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<document id="800CED90FF343C3B0EFCF48252E5916F" ID-CLB-Dataset="45587" ID-DOI="10.11646/zootaxa.3811.4.2" ID-GBIF-Dataset="dad75c9b-1930-47fb-ba79-eb519a90c45a" ID-ISSN="1175-5326" ID-Zenodo-Dep="253023" ID-ZooBank="99629EFB-F2F6-4DE8-AABA-D71069106912" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460432064619" checkinUser="plazi" docAuthor="Bilton, David T." docDate="2014" docId="03D687DBE34F0C02BBB98FA4FD169F70" docLanguage="en" docName="zt03811p462.pdf" docOrigin="Zootaxa 3811 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterosthetops hawequas Perkins 2008" docType="treatment" docVersion="11" lastPageNumber="458" masterDocId="FFEFFFA3E35B0C17BB2E8A1EFFE99E6C" masterDocTitle="New species and new records of Pterosthetops: eumadicolous water beetles of the South African Cape (Coleoptera, Hydraenidae)" masterLastPageNumber="462" masterPageNumber="438" pageNumber="458" updateTime="1698331499332" updateUser="ExternalLinkService">
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<mods:title id="41C3ACD3B797007EFC707F1027A2F4D3">New species and new records of Pterosthetops: eumadicolous water beetles of the South African Cape (Coleoptera, Hydraenidae)</mods:title>
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<mods:namePart id="6B8A3C4DBAC18A1124765D5B9A0D7F08">Bilton, David T.</mods:namePart>
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<mods:date id="B4410C93712204273F512DA123DFBFDE">2014</mods:date>
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<treatment id="03D687DBE34F0C02BBB98FA4FD169F70" ID-DOI="http://doi.org/10.5281/zenodo.6129835" ID-GBIF-Taxon="119430547" ID-Zenodo-Dep="6129835" LSID="urn:lsid:plazi:treatment:03D687DBE34F0C02BBB98FA4FD169F70" httpUri="http://treatment.plazi.org/id/03D687DBE34F0C02BBB98FA4FD169F70" lastPageId="21" lastPageNumber="458" pageId="20" pageNumber="458">
<subSubSection id="C3656546E34F0C02BBB98FA4FD169F70" lastPageId="21" lastPageNumber="459" pageId="20" pageNumber="458" type="nomenclature">
<paragraph id="8BC036CDE34F0C03BBB98FA4FD8A9BB9" blockId="20.[151,611,1466,1493]" box="[151,611,1466,1493]" pageId="20" pageNumber="458">
<heading id="D08881A1E34F0C03BBB98FA4FD8A9BB9" bold="true" box="[151,611,1466,1493]" fontSize="11" level="1" pageId="20" pageNumber="458" reason="1">
<taxonomicName id="4C7F4D4EE34F0C03BBB98FA4FD8A9BB9" ID-CoL="78LS2" authority="Perkins 2008" authorityName="Perkins" authorityYear="2008" box="[151,611,1466,1493]" class="Insecta" family="Hydraenidae" genus="Pterosthetops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="458" phylum="Arthropoda" rank="species" species="hawequas">
<emphasis id="B90BEADFE34F0C03BBB98FA4FD8A9BB9" bold="true" box="[151,611,1466,1493]" pageId="20" pageNumber="458">
<emphasis id="B90BEADFE34F0C03BBB98FA4FE509BB8" bold="true" box="[151,441,1466,1492]" italics="true" pageId="20" pageNumber="458">Pterosthetops hawequas</emphasis>
<bibRefCitation id="EFEE4B3CE34F0C03BAEF8FA5FD8A9BB9" author="Perkins" box="[449,611,1467,1493]" pageId="20" pageNumber="458" refString="Perkins, P. D. (2008) New species and new collection records of Prosthetopine water beetles from southern Africa (Coleoptera: Hydraenidae). Zootaxa, 1864, 1 - 124." type="journal article" year="2008">Perkins 2008</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC036CDE34F0C03BBB98FE2FCCF98C9" blockId="20.[151,1437,1532,2025]" pageId="20" pageNumber="458">
Described from a single male from
<taxonomicName id="4C7F4D4EE34F0C03B90F8FE3FD709878" box="[545,665,1533,1556]" class="Insecta" family="Hydraenidae" genus="Pterosthetops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="461" phylum="Arthropoda" rank="species" species="hawequas">Hawequas</taxonomicName>
, close to Du Toit’s Kloof Pass in the southwest of the Weastern Cape. New records show that this species is actually widespread in the Western Cape, its known range now extending from Franschhoek Pass, north to the edge of the Great Escarpment at Vanrhyns Pass (
<figureCitation id="13442A48E34F0C03BF538C5AFB2D9831" box="[1149,1220,1604,1629]" captionStart="FIGURE 6" captionStartId="18.[151,250,1906,1928]" captionTargetBox="[213,1385,202,1868]" captionTargetId="figure@18.[204,1394,193,1885]" captionTargetPageId="18" captionText="FIGURE 6. Habitats of Pterosthetops species. A) Middelberg Pass, seepages over rock with algae on R 303 road above Blinkwater— P. hawequas, P. tuberculatus sp. nov.; B) Franschhoekpas, 500 m wet rock face formed by stream beside R 45 road— P. hawequas, P. impressus, P. tuberculatus sp. nov.; C) View along Great Escarpment from Vanrhyns Pass— P. hawequas was abundant on seep below tree at bottom left of photo. Photos D T Bilton." httpUri="https://zenodo.org/record/253029/files/figure.png" pageId="20" pageNumber="458">Fig. 6</figureCitation>
C). Often the most abundant species of the genus, particularly on seepages with flocculated clay deposits (e.g. at Middelberg pass,
<figureCitation id="13442A48E34F0C03BE5E8C76FF4F98C9" captionStart="FIGURE 6" captionStartId="18.[151,250,1906,1928]" captionTargetBox="[213,1385,202,1868]" captionTargetId="figure@18.[204,1394,193,1885]" captionTargetPageId="18" captionText="FIGURE 6. Habitats of Pterosthetops species. A) Middelberg Pass, seepages over rock with algae on R 303 road above Blinkwater— P. hawequas, P. tuberculatus sp. nov.; B) Franschhoekpas, 500 m wet rock face formed by stream beside R 45 road— P. hawequas, P. impressus, P. tuberculatus sp. nov.; C) View along Great Escarpment from Vanrhyns Pass— P. hawequas was abundant on seep below tree at bottom left of photo. Photos D T Bilton." httpUri="https://zenodo.org/record/253029/files/figure.png" pageId="20" pageNumber="458">Fig. 6</figureCitation>
A). All additional records from rock faces below springs.
</paragraph>
<paragraph id="8BC036CDE34F0C02BBE98CAEFD169F70" blockId="20.[151,1437,1532,2025]" lastBlockId="21.[151,1436,151,285]" lastPageId="21" lastPageNumber="459" pageId="20" pageNumber="458">
Additional records:
<date id="FFC1100DE34F0C03BA878CAEFD8998A4" box="[425,608,1712,1737]" pageId="20" pageNumber="458" value="2002-09">September 2002</date>
Western Cape, Cederberg, seepages on rock beside R364 ca.
<quantity id="4C879B28E34F0C03BE328CAEFA8D98A5" box="[1308,1380,1712,1737]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="458" unit="km" value="20.0">20 km</quantity>
E of Clanwilliam—
<specimenCount id="9D79FD44E34F0C03BA6E8CCAFE879880" box="[320,366,1748,1773]" pageId="20" pageNumber="458" type="generic">3 ♂</specimenCount>
;
<date id="FFC1100DE34F0C03BAAE8CCAFDD59880" box="[384,572,1748,1773]" pageId="20" pageNumber="458" value="2002-09">September 2002</date>
Western Cape, Mitchell’s Pass nr. Ceres, seepages over rock—
<specimenCount id="9D79FD44E34F0C03BE1A8CCAFAB79880" box="[1332,1374,1748,1773]" pageId="20" pageNumber="458" type="generic">1 ♂</specimenCount>
, 2♀;
<date id="FFC1100DE34F0C03BBB98CE6FEA7997C" box="[151,334,1784,1809]" pageId="20" pageNumber="458" value="2002-09">September 2002</date>
Western Cape, Bainskloof Pass above Wellington, seepages over rock beside RXX road on north side of pass—
<specimenCount id="9D79FD44E34F0C03BA138D02FE8F9958" box="[317,358,1820,1844]" pageId="20" pageNumber="458" type="generic">1 ♂</specimenCount>
;
<date id="FFC1100DE34F0C03BA5D8D02FE049959" box="[371,493,1820,1845]" pageId="20" pageNumber="458" value="2008-09-26">26/ix/2008</date>
Western Cape, Du Toitskloof Pass, wet rock face—
<specimenCount id="9D79FD44E34F0C03BF6F8D02FB839958" box="[1089,1130,1820,1845]" pageId="20" pageNumber="458" type="generic">1 ♂</specimenCount>
;
<date id="FFC1100DE34F0C03BF598D02FB199959" box="[1143,1264,1820,1845]" pageId="20" pageNumber="458" value="2009-09-25">25/ix/2009</date>
Western Cape, Franschhoek Pass, wet rock faces beside R45 road (
<figureCitation id="13442A48E34F0C03B9DE8D5EFCDD9935" box="[752,820,1856,1881]" captionStart="FIGURE 6" captionStartId="18.[151,250,1906,1928]" captionTargetBox="[213,1385,202,1868]" captionTargetId="figure@18.[204,1394,193,1885]" captionTargetPageId="18" captionText="FIGURE 6. Habitats of Pterosthetops species. A) Middelberg Pass, seepages over rock with algae on R 303 road above Blinkwater— P. hawequas, P. tuberculatus sp. nov.; B) Franschhoekpas, 500 m wet rock face formed by stream beside R 45 road— P. hawequas, P. impressus, P. tuberculatus sp. nov.; C) View along Great Escarpment from Vanrhyns Pass— P. hawequas was abundant on seep below tree at bottom left of photo. Photos D T Bilton." httpUri="https://zenodo.org/record/253029/files/figure.png" pageId="20" pageNumber="458">Fig. 6</figureCitation>
B)—
<specimenCount id="9D79FD44E34F0C03B8438D5EFC709934" box="[877,921,1856,1881]" pageId="20" pageNumber="458" type="generic">4 ♂</specimenCount>
, 4 ♀;
<date id="FFC1100DE34F0C03B8C68D5EFB889935" box="[1000,1121,1856,1881]" pageId="20" pageNumber="458" value="2010-09-20">20/ix/2010</date>
Western Cape, Middelberg Pass, seepages over rock with algae on R303 road above Blinkwater (
<figureCitation id="13442A48E34F0C03B8EC8D7AFBEE9911" box="[962,1031,1892,1917]" captionStart="FIGURE 6" captionStartId="18.[151,250,1906,1928]" captionTargetBox="[213,1385,202,1868]" captionTargetId="figure@18.[204,1394,193,1885]" captionTargetPageId="18" captionText="FIGURE 6. Habitats of Pterosthetops species. A) Middelberg Pass, seepages over rock with algae on R 303 road above Blinkwater— P. hawequas, P. tuberculatus sp. nov.; B) Franschhoekpas, 500 m wet rock face formed by stream beside R 45 road— P. hawequas, P. impressus, P. tuberculatus sp. nov.; C) View along Great Escarpment from Vanrhyns Pass— P. hawequas was abundant on seep below tree at bottom left of photo. Photos D T Bilton." httpUri="https://zenodo.org/record/253029/files/figure.png" pageId="20" pageNumber="458">Fig. 6</figureCitation>
A)—
<specimenCount id="9D79FD44E34F0C03BF6C8D7AFB879910" box="[1090,1134,1892,1917]" pageId="20" pageNumber="458" type="generic">2 ♂</specimenCount>
, 4 ♀;
<date id="FFC1100DE34F0C03BF938D7AFADE9911" box="[1213,1335,1892,1917]" pageId="20" pageNumber="458" value="2010-09-21">21/ix/2010</date>
Western Cape, Cederberg, Uitkyk Pass, seepages over rock face beside road (
<figureCitation id="13442A48E34F0C03B8C58D96FBDF99CD" box="[1003,1078,1928,1953]" captionStart="FIGURE 5" captionStartId="17.[151,250,1843,1865]" captionTargetBox="[193,1381,200,1808]" captionTargetId="figure@17.[186,1391,193,1814]" captionTargetPageId="17" captionText="FIGURE 5. Habitats of Pterosthetops species. A) Cederberg, seepage over sloping rock beside stream @ 1,200 m below Wolfberg Arch, beside Matjiesrivier-Wupperthal road— P. coriaceus sp. nov., P. hawequas, P. impressus; B) Witzenberg, Michell’s Pass, wet rock face along R 46 road - P. baini sp. nov., P. hawequas, P. impressus (arrow indicates larva of Aspidytes niobe Ribera, Beutel, Balke &amp; Vogler, 2002); C) Cederberg, Uitkyk Pass, seepages over rock face beside road— P. harrisoni, P. hawequas, P. impressus, P. pulcherrimus sp. nov., P. tuberculatus sp. nov., P. uitkyki sp. nov.; D) Table Mountain, seepage over rock in Echo Valley— P. equestrius, P. impressus. Photos D T &amp; R Bilton." httpUri="https://zenodo.org/record/253028/files/figure.png" pageId="20" pageNumber="458">Fig. 5</figureCitation>
C);
<date id="FFC1100DE34F0C03BF478D96FB0B99CD" box="[1129,1250,1928,1953]" pageId="20" pageNumber="458" value="2011-09-23">23/ix/2011</date>
Western Cape, Cederberg, seepages over exposed rock @
<quantity id="4C879B28E34F0C03B9AC8DB2FD0B99A8" box="[642,738,1964,1988]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="458" unit="m" value="1200.0">1,200 m</quantity>
, below Wolfberg Arch, beside Matjiesrivier-Wupperthal road (
<figureCitation id="13442A48E34F0C03BBB18DCEFF089985" box="[159,225,2000,2025]" captionStart="FIGURE 5" captionStartId="17.[151,250,1843,1865]" captionTargetBox="[193,1381,200,1808]" captionTargetId="figure@17.[186,1391,193,1814]" captionTargetPageId="17" captionText="FIGURE 5. Habitats of Pterosthetops species. A) Cederberg, seepage over sloping rock beside stream @ 1,200 m below Wolfberg Arch, beside Matjiesrivier-Wupperthal road— P. coriaceus sp. nov., P. hawequas, P. impressus; B) Witzenberg, Michell’s Pass, wet rock face along R 46 road - P. baini sp. nov., P. hawequas, P. impressus (arrow indicates larva of Aspidytes niobe Ribera, Beutel, Balke &amp; Vogler, 2002); C) Cederberg, Uitkyk Pass, seepages over rock face beside road— P. harrisoni, P. hawequas, P. impressus, P. pulcherrimus sp. nov., P. tuberculatus sp. nov., P. uitkyki sp. nov.; D) Table Mountain, seepage over rock in Echo Valley— P. equestrius, P. impressus. Photos D T &amp; R Bilton." httpUri="https://zenodo.org/record/253028/files/figure.png" pageId="20" pageNumber="458">Fig. 5</figureCitation>
A)—1 ♀;
<date id="FFC1100DE34F0C03BA798DCEFE399985" box="[343,464,2000,2025]" pageId="20" pageNumber="458" value="2012-09-24">24/ix/2012</date>
Western Cape, Cederberg, Uitkyk Pass, seepages over rock face beside road at top of pass (
<figureCitation id="13442A48E34E0C02BBF28A89FECA9EC3" box="[220,291,151,176]" captionStart="FIGURE 7" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[157,1429,200,1895]" captionTargetId="figure@19.[151,1436,193,1901]" captionTargetPageId="19" captionText="FIGURE 7. Habitats of Pterosthetops species. A) Swartberg, seepage on sloping rock face beside R 328 road, north of Swartberg Pass, 2 km N of junction with Gamkaskloof road— P. br i nc k i, P. swartbergensis sp. nov. Seepage dry, January 2014; B) Cederberg, seepage at top of Uitkyk pass— P. hawequas, P. tuberculatus sp. nov. C) Cederberg, view from summit of Uitkyk Pass. Photos D T Bilton &amp; A Foggo." httpUri="https://zenodo.org/record/253030/files/figure.png" pageId="21" pageNumber="459">Fig. 7</figureCitation>
B &amp; C);
<date id="FFC1100DE34E0C02BABA8A86FE119EDC" box="[404,504,152,176]" pageId="21" pageNumber="459" value="2013-10-03">3/x/2013</date>
Western Cape, Cederberg, wet rock face in permanent spring below Wolfberg Cracks—abundant;
<date id="FFC1100DE34E0C02BA508AA2FE119EB9" box="[382,504,188,213]" pageId="21" pageNumber="459" value="2010-09-18">18/ix/2010</date>
Northern Cape, Vanrhyns Pass, seepage on rock face along R27 road (
<figureCitation id="13442A48E34E0C02BE418AA2FF4D9E94" captionStart="FIGURE 6" captionStartId="18.[151,250,1906,1928]" captionTargetBox="[213,1385,202,1868]" captionTargetId="figure@18.[204,1394,193,1885]" captionTargetPageId="18" captionText="FIGURE 6. Habitats of Pterosthetops species. A) Middelberg Pass, seepages over rock with algae on R 303 road above Blinkwater— P. hawequas, P. tuberculatus sp. nov.; B) Franschhoekpas, 500 m wet rock face formed by stream beside R 45 road— P. hawequas, P. impressus, P. tuberculatus sp. nov.; C) View along Great Escarpment from Vanrhyns Pass— P. hawequas was abundant on seep below tree at bottom left of photo. Photos D T Bilton." httpUri="https://zenodo.org/record/253029/files/figure.png" pageId="21" pageNumber="459">Fig. 6</figureCitation>
C)—abundant;
<date id="FFC1100DE34E0C02BA7B8AC1FE279E94" box="[341,462,223,248]" pageId="21" pageNumber="459" value="2011-09-27">27/ix/2011</date>
Northern Cape,
<quantity id="4C879B28E34E0C02B9BC8AFEFD269E94" box="[658,719,223,248]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="459" unit="km" value="1.0">1 km</quantity>
E. of Vanrhyns Pass, seepage on almost horizontal sandstone beside temporary pond—abundant. Distribution
<figureCitation id="13442A48E34E0C02B9998B1AFD139F70" box="[695,762,260,285]" captionStart="FIGURE 8" captionStartId="20.[151,250,1108,1130]" captionTargetBox="[216,1370,177,1086]" captionTargetId="figure@20.[216,1370,177,1086]" captionTargetPageId="20" captionText="FIGURE 8. Distribution of Pterosthetops species in the Western Cape: red symbols P. br i nc k i, green symbols P. equestrius, yellow symbols P. hawequas, white symbols P. indwei sp. nov., blue symbol P. swartbergensis sp. nov. Basemap source: ESri, GigitalGlobe, GeoEye, i-cubed, USDA, USGS, AEX, Getmapping, Aerogrid, IGN, IGP, Swisstopo and the GIS User Community." httpUri="https://zenodo.org/record/253031/files/figure.png" pageId="21" pageNumber="459">Fig. 8</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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