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<document id="72D46371748305ABED0DC4DB9DF6C880" ID-DOI="10.5281/zenodo.5353544" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353544" ID-ZooBank="772BDDF2-AD14-4950-B3B3-D6A2C9E403C2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630210667410" checkinUser="felipe" docAuthor="Gordon, Robert D., F, Guillermo González &amp; Hanley, Guy A." docDate="2020" docId="03D687B4FF876A0AFF0AFF54FE25FB25" docLanguage="en" docName="0766.pdf" docOrigin="Insecta Mundi 2020 (766)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Calloeneis bennetti Gordon" docType="treatment" docVersion="4" lastPageNumber="15" masterDocId="FFEFFFCCFF976A1AFFCAFF83FFB6FF8F" masterDocTitle="South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="15" updateTime="1699187120983" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="35823A922087A86E70969FA81B935D93">South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini)</mods:title>
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<mods:namePart id="D141B27121610F86370B57228E13C932">Gordon, Robert D.</mods:namePart>
<mods:affiliation id="E510BB4F4F53B5ABA5B50C0962C7883F">Northern Plains Entomology PO Box 65 Willow City, ND 58384, USA</mods:affiliation>
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<mods:namePart id="6A19A5EB4695114E123BA172E733612A">F, Guillermo González</mods:namePart>
<mods:affiliation id="823DEAE0C909AC4F42798C6E3D08E600">Nocedal 6455 La Reina Santiago, Chile</mods:affiliation>
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<mods:namePart id="7790FB29A2291FE584E4ECED80223638">Hanley, Guy A.</mods:namePart>
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<mods:date id="79B4E2CC932E5E75A8CDEBE9F56EF3E9">2020</mods:date>
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<mods:identifier id="80F780C358FC5431DE756D71C0A50CE9" type="DOI">10.5281/zenodo.5353544</mods:identifier>
<mods:identifier id="A20A44FBB29B7FE5F3F7A743769E5C73" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03D687B4FF876A0AFF0AFF54FE25FB25" ID-DOI="http://doi.org/10.5281/zenodo.5451068" ID-Zenodo-Dep="5451068" LSID="urn:lsid:plazi:treatment:03D687B4FF876A0AFF0AFF54FE25FB25" httpUri="http://treatment.plazi.org/id/03D687B4FF876A0AFF0AFF54FE25FB25" lastPageNumber="15" pageId="16" pageNumber="15">
<subSubSection id="C3656529FF876A0AFF0AFF54FDD5FF60" box="[192,611,214,239]" pageId="16" pageNumber="15" type="nomenclature">
<paragraph id="8BC036A2FF876A0AFF0AFF54FDD5FF60" blockId="16.[192,611,214,239]" box="[192,611,214,239]" pageId="16" pageNumber="15">
<heading id="D08881CEFF876A0AFF0AFF54FDD5FF60" bold="true" box="[192,611,214,239]" fontSize="10" level="2" pageId="16" pageNumber="15" reason="3">
<emphasis id="B90BEAB0FF876A0AFF0AFF54FDD5FF60" bold="true" box="[192,611,214,239]" pageId="16" pageNumber="15">
18.
<taxonomicName id="4C7F4D21FF876A0AFF24FF55FDD5FF60" authority="Gordon" authorityName="Gordon" box="[238,611,214,239]" class="Insecta" family="Coccinellidae" genus="Calloeneis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="bennetti">
<emphasis id="B90BEAB0FF876A0AFF24FF55FE42FF61" bold="true" box="[238,500,214,238]" italics="true" pageId="16" pageNumber="15">Calloeneis bennetti</emphasis>
Gordon
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656529FF876A0AFF0AFE85FDD4FE93" box="[192,610,262,284]" pageId="16" pageNumber="15" type="reference_group">
<paragraph id="8BC036A2FF876A0AFF0AFE85FDD4FE93" blockId="16.[192,610,262,284]" box="[192,610,262,284]" pageId="16" pageNumber="15">
<treatmentCitationGroup id="AB6F118CFF876A0AFF0AFE85FDD4FE93" box="[192,610,262,284]" pageId="16" pageNumber="15">
<treatmentCitation id="0ADE10B3FF876A0AFF0AFE85FDEBFE93" author="Gordon, R. D." box="[192,605,262,284]" page="216" pageId="16" pageNumber="15" year="1978">
<taxonomicName id="4C7F4D21FF876A0AFF0AFE85FDEBFE93" authority="Gordon 1978: 216" authorityName="Gordon" authorityPageNumber="216" authorityYear="1978" box="[192,605,262,284]" class="Insecta" family="Coccinellidae" genus="Calloeneis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="bennetti">
<emphasis id="B90BEAB0FF876A0AFF0AFE85FE25FE94" box="[192,403,262,283]" italics="true" pageId="16" pageNumber="15">Calloeneis bennetti</emphasis>
<bibRefCitation id="EFEE4B53FF876A0AFE53FE85FDEBFE93" author="Gordon, R. D." box="[409,605,262,284]" pageId="16" pageNumber="15" pagination="205 - 218" refId="ref14846" refString="Gordon, R. D. 1978. West Indian Coccinellidae II (Coleoptera): Some scale predators with keys to genera and species. The Coleopterists Bulletin 32: 205 - 218." type="journal article" year="1978">Gordon 1978: 216</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3656529FF876A0AFF0AFEB7FE68FC28" pageId="16" pageNumber="15" type="description">
<paragraph id="8BC036A2FF876A0AFF0AFEB7FD96FC83" blockId="16.[190,1441,308,781]" pageId="16" pageNumber="15">
<emphasis id="B90BEAB0FF876A0AFF0AFEB7FE04FEC3" bold="true" box="[192,434,308,332]" pageId="16" pageNumber="15">Description. Male</emphasis>
. Length
<quantity id="4C879B47FF876A0AFDD1FEB7FDC1FEC4" box="[539,631,308,332]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="16" pageNumber="15" unit="mm" value="1.7">1.7 mm</quantity>
, width
<quantity id="4C879B47FF876A0AFD07FEB7FC9CFEC4" box="[717,810,308,332]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="16" pageNumber="15" unit="mm" value="1.6">1.6 mm</quantity>
; dorsal surface shiny except head faintly alutaceous. Color black (
<figureCitation id="13442A27FF876A0AFE90FED7FE04FEE3" box="[346,434,340,364]" captionStart="Figures 6980" captionStartId="25.[192,287,1678,1700]" captionTargetBox="[209,1403,231,1603]" captionTargetId="figure-78@25.[192,1440,213,1646]" captionTargetPageId="25" captionText="Figures 6980. Calloeneis spp.6971) Calloeneis appropinquans. 69) Habitus.70) Frons.71) Penis guide ventral. 7276) Calloeneis rosalie. 72) Habitus.73) Frons.74) Penis guide ventral.75) Penis guide lateral.76) Penis.7780) Calloeneis bennetti. 77) Habitus. 78) Frons. 79) Penis guide ventral. 80) Penis guide lateral." figureDoi="http://doi.org/10.5281/zenodo.5353558" httpUri="https://zenodo.org/record/5353558/files/figure.png" pageId="16" pageNumber="15">Fig. 77</figureCitation>
); head with vertex, median area of frons dark brown, lateral area of frons next to eye, clypeus yellowish brown; ventral surface yellow except mesosternum reddish brown, prosternum, metasternum dark brown; abdomen yellow with median 1/3 of basal ventrite brown. Head with dense punctures separated by less than a diameter; pronotal punctures small, separated by less than a diameter; elytral punctures larger than on pronotum, separated by a diameter or less; prosternal, meso-, metasternal punctures distinct, separated by about a diameter or less; basal abdominal ventrite without visible punctures, ventrites 2, 3 finely punctured, ventrites 3, 4 finely, densely punctured medially; ventrite 5 finely, densely punctured. Head slightly widened from middle of frons to clypeal apex, clypeal apex slightly rounded (
<figureCitation id="13442A27FF876A0AFE5CFDD7FE5DFDE3" box="[406,491,596,620]" captionStart="Figures 6980" captionStartId="25.[192,287,1678,1700]" captionTargetBox="[209,1403,231,1603]" captionTargetId="figure-78@25.[192,1440,213,1646]" captionTargetPageId="25" captionText="Figures 6980. Calloeneis spp.6971) Calloeneis appropinquans. 69) Habitus.70) Frons.71) Penis guide ventral. 7276) Calloeneis rosalie. 72) Habitus.73) Frons.74) Penis guide ventral.75) Penis guide lateral.76) Penis.7780) Calloeneis bennetti. 77) Habitus. 78) Frons. 79) Penis guide ventral. 80) Penis guide lateral." figureDoi="http://doi.org/10.5281/zenodo.5353558" httpUri="https://zenodo.org/record/5353558/files/figure.png" pageId="16" pageNumber="15">Fig. 78</figureCitation>
); eye canthus short, about half width of eye. Prosternum with lateral carina on each side curved, extended from apex of intercoxal process to lateral margin of prosternum. Epipleuron not descending externally. Postcoxal line on basal abdominal ventrite curved, extended 3/4 distance to rear margin of ventrite. Genitalia with penis guide 5/8 as long as paramere, widened anterior to base, then abruptly tapered to acute, slightly emarginate apex; paramere gradually widened from base to apex (
<figureCitation id="13442A27FF876A0AFEC6FD77FE39FC82" box="[268,399,756,781]" captionStart="Figures 6980" captionStartId="25.[192,287,1678,1700]" captionTargetBox="[209,1403,231,1603]" captionTargetId="figure-78@25.[192,1440,213,1646]" captionTargetPageId="25" captionText="Figures 6980. Calloeneis spp.6971) Calloeneis appropinquans. 69) Habitus.70) Frons.71) Penis guide ventral. 7276) Calloeneis rosalie. 72) Habitus.73) Frons.74) Penis guide ventral.75) Penis guide lateral.76) Penis.7780) Calloeneis bennetti. 77) Habitus. 78) Frons. 79) Penis guide ventral. 80) Penis guide lateral." figureDoi="http://doi.org/10.5281/zenodo.5353558" httpUri="https://zenodo.org/record/5353558/files/figure.png" pageId="16" pageNumber="15">Fig. 79, 80</figureCitation>
); penis lost.
</paragraph>
<paragraph id="8BC036A2FF876A0AFF0AFCA1FAB8FCD5" blockId="16.[192,1440,802,858]" pageId="16" pageNumber="15">
<emphasis id="B90BEAB0FF876A0AFF0AFCA1FE9BFCB5" bold="true" box="[192,301,802,826]" pageId="16" pageNumber="15">Female.</emphasis>
Similar to male except head entirely brown, genitalia with ramus of spermathecal capsule bulbous, cornu narrowed from base to abruptly rounded apex (
<figureCitation id="13442A27FF876A0AFC76FCC1FB85FCD5" box="[956,1075,834,858]" captionStart="Figures 2637" captionStartId="22.[192,287,1670,1692]" captionTargetBox="[206,1414,231,1600]" captionTargetId="figure-86@22.[192,1440,213,1646]" captionTargetPageId="22" captionText="Figures 2637. Calloeneis spp. 2629) Calloeneis myra. 26) Habitus.27) Frons.28) Penis guide ventral. 29) Penis guide lateral. 3031) Calloeneis leticia. 30) Habitus. 31) Frons. 3237) Calloeneis signata. 32) Habitus. 33) Frons. 34) Penis guide ventral. 35) Penis guide lateral. 36) Penis. 37) Spermathecal capsule." figureDoi="http://doi.org/10.5281/zenodo.5353552" httpUri="https://zenodo.org/record/5353552/files/figure.png" pageId="16" pageNumber="15">
Figure
<quantity id="4C879B47FF876A0AFBDFFCC1FB85FCD5" box="[1045,1075,834,858]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" pageId="16" pageNumber="15" unit="in" value="33.0">33</quantity>
</figureCitation>
in
<bibRefCitation id="EFEE4B53FF876A0AFB90FCC1FAB4FCD5" author="Gordon, R. D." box="[1114,1282,834,858]" pageId="16" pageNumber="15" pagination="205 - 218" refId="ref14846" refString="Gordon, R. D. 1978. West Indian Coccinellidae II (Coleoptera): Some scale predators with keys to genera and species. The Coleopterists Bulletin 32: 205 - 218." type="journal article" year="1978">Gordon, 1978</bibRefCitation>
).
</paragraph>
<paragraph id="8BC036A2FF876A0AFF0AFCECFE68FC28" blockId="16.[192,1440,879,935]" pageId="16" pageNumber="15">
<emphasis id="B90BEAB0FF876A0AFF0AFCECFEFAFC08" bold="true" box="[192,332,879,903]" pageId="16" pageNumber="15">Variation.</emphasis>
Length
<quantity id="4C879B47FF876A0AFE7BFCECFDECFC08" box="[433,602,879,903]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="16" pageNumber="15" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5 to 1.7 mm</quantity>
, width
<quantity id="4C879B47FF876A0AFD79FCECFCE9FC08" box="[691,863,879,903]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="16" pageNumber="15" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4 to 1.6 mm</quantity>
., male head may be entirely yellow with lateral 1/3 of pronotum yellow.
</paragraph>
</subSubSection>
<subSubSection id="C3656529FF876A0AFF0AFC3EFA9DFBD2" pageId="16" pageNumber="15" type="materials_examined">
<paragraph id="8BC036A2FF876A0AFF0AFC3EFDD7FC5A" blockId="16.[192,609,956,981]" box="[192,609,956,981]" pageId="16" pageNumber="15">
<emphasis id="B90BEAB0FF876A0AFF0AFC3EFECDFC5B" bold="true" box="[192,379,956,981]" pageId="16" pageNumber="15">
<typeStatus id="54C48800FF876A0AFF0AFC3EFEB2FC5A" box="[192,260,957,981]" pageId="16" pageNumber="15">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3687632FF876A0AFE48FC3EFE46FC5A" box="[386,496,957,981]" name="Trinidad and Tobago" pageId="16" pageNumber="15">Trinidad</collectingCountry>
,
<collectingRegion id="49BBF840FF876A0AFE36FC3EFDE8FC5A" box="[508,606,957,981]" country="Trinidad and Tobago" name="Mayaro/Rio Claro" pageId="16" pageNumber="15">Mayaro</collectingRegion>
.
</paragraph>
<paragraph id="8BC036A2FF876A0AFF0AFC69FDBDFB8D" blockId="16.[192,523,1002,1026]" box="[192,523,1002,1026]" pageId="16" pageNumber="15">
<emphasis id="B90BEAB0FF876A0AFF0AFC69FE11FB8D" bold="true" box="[192,423,1002,1026]" pageId="16" pageNumber="15">
<typeStatus id="54C48800FF876A0AFF0AFC69FEB2FB8D" box="[192,260,1002,1026]" pageId="16" pageNumber="15">Type</typeStatus>
depository.
</emphasis>
USNM.
</paragraph>
<paragraph id="8BC036A2FF876A0AFF0AFB94FD1CFBA0" blockId="16.[192,682,1047,1071]" box="[192,682,1047,1071]" pageId="16" pageNumber="15">
<emphasis id="B90BEAB0FF876A0AFF0AFB94FD86FBA0" bold="true" box="[192,560,1047,1071]" pageId="16" pageNumber="15">Geographical distribution.</emphasis>
<collectingCountry id="F3687632FF876A0AFDFCFB94FD13FBA0" box="[566,677,1047,1071]" name="Trinidad and Tobago" pageId="16" pageNumber="15">Trinidad</collectingCountry>
.
</paragraph>
<paragraph id="8BC036A2FF876A0AFF0AFBC6FA9DFBD2" blockId="16.[192,1323,1093,1117]" box="[192,1323,1093,1117]" pageId="16" pageNumber="15">
<emphasis id="B90BEAB0FF876A0AFF0AFBC6FE5FFBD2" bold="true" box="[192,489,1093,1117]" pageId="16" pageNumber="15">Specimens examined.</emphasis>
6.
<emphasis id="B90BEAB0FF876A0AFDC7FBC6FD39FBD2" bold="true" box="[525,655,1093,1117]" pageId="16" pageNumber="15">
<collectingCountry id="F3687632FF876A0AFDC7FBC6FD3FFBD2" box="[525,649,1093,1117]" name="Trinidad and Tobago" pageId="16" pageNumber="15">Trinidad</collectingCountry>
.
</emphasis>
Imperial College,
<collectingRegion id="49BBF840FF876A0AFCBAFBC6FC67FBD2" box="[880,977,1093,1117]" country="Trinidad and Tobago" name="Mayaro/Rio Claro" pageId="16" pageNumber="15">Mayaro</collectingRegion>
; Morne Bleu, St. Augustine.
</paragraph>
</subSubSection>
<subSubSection id="C3656529FF876A0AFF0AFBF1FE25FB25" pageId="16" pageNumber="15" type="discussion">
<paragraph id="8BC036A2FF876A0AFF0AFBF1FE25FB25" blockId="16.[190,1440,1138,1194]" pageId="16" pageNumber="15">
<emphasis id="B90BEAB0FF876A0AFF0AFBF1FEF5FB05" bold="true" box="[192,323,1138,1162]" pageId="16" pageNumber="15">Remarks.</emphasis>
This is another dorsally black species separable from other such species only by examination of male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>