goding" pageId="15" pageNumber="354" pagination="1 - 34" refId="ref13121" refString="Goding, F. W. (1925) Synopsis of the Cicadidae of Ecuador. Revista del Colegio Nacional Rocafuerte, 7, 1 - 34." type="journal article" year="1925">Goding, 1925</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F4FFE55922FBA6FB40FAD5FEFF" box="[1121,1313,259,285]" pageId="15" pageNumber="354">
|
||
<emphasis id="B93FEA8FFFE55922FBA6FB40FB32FEFF" bold="true" box="[1121,1222,259,285]" pageId="15" pageNumber="354">n. comb.</emphasis>
|
||
,
|
||
<emphasis id="B93FEA8FFFE55922FB13FB40FAD5FEFF" bold="true" box="[1236,1313,259,285]" pageId="15" pageNumber="354">n. syn.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C4B4D1EFFE55922FAE8FB40FE76FEA3" authority="(Goding, 1925)" authorityName="Sanborn" authorityYear="2023" baseAuthorityName="Goding" baseAuthorityYear="1925" class="Insecta" family="Cicadidae" genus="Zammara" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="354" phylum="Arthropoda" rank="species" species="pulla" status="n. comb., n. syn.">
|
||
<emphasis id="B93FEA8FFFE55922FAE8FB40FF25FEA3" italics="true" pageId="15" pageNumber="354">Zammara pulla</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDA4B6CFFE55922FF18FB64FE8EFEA3" author="Goding" box="[223,378,294,321]" firstAuthor="Goding" pageId="15" pageNumber="354" pagination="1 - 34" refId="ref13121" refString="Goding, F. W. (1925) Synopsis of the Cicadidae of Ecuador. Revista del Colegio Nacional Rocafuerte, 7, 1 - 34." type="journal article" year="1925">Goding, 1925</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F4FFE55922FE4EFB64FDB2FEA3" box="[393,582,295,321]" pageId="15" pageNumber="354">
|
||
<emphasis id="B93FEA8FFFE55922FE4EFB64FE19FEA3" bold="true" box="[393,493,295,321]" pageId="15" pageNumber="354">n. comb.</emphasis>
|
||
,
|
||
<emphasis id="B93FEA8FFFE55922FE3CFB64FDB2FEA3" bold="true" box="[507,582,295,321]" pageId="15" pageNumber="354">n. syn.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C4B4D1EFFE55922FD45FB64FC25FEA3" authority="Schmidt, 1919" authorityName="Schmidt" authorityYear="1919" box="[642,977,295,321]" class="Insecta" family="Cicadidae" genus="Zammara" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="354" phylum="Arthropoda" rank="species" species="erna">
|
||
<emphasis id="B93FEA8FFFE55922FD45FB64FCDCFEA2" box="[642,808,295,320]" italics="true" pageId="15" pageNumber="354">Zammara erna</emphasis>
|
||
<bibRefCitation id="EFDA4B6CFFE55922FCE8FB64FC25FEA3" author="Schmidt" box="[815,977,295,321]" firstAuthor="Schmidt" pageId="15" pageNumber="354" pagination="383 - 394" refId="ref16387" refString="Schmidt, E. (1919) Beitrag zur kenntnie der genera Zammara Amyot et Serville und Orellana Distant. (Rhynchota-Homoptera.). Stettin Entomologische Zeitung. Herausgegeben von dem entomologischen Vereine zu Stettin, 80, 383 - 394. https: // doi. org / 10.5962 / bhl. title. 8515" type="journal article" year="1919">Schmidt, 1919</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA8FFFE55922FC10FB64FBD7FEA3" bold="true" box="[983,1059,295,321]" pageId="15" pageNumber="354">
|
||
<taxonomicNameLabel id="A20C57F4FFE55922FC10FB64FBD7FEA3" box="[983,1059,295,321]" pageId="15" pageNumber="354">n. syn.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are junior synonyms of
|
||
<taxonomicName id="4C4B4D1EFFE55922FAE8FB64FE6AFE87" authority="Walker, 1850" authorityName="Walker" authorityYear="1850" class="Insecta" family="Cicadidae" genus="Zammara" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="354" phylum="Arthropoda" rank="species" species="intricata">
|
||
<emphasis id="B93FEA8FFFE55922FAE8FB64FF03FE86" italics="true" pageId="15" pageNumber="354">Zammara intricata</emphasis>
|
||
<bibRefCitation id="EFDA4B6CFFE55922FEC5FB08FE6AFE87" author="Walker" box="[258,414,330,357]" firstAuthor="Walker" pageId="15" pageNumber="354" refId="ref16857" refString="Walker, F. (1850) List of the Specimens of Homopterous Insects in the Collection of the British Museum. Part I. British Museum Trustees, London, 260 pp. https: // doi. org / 10.5962 / bhl. title. 9063" type="book" year="1850">Walker, 1850</bibRefCitation>
|
||
</taxonomicName>
|
||
. Finally,
|
||
<taxonomicName id="4C4B4D1EFFE55922FDCBFB08FC47FE87" authority="Sanborn, 2004" authorityName="Sanborn" authorityYear="2004" box="[524,947,331,357]" class="Insecta" family="Cicadidae" genus="Zammara" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="354" phylum="Arthropoda" rank="species" species="medialinea">
|
||
<emphasis id="B93FEA8FFFE55922FDCBFB08FCF4FE87" box="[524,768,331,357]" italics="true" pageId="15" pageNumber="354">Zammara medialinea</emphasis>
|
||
<bibRefCitation id="EFDA4B6CFFE55922FCCCFB08FC47FE87" author="Sanborn" box="[779,947,331,357]" firstAuthor="Sanborn" pageId="15" pageNumber="354" pagination="365 - 371" refId="ref14433" refString="Sanborn, A. F. (2004) Two new Zammara species from South America (Hemiptera: Cicadoidea: Cicadidae). Florida Entomologist, 87, 365 - 371. https: // doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0365: tnzsfs] 2.0. co; 2" type="journal article" year="2004">Sanborn, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA8FFFE55922FC7AFB08FBFAFE87" bold="true" box="[957,1038,331,357]" pageId="15" pageNumber="354">
|
||
<taxonomicNameLabel id="A20C57F4FFE55922FC7AFB08FBFAFE87" box="[957,1038,331,357]" pageId="15" pageNumber="354">n. syn.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is a junior synonym of
|
||
<taxonomicName id="4C4B4D1EFFE55922FAE8FB08FE55FE6B" authority="Distant, 1883" authorityName="Distant" authorityYear="1883" class="Insecta" family="Cicadidae" genus="Zammara" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="354" phylum="Arthropoda" rank="species" species="luculenta">
|
||
<emphasis id="B93FEA8FFFE55922FAE8FB08FF0BFE6B" italics="true" pageId="15" pageNumber="354">Zammara luculenta</emphasis>
|
||
<bibRefCitation id="EFDA4B6CFFE55922FEC0FB2CFE55FE6B" author="Distant" box="[263,417,367,393]" firstAuthor="Distant" pageId="15" pageNumber="354" pagination="187 - 194" refId="ref12182" refString="Distant, W. L. (1883) Contributions to a proposed monograph of the homopterous family Cicadidae. Part I. Proceedings of the Zoological Society of London, 1883, 187 - 194, pl. 25. https: // doi. org / 10 / 1111 / j / 1469 - 7998.1883. tb 06649. x" type="journal article" year="1883">Distant, 1883</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFDA4B6CFFE55922FE6BFB2CFD9CFE6B" author="Goemans" box="[428,616,367,393]" firstAuthor="Goemans" pageId="15" pageNumber="354" refId="ref13149" refString="Goemans, G. (2016) The classification and phylogeny of the Neotropical cicada tribe Zammarini (Hemiptera, Cicadidae) and a revision of its type genus Zammara Amyot & Audinet-Serville, 1843 and its sister genus Zammaralna Boulard & Sueur, 1996. Ph. D. dissertation, University of Connecticut, Storrs, 206 pp." type="book" year="2016">Goemans (2016)</bibRefCitation>
|
||
illustrated the variability in coloration and wing patterns in these species which led to the description of multiple taxa for the highly variable species involved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |