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<mods:title id="D5F6310EA2024DAF5FAE5CD36AEA232A">Aquatic Coleoptera Of Singapore: Species Richness, Ecology And Conservation #</mods:title>
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<heading id="D08881CDA739FFB33AC4F9E0BA214E0E" bold="true" box="[967,1343,1639,1663]" centered="true" fontSize="10" level="2" pageId="22" pageNumber="119" reason="2">
<taxonomicName id="4C7F4D22A739FFB33AC4F9E0BA214E0E" ID-CoL="7VG4V" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[967,1343,1639,1663]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="119" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B90BEAB3A739FFB33AC4F9E0BA214E0E" bold="true" box="[967,1343,1639,1663]" pageId="22" pageNumber="119">
<emphasis id="B90BEAB3A739FFB33AC4F9E0BB294E0E" bold="true" box="[967,1079,1639,1663]" italics="true" pageId="22" pageNumber="119">Hydaticus</emphasis>
?
<emphasis id="B90BEAB3A739FFB33D4FF9E0BBB34E0E" bold="true" box="[1100,1197,1639,1663]" italics="true" pageId="22" pageNumber="119">concolor</emphasis>
<bibRefCitation id="EFEE4B50A739FFB33DB7F9E0BA214E0E" author="Sharp, D" box="[1204,1343,1639,1663]" pageId="22" pageNumber="119" pagination="6 - 18" refId="ref41075" refString="Sharp, D., 1882. On aquatic carnivorous Coleoptera or Dytiscidae. Transactions of the Royal Dublin Society, (2) 2: 179 - 1003, pl. 6 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</emphasis>
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<treatmentCitation id="0ADE10B0A739FFB33A40F923BBD54EC8" author="Sharp, D" box="[835,1227,1700,1721]" page="662" pageId="22" pageNumber="119" year="1882">
<taxonomicName id="4C7F4D22A739FFB33A40F923BBD54EC8" ID-CoL="7VG4V" authority="Sharp, 1882: 662" authorityName="Sharp" authorityPageNumber="662" authorityYear="1882" box="[835,1227,1700,1721]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="119" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B90BEAB3A739FFB33A40F923BB124EC8" box="[835,1036,1700,1721]" italics="true" pageId="22" pageNumber="119">Hydaticus concolor</emphasis>
<bibRefCitation id="EFEE4B50A739FFB33D15F923BBD54EC8" author="Sharp, D" box="[1046,1227,1700,1721]" pageId="22" pageNumber="119" pagination="6 - 18" refId="ref41075" refString="Sharp, D., 1882. On aquatic carnivorous Coleoptera or Dytiscidae. Transactions of the Royal Dublin Society, (2) 2: 179 - 1003, pl. 6 - 18." type="journal article" year="1882">Sharp, 1882: 662</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="0ADE10B0A739FFB33DDAF923BAA04EC8" author="Regimbart, M" box="[1241,1470,1700,1721]" page="327" pageId="22" pageNumber="119" year="1899">
<bibRefCitation id="EFEE4B50A739FFB33DDAF923BAA04EC8" author="Regimbart, M" box="[1241,1470,1700,1721]" pageId="22" pageNumber="119" pagination="186 - 367" refId="ref40540" refString="Regimbart, M., 1899. Revision des Dytiscidae de la region indosino-malaise. Annales de la Societe Entomologique de France, 68: 186 - 367." type="journal article" year="1899">Régimbart, 1899: 327</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A739FFB33A6AF947BB4D4EA4" author="Zimmermann, A" box="[873,1107,1728,1749]" page="219" pageId="22" pageNumber="119" year="1920">
<bibRefCitation id="EFEE4B50A739FFB33A6AF947BB4D4EA4" author="Zimmermann, A" box="[873,1107,1728,1749]" pageId="22" pageNumber="119" pagination="1 - 326" refId="ref42405" refString="Zimmermann, A., 1920. Dytiscidae. In: Junk, W. &amp; S. Schenkling (eds.), Coleopterorum Catatalogus, Berlin, 4, pars 71, 1 - 326." type="journal article" year="1920">Zimmermann, 1920: 219</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A739FFB33D58F947BBBD4EA4" author="Zimmermann, A" box="[1115,1187,1728,1749]" page="6" pageId="22" pageNumber="119" year="1927">
<bibRefCitation id="EFEE4B50A739FFB33D58F947BBBD4EA4" author="Zimmermann, A" box="[1115,1187,1728,1749]" pageId="22" pageNumber="119" pagination="1 - 44" refId="ref42443" refString="Zimmermann, A., 1927. Fauna sumatrensis (Beitrag 45): Revision der Haliplidae et Dytiscidae von Sumatra. Supplementa entomologica, 16: 1 - 44." type="journal article" year="1927">1927: 6</bibRefCitation>
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;
<treatmentCitation id="0ADE10B0A739FFB33DAFF947BA454EA4" author="Vazirani, T" box="[1196,1371,1728,1749]" page="76" pageId="22" pageNumber="119" year="1977">
<bibRefCitation id="EFEE4B50A739FFB33DAFF947BA454EA4" author="Vazirani, T" box="[1196,1371,1728,1749]" pageId="22" pageNumber="119" refId="ref41335" refString="Vazirani, T. G., 1977. Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India, Occasional Paper, 6: ii + 111 pp." type="book" year="1977">Vazirani, 1977: 76</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A739FFB33C60F947BCD04E83" author="Wewalka, G" page="135" pageId="22" pageNumber="119" year="1979">
<bibRefCitation id="EFEE4B50A739FFB33C60F947BCD04E83" author="Wewalka, G" pageId="22" pageNumber="119" pagination="119 - 139" refId="ref41628" refString="Wewalka, G., 1979. Revision der Artengruppe des Hydaticus (Guignotites) fabricii (MacLeay), (Coleoptera, Dytiscidae). Koleopterologische Rundschau, 54: 119 - 139." type="journal article" year="1979">Wewalka, 1979: 135</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A739FFB33ADAF95ABB884E83" author="Nilsson, A" box="[985,1174,1757,1778]" page="101" pageId="22" pageNumber="119" year="2001">
<bibRefCitation id="EFEE4B50A739FFB33ADAF95ABB884E83" author="Nilsson, A" box="[985,1174,1757,1778]" pageId="22" pageNumber="119" refId="ref39046" refString="Nilsson, A. N., 2001. Dytiscidae (Coleoptera). In: World Catalogue of Insects Volume 3. Apollo Books, Stenstrup, Denmark, 395 pp." type="book" year="2001">Nilsson, 2001: 101</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C365652AA739FFB33A40F890BA8C4F38" pageId="22" pageNumber="119" type="materials_examined">
<paragraph id="8BC036A1A739FFB33A40F890BA8C4F38" blockId="22.[835,1474,1814,1865]" pageId="22" pageNumber="119">
<emphasis id="B90BEAB3A739FFB33A40F890BB0C4F5D" bold="true" box="[835,1042,1815,1836]" italics="true" pageId="22" pageNumber="119">Material examined. </emphasis>
<specimenCount id="9D79FD28A739FFB33D14F890BB9C4F5D" box="[1047,1154,1815,1836]" count="1" pageId="22" pageNumber="119" type="generic">1 specimen</specimenCount>
<emphasis id="B90BEAB3A739FFB33D94F891BA054F5A" bold="true" box="[1175,1307,1814,1835]" pageId="22" pageNumber="119">Other areas -</emphasis>
<materialsCitation id="3B173CFCA739FFB33C23F890BA904F38" collectingDate="1990-06-16" collectionCode="ZRC" collectorName="Lorong Banir &amp; Lua" pageId="22" pageNumber="119" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D79FD28A739FFB33C23F890BA6A4F5D" box="[1312,1396,1815,1836]" count="1" pageId="22" pageNumber="119" type="female">1 female</specimenCount>
,
<collectorName id="268A5377A739FFB33C7FF890BC624F38" pageId="22" pageNumber="119">Lorong Banir</collectorName>
, pond 1,
<date id="FFC11061A739FFB33ADBF8B3BB4C4F38" box="[984,1106,1844,1865]" pageId="22" pageNumber="119" value="1990-06-16">
<collectingDate id="EF85E989A739FFB33ADBF8B3BB4C4F38" box="[984,1106,1844,1865]" pageId="22" pageNumber="119" value="1990-06-16">16 Jun.1990</collectingDate>
</date>
, NS 190, coll.
<collectorName id="268A5377A739FFB33DE4F8B3BA134F38" box="[1255,1293,1844,1865]" pageId="22" pageNumber="119">Lua</collectorName>
et al. (
<collectionCode id="ED6EAE64A739FFB33C51F8B3BA994F38" box="[1362,1415,1844,1865]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="22" pageNumber="119">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C365652AA739FFB33A40F8EABB514FD4" pageId="22" pageNumber="119" type="biology_ecology">
<paragraph id="8BC036A1A739FFB33A40F8EABB514FD4" blockId="22.[835,1474,1901,1957]" pageId="22" pageNumber="119">
<emphasis id="B90BEAB3A739FFB33A40F8EABCA74FF4" bold="true" box="[835,953,1901,1925]" italics="true" pageId="22" pageNumber="119">Ecology. </emphasis>
Unknown. Probably a freshwater swamp forest species like
<taxonomicName id="4C7F4D22A739FFB33ACAF809BB564FD4" authorityName="Regimbart" authorityYear="1899" box="[969,1096,1934,1957]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="119" phylum="Arthropoda" rank="species" species="reductus">
<emphasis id="B90BEAB3A739FFB33ACAF809BB564FD4" box="[969,1096,1934,1957]" italics="true" pageId="22" pageNumber="119">H. reductus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C365652AA739FFB23A40F84ABE0A4929" lastPageId="23" lastPageNumber="120" pageId="22" pageNumber="119" type="discussion">
<paragraph id="8BC036A1A739FFB23A40F84ABE0A4929" blockId="22.[835,1474,1997,2117]" lastBlockId="23.[113,753,193,344]" lastPageId="23" lastPageNumber="120" pageId="22" pageNumber="119">
<emphasis id="B90BEAB3A739FFB33A40F84ABBBC4F94" box="[835,1186,1997,2021]" italics="true" pageId="22" pageNumber="119">
<emphasis id="B90BEAB3A739FFB33A40F84ABCDD4F94" bold="true" box="[835,963,1997,2021]" italics="true" pageId="22" pageNumber="119">Remarks. </emphasis>
<taxonomicName id="4C7F4D22A739FFB33ACFF849BBBC4F94" authorityName="Sharp" authorityYear="1882" box="[972,1186,1998,2021]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="119" phylum="Arthropoda" rank="species" species="concolor">Hydaticus concolor</taxonomicName>
</emphasis>
and other species such as
<taxonomicName id="4C7F4D22A739FFB33A40F869BBEE4074" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[835,1264,2030,2053]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="119" phylum="Arthropoda" rank="species" species="sexguttatus">
<emphasis id="B90BEAB3A739FFB33A40F869BB514074" box="[835,1103,2030,2053]" italics="true" pageId="22" pageNumber="119">Hydaticus sexguttatus</emphasis>
<bibRefCitation id="EFEE4B50A739FFB33D5FF869BBEE4074" author="Sharp, D" box="[1116,1264,2030,2053]" pageId="22" pageNumber="119" pagination="6 - 18" refId="ref41075" refString="Sharp, D., 1882. On aquatic carnivorous Coleoptera or Dytiscidae. Transactions of the Royal Dublin Society, (2) 2: 179 - 1003, pl. 6 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C7F4D22A739FFB33C30F869BCE74054" authority="Regimbart, 1899" authorityName="Regimbart" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="119" phylum="Arthropoda" rank="species" species="reductus">
<emphasis id="B90BEAB3A739FFB33C30F869BADC4074" box="[1331,1474,2030,2053]" italics="true" pageId="22" pageNumber="119">H. reductus</emphasis>
<bibRefCitation id="EFEE4B50A739FFB33A40F789BCE74054" author="Regimbart, M" box="[835,1017,2062,2085]" pageId="22" pageNumber="119" pagination="186 - 367" refId="ref40540" refString="Regimbart, M., 1899. Revision des Dytiscidae de la region indosino-malaise. Annales de la Societe Entomologique de France, 68: 186 - 367." type="journal article" year="1899">Régimbart, 1899</bibRefCitation>
</taxonomicName>
form an assemblage of small, comparably dark species that remain to be revised. Our specimens were compared with identified beetles in CGW and MNHN.
<taxonomicName id="4C7F4D22A738FFB23972FF66BE574889" authorityName="Sharp" authorityYear="1882" box="[113,329,225,248]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="120" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B90BEAB3A738FFB23972FF66BE574889" box="[113,329,225,248]" italics="true" pageId="23" pageNumber="120">Hydaticus concolor</emphasis>
</taxonomicName>
is recorded from
<collectingCountry id="F3687631A738FFB23B1BFF66BD664889" box="[536,632,225,248]" name="Vietnam" pageId="23" pageNumber="120">Vietnam</collectingCountry>
,
<collectingCountry id="F3687631A738FFB23B84FF66BDEE4889" box="[647,752,225,248]" name="Indonesia" pageId="23" pageNumber="120">Indonesia</collectingCountry>
(
<bibRefCitation id="EFEE4B50A738FFB2397AFE86BE3F4969" author="Vazirani, T" box="[121,289,257,280]" pageId="23" pageNumber="120" refId="ref41335" refString="Vazirani, T. G., 1977. Catalogue of Oriental Dytiscidae. Records of the Zoological Survey of India, Occasional Paper, 6: ii + 111 pp." type="book" year="1977">Vazirani, 1977</bibRefCitation>
;
<bibRefCitation id="EFEE4B50A738FFB23833FE86BEFF4969" author="Wewalka, G" box="[304,481,257,280]" pageId="23" pageNumber="120" pagination="119 - 139" refId="ref41628" refString="Wewalka, G., 1979. Revision der Artengruppe des Hydaticus (Guignotites) fabricii (MacLeay), (Coleoptera, Dytiscidae). Koleopterologische Rundschau, 54: 119 - 139." type="journal article" year="1979">Wewalka, 1979</bibRefCitation>
). If the determination is correct it will be the first record for the Malaysian Peninsula and
<collectingCountry id="F3687631A738FFB2399CFEC6BE0E4929" box="[159,272,321,344]" name="Singapore" pageId="23" pageNumber="120">Singapore</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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