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<document id="471693CCDCFDF359CCAAF93AC065E460" ID-DOI="10.37520/aemnp.2021.022" ID-ISSN="1804-6487" ID-Zenodo-Dep="5820961" ID-ZooBank="32519CC9-3658-469A-926D-6A1EBEE0FC59" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641388598144" checkinUser="plazi" docAuthor="Yamada, Kazutaka &amp; Yasunaga, Tomohide" docDate="2021" docId="03D687AFA11BFFD96B63FF3BFDFF8044" docLanguage="en" docName="ActaEntMusNatPra.61.2.375-426.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 61 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Anthocoris venustus Yamada &amp; Yasunaga 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="416" masterDocId="FFEFFFD7A13FFFF06A4AFF85FF9D8251" masterDocTitle="The tribe Anthocorini in Japan (Hemiptera: Anthocoridae): descriptions of new species, review of distribution and bionomics" masterLastPageNumber="426" masterPageNumber="375" pageNumber="411" updateTime="1699285380998" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
<mods:mods id="D6B63DEB4282580E72B031527C5E0558" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CB534AEA96D21DE133C9BA2AF68DF9EF">
<mods:title id="411D87F2DCFBE70280077454C034A2EE">The tribe Anthocorini in Japan (Hemiptera: Anthocoridae): descriptions of new species, review of distribution and bionomics</mods:title>
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<mods:name id="EB779C7BD5BF0A43596CA70A33722A26" type="personal">
<mods:role id="9A54B50AEFAE78E08C2462E9549DCAFA">
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</mods:role>
<mods:namePart id="550B262969C2A7A0F7567C85D19DF8D0">Yamada, Kazutaka</mods:namePart>
<mods:affiliation id="322982D08390E03659334D945D493D41">Tokushima Prefectural Museum, Bunka-no-Mori Park, Mukôterayama, Hachiman-chô, Tokushima 770 - 8070, Japan;</mods:affiliation>
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</mods:role>
<mods:namePart id="C1F6BA390F4049C3EAA4208F5E6E65A1">Yasunaga, Tomohide</mods:namePart>
<mods:affiliation id="4AD9F0F7BF37881CEAD51F6F673500D6">Research Associate, Division of Invertebrate Zoology, American Museum of Natural History, New York, c / o Nameshi</mods:affiliation>
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<mods:title id="573C04DCC324F4C5E085C321011245E4">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="2FADA0AD49074E87AF2D457E86C4F028">
<mods:date id="8692234A23E1247EC885FA882D2471AA">2021</mods:date>
<mods:detail id="D90915780CBE263077FA5F73EAA23B19" type="pubDate">
<mods:number id="43129504DE5F08267039552EA506AB0F">2021-10-19</mods:number>
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<mods:number id="5CAACDB42495A621995CD0E2E6F2AD0F">61</mods:number>
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<mods:identifier id="B1A6AF85E0524E4B9F655569BF57D571" type="DOI">10.37520/aemnp.2021.022</mods:identifier>
<mods:identifier id="71A5693C0A330492219089E8831042DA" type="ISSN">1804-6487</mods:identifier>
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<treatment id="03D687AFA11BFFD96B63FF3BFDFF8044" LSID="urn:lsid:plazi:treatment:03D687AFA11BFFD96B63FF3BFDFF8044" httpUri="http://treatment.plazi.org/id/03D687AFA11BFFD96B63FF3BFDFF8044" lastPageId="41" lastPageNumber="416" pageId="36" pageNumber="411">
<subSubSection id="C3656532A11BFFD46B63FF3BFDFB8284" box="[297,614,190,213]" pageId="36" pageNumber="411" type="nomenclature">
<paragraph id="8BC036B9A11BFFD46B63FF3BFDFB8284" blockId="36.[297,614,190,213]" box="[297,614,190,213]" pageId="36" pageNumber="411">
<heading id="D08881D5A11BFFD46B63FF3BFDFB8284" box="[297,614,190,213]" centered="true" fontSize="10" level="2" pageId="36" pageNumber="411" reason="3">
<emphasis id="B90BEAABA11BFFD46B63FF3BFDFB8284" bold="true" box="[297,614,190,213]" pageId="36" pageNumber="411">
<taxonomicName id="4C7F4D3AA11BFFD46B63FF3BFD948284" authority="Yamada &amp; Yasunaga, 2021" authorityName="Yamada &amp; Yasunaga" authorityYear="2021" box="[297,521,190,213]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="411" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="B90BEAABA11BFFD46B63FF3BFD948284" bold="true" box="[297,521,190,213]" italics="true" pageId="36" pageNumber="411">Anthocoris venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857D0A11BFFD4685AFF3BFDFB8284" box="[528,614,190,213]" pageId="36" pageNumber="411" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3656532A11BFFD46AACFF59FD1B804D" pageId="36" pageNumber="411" type="description">
<paragraph id="8BC036B9A11BFFD46AACFF59FD1B804D" blockId="36.[151,761,220,540]" pageId="36" pageNumber="411">
(
<figureCitation id="13442A3CA11BFFD46AA6FF59FED782BE" box="[236,330,220,239]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="36" pageNumber="411">Figs 8E–H</figureCitation>
,
<figureCitation id="13442A3CA11BFFD46B19FF59FEF282BE" box="[339,367,220,239]" captionStart="Fig" captionStartId="18.[151,182,1972,1991]" captionTargetBox="[204,1383,189,1959]" captionTargetId="figure-13@18.[204,1383,189,1962]" captionTargetPageId="18" captionText="Fig. 9. Scanning electron micrographs of ostiolar peritreme and evaporatorium of Anthocoris spp, left lateroventral view. A – A. chibi Hiura, 1959, female; B – A. confusus Reuter, 1884, male (from India); C – A. japonicus Poppius, 1909, female; D – A. kalopanacis Kerzhner, 1977, female; E – A. takahashii Hiura, 1959, female; F – A. miyamotoi Hiura, 1959, female; G – A. venustus sp. nov., paratype, male. Abbreviations: eva ‒ evaporatorium; mf ‒ median furrow; op ‒ ostiolar peritreme; sca ‒ supracoxal area." figureDoi="http://doi.org/10.5281/zenodo.5820987" httpUri="https://zenodo.org/record/5820987/files/figure.png" pageId="36" pageNumber="411">9G</figureCitation>
,
<figureCitation id="13442A3CA11BFFD46B32FF59FE2B82BE" box="[376,438,220,239]" captionStart="Fig" captionStartId="20.[150,181,2004,2023]" captionTargetBox="[151,1433,1025,1988]" captionTargetId="figure-684@20.[151,1436,1025,1991]" captionTargetPageId="20" captionText="Fig. 11. Scanning electron micrographs of male genitalia of Anthocoris spp. A–B – A. miyamotoi Hiura, 1959; C–D – A. venustus sp. nov., paratype. A, C – pygophore with paramere, dorsal (A) and lateral (C) views; B, D – paramere, dorsal (B) and lateral (D) views. Abbreviations: prm ‒ paramere; pyg ‒ pygophore." figureDoi="http://doi.org/10.5281/zenodo.5820997" httpUri="https://zenodo.org/record/5820997/files/figure.png" pageId="36" pageNumber="411">11C–D</figureCitation>
,
<figureCitation id="13442A3CA11BFFD46BF5FF59FE7B82BE" box="[447,486,220,239]" captionStart="Fig" captionStartId="22.[151,182,1996,2015]" captionTargetBox="[236,1400,657,1963]" captionTargetId="figure-13@22.[151,1436,648,1972]" captionTargetPageId="22" captionText="Fig. 12. Abdominal sterna II–III of Anthocoris spp., male, ventral view (setae omitted). A – A. chibi Hiura, 1959; B – A. confusus Reuter, 1884; C – A. japonicus Poppius, 1909; D – A. nemoralis (Fabricius, 1794); E – A. takahashii Hiura, 1959; F – A. miyamotoi Hiura, 1959; G – A. venustus sp. nov., paratype. Abbreviations: ma ‒ membranous area; st2–3 ‒ abdominal sternum II to III. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5820999" httpUri="https://zenodo.org/record/5820999/files/figure.png" pageId="36" pageNumber="411">12G</figureCitation>
,
<figureCitation id="13442A3CA11BFFD46BA5FF59FDB782BE" box="[495,554,220,239]" captionStart="Fig" captionStartId="26.[151,182,887,906]" captionTargetBox="[161,1401,197,863]" captionTargetId="figure-13@26.[151,1436,186,876]" captionTargetPageId="26" captionText="Fig. 14. Male genitalia of Anthocoris spp.A–C – A. miyamotoi Hiura, 1959; D–F – A. venustus sp. nov., paratype.A, D – pygophore with paramere (ejaculatory bulb omitted), dorsal view; B–C, E–F – paramere, two different views.Abbreviations: prm ‒ paramere; pyg ‒ pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821005" httpUri="https://zenodo.org/record/5821005/files/figure.png" pageId="36" pageNumber="411">14D–F</figureCitation>
,
<figureCitation id="13442A3CA11BFFD46879FF59FDC482BE" box="[563,601,220,239]" captionStart="Fig" captionStartId="30.[151,182,1991,2010]" captionTargetBox="[211,1367,202,1897]" captionTargetId="figure-13@30.[172,1416,189,1909]" captionTargetPageId="30" captionText="Fig. 15. Female genitalia of Anthocoris spp., dorsal view.A – A. chibi Hiura, 1959; B – A. confusus Reuter, 1884; C – A. japonicus Poppius, 1909; D – A. nemoralis (Fabricius,1794); E – A.takahashii Hiura, 1959; F – A. miyamotoi Hiura, 1959; G – A. venustus sp. nov., paratype.Abbreviations: ct ‒ copulatory tube; ism ‒ intersegmental membrane; sp ‒ sperm pouch; t ‒ trunk of conductive tissue (C–D, trunk of conductive tissue dissolved). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821009" httpUri="https://zenodo.org/record/5821009/files/figure.png" pageId="36" pageNumber="411">15G</figureCitation>
,
<figureCitation id="13442A3CA11BFFD46829FF59FD1982BE" box="[611,644,220,239]" captionStart="Fig" captionStartId="32.[151,182,1628,1647]" captionTargetBox="[151,1436,193,1607]" captionTargetId="figure-13@32.[151,1437,193,1607]" captionTargetPageId="32" captionText="Fig. 16. Habitus images of living individuals of Japanese Anthocorini spp. A – Acompocoris brevirostris Kerzhner, 1979, male; B – same, female; C – Tetraphleps aterrima (J. Sahlberg, 1878); D – Anthocoris confusus Reuter, 1884; E – A. takahashii Hiura, 1959; F – A. venustus sp. nov. (courtesy of M. Takai)." figureDoi="http://doi.org/10.5281/zenodo.5821011" httpUri="https://zenodo.org/record/5821011/files/figure.png" pageId="36" pageNumber="411">16F</figureCitation>
,
<figureCitation id="13442A3CA11BFFD468C7FF59FD3E82BE" box="[653,675,220,239]" captionStart="Fig" captionStartId="39.[153,184,2034,2053]" captionTargetBox="[165,1422,635,1952]" captionTargetId="figure-204@39.[151,1437,616,1970]" captionTargetPageId="39" captionText="Fig. 22. Distribution of Anthocoris miyamotoi Hiura, 1959 (circles) and A. venustus sp. nov. (triangles) in Japan. Filled symbols represent records based on specimens examined by us, open symbols represent literature records." figureDoi="http://doi.org/10.5281/zenodo.5821024" httpUri="https://zenodo.org/record/5821024/files/figure.png" pageId="36" pageNumber="411">22</figureCitation>
)
<taxonomicName id="4C7F4D3AA11BFFD46ADDFF7BFED38340" authorityName="Hiura" authorityYear="1959" box="[151,334,254,273]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="411" phylum="Arthropoda" rank="species" species="miyamotoi">
<emphasis id="B90BEAABA11BFFD46ADDFF7BFED38340" box="[151,334,254,273]" italics="true" pageId="36" pageNumber="411">Anthocoris miyamotoi</emphasis>
</taxonomicName>
(misidentification): Hංඎඋൺ (1959): 3 (description,in part);KൾඋඓHඇൾඋ (1988):774 (in key); Mංඒൺආඈඍඈ &amp; YൺඌඎඇൺGൺ (1989): 165 (listed, distribution); TඈආඈKඎඇං (1992): 158 (record); TඈආඈKඎඇං (1993): 168 (diagnosis, habitat, distribution); YඈඌHංඍඈආං (1994): 26 (record); Pඣඋංർൺඋඍ (1996): 113 (catalogue,distribution);Iංඃංආൺ (1997): 15 (record); Yൺආൺආඈඍඈ (1997): 15 (record); YൺඌඎඇൺGൺ (1997b): 224 (record); TൺKൾඇඈ (1998): 44 (listed, associated plant); Yൺආൺආඈඍඈ (1999): 94 (record); Yൺආൺආඈඍඈ (2001): 5 (record); KൺඇඒඎKඈඏൺ &amp; MൺඋඎඌංK (2006):168 (record);VංඇඈKඎඋඈඏ et al.(2010):57 (catalogue, distribution);Yൺආൺආඈඍඈ (2011):4 (record);Yൺආൺൽൺ et al. (2016):422 (catalogue, distribution); NൺKൺඍൺඇං (2019): 94 (record).
</paragraph>
</subSubSection>
<subSubSection id="C3656532A11BFFD66ADDFDB7FBA18310" lastPageId="38" lastPageNumber="413" pageId="36" pageNumber="411" type="materials_examined">
<paragraph id="8BC036B9A11BFFD66ADDFDB7FBA18310" blockId="36.[151,761,562,661]" lastBlockId="38.[828,1437,189,321]" lastPageId="38" lastPageNumber="413" pageId="36" pageNumber="411">
<emphasis id="B90BEAABA11BFFD46ADDFDB7FE908014" bold="true" box="[151,269,562,581]" pageId="36" pageNumber="411">
<typeStatus id="54C4881BA11BFFD46ADDFDB7FF5E8014" box="[151,195,562,581]" pageId="36" pageNumber="411">Type</typeStatus>
locality.
</emphasis>
Japan, Hokkaido,Hiyama-gun, Kaminokuni-cho,Yunotai.
<emphasis id="B90BEAABA11BFFD46ADDFDC8FE818031" bold="true" box="[151,284,589,608]" pageId="36" pageNumber="411">
<typeStatus id="54C4881BA11BFFD46ADDFDC8FF598031" box="[151,196,589,608]" pageId="36" pageNumber="411">Type</typeStatus>
material.
</emphasis>
Hඈඅඈඍඒඉൾ: J (
<figureCitation id="13442A3CA11BFFD46BEBFDC8FE608031" box="[417,509,589,608]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="36" pageNumber="411">Figs 8E–F</figureCitation>
), ‘JAPAN/ Hokkaido \ Yunotai, Hiyama \
<date id="FFC11079A11BFFD46B44FDEDFE1C802A" box="[270,385,616,635]" pageId="36" pageNumber="411" value="1994-08-19">19. viii. 1994</date>
\ T. Yasunaga leg.’ [printed], ‘On
<taxonomicName id="4C7F4D3AA11BFFD468E3FDEDFD68802A" box="[681,757,616,635]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="36" pageNumber="411" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEAABA11BFFD468E3FDEDFD49802A" box="[681,724,616,635]" italics="true" pageId="36" pageNumber="411">Salix</emphasis>
sp.
</taxonomicName>
’ [printed]; mounted on a triangular card, in intact condition (
<collectionCode id="ED6EAE7CA11BFFD468E5FD07FD7280C4" box="[687,751,642,661]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="36" pageNumber="411" type="Herbarium">TKPM</collectionCode>
). Pൺඋൺඍඒඉൾඌ:
<emphasis id="B90BEAABA11BFFD469EAFF3BFBC48280" bold="true" box="[928,1113,190,209]" pageId="36" pageNumber="411">
<collectingCountry id="F3687629A11BFFD469EAFF3BFC758280" box="[928,1000,190,209]" name="Japan" pageId="36" pageNumber="411">JAPAN</collectingCountry>
: HඈKKൺංൽඈ:
</emphasis>
2 JJ
<specimenCount id="9D79FD30A11BFFD46ED9FF3BFB2F8281" box="[1171,1202,183,209]" pageId="36" pageNumber="411" type="female">1 ♀</specimenCount>
, Wakkanai-shi,
<date id="FFC11079A11BFFD46F76FF3BFA048280" box="[1340,1433,190,209]" pageId="36" pageNumber="411" value="1974-09-15">15.ix.1974</date>
, Y. Furuki (
<collectionCode id="ED6EAE7CA11BFFD469D0FF5CFC5382BD" box="[922,974,217,236]" pageId="36" pageNumber="411">EUM</collectionCode>
); 4 JJ
<specimenCount id="9D79FD30A11BFFD46E5CFF5CFBAA82BA" box="[1046,1079,210,236]" pageId="36" pageNumber="411" type="female">1 ♀</specimenCount>
, Horokanai-cho, Moshiri,
<date id="FFC11079A11BFFD46F54FF5CFAE082BD" box="[1310,1405,217,236]" pageId="36" pageNumber="411" value="1994-08-03">3.viii.1994</date>
, on
<taxonomicName id="4C7F4D3AA11BFFD46976FF76FC1A8357" box="[828,903,243,262]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="36" pageNumber="411" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEAABA11BFFD46976FF76FCFA8357" box="[828,871,243,262]" italics="true" pageId="36" pageNumber="411">Salix</emphasis>
sp.
</taxonomicName>
, Y. Tôdo (1 J
<specimenCount id="9D79FD30A11BFFD46E4FFF76FBBB8357" box="[1029,1062,237,262]" pageId="36" pageNumber="411" type="female">1 ♀</specimenCount>
TKPM, 2 JJ
<collectionCode id="ED6EAE7CA11BFFD46EECFF76FB7B8357" box="[1190,1254,243,262]" country="Czech Republic" name="National Museum Prague" pageId="36" pageNumber="411">NMPC</collectionCode>
); 1 J
<specimenCount id="9D79FD30A11BFFD46F51FF76FAD78357" box="[1307,1354,237,262]" pageId="36" pageNumber="411" type="female">2 ♀♀</specimenCount>
, Horokanai-cho, Moshiri-Shirakaba,
<date id="FFC11079A11BFFD46E7DFE8BFB2C8370" box="[1079,1201,270,289]" pageId="36" pageNumber="411" value="1994-08-02" valueMax="1994-08-07" valueMin="1994-08-02">2.–7.viii.1994</date>
, on
<taxonomicName id="4C7F4D3AA11BFFD46E90FE8BFABA8370" box="[1242,1319,270,289]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="36" pageNumber="411" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEAABA11BFFD46E90FE8BFA988370" box="[1242,1285,270,289]" italics="true" pageId="36" pageNumber="411">Salix</emphasis>
sp.
</taxonomicName>
, T. Yasunaga (
<specimenCount id="9D79FD30A11BFFD46908FEACFCEF836A" box="[834,882,290,316]" pageId="36" pageNumber="411" type="female">2 ♀♀</specimenCount>
TKPM, 1 J ZMAS); 1 J, Monbetsu-shi, Kônomai,
<date id="FFC11079A11BFFD46F70FEACFA04836D" box="[1338,1433,297,316]" pageId="36" pageNumber="411" value="1997-08-03">3.viii.1997</date>
, S. Yamashita (
<collectionCode id="ED6EAE7CA11BFFD469F3FEC6FC648307" box="[953,1017,323,342]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="36" pageNumber="411" type="Herbarium">TKPM</collectionCode>
); 1 J
<specimenCount id="9D79FD30A11BFFD46E78FEC6FBFE8307" box="[1074,1123,317,342]" pageId="36" pageNumber="411" type="female">2 ♀♀</specimenCount>
, Higashikawa-cho, Mt. Asahi-dake,
<quantity id="4C879B5CA11BFFD46976FEDBFC028320" box="[828,927,350,369]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="36" pageNumber="411" unit="m" value="500.0" valueMax="800.0" valueMin="200.0">200–800 m</quantity>
,
<date id="FFC11079A11BFFD469EDFEDBFBB68320" box="[935,1067,350,369]" pageId="36" pageNumber="411" value="1994-08-09" valueMax="1994-08-10" valueMin="1994-08-09">9.–10.viii.1994</date>
, on
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<emphasis id="B90BEAABA11BFFD46E05FEDBFBE78320" box="[1103,1146,350,369]" italics="true" pageId="36" pageNumber="411">Salix</emphasis>
sp.
</taxonomicName>
, Y. Tôdo (1 J
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TKPM,
<specimenCount id="9D79FD30A11BFFD46FDBFEDBFCD683DA" pageId="36" pageNumber="411" type="female">1 ♀</specimenCount>
ZMAS); 1 J, Higashikawa-cho, Mt. Asahi-dake,
<quantity id="4C879B5CA11BFFD46EB3FEFCFAAF83DD" box="[1273,1330,377,396]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="36" pageNumber="411" unit="m" value="800.0">800 m</quantity>
,
<date id="FFC11079A11BFFD46F70FEFCFA0583DD" box="[1338,1432,377,396]" pageId="36" pageNumber="411" value="1994-08-09">9.viii.1994</date>
, R. Matsumoto (
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); 1 J
<specimenCount id="9D79FD30A11BFFD46E79FE16FBCF83F7" box="[1075,1106,397,422]" pageId="36" pageNumber="411" type="female">1 ♀</specimenCount>
, Higashikawa-cho, Tennin-kyô Valley,
<date id="FFC11079A11BFFD46976FE2BFC3E8390" box="[828,931,430,449]" pageId="36" pageNumber="411" value="1994-08-11">11.viii.1994</date>
, on
<taxonomicName id="4C7F4D3AA11BFFD46989FE2BFB918390" box="[963,1036,430,449]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="36" pageNumber="411" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEAABA11BFFD46989FE2BFC708390" box="[963,1005,430,449]" italics="true" pageId="36" pageNumber="411">Salix</emphasis>
sp.
</taxonomicName>
, Y.Tôdo (
<collectionCode id="ED6EAE7CA11BFFD46E14FE2BFB038390" box="[1118,1182,430,449]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="36" pageNumber="411" type="Herbarium">TKPM</collectionCode>
); 1J
<specimenCount id="9D79FD30A11BFFD46E87FE2BFB778391" box="[1229,1258,423,449]" pageId="36" pageNumber="411" type="female">1♀</specimenCount>
, Shintotsukawa-cho,
<date id="FFC11079A11BFFD46976FE4CFC03838D" box="[828,926,457,476]" pageId="36" pageNumber="411" value="1998-07-26">26.vii.1998</date>
, S. Gotoh (
<collectionCode id="ED6EAE7CA11BFFD469B0FE4CFBA7838D" box="[1018,1082,457,476]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="36" pageNumber="411" type="Herbarium">TKPM</collectionCode>
); 3JJ
<specimenCount id="9D79FD30A11BFFD46E33FE4CFB38838A" box="[1145,1189,450,476]" pageId="36" pageNumber="411" type="female">4♀♀</specimenCount>
(one in
<figureCitation id="13442A3CA11BFFD46EA3FE4CFAAB838D" box="[1257,1334,457,476]" captionStart="Fig" captionStartId="30.[151,182,1991,2010]" captionTargetBox="[211,1367,202,1897]" captionTargetId="figure-13@30.[172,1416,189,1909]" captionTargetPageId="30" captionText="Fig. 15. Female genitalia of Anthocoris spp., dorsal view.A – A. chibi Hiura, 1959; B – A. confusus Reuter, 1884; C – A. japonicus Poppius, 1909; D – A. nemoralis (Fabricius,1794); E – A.takahashii Hiura, 1959; F – A. miyamotoi Hiura, 1959; G – A. venustus sp. nov., paratype.Abbreviations: ct ‒ copulatory tube; ism ‒ intersegmental membrane; sp ‒ sperm pouch; t ‒ trunk of conductive tissue (C–D, trunk of conductive tissue dissolved). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821009" httpUri="https://zenodo.org/record/5821009/files/figure.png" pageId="36" pageNumber="411">Fig. 15G</figureCitation>
), Shari-cho, Mt. On-nebetsu-dake, Opekepu-rindô,
<date id="FFC11079A11BFFD46EC4FE66FABA83A7" box="[1166,1319,483,502]" pageId="36" pageNumber="411" value="1995-08-31" valueMax="1995-09-01" valueMin="1995-08-31">31.viii.–1.ix.1995</date>
, T. Yasunaga &amp; Y. Tôdo (1 J
<specimenCount id="9D79FD30A11BFFD46987FE7BFC628041" box="[973,1023,503,529]" pageId="36" pageNumber="411" type="female">3 ♀♀</specimenCount>
TKPM, 2 JJ
<specimenCount id="9D79FD30A11BFFD46EC9FE7BFB3B8041" box="[1155,1190,503,529]" pageId="36" pageNumber="411" type="female">1 ♀</specimenCount>
<collectionCode id="ED6EAE7CA11BFFD46EE3FE7BFB738040" box="[1193,1262,510,529]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="36" pageNumber="411">AMNH</collectionCode>
); 1 J, Akkeshi-cho, Suminoe,
<date id="FFC11079A11BFFD469DBFD9CFC6D807D" box="[913,1008,537,556]" pageId="36" pageNumber="411" value="2004-08-04">4.viii.2004</date>
, R. Matsumoto (
<collectionCode id="ED6EAE7CA11BFFD46E36FD9CFB5F807D" box="[1148,1218,537,556]" country="Japan" name="Osaka Museum of Natural History" pageId="36" pageNumber="411">OMNH</collectionCode>
); 1 J
<specimenCount id="9D79FD30A11BFFD46EBEFD9CFABC807A" box="[1268,1313,530,556]" pageId="36" pageNumber="411" type="female">2 ♀♀</specimenCount>
, Kushiro-cho, Takkobu,
<date id="FFC11079A11BFFD469DAFDB6FC648017" box="[912,1017,563,582]" pageId="36" pageNumber="411" value="1995-08-30">30.viii.1995</date>
, on
<taxonomicName id="4C7F4D3AA11BFFD46E50FDB6FBF98017" box="[1050,1124,563,582]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="36" pageNumber="411" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEAABA11BFFD46E50FDB6FBD88017" box="[1050,1093,563,582]" italics="true" pageId="36" pageNumber="411">Salix</emphasis>
sp.
</taxonomicName>
, Y.Tôdo (1 J TKPM,
<specimenCount id="9D79FD30A11BFFD46F6AFDB6FAD18017" box="[1312,1356,557,582]" pageId="36" pageNumber="411" type="female">2♀♀</specimenCount>
<collectionCode id="ED6EAE7CA11BFFD46F19FDB6FA0E8017" box="[1363,1427,563,582]" country="Czech Republic" name="National Museum Prague" pageId="36" pageNumber="411">NMPC</collectionCode>
);
<specimenCount id="9D79FD30A11BFFD46976FDCBFCC78031" box="[828,858,583,609]" pageId="36" pageNumber="411" type="female">1 ♀</specimenCount>
, Kushiro-cho, Iwabokki-suimon,
<date id="FFC11079A11BFFD46E3FFDCBFB408030" box="[1141,1245,590,609]" pageId="36" pageNumber="411" value="1995-08-28">28.viii.1995</date>
, on
<taxonomicName id="4C7F4D3AA11BFFD46EB4FDCBFADB8030" box="[1278,1350,590,609]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="36" pageNumber="411" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEAABA11BFFD46EB4FDCBFAB58030" box="[1278,1320,590,609]" italics="true" pageId="36" pageNumber="411">Salix</emphasis>
sp.
</taxonomicName>
, M.Ozaki (
<collectionCode id="ED6EAE7CA11BFFD46908FDECFC1F802D" box="[834,898,617,636]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="36" pageNumber="411" type="Herbarium">TKPM</collectionCode>
); 4 JJ (one in
<figureCitation id="13442A3CA11BFFD46E4CFDECFBD1802D" box="[1030,1100,617,636]" captionStart="Fig" captionStartId="18.[151,182,1972,1991]" captionTargetBox="[204,1383,189,1959]" captionTargetId="figure-13@18.[204,1383,189,1962]" captionTargetPageId="18" captionText="Fig. 9. Scanning electron micrographs of ostiolar peritreme and evaporatorium of Anthocoris spp, left lateroventral view. A – A. chibi Hiura, 1959, female; B – A. confusus Reuter, 1884, male (from India); C – A. japonicus Poppius, 1909, female; D – A. kalopanacis Kerzhner, 1977, female; E – A. takahashii Hiura, 1959, female; F – A. miyamotoi Hiura, 1959, female; G – A. venustus sp. nov., paratype, male. Abbreviations: eva ‒ evaporatorium; mf ‒ median furrow; op ‒ ostiolar peritreme; sca ‒ supracoxal area." figureDoi="http://doi.org/10.5281/zenodo.5820987" httpUri="https://zenodo.org/record/5820987/files/figure.png" pageId="36" pageNumber="411">Figs 9G</figureCitation>
,
<figureCitation id="13442A3CA11BFFD46E1FFDECFB0E802D" box="[1109,1171,617,636]" captionStart="Fig" captionStartId="20.[150,181,2004,2023]" captionTargetBox="[151,1433,1025,1988]" captionTargetId="figure-684@20.[151,1436,1025,1991]" captionTargetPageId="20" captionText="Fig. 11. Scanning electron micrographs of male genitalia of Anthocoris spp. A–B – A. miyamotoi Hiura, 1959; C–D – A. venustus sp. nov., paratype. A, C – pygophore with paramere, dorsal (A) and lateral (C) views; B, D – paramere, dorsal (B) and lateral (D) views. Abbreviations: prm ‒ paramere; pyg ‒ pygophore." figureDoi="http://doi.org/10.5281/zenodo.5820997" httpUri="https://zenodo.org/record/5820997/files/figure.png" pageId="36" pageNumber="411">11C–D</figureCitation>
, one in
<figureCitation id="13442A3CA11BFFD46E9CFDECFABA802D" box="[1238,1319,617,636]" captionStart="Fig" captionStartId="22.[151,182,1996,2015]" captionTargetBox="[236,1400,657,1963]" captionTargetId="figure-13@22.[151,1436,648,1972]" captionTargetPageId="22" captionText="Fig. 12. Abdominal sterna II–III of Anthocoris spp., male, ventral view (setae omitted). A – A. chibi Hiura, 1959; B – A. confusus Reuter, 1884; C – A. japonicus Poppius, 1909; D – A. nemoralis (Fabricius, 1794); E – A. takahashii Hiura, 1959; F – A. miyamotoi Hiura, 1959; G – A. venustus sp. nov., paratype. Abbreviations: ma ‒ membranous area; st2–3 ‒ abdominal sternum II to III. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5820999" httpUri="https://zenodo.org/record/5820999/files/figure.png" pageId="36" pageNumber="411">Figs 12G</figureCitation>
,
<figureCitation id="13442A3CA11BFFD46F7AFDECFAF1802D" box="[1328,1388,617,636]" captionStart="Fig" captionStartId="26.[151,182,887,906]" captionTargetBox="[161,1401,197,863]" captionTargetId="figure-13@26.[151,1436,186,876]" captionTargetPageId="26" captionText="Fig. 14. Male genitalia of Anthocoris spp.A–C – A. miyamotoi Hiura, 1959; D–F – A. venustus sp. nov., paratype.A, D – pygophore with paramere (ejaculatory bulb omitted), dorsal view; B–C, E–F – paramere, two different views.Abbreviations: prm ‒ paramere; pyg ‒ pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821005" httpUri="https://zenodo.org/record/5821005/files/figure.png" pageId="36" pageNumber="411">14D–F</figureCitation>
)
<specimenCount id="9D79FD30A11BFFD46F3DFDECFA0A802A" box="[1399,1431,610,636]" pageId="36" pageNumber="411" type="female">1 ♀</specimenCount>
, Makubetsu-cho, Chûrui, Kimontô,
<date id="FFC11079A11BFFD46E2AFD06FB5580C7" box="[1120,1224,643,662]" pageId="36" pageNumber="411" value="1995-08-21">21.viii.1995</date>
, on
<taxonomicName id="4C7F4D3AA11BFFD46EA2FD06FAAC80C7" box="[1256,1329,643,662]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="36" pageNumber="411" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEAABA11BFFD46EA2FD06FA8F80C7" box="[1256,1298,643,662]" italics="true" pageId="36" pageNumber="411">Salix</emphasis>
sp.
</taxonomicName>
, T.Yasunaga &amp; Y. Tôdo (
<collectionCode id="ED6EAE7CA11AFFD56B4EFF3BFED98280" box="[260,324,190,209]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56B1CFF3BFEE48281" box="[342,377,183,209]" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Nakasatsunai-mura, Pyoutan-no-Taki Fall,
<date id="FFC11079A11AFFD56ADDFF5CFF6482BD" box="[151,249,217,236]" pageId="37" pageNumber="412" value="2000-07-16">16.vii.2000</date>
, light trap, K.Yamada (
<collectionCode id="ED6EAE7CA11AFFD56BF3FF5CFE6482BD" box="[441,505,217,236]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
); 1J, Urakawa-cho,Nishicha, Hidaka-horobetsu-gawa Riv.,
<date id="FFC11079A11AFFD56BC5FF76FE6B8357" box="[399,502,243,262]" pageId="37" pageNumber="412" value="1995-08-18">18.viii.1995</date>
, on
<taxonomicName id="4C7F4D3AA11AFFD5685CFF76FDC38357" box="[534,606,243,262]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="37" pageNumber="412" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEAABA11AFFD5685CFF76FDDD8357" box="[534,576,243,262]" italics="true" pageId="37" pageNumber="412">Salix</emphasis>
sp.
</taxonomicName>
, Y.Tôdo (
<collectionCode id="ED6EAE7CA11AFFD568E5FF76FD738357" box="[687,750,243,262]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56ADDFE8BFF2B8371" box="[151,182,263,289]" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Tobetsu-cho, Aoyama,
<date id="FFC11079A11AFFD56BCEFE8BFE7A8370" box="[388,487,270,289]" pageId="37" pageNumber="412" value="1997-07-14">14.vii.1997</date>
, A. Makino (
<collectionCode id="ED6EAE7CA11AFFD5681FFE8BFD088370" box="[597,661,270,289]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
); 1 J, same locality, on
<taxonomicName id="4C7F4D3AA11AFFD56AB4FEACFE5C836D" box="[254,449,297,316]" class="Magnoliopsida" family="Oleaceae" genus="Fraxinus" kingdom="Plantae" order="Lamiales" pageId="37" pageNumber="412" phylum="Tracheophyta" rank="species" species="mandshurica">
<emphasis id="B90BEAABA11AFFD56AB4FEACFE5C836D" box="[254,449,297,316]" italics="true" pageId="37" pageNumber="412">Fraxinus mandshurica</emphasis>
</taxonomicName>
,
<date id="FFC11079A11AFFD56B87FEACFDAC836D" box="[461,561,297,316]" pageId="37" pageNumber="412" value="1997-07-26">26.vii.1997</date>
, T. Yasunaga (
<collectionCode id="ED6EAE7CA11AFFD568E5FEACFD73836D" box="[687,750,297,316]" pageId="37" pageNumber="412">TYCN</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56ADDFEC6FF258307" box="[151,184,317,342]" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Atsuta Village, Hatsutari, on
<taxonomicName id="4C7F4D3AA11AFFD56BF6FEC6FDE98307" box="[444,628,323,342]" class="Magnoliopsida" family="Juglandaceae" genus="Juglans" kingdom="Plantae" order="Fagales" pageId="37" pageNumber="412" phylum="Tracheophyta" rank="species" species="mandshurica">
<emphasis id="B90BEAABA11AFFD56BF6FEC6FDE98307" box="[444,628,323,342]" italics="true" pageId="37" pageNumber="412">Juglans mandshurica</emphasis>
</taxonomicName>
,
<date id="FFC11079A11AFFD56835FEC6FD438307" box="[639,734,323,342]" pageId="37" pageNumber="412" value="1998-08-07">7.viii.1998</date>
, T. Yasunaga (
<collectionCode id="ED6EAE7CA11AFFD56AB8FEDBFEAC8320" box="[242,305,350,369]" pageId="37" pageNumber="412">TYCN</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56B74FEDBFEC68321" box="[318,347,343,369]" pageId="37" pageNumber="412" type="female">1♀</specimenCount>
, Tobetsu-cho, Ishikari-gawa Riv.,
<date id="FFC11079A11AFFD56831FEDBFD418320" box="[635,732,350,369]" pageId="37" pageNumber="412" value="1997-07-17">17.vii.1997</date>
,A. Makino (
<collectionCode id="ED6EAE7CA11AFFD56AA9FEFCFEBE83DD" box="[227,291,377,396]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56B65FEFCFED183DA" box="[303,332,370,396]" pageId="37" pageNumber="412" type="female">1♀</specimenCount>
, Sapporo-shi,Mt.Moiwa,
<date id="FFC11079A11AFFD56866FEFCFD1483DD" box="[556,649,377,396]" pageId="37" pageNumber="412" value="1970-08-03">3.viii.1970</date>
, S. Kinoshita (
<collectionCode id="ED6EAE7CA11AFFD56AD7FE16FF4D83F7" box="[157,208,403,422]" pageId="37" pageNumber="412">EUM</collectionCode>
); 1J, Sapporo-shi, Otarunai-gawa-oku-rindô,
<geoCoordinate id="EE4B507EA11AFFD5681AFE16FD0683F7" box="[592,667,403,422]" degrees="43.082" direction="north" orientation="latitude" pageId="37" pageNumber="412" precision="55" value="43.082">43.082N</geoCoordinate>
,
<geoCoordinate id="EE4B507EA11AFFD568E8FE16FD6883F7" box="[674,757,403,422]" degrees="141.106" direction="east" orientation="longitude" pageId="37" pageNumber="412" precision="55" value="141.106">141.106E</geoCoordinate>
,
<date id="FFC11079A11AFFD56ADDFE2BFF678390" box="[151,250,430,449]" pageId="37" pageNumber="412" value="2019-07-28">28.vii.2019</date>
, A. Yamamoto (
<collectionCode id="ED6EAE7CA11AFFD56BCEFE2BFE598390" box="[388,452,430,449]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56B99FE2BFE6E8391" box="[467,499,423,449]" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Otaru-shi, Nagahashi-Naebo,
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,
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,
<date id="FFC11079A11AFFD56B0FFE4CFE06838D" box="[325,411,457,476]" pageId="37" pageNumber="412" value="2018-05-18">18.v.2018</date>
,A.Yamamoto (
<collectionCode id="ED6EAE7CA11AFFD56854FE4CFDC3838D" box="[542,606,457,476]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56821FE4CFD0A838A" box="[619,663,450,476]" pageId="37" pageNumber="412" type="female">2♀♀</specimenCount>
, Otaru-shi, Asari-tôge,
<date id="FFC11079A11AFFD56AB3FE66FEC783A7" box="[249,346,483,502]" pageId="37" pageNumber="412" value="1995-07-23">23.vii.1995</date>
, on
<taxonomicName id="4C7F4D3AA11AFFD56B30FE66FE5F83A7" box="[378,450,483,502]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="37" pageNumber="412" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEAABA11AFFD56B30FE66FE3983A7" box="[378,420,483,502]" italics="true" pageId="37" pageNumber="412">Salix</emphasis>
sp.
</taxonomicName>
, Y.Tôdo (
<collectionCode id="ED6EAE7CA11AFFD56859FE66FDCF83A7" box="[531,594,483,502]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
); 1J, same locality,
<date id="FFC11079A11AFFD56ADDFE7BFE9C8040" box="[151,257,510,529]" pageId="37" pageNumber="412" value="1996-08-18">18.viii.1996</date>
, on Alder, collector unknown (
<collectionCode id="ED6EAE7CA11AFFD5685BFE7BFDCC8040" box="[529,593,510,529]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56828FE7BFD198041" box="[610,644,503,529]" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Niseko, Mt. Chise-nupuri,
<quantity id="4C879B5CA11AFFD56B5BFD9CFEEE807D" box="[273,371,537,556]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="37" pageNumber="412" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">500–600 m</quantity>
,
<date id="FFC11079A11AFFD56B31FD9CFE4E807D" box="[379,467,537,556]" pageId="37" pageNumber="412" value="1994-07-09">9.vii.1994</date>
, on
<taxonomicName id="4C7F4D3AA11AFFD56BBFFD9CFF798017" authority=", T. Yasunaga (TKPM)" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="37" pageNumber="412" phylum="Tracheophyta" rank="species" species="crispula">
<emphasis id="B90BEAABA11AFFD56BBFFD9CFD1A807D" box="[501,647,537,556]" italics="true" pageId="37" pageNumber="412">Quercus crispula</emphasis>
, T. Yasunaga (TKPM)
</taxonomicName>
;
<specimenCount id="9D79FD30A11AFFD56AA5FDB6FE8F8017" box="[239,274,557,582]" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Chitose-shi, Lake side of Shikotsu-ko,
<date id="FFC11079A11AFFD56831FDB6FD698017" box="[635,756,563,582]" pageId="37" pageNumber="412" value="1984-08-05" valueMax="1984-08-06" valueMin="1984-08-05">5.–6.viii.1984</date>
, Y. Honda (
<collectionCode id="ED6EAE7CA11AFFD56ABCFDCBFEB78030" box="[246,298,590,609]" pageId="37" pageNumber="412">EUM</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56B70FDCBFEC68031" box="[314,347,583,609]" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Toyo-ura-cho, Iburi,
<date id="FFC11079A11AFFD56856FDCBFD188030" box="[540,645,590,609]" pageId="37" pageNumber="412" value="1997-08-21">21.viii.1997</date>
, T. Yasunaga (
<collectionCode id="ED6EAE7CA11AFFD56908FF41FC1F8286" box="[834,898,196,215]" pageId="37" pageNumber="412">TYCN</collectionCode>
). Rebun-tô Is.: 2 JJ, Hamanaka,
<date id="FFC11079A11AFFD56EF3FF41FA858286" box="[1209,1304,196,215]" pageId="37" pageNumber="412" value="1991-09-22">22.ix.1991</date>
, M. Tomokuni (
<collectionCode id="ED6EAE7CA11AFFD56908FF5BFC1C82A0" box="[834,897,222,241]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="37" pageNumber="412">NSMT</collectionCode>
); 1J, Kabukai,Rebun-rindô,
<date id="FFC11079A11AFFD56E3DFF5BFB4E82A0" box="[1143,1235,222,241]" pageId="37" pageNumber="412" value="1991-09-23">23.ix.1991</date>
, M.Tomokuni (
<collectionCode id="ED6EAE7CA11AFFD56F1EFF5BFA0E82A0" box="[1364,1427,222,241]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="37" pageNumber="412">NSMT</collectionCode>
). Rishiri-tô Is.: 2JJ
<specimenCount id="9D79FD30A11AFFD569AAFF7CFC60835A" box="[992,1021,242,268]" pageId="37" pageNumber="412" type="female">1♀</specimenCount>
, Oshidomari,
<date id="FFC11079A11AFFD56E3EFF7CFB4D835D" box="[1140,1232,249,268]" pageId="37" pageNumber="412" value="1991-09-17">17.ix.1991</date>
, M.Tomokuni (
<collectionCode id="ED6EAE7CA11AFFD56F1EFF7CFA09835D" box="[1364,1428,249,268]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="37" pageNumber="412">NSMT</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56976FE91FCC18377" box="[828,860,269,295]" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Oniwaki,
<date id="FFC11079A11AFFD569F6FE91FBD98376" box="[956,1092,276,295]" pageId="37" pageNumber="412" value="1994-07-28" valueMax="1994-07-29" valueMin="1994-07-28">28.–29.vii.1994</date>
, on
<taxonomicName id="4C7F4D3AA11AFFD56E23FE91FB298376" box="[1129,1204,276,295]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="37" pageNumber="412" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEAABA11AFFD56E23FE91FB098376" box="[1129,1172,276,295]" italics="true" pageId="37" pageNumber="412">Salix</emphasis>
sp.
</taxonomicName>
, Y. Tôdo (
<collectionCode id="ED6EAE7CA11AFFD56F46FE91FAD18376" box="[1292,1356,276,295]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
); 1 J, no detailed locality,
<date id="FFC11079A11AFFD5699BFEABFBCE8310" box="[977,1107,302,321]" pageId="37" pageNumber="412" value="1974-09-16" valueMax="1974-09-17" valueMin="1974-09-16">16.–17.ix.1974</date>
, Y. Furuki (
<collectionCode id="ED6EAE7CA11AFFD56EF7FEABFB6C8310" box="[1213,1265,302,321]" pageId="37" pageNumber="412">EUM</collectionCode>
).
<emphasis id="B90BEAABA11AFFD56F4AFEABFAC98310" bold="true" box="[1280,1364,302,321]" pageId="37" pageNumber="412">HඈඇඌHඎ:</emphasis>
<collectingRegion id="49BBF85BA11AFFD56F10FEABFCF4830D" country="Japan" name="Aomori" pageId="37" pageNumber="412">Aomori Pref.</collectingRegion>
:
<specimenCount id="9D79FD30A11AFFD56924FECCFC17830A" box="[878,906,322,348]" pageId="37" pageNumber="412" type="female">1♀</specimenCount>
, Mutsu-shi,Sekine,
<date id="FFC11079A11AFFD56E7CFECCFB16830D" box="[1078,1163,329,348]" pageId="37" pageNumber="412" value="1982-05-31">31.v.1982</date>
, B. Tanaka (
<collectionCode id="ED6EAE7CA11AFFD56EA5FECCFAB2830D" box="[1263,1327,329,348]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
).
<collectingRegion id="49BBF85BA11AFFD56F71FECCFA06830D" box="[1339,1435,329,348]" country="Japan" name="Iwate" pageId="37" pageNumber="412">Iwate Pref.</collectingRegion>
:
<specimenCount id="9D79FD30A11AFFD56976FEE1FCC28327" box="[828,863,349,375]" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Yahaba-cho, Mt. Nansyou-zan,
<date id="FFC11079A11AFFD56EC9FEE1FB418326" box="[1155,1244,356,375]" pageId="37" pageNumber="412" value="2004-07-01">1.vii.2004</date>
, T. Ishizaki (
<collectionCode id="ED6EAE7CA11AFFD56F19FEE1FA098326" box="[1363,1428,356,375]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
).
<collectingRegion id="49BBF85BA11AFFD56976FEFBFC3183C0" box="[828,940,382,401]" country="Japan" name="Tochigi" pageId="37" pageNumber="412">Tochigi Pref.</collectingRegion>
: 1J, Nasushiobara-shi, Nakagawa-kahan-kôen,
<date id="FFC11079A11AFFD56F76FEFBFA0483C0" box="[1340,1433,382,401]" pageId="37" pageNumber="412" value="2016-08-04">4.viii.2016</date>
, S. Maehara (
<collectionCode id="ED6EAE7CA11AFFD569EEFE1CFC7983FD" box="[932,996,409,428]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
); 1J, Nikko-shi, Kawamata,
<date id="FFC11079A11AFFD56E90FE1CFAAB83FD" box="[1242,1334,409,428]" pageId="37" pageNumber="412" value="2019-09-27">27.ix.2019</date>
, S. Maehara (
<collectionCode id="ED6EAE7CA11AFFD56908FE31FC1F8396" box="[834,898,436,455]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD569D8FE31FC2E8397" box="[914,947,429,455]" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Nikko-shi, Yumoto,
<date id="FFC11079A11AFFD56E3BFE31FB5A8396" box="[1137,1223,436,455]" pageId="37" pageNumber="412" value="2007-05-21">21.v.2007</date>
, S. Maehara (
<collectionCode id="ED6EAE7CA11AFFD56F0BFE31FA1C8396" box="[1345,1409,436,455]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
); 1 J(
<figureCitation id="13442A3CA11AFFD5691DFE4BFC0683B0" box="[855,923,462,481]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="37" pageNumber="412">Fig. 8G</figureCitation>
), Nikko-shi, Ryû-oh-kyô,
<date id="FFC11079A11AFFD56E31FE4BFB4383B0" box="[1147,1246,462,481]" pageId="37" pageNumber="412" value="2010-07-19">19.vii.2010</date>
, S. Maehara (
<collectionCode id="ED6EAE7CA11AFFD56F19FE4BFA0E83B0" box="[1363,1427,462,481]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
); 1 J
<specimenCount id="9D79FD30A11AFFD5692BFE6CFC1C83AA" box="[865,897,482,508]" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Nikko-shi, Shôbugahama,
<date id="FFC11079A11AFFD56E25FE6CFB5083AD" box="[1135,1229,489,508]" pageId="37" pageNumber="412" value="2010-08-05">5.viii.2010</date>
, S. Maehara (
<collectionCode id="ED6EAE7CA11AFFD56F09FE6CFA1E83AD" box="[1347,1411,489,508]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56FD8FE6CFCD18047" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Nikko-shi, Chȗgȗshi, Asegata,
<date id="FFC11079A11AFFD56E16FD81FB278046" box="[1116,1210,516,535]" pageId="37" pageNumber="412" value="2020-09-18">18.ix.2020</date>
, S. Maehara (
<collectionCode id="ED6EAE7CA11AFFD56F61FD81FAF68046" box="[1323,1387,516,535]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56F32FD81FA0B8047" box="[1400,1430,509,535]" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Shimotsuke-shi,Hosoya,
<date id="FFC11079A11AFFD56E44FD9BFBC08060" box="[1038,1117,542,561]" pageId="37" pageNumber="412" value="2000-11-04">4.xi.2000</date>
, S. Maehara (
<collectionCode id="ED6EAE7CA11AFFD56E81FD9BFA978060" box="[1227,1290,542,561]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56F5CFD9BFAAF8060" box="[1302,1330,536,561]" pageId="37" pageNumber="412" type="female">1♀</specimenCount>
, Mooka-shi, Isagahara,
<date id="FFC11079A11AFFD569D3FDBCFC71801D" box="[921,1004,569,588]" pageId="37" pageNumber="412" value="2018-06-06">6.vi.2018</date>
, S. Maehara (
<collectionCode id="ED6EAE7CA11AFFD56E2EFDBCFB39801D" box="[1124,1188,569,588]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="37" pageNumber="412" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56EFEFDBCFB49801A" box="[1204,1236,562,588]" pageId="37" pageNumber="412" type="female">1 ♀</specimenCount>
, Tochigi-shi, Hoshino- -machi,
<date id="FFC11079A11AFFD569CAFDD1FC4E8036" box="[896,979,596,615]" pageId="37" pageNumber="412" value="1991-03-06">6.iii.1991</date>
, H.Yoshitomi (
<collectionCode id="ED6EAE7CA11AFFD56E1EFDD1FB158036" box="[1108,1160,596,615]" pageId="37" pageNumber="412">EUM</collectionCode>
);
<specimenCount id="9D79FD30A11AFFD56EDFFDD1FB2F8037" box="[1173,1202,589,615]" pageId="37" pageNumber="412" type="female">1♀</specimenCount>
, Tochigi-shi, Oominagawa,
<date id="FFC11079A119FFD66ADDFF41FF438286" box="[151,222,196,215]" pageId="38" pageNumber="413" value="1994-01-06">6.i.1994</date>
, H. Yoshitomi (
<collectionCode id="ED6EAE7CA119FFD66B23FF41FE008286" box="[361,413,196,215]" pageId="38" pageNumber="413">EUM</collectionCode>
).
<collectingRegion id="49BBF85BA119FFD66BE7FF41FDB98286" box="[429,548,196,215]" country="Japan" name="Nagano" pageId="38" pageNumber="413">Nagano Pref.</collectingRegion>
:
<specimenCount id="9D79FD30A119FFD66866FF41FDD38287" box="[556,590,189,215]" pageId="38" pageNumber="413" type="female">1 ♀</specimenCount>
, Kijimadaira-mura, Kayanodaira,
<date id="FFC11079A119FFD66B44FF5BFEC482A0" box="[270,345,222,241]" pageId="38" pageNumber="413" value="2000-05-01">1.v.2000</date>
, Y. Kita (
<collectionCode id="ED6EAE7CA119FFD66BECFF5BFE7B82A0" box="[422,486,222,241]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="38" pageNumber="413" type="Herbarium">TKPM</collectionCode>
).
<collectingRegion id="49BBF85BA119FFD66BBEFF5BFDD082A0" box="[500,589,222,241]" country="Japan" name="Gifu" pageId="38" pageNumber="413">Gifu Pref.</collectingRegion>
:
<specimenCount id="9D79FD30A119FFD66819FF5BFDEC82A0" box="[595,625,216,241]" pageId="38" pageNumber="413" type="female">1 ♀</specimenCount>
, Takayama-shi, Hirayu,
<date id="FFC11079A119FFD66A90FF7CFEA1835D" box="[218,316,249,268]" pageId="38" pageNumber="413" value="1959-07-28">28.vii.1959</date>
, M. Miyatake (
<collectionCode id="ED6EAE7CA119FFD66BF2FF7CFE76835D" box="[440,491,249,268]" pageId="38" pageNumber="413">EUM</collectionCode>
).
<collectingRegion id="49BBF85BA119FFD66BBDFF7CFDCB835D" box="[503,598,249,268]" country="Japan" name="Aichi" pageId="38" pageNumber="413">Aichi Pref.</collectingRegion>
:
<specimenCount id="9D79FD30A119FFD66811FF7CFDEA835A" box="[603,631,242,268]" pageId="38" pageNumber="413" type="female">1♀</specimenCount>
, Toyone-mura, Mt.Chausu-yama,
<date id="FFC11079A119FFD66B7EFE91FE248376" box="[308,441,276,295]" pageId="38" pageNumber="413" value="1974-07-15" valueMax="1974-07-16" valueMin="1974-07-15">15.–16.vii.1974</date>
,Y.Hori (
<collectionCode id="ED6EAE7CA119FFD6684FFE91FDD98376" box="[517,580,276,295]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="38" pageNumber="413" type="Herbarium">TKPM</collectionCode>
).
<collectingRegion id="49BBF85BA119FFD6681AFE91FD348376" box="[592,681,276,295]" country="Japan" name="Nara" pageId="38" pageNumber="413">Nara Pref.</collectingRegion>
:1J (
<figureCitation id="13442A3CA119FFD6689CFE91FF2E8310" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="38" pageNumber="413">Fig. 8H</figureCitation>
),
<specimenCount id="9D79FD30A119FFD66A89FEABFF6E8310" box="[195,243,296,321]" pageId="38" pageNumber="413" type="female">2 ♀♀</specimenCount>
, Kamikitayama-mura, Ôdaigahara,
<date id="FFC11079A119FFD6686DFEABFD0D8310" box="[551,656,302,321]" pageId="38" pageNumber="413" value="2004-08-16">16.viii.2004</date>
, K. Yamada (
<collectionCode id="ED6EAE7CA119FFD66AD7FECCFF40830D" box="[157,221,329,348]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="38" pageNumber="413" type="Herbarium">TKPM</collectionCode>
); 1J
<specimenCount id="9D79FD30A119FFD66B47FECCFEA7830A" box="[269,314,322,348]" pageId="38" pageNumber="413" type="female">2 ♀♀</specimenCount>
, same locality,
<date id="FFC11079A119FFD66BF7FECCFDDD830D" box="[445,576,329,348]" pageId="38" pageNumber="413" value="2004-11-14" valueMax="2004-11-15" valueMin="2004-11-14">14.–15.xi.2004</date>
, K.Yamada (
<collectionCode id="ED6EAE7CA119FFD668E4FECCFD73830D" box="[686,750,329,348]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="38" pageNumber="413" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A119FFD66ADDFEE1FF2B8327" box="[151,182,349,375]" pageId="38" pageNumber="413" type="female">1 ♀</specimenCount>
, Nosegawa-mura, Mt. Obako-dake,
<date id="FFC11079A119FFD66BA0FEE1FDA28326" box="[490,575,356,375]" pageId="38" pageNumber="413" value="2000-05-25">25.v.2000</date>
, K. Yamada (
<collectionCode id="ED6EAE7CA119FFD668E5FEE1FD728326" box="[687,751,356,375]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="38" pageNumber="413" type="Herbarium">TKPM</collectionCode>
).
<emphasis id="B90BEAABA119FFD66ADDFEFBFF7283C0" bold="true" box="[151,239,382,401]" pageId="38" pageNumber="413">SHංKඈKඎ:</emphasis>
<collectingRegion id="49BBF85BA119FFD66AB9FEFBFE1983C0" box="[243,388,382,401]" country="Japan" name="Tokushima" pageId="38" pageNumber="413">Tokushima Pref.</collectingRegion>
: 2JJ
<specimenCount id="9D79FD30A119FFD66BF6FEFBFE4783C0" box="[444,474,376,401]" pageId="38" pageNumber="413" type="female">1 ♀</specimenCount>
, Naka-cho, Kisawa,
<date id="FFC11079A119FFD668C1FEFBFD6983C0" box="[651,756,382,401]" pageId="38" pageNumber="413" value="1998-08-16">16.viii.1998</date>
, M. Takai (
<collectionCode id="ED6EAE7CA119FFD66AA1FE1CFEB783FD" box="[235,298,409,428]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="38" pageNumber="413" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A119FFD66B7DFE1CFECE83FA" box="[311,339,402,428]" pageId="38" pageNumber="413" type="female">1♀</specimenCount>
, Naka-cho, Kawanari-tôge,
<date id="FFC11079A119FFD66875FE1CFD0883FD" box="[575,661,409,428]" pageId="38" pageNumber="413" value="2011-07-06">6.vii.2011</date>
, K.Yamada (
<collectionCode id="ED6EAE7CA119FFD66AD7FE31FF408396" box="[157,221,436,455]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="38" pageNumber="413" type="Herbarium">TKPM</collectionCode>
); 1 J, Naka-cho, Iwakura,
<date id="FFC11079A119FFD66B8EFE31FDB18396" box="[452,556,436,455]" pageId="38" pageNumber="413" value="2010-08-23">23.viii.2010</date>
, K. Yamada (
<collectionCode id="ED6EAE7CA119FFD668D4FE31FD438396" box="[670,734,436,455]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="38" pageNumber="413" type="Herbarium">TKPM</collectionCode>
); 1 unsexed, Naka-cho, Ichinomori to Nishijima,
<date id="FFC11079A119FFD66854FE4BFD1C83B0" box="[542,641,462,481]" pageId="38" pageNumber="413" value="1984-07-16">16.vii.1984</date>
, M. Miyatake (
<collectionCode id="ED6EAE7CA119FFD66AD7FE6CFF4C83AD" box="[157,209,489,508]" pageId="38" pageNumber="413">EUM</collectionCode>
); 1 J, Mt. Tsurugi,
<date id="FFC11079A119FFD66B33FE6CFE7C83AD" box="[377,481,489,508]" pageId="38" pageNumber="413" value="1959-08-14">14.viii.1959</date>
, I. Hiura (
<collectionCode id="ED6EAE7CA119FFD66872FE6CFDE383AD" box="[568,638,489,508]" country="Japan" name="Osaka Museum of Natural History" pageId="38" pageNumber="413">OMNH</collectionCode>
).
<collectingRegion id="49BBF85BA119FFD668C6FE6CFD6B83AD" box="[652,758,489,508]" country="Japan" name="Ehime" pageId="38" pageNumber="413">Ehime Pref.</collectingRegion>
:
<specimenCount id="9D79FD30A119FFD66ADDFD81FF278047" box="[151,186,509,535]" pageId="38" pageNumber="413" type="female">1 ♀</specimenCount>
, Kumakôgen-cho, Omogo-kei,
<date id="FFC11079A119FFD66B9BFD81FDB08046" box="[465,557,516,535]" pageId="38" pageNumber="413" value="1958-04-26">26.iv.1958</date>
, Y. Takaishi (
<collectionCode id="ED6EAE7CA119FFD668EDFD81FD468046" box="[679,731,516,535]" pageId="38" pageNumber="413">EUM</collectionCode>
);
<specimenCount id="9D79FD30A119FFD668A7FD81FF3A8060" pageId="38" pageNumber="413" type="female">1 ♀</specimenCount>
, Kumakôgen-cho, Ishizuchi-san,
<date id="FFC11079A119FFD66B86FD9BFDB28060" box="[460,559,542,561]" pageId="38" pageNumber="413" value="2021-07-13">13.vii.2021</date>
, H. Yoshitomi (
<collectionCode id="ED6EAE7CA119FFD668F1FD9BFD728060" box="[699,751,542,561]" pageId="38" pageNumber="413">EUM</collectionCode>
).
<collectingRegion id="49BBF85BA119FFD66ADDFDBCFE9D801D" box="[151,256,569,588]" country="Japan" name="Kochi" pageId="38" pageNumber="413">Kochi Pref.</collectingRegion>
: 1 J, Ôtoyo-cho, Mt. Kajigamori,
<date id="FFC11079A119FFD6687FFDBCFD03801D" box="[565,670,569,588]" pageId="38" pageNumber="413" value="1959-08-20">20.viii.1959</date>
, K. Morimoto (
<collectionCode id="ED6EAE7CA119FFD66A84FDD1FE918036" box="[206,268,596,615]" pageId="38" pageNumber="413">ELKU</collectionCode>
).
<emphasis id="B90BEAABA119FFD66B53FDD1FEF68036" bold="true" box="[281,363,596,615]" pageId="38" pageNumber="413">KඒඎඌHඎ:</emphasis>
<collectingRegion id="49BBF85BA119FFD66B3AFDD1FE708036" box="[368,493,596,615]" country="Japan" name="Fukuoka" pageId="38" pageNumber="413">Fukuoka Pref.</collectingRegion>
: 2 JJ
<specimenCount id="9D79FD30A119FFD6686DFDD1FDC88037" box="[551,597,589,615]" pageId="38" pageNumber="413" type="female">2 ♀♀</specimenCount>
, Soeda-machi, Mt. Hikosan,
<date id="FFC11079A119FFD669C5FF41FB9A8286" box="[911,1031,196,215]" pageId="38" pageNumber="413" value="1997-09-08" valueMax="1997-09-11" valueMin="1997-09-08">8.–11.ix.1997</date>
, T.
<collectingRegion id="49BBF85BA119FFD66E63FF41FBEB8286" box="[1065,1142,196,215]" country="Japan" name="Ishikawa" pageId="38" pageNumber="413">Ishikawa</collectingRegion>
(
<collectionCode id="ED6EAE7CA119FFD66ECEFF41FB598286" box="[1156,1220,196,215]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="38" pageNumber="413" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A119FFD66E9EFF41FB6B8287" box="[1236,1270,189,215]" pageId="38" pageNumber="413" type="female">1 ♀</specimenCount>
, same locality and date, T. Shimada (
<collectionCode id="ED6EAE7CA119FFD6699CFF5BFB8B82A0" box="[982,1046,222,241]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="38" pageNumber="413" type="Herbarium">TKPM</collectionCode>
).
<collectingRegion id="49BBF85BA119FFD66E6FFF5BFBE182A0" box="[1061,1148,222,241]" country="Japan" name="Oita" pageId="38" pageNumber="413">Oita Pref.</collectingRegion>
:
<specimenCount id="9D79FD30A119FFD66EC9FF5BFB3E82A0" box="[1155,1187,216,241]" pageId="38" pageNumber="413" type="female">1 ♀</specimenCount>
, Yufu-shi, Syônai-cho, Oike,
<date id="FFC11079A119FFD66976FF7CFC00835D" box="[828,925,249,268]" pageId="38" pageNumber="413" value="1995-07-27">27.vii.1995</date>
,Y.Nakatani (
<collectionCode id="ED6EAE7CA119FFD66E45FF7CFBD3835D" box="[1039,1102,249,268]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="38" pageNumber="413" type="Herbarium">TKPM</collectionCode>
);
<specimenCount id="9D79FD30A119FFD66E11FF7CFBEA835A" box="[1115,1143,242,268]" pageId="38" pageNumber="413" type="female">1♀</specimenCount>
, Mt.Kujyû,
<date id="FFC11079A119FFD66EABFF7CFAAC835D" box="[1249,1329,249,268]" pageId="38" pageNumber="413" value="1959-04-09">9.iv.1959</date>
,Y. Miyatake (
<collectionCode id="ED6EAE7CA119FFD66908FE91FC158376" box="[834,904,276,295]" country="Japan" name="Osaka Museum of Natural History" pageId="38" pageNumber="413">OMNH</collectionCode>
).
<collectingRegion id="49BBF85BA119FFD669DFFE91FB8A8376" box="[917,1047,276,295]" country="Japan" name="Miyazaki" pageId="38" pageNumber="413">Miyazaki Pref.</collectingRegion>
:
<specimenCount id="9D79FD30A119FFD66E57FE91FBA68377" box="[1053,1083,269,295]" pageId="38" pageNumber="413" type="female">1 ♀</specimenCount>
, Gokase-cho, Mt. Shiraiwayama,
<date id="FFC11079A119FFD66F16FE91FC198310" pageId="38" pageNumber="413" value="2016-07-22" valueMax="2016-07-24" valueMin="2016-07-22">22.–24. vii.2016</date>
, T. Yoshida (
<collectionCode id="ED6EAE7CA119FFD669B8FEABFBAF8310" box="[1010,1074,302,321]" country="Japan" lsid="urn:lsid:biocol.org:col:15119" name="Tokushima Prefectural Museum" pageId="38" pageNumber="413" type="Herbarium">TKPM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DF006631A11BFFD46ADCF871FBBC8A73" ID-DOI="http://doi.org/10.5281/zenodo.5821017" ID-Zenodo-Dep="5821017" httpUri="https://zenodo.org/record/5821017/files/figure.png" pageId="36" pageNumber="411" startId="36.[150,181,2036,2055]" targetBox="[187,1400,725,1997]" targetPageId="36">
<paragraph id="8BC036B9A11BFFD46ADCF871FBBC8A73" blockId="36.[150,1436,2036,2082]" pageId="36" pageNumber="411">
Fig. 19. Distribution of
<taxonomicName id="4C7F4D3AA11BFFD46B20F871FD4C8A56" authority="Kerzhner, 1979" authorityName="Kerzhner" authorityYear="1979" box="[362,721,2036,2055]" class="Insecta" family="Anthocoridae" genus="Acompocoris" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="411" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="B90BEAABA11BFFD46B20F871FDD98A56" box="[362,580,2036,2055]" italics="true" pageId="36" pageNumber="411">Acompocoris brevirostris</emphasis>
<bibRefCitation id="EFEE4B48A11BFFD46801F871FD4C8A56" author="KERZHNER I. M." box="[587,721,2036,2055]" pageId="36" pageNumber="411" pagination="14 - 65" refId="ref53445" refString="KERZHNER I. M. 1979: Novye i poluzhestkokrylye (Heteroptera) s Dal'nego Vostoka SSSR. (New Heteroptera from the Far East of the USSR). Trudy Zoologicheskogo Instituta Akademiya Nauk SSSR 81: 14 - 65 (in Russian, English title)." type="journal article" year="1979">Kerzhner, 1979</bibRefCitation>
</taxonomicName>
(circles) and
<taxonomicName id="4C7F4D3AA11BFFD46906F871FB328A56" authority="(J. Sahlberg, 1878)" baseAuthorityName="J. Sahlberg" baseAuthorityYear="1878" box="[844,1199,2036,2055]" class="Insecta" family="Anthocoridae" genus="Tetraphleps" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="411" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B90BEAABA11BFFD46906F871FB9F8A56" box="[844,1026,2036,2055]" italics="true" pageId="36" pageNumber="411">Tetraphleps aterrima</emphasis>
(J.
<bibRefCitation id="EFEE4B48A11BFFD46E6FF871FB348A56" author="SAHLBERG J." box="[1061,1193,2036,2055]" pageId="36" pageNumber="411" pagination="1 - 39" refId="ref59232" refString="SAHLBERG J. 1878: Bidrag till Nordvestra Sibiriens lnsektfauna. Hemiptera Heteroptera insamlade under expeditionema till Obi och Jenesej 1876 och 1877. Kungliga Svenska Vetenskapsakademiens Handlingar (Ny Foldj) 16 (4): 1 - 39." type="journal article" year="1878">Sahlberg, 1878</bibRefCitation>
)
</taxonomicName>
(triangles) in Japan. Filled symbols represent records based on specimens examined by us, open symbols represent literature records.
</paragraph>
</caption>
<caption id="DF006631A11AFFD56ADFF877FC9A8A71" ID-DOI="http://doi.org/10.5281/zenodo.5821020" ID-Zenodo-Dep="5821020" httpUri="https://zenodo.org/record/5821020/files/figure.png" pageId="37" pageNumber="412" startId="37.[149,180,2034,2053]" targetBox="[166,1422,674,1990]" targetPageId="37">
<paragraph id="8BC036B9A11AFFD56ADFF877FC9A8A71" blockId="37.[149,1437,2034,2080]" pageId="37" pageNumber="412">
Fig. 20. Distribution of
<taxonomicName id="4C7F4D3AA11AFFD56B28F877FDC18A54" authority="Hiura, 1959" authorityName="Hiura" authorityYear="1959" box="[354,604,2034,2053]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="412" phylum="Arthropoda" rank="species" species="chibi">
<emphasis id="B90BEAABA11AFFD56B28F877FE6D8A54" box="[354,496,2034,2053]" italics="true" pageId="37" pageNumber="412">Anthocoris chibi</emphasis>
<bibRefCitation id="EFEE4B48A11AFFD56BBFF877FDC18A54" author="HIURA I." box="[501,604,2034,2053]" pageId="37" pageNumber="412" pagination="1 - 10" refId="ref51208" refString="HIURA I. 1959: Contribution to the knowledge of Anthocoridae from Japan and its adjacent territories (Hemiptera-Heteroptera) 1. Bulletin of the Osaka Museum of Natural History 11: 1 - 10, plates 5." type="journal article" year="1959">Hiura, 1959</bibRefCitation>
</taxonomicName>
(circles) and
<taxonomicName id="4C7F4D3AA11AFFD5689AF877FC3B8A54" authority="Reuter, 1884" authorityName="Reuter" authorityYear="1884" box="[720,934,2034,2053]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="412" phylum="Arthropoda" rank="species" species="confusus">
<emphasis id="B90BEAABA11AFFD5689AF877FCAE8A54" box="[720,819,2034,2053]" italics="true" pageId="37" pageNumber="412">A. confusus</emphasis>
<bibRefCitation id="EFEE4B48A11AFFD56972F877FC3B8A54" author="REUTER O. M." box="[824,934,2034,2053]" pageId="37" pageNumber="412" pagination="555 - 758" refId="ref58799" refString="REUTER O. M. 1884: Monographia Anthocoridarum Orbis terrestris. Helsingforsiae, 204 pp. [Also published in Acta Societatis Scientiarum Fennicae 14 [1885]: 555 - 758.]" type="book chapter" year="1884">Reuter, 1884</bibRefCitation>
</taxonomicName>
(squares) in Japan. Filled symbols represent records based on specimens examined by us, open symbols represent literature records.
</paragraph>
</caption>
<subSubSection id="C3656532A119FFD76976FEDFFD888044" lastPageId="39" lastPageNumber="414" pageId="38" pageNumber="413" type="diagnosis">
<paragraph id="8BC036B9A119FFD76976FEDFFD888044" blockId="38.[828,1437,346,593]" lastBlockId="39.[151,760,190,533]" lastPageId="39" lastPageNumber="414" pageId="38" pageNumber="413">
<emphasis id="B90BEAABA119FFD66976FEDFFBD88320" bold="true" box="[828,1093,346,369]" pageId="38" pageNumber="413">Differential diagnosis.</emphasis>
Recognized by the following combination of characters: head and pronotum uniformly black to blackish brown (
<figureCitation id="13442A3CA119FFD66E1BFE1FFB5183E0" box="[1105,1228,410,433]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="38" pageNumber="413">Figs 8E–H</figureCitation>
); clavus with basal and apical portions blackish brown, inner margin narrowly darkened; endocorium with basal and apical portions mostly blackish brown; embolium with basal and apical portions blackish brown; median portion of clavus and outer part of median portion of endocorium and remaining area of embolium whitish and subhyaline; area in front of cuneal fracture whitish or subhyaline; outer (posterior) margin of ostiolar peritreme not close to supracoxal area (
<figureCitation id="13442A3CA118FFD768DFFF7BFD768344" box="[661,747,254,277]" captionStart="Fig" captionStartId="18.[151,182,1972,1991]" captionTargetBox="[204,1383,189,1959]" captionTargetId="figure-13@18.[204,1383,189,1962]" captionTargetPageId="18" captionText="Fig. 9. Scanning electron micrographs of ostiolar peritreme and evaporatorium of Anthocoris spp, left lateroventral view. A – A. chibi Hiura, 1959, female; B – A. confusus Reuter, 1884, male (from India); C – A. japonicus Poppius, 1909, female; D – A. kalopanacis Kerzhner, 1977, female; E – A. takahashii Hiura, 1959, female; F – A. miyamotoi Hiura, 1959, female; G – A. venustus sp. nov., paratype, male. Abbreviations: eva ‒ evaporatorium; mf ‒ median furrow; op ‒ ostiolar peritreme; sca ‒ supracoxal area." figureDoi="http://doi.org/10.5281/zenodo.5820987" httpUri="https://zenodo.org/record/5820987/files/figure.png" pageId="39" pageNumber="414">Fig. 9G</figureCitation>
); paramere (
<figureCitation id="13442A3CA118FFD76B43FE9BFE138364" box="[265,398,286,309]" captionStart="Fig" captionStartId="20.[150,181,2004,2023]" captionTargetBox="[151,1433,1025,1988]" captionTargetId="figure-684@20.[151,1436,1025,1991]" captionTargetPageId="20" captionText="Fig. 11. Scanning electron micrographs of male genitalia of Anthocoris spp. A–B – A. miyamotoi Hiura, 1959; C–D – A. venustus sp. nov., paratype. A, C – pygophore with paramere, dorsal (A) and lateral (C) views; B, D – paramere, dorsal (B) and lateral (D) views. Abbreviations: prm ‒ paramere; pyg ‒ pygophore." figureDoi="http://doi.org/10.5281/zenodo.5820997" httpUri="https://zenodo.org/record/5820997/files/figure.png" pageId="39" pageNumber="414">Figs 11C–D</figureCitation>
,
<figureCitation id="13442A3CA118FFD76BD3FE9BFE7F8364" box="[409,482,286,309]" captionStart="Fig" captionStartId="26.[151,182,887,906]" captionTargetBox="[161,1401,197,863]" captionTargetId="figure-13@26.[151,1436,186,876]" captionTargetPageId="26" captionText="Fig. 14. Male genitalia of Anthocoris spp.A–C – A. miyamotoi Hiura, 1959; D–F – A. venustus sp. nov., paratype.A, D – pygophore with paramere (ejaculatory bulb omitted), dorsal view; B–C, E–F – paramere, two different views.Abbreviations: prm ‒ paramere; pyg ‒ pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821005" httpUri="https://zenodo.org/record/5821005/files/figure.png" pageId="39" pageNumber="414">14E–F</figureCitation>
) slender and very slightly curved, with longitudinal groove in form of shallow depression; copulatory tube (
<figureCitation id="13442A3CA118FFD76BC8FEDBFE7C8324" box="[386,481,350,373]" captionStart="Fig" captionStartId="30.[151,182,1991,2010]" captionTargetBox="[211,1367,202,1897]" captionTargetId="figure-13@30.[172,1416,189,1909]" captionTargetPageId="30" captionText="Fig. 15. Female genitalia of Anthocoris spp., dorsal view.A – A. chibi Hiura, 1959; B – A. confusus Reuter, 1884; C – A. japonicus Poppius, 1909; D – A. nemoralis (Fabricius,1794); E – A.takahashii Hiura, 1959; F – A. miyamotoi Hiura, 1959; G – A. venustus sp. nov., paratype.Abbreviations: ct ‒ copulatory tube; ism ‒ intersegmental membrane; sp ‒ sperm pouch; t ‒ trunk of conductive tissue (C–D, trunk of conductive tissue dissolved). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821009" httpUri="https://zenodo.org/record/5821009/files/figure.png" pageId="39" pageNumber="414">Fig. 15G</figureCitation>
) approximately
<quantity id="4C879B5CA118FFD768C4FEDBFD408324" box="[654,733,350,373]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="39" pageNumber="414" unit="mm" value="0.8">0.8 mm</quantity>
in length, basally thickened, with a few twists. Closely related to
<taxonomicName id="4C7F4D3AA118FFD76AFBFE1BFEA383E4" authorityName="Hiura" authorityYear="1959" box="[177,318,414,437]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="414" phylum="Arthropoda" rank="species" species="miyamotoi">
<emphasis id="B90BEAABA118FFD76AFBFE1BFEA383E4" box="[177,318,414,437]" italics="true" pageId="39" pageNumber="414">A. miyamotoi</emphasis>
</taxonomicName>
but distinguished from that species by the uniformly blackish head and pronotum, slender and very slightly curved paramere, and copulatory tube longer and basally thickened with a few twists.
</paragraph>
</subSubSection>
<caption id="DF006631A119FFD66AD3F877FB378A71" ID-DOI="http://doi.org/10.5281/zenodo.5821022" ID-Zenodo-Dep="5821022" httpUri="https://zenodo.org/record/5821022/files/figure.png" pageId="38" pageNumber="413" startId="38.[153,184,2034,2053]" targetBox="[168,1419,673,1984]" targetPageId="38">
<paragraph id="8BC036B9A119FFD66AD3F877FB378A71" blockId="38.[151,1436,2034,2080]" pageId="38" pageNumber="413">
Fig. 21. Distribution of
<taxonomicName id="4C7F4D3AA119FFD66B14F877FD1F8A54" authority="Hiura, 1959" authorityName="Hiura" authorityYear="1959" box="[350,642,2034,2053]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="413" phylum="Arthropoda" rank="species" species="takahashii">
<emphasis id="B90BEAABA119FFD66B14F877FD858A54" box="[350,536,2034,2053]" italics="true" pageId="38" pageNumber="413">Anthocoris takahashii</emphasis>
<bibRefCitation id="EFEE4B48A119FFD66857F877FD1F8A54" author="HIURA I." box="[541,642,2034,2053]" pageId="38" pageNumber="413" pagination="1 - 10" refId="ref51208" refString="HIURA I. 1959: Contribution to the knowledge of Anthocoridae from Japan and its adjacent territories (Hemiptera-Heteroptera) 1. Bulletin of the Osaka Museum of Natural History 11: 1 - 10, plates 5." type="journal article" year="1959">Hiura, 1959</bibRefCitation>
</taxonomicName>
(circles),
<taxonomicName id="4C7F4D3AA119FFD6689FF877FC208A54" authority="Poppius, 1909" authorityName="Poppius" authorityYear="1909" box="[725,957,2034,2053]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="413" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90BEAABA119FFD6689FF877FCA28A54" box="[725,831,2034,2053]" italics="true" pageId="38" pageNumber="413">A. japonicus</emphasis>
<bibRefCitation id="EFEE4B48A119FFD66909F877FC208A54" author="POPPIUS B." box="[835,957,2034,2053]" pageId="38" pageNumber="413" pagination="1 - 43" refId="ref58375" refString="POPPIUS B. 1909: Beitrage zur Kenntnis der Anthocoriden. Acta Societatis Scientiarum Fennicae 37 (9): 1 - 43." type="journal article" year="1909">Poppius, 1909</bibRefCitation>
</taxonomicName>
(triangles), and
<taxonomicName id="4C7F4D3AA119FFD66E0FF877FAD78A54" authority="Kerzhner, 1977" authorityName="Kerzhner" authorityYear="1977" box="[1093,1354,2034,2053]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="413" phylum="Arthropoda" rank="species" species="kalopanacis">
<emphasis id="B90BEAABA119FFD66E0FF877FB5E8A54" box="[1093,1219,2034,2053]" italics="true" pageId="38" pageNumber="413">A. kalopanacis</emphasis>
<bibRefCitation id="EFEE4B48A119FFD66E8DF877FAD78A54" author="KERZHNER I. M." box="[1223,1354,2034,2053]" pageId="38" pageNumber="413" pagination="6 - 35" refId="ref53334" refString="KERZHNER I. M. 1977: Novye i maloizvestnye vidy poluzhestkokrylykh (Heteroptera) s Dal'nego Vostoka SSSR. (New and little-known species of Heteroptera from the Far East of the USSR). Trudy Zoologicheskogo Instituta Akademiya Nauk SSSR 62 [1976]: 6 - 35 (in Russian, English title)." type="journal article" year="1977">Kerzhner, 1977</bibRefCitation>
</taxonomicName>
(squares) in Japan. Filled symbols represent records based on specimens examined by us, open symbols represent literature records.
</paragraph>
</caption>
<subSubSection id="C3656532A118FFD86976FF3BFAB384E4" lastPageId="40" lastPageNumber="415" pageId="39" pageNumber="414" type="description">
<paragraph id="8BC036B9A118FFD86976FF3BFD358024" blockId="39.[828,1437,190,533]" lastBlockId="40.[151,760,190,2069]" lastPageId="40" lastPageNumber="415" pageId="39" pageNumber="414">
<emphasis id="B90BEAABA118FFD76976FF3BFBDC8284" bold="true" box="[828,1089,190,213]" pageId="39" pageNumber="414">
Description.
<emphasis id="B90BEAABA118FFD7698DFF3BFBDC8284" bold="true" box="[967,1089,190,213]" italics="true" pageId="39" pageNumber="414">Coloration.</emphasis>
</emphasis>
Body (
<figureCitation id="13442A3CA118FFD76EC1FF3BFB608284" box="[1163,1277,190,213]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="39" pageNumber="414">Figs 8E–H</figureCitation>
,
<figureCitation id="13442A3CA118FFD76F4FFF3BFAB28284" box="[1285,1327,190,213]" captionStart="Fig" captionStartId="32.[151,182,1628,1647]" captionTargetBox="[151,1436,193,1607]" captionTargetId="figure-13@32.[151,1437,193,1607]" captionTargetPageId="32" captionText="Fig. 16. Habitus images of living individuals of Japanese Anthocorini spp. A – Acompocoris brevirostris Kerzhner, 1979, male; B – same, female; C – Tetraphleps aterrima (J. Sahlberg, 1878); D – Anthocoris confusus Reuter, 1884; E – A. takahashii Hiura, 1959; F – A. venustus sp. nov. (courtesy of M. Takai)." figureDoi="http://doi.org/10.5281/zenodo.5821011" httpUri="https://zenodo.org/record/5821011/files/figure.png" pageId="39" pageNumber="414">16F</figureCitation>
) generally black to blackish brown. Head and pronotum (
<figureCitation id="13442A3CA118FFD76F75FF5BFCEC8344" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="39" pageNumber="414">Figs 8E, G–H</figureCitation>
) blackish brown, sometimes vertex and callus tinged with reddish brown; eyes reddish black; margin of ocellus red to reddish brown. Antennal segment I dark brown; segment II yellowish brown, dark brown on base and apical 1/4 to 1/3; segment III yellowish brown, dark brown on apical half; segment IV dark brown. Labium (
<figureCitation id="13442A3CA118FFD76E9DFE1BFAB983E4" box="[1239,1316,414,437]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="39" pageNumber="414">Fig. 8F</figureCitation>
) uniformly black to blackish brown. Scutellum overall black to blackish brown. Clavus with basal portion and apex blackish brown, inner margin narrowly darkened (
<figureCitation id="13442A3CA118FFD76EB6FE7BFA0D8044" box="[1276,1424,510,533]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="39" pageNumber="414">Figs 8E,G–H</figureCitation>
); endocorium with basal and apical portions mostly blackish brown (
<figureCitation id="13442A3CA117FFD86AA7FF5BFE1F82A4" box="[237,386,222,245]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="40" pageNumber="415">Figs 8E,G–H</figureCitation>
); embolium with basal and apical portions blackish brown (
<figureCitation id="13442A3CA117FFD86BE5FF7BFDD98344" box="[431,580,254,277]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="40" pageNumber="415">Figs 8E,G–H</figureCitation>
); cuneus wholly blackish brown; median portion of clavus and outer part of median portion of endocorium and remaining area of embolium whitish and subhyaline (
<figureCitation id="13442A3CA117FFD86846FEDBFD038324" box="[524,670,350,373]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="40" pageNumber="415">Figs 8E,G–H</figureCitation>
); area in front of cuneal fracture whitish or subhyaline; membrane smoky dark brown, with basal and innermost portions and area behind apex of cuneus off- white (
<figureCitation id="13442A3CA117FFD8686EFE3BFD298384" box="[548,692,446,469]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="40" pageNumber="415">Figs 8E,G–H</figureCitation>
). Legs dark reddish brown; trochanters tinged with yellowish brown; apical half or apical 2/3 of protibiae sometimes tinged with yellowish brown. Venter of thorax uniformly blackish brown.Abdomen black to blackish brown. Pygophore blackish brown with pale yellow paramere.
</paragraph>
<caption id="DF006631A118FFD76AD3F877FC9A8A71" ID-DOI="http://doi.org/10.5281/zenodo.5821024" ID-Zenodo-Dep="5821024" httpUri="https://zenodo.org/record/5821024/files/figure.png" pageId="39" pageNumber="414" startId="39.[153,184,2034,2053]" targetBox="[165,1422,635,1952]" targetPageId="39">
<paragraph id="8BC036B9A118FFD76AD3F877FC9A8A71" blockId="39.[151,1437,2034,2080]" pageId="39" pageNumber="414">
Fig. 22. Distribution of
<taxonomicName id="4C7F4D3AA118FFD76B2AF877FD1A8A54" authority="Hiura, 1959" authorityName="Hiura" authorityYear="1959" box="[352,647,2034,2053]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="414" phylum="Arthropoda" rank="species" species="miyamotoi">
<emphasis id="B90BEAABA118FFD76B2AF877FD818A54" box="[352,540,2034,2053]" italics="true" pageId="39" pageNumber="414">Anthocoris miyamotoi</emphasis>
<bibRefCitation id="EFEE4B48A118FFD7686BF877FD1A8A54" author="HIURA I." box="[545,647,2034,2053]" pageId="39" pageNumber="414" pagination="1 - 10" refId="ref51208" refString="HIURA I. 1959: Contribution to the knowledge of Anthocoridae from Japan and its adjacent territories (Hemiptera-Heteroptera) 1. Bulletin of the Osaka Museum of Natural History 11: 1 - 10, plates 5." type="journal article" year="1959">Hiura, 1959</bibRefCitation>
</taxonomicName>
(circles) and
<taxonomicName id="4C7F4D3AA118FFD768B0F877FCC78A54" authority="Yamada &amp; Yasunaga, 2021" authorityName="Yamada &amp; Yasunaga" authorityYear="2021" box="[762,858,2034,2053]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="414" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="B90BEAABA118FFD768B0F877FCC78A54" box="[762,858,2034,2053]" italics="true" pageId="39" pageNumber="414">A. venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857D0A118FFD7692AF877FC3F8A54" box="[864,930,2034,2053]" pageId="39" pageNumber="414" rank="species">sp. nov.</taxonomicNameLabel>
(triangles) in Japan. Filled symbols represent records based on specimens examined by us, open symbols represent literature records.
</paragraph>
</caption>
<paragraph id="8BC036B9A117FFD86AFFFDFBFD6587E4" blockId="40.[151,760,190,2069]" pageId="40" pageNumber="415">
<emphasis id="B90BEAABA117FFD86AFFFDFBFEB980C4" bold="true" box="[181,292,638,661]" italics="true" pageId="40" pageNumber="415">Structure.</emphasis>
Body (
<figureCitation id="13442A3CA117FFD86B3CFDFBFE7080C4" box="[374,493,638,661]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="40" pageNumber="415">Figs 8E–H</figureCitation>
) elongate, shiny on dorsal and ventral surfaces, sparsely covered with suberect, simple, yellowish setae. Head (
<figureCitation id="13442A3CA117FFD86BB9FD3BFDF18084" box="[499,620,702,725]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="40" pageNumber="415">Figs 8E–H</figureCitation>
) cylindrical, impunctate, approximately as long as width across eyes; three pairs of long, erect trichobothria on the dorsal surface of head, one on anterior clypeus, one in front of eye, and one behind ocellus; anteocular region as long as length of eye in dorsal view in male, slightly longer than length of eye in female; vertex approximately twice as wide as eye in dorsal view in male, approximately 2.5 times as wide as eye in female; postocular region constricted, demarcated by transverse shallow furrow; neck long, smooth, highly polished; eye oblong, not exceeding level of dorsal surface of head in lateral view but exceeding level of ventral surface of head. Antennae (
<figureCitation id="13442A3CA117FFD86BDFFBBBFD978604" box="[405,522,1086,1109]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="40" pageNumber="415">Figs 8E–H</figureCitation>
) densely covered with short, decumbent, yellowish setae interspersed with long, erect setae, of which the longest are as long as or slightly shorter than width of corresponding segment; segment I just reaching apex of head, sparsely covered with short setae; segment II gradually thickened toward apex, slightly thicker in the male than in the female, approximately as long as head width across eyes; segment III approximately 0.7 times as long as segment II; segment IV longer than segment III. Labium (
<figureCitation id="13442A3CA117FFD86B33FADBFE5B8724" box="[377,454,1374,1397]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="40" pageNumber="415">Fig. 8F</figureCitation>
) just reaching procoxae; segment III approximately 4.0–5.0 times as long as segment II; segment IV approximately half as long as segment III.
</paragraph>
<paragraph id="8BC036B9A117FFD86AFFFA3BFB9080E4" blockId="40.[151,760,190,2069]" lastBlockId="40.[828,1437,190,2069]" pageId="40" pageNumber="415">
Pronotum (
<figureCitation id="13442A3CA117FFD86B7FFA3BFE508784" box="[309,461,1470,1493]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="40" pageNumber="415">Figs 8E,G–H</figureCitation>
) with pairs of long, erect trichobothria on midline of collar and on each anterior and posterior angle; anterior margin very slightly curved, approximately 0.7 times as long as mesal length; lateral margin very slightly curved; lateral carina slightly expanded anteriorly, more obscure posteriad; posterior margin concave inwardly, approximately 2.5 times as wide as anterior margin; collar long, transversely rugose, with scattered long setae; callus extremely swollen, polished, impunctate, sides strongly bulging, demarcated posteriorly by deep transverse impression; deep impression with short parallel carinae and coarse punctures; posterior lobe behind callus densely covered with minute punctures. Scutellum (
<figureCitation id="13442A3CA117FFD86AD5F8DBFEB68524" box="[159,299,1886,1909]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="40" pageNumber="415">Figs 8E,G–H</figureCitation>
) smooth, sub-equilateral, wider at base than long, depressed through middle. Hemelytra (
<figureCitation id="13442A3CA117FFD86833F8FBFD6D85C4" box="[633,752,1918,1941]" captionStart="Fig" captionStartId="16.[151,182,1562,1581]" captionTargetBox="[206,1412,195,1511]" captionTargetId="figure-13@16.[157,1440,183,1533]" captionTargetPageId="16" captionText="Fig. 8. Habitus of Anthocoris spp., male, dorsal (A, C–E, G–H) and lateral (B, F) views. A–B – A. miyamotoi Hiura, 1959, holotype (Nakanoshima Is., Tokara Isls.); C – same (Wakayama, Honshu); D – same (Kagoshima, Kyushu); E–F – A. venustus sp. nov., holotype (Hokkaido); G – same, paratype (Tochigi, Honshu); H – same, paratype (Nara, Honshu). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5820985" httpUri="https://zenodo.org/record/5820985/files/figure.png" pageId="40" pageNumber="415">Figs 8E–H</figureCitation>
) sparsely covered with long, erect, simple, yellowish setae, and with minute punctures; costal margin slightly sinuate; endocorium approximately 1.6 times as wide as maximum width of embolium; cuneal margin approximately 0.7 times as long as embolial margin; membrane with three veins, outermost vein distinct and slightly curved, inner two veins obscure and very slightly curved. Ostiolar peritreme (
<figureCitation id="13442A3CA117FFD869F9FE9BFB948364" box="[947,1033,286,309]" captionStart="Fig" captionStartId="18.[151,182,1972,1991]" captionTargetBox="[204,1383,189,1959]" captionTargetId="figure-13@18.[204,1383,189,1962]" captionTargetPageId="18" captionText="Fig. 9. Scanning electron micrographs of ostiolar peritreme and evaporatorium of Anthocoris spp, left lateroventral view. A – A. chibi Hiura, 1959, female; B – A. confusus Reuter, 1884, male (from India); C – A. japonicus Poppius, 1909, female; D – A. kalopanacis Kerzhner, 1977, female; E – A. takahashii Hiura, 1959, female; F – A. miyamotoi Hiura, 1959, female; G – A. venustus sp. nov., paratype, male. Abbreviations: eva ‒ evaporatorium; mf ‒ median furrow; op ‒ ostiolar peritreme; sca ‒ supracoxal area." figureDoi="http://doi.org/10.5281/zenodo.5820987" httpUri="https://zenodo.org/record/5820987/files/figure.png" pageId="40" pageNumber="415">Fig. 9G</figureCitation>
) wide, gently curved anteriorly, gradually narrowing toward apex and then continued by a fine carina that reaches the anterior margin of metapleura; outer (posterior) margin of ostiolar peritreme not close to supracoxal area; evaporatorium bearing a small number of long, recumbent setae. Fossula spongiosa present on apex of all tibiae, enlarged on protibiae and small on meso- and metatibiae; apical 1/4 of metatibiae bearing small spines on ventral side. Abdominal sternum III (
<figureCitation id="13442A3CA117FFD86EA0FD9BFAD18064" box="[1258,1356,542,565]" captionStart="Fig" captionStartId="22.[151,182,1996,2015]" captionTargetBox="[236,1400,657,1963]" captionTargetId="figure-13@22.[151,1436,648,1972]" captionTargetPageId="22" captionText="Fig. 12. Abdominal sterna II–III of Anthocoris spp., male, ventral view (setae omitted). A – A. chibi Hiura, 1959; B – A. confusus Reuter, 1884; C – A. japonicus Poppius, 1909; D – A. nemoralis (Fabricius, 1794); E – A. takahashii Hiura, 1959; F – A. miyamotoi Hiura, 1959; G – A. venustus sp. nov., paratype. Abbreviations: ma ‒ membranous area; st2–3 ‒ abdominal sternum II to III. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5820999" httpUri="https://zenodo.org/record/5820999/files/figure.png" pageId="40" pageNumber="415">Fig. 12G</figureCitation>
) with a pair of banana-shaped membranous areas on anterior half, membranous area somewhat widened near base; dorsal laterotergites not fused with mediotergites on abdominal segments II and III.
</paragraph>
<paragraph id="8BC036B9A117FFD86910FD3BFA8C8644" blockId="40.[828,1437,190,2069]" pageId="40" pageNumber="415">
Male genitalia (
<figureCitation id="13442A3CA117FFD86E4AFD3BFB188084" box="[1024,1157,702,725]" captionStart="Fig" captionStartId="20.[150,181,2004,2023]" captionTargetBox="[151,1433,1025,1988]" captionTargetId="figure-684@20.[151,1436,1025,1991]" captionTargetPageId="20" captionText="Fig. 11. Scanning electron micrographs of male genitalia of Anthocoris spp. A–B – A. miyamotoi Hiura, 1959; C–D – A. venustus sp. nov., paratype. A, C – pygophore with paramere, dorsal (A) and lateral (C) views; B, D – paramere, dorsal (B) and lateral (D) views. Abbreviations: prm ‒ paramere; pyg ‒ pygophore." figureDoi="http://doi.org/10.5281/zenodo.5820997" httpUri="https://zenodo.org/record/5820997/files/figure.png" pageId="40" pageNumber="415">Figs 11C–D</figureCitation>
,
<figureCitation id="13442A3CA117FFD86EC5FD3BFB408084" box="[1167,1245,702,725]" captionStart="Fig" captionStartId="26.[151,182,887,906]" captionTargetBox="[161,1401,197,863]" captionTargetId="figure-13@26.[151,1436,186,876]" captionTargetPageId="26" captionText="Fig. 14. Male genitalia of Anthocoris spp.A–C – A. miyamotoi Hiura, 1959; D–F – A. venustus sp. nov., paratype.A, D – pygophore with paramere (ejaculatory bulb omitted), dorsal view; B–C, E–F – paramere, two different views.Abbreviations: prm ‒ paramere; pyg ‒ pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821005" httpUri="https://zenodo.org/record/5821005/files/figure.png" pageId="40" pageNumber="415">14D–F</figureCitation>
): Pygophore (
<figureCitation id="13442A3CA117FFD86F3BFD3BFCF680A4" captionStart="Fig" captionStartId="26.[151,182,887,906]" captionTargetBox="[161,1401,197,863]" captionTargetId="figure-13@26.[151,1436,186,876]" captionTargetPageId="26" captionText="Fig. 14. Male genitalia of Anthocoris spp.A–C – A. miyamotoi Hiura, 1959; D–F – A. venustus sp. nov., paratype.A, D – pygophore with paramere (ejaculatory bulb omitted), dorsal view; B–C, E–F – paramere, two different views.Abbreviations: prm ‒ paramere; pyg ‒ pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821005" httpUri="https://zenodo.org/record/5821005/files/figure.png" pageId="40" pageNumber="415">Fig. 14D</figureCitation>
) rather pointed apically, asymmetrical, covered with 6–8 long, stout setae intermixed with short, suberect setae along outer margin and on posteroventral surface, of which the longest are approximately half length of pygophore; mid-dorsal surface hirsute with suberect setae; paramere (
<figureCitation id="13442A3CA117FFD86909FCFBFC3481C4" box="[835,937,894,917]" captionStart="Fig" captionStartId="20.[150,181,2004,2023]" captionTargetBox="[151,1433,1025,1988]" captionTargetId="figure-684@20.[151,1436,1025,1991]" captionTargetPageId="20" captionText="Fig. 11. Scanning electron micrographs of male genitalia of Anthocoris spp. A–B – A. miyamotoi Hiura, 1959; C–D – A. venustus sp. nov., paratype. A, C – pygophore with paramere, dorsal (A) and lateral (C) views; B, D – paramere, dorsal (B) and lateral (D) views. Abbreviations: prm ‒ paramere; pyg ‒ pygophore." figureDoi="http://doi.org/10.5281/zenodo.5820997" httpUri="https://zenodo.org/record/5820997/files/figure.png" pageId="40" pageNumber="415">Figs 11D</figureCitation>
,
<figureCitation id="13442A3CA117FFD869FFFCFBFC6381C4" box="[949,1022,894,917]" captionStart="Fig" captionStartId="26.[151,182,887,906]" captionTargetBox="[161,1401,197,863]" captionTargetId="figure-13@26.[151,1436,186,876]" captionTargetPageId="26" captionText="Fig. 14. Male genitalia of Anthocoris spp.A–C – A. miyamotoi Hiura, 1959; D–F – A. venustus sp. nov., paratype.A, D – pygophore with paramere (ejaculatory bulb omitted), dorsal view; B–C, E–F – paramere, two different views.Abbreviations: prm ‒ paramere; pyg ‒ pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821005" httpUri="https://zenodo.org/record/5821005/files/figure.png" pageId="40" pageNumber="415">14E–F</figureCitation>
) slender, very slightly curved but bent at basal 1/3, with a few very short, erect setae on sub-basal portion, the apex hooked and reaching nearly half of the left outer margin of pygophore; longitudinal groove in form of shallow depression, twisted basally.
</paragraph>
<paragraph id="8BC036B9A117FFD86910FB9BFA0086E4" blockId="40.[828,1437,190,2069]" pageId="40" pageNumber="415">
Female genitalia (
<figureCitation id="13442A3CA117FFD86E52FB9BFBE68664" box="[1048,1147,1054,1077]" captionStart="Fig" captionStartId="30.[151,182,1991,2010]" captionTargetBox="[211,1367,202,1897]" captionTargetId="figure-13@30.[172,1416,189,1909]" captionTargetPageId="30" captionText="Fig. 15. Female genitalia of Anthocoris spp., dorsal view.A – A. chibi Hiura, 1959; B – A. confusus Reuter, 1884; C – A. japonicus Poppius, 1909; D – A. nemoralis (Fabricius,1794); E – A.takahashii Hiura, 1959; F – A. miyamotoi Hiura, 1959; G – A. venustus sp. nov., paratype.Abbreviations: ct ‒ copulatory tube; ism ‒ intersegmental membrane; sp ‒ sperm pouch; t ‒ trunk of conductive tissue (C–D, trunk of conductive tissue dissolved). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5821009" httpUri="https://zenodo.org/record/5821009/files/figure.png" pageId="40" pageNumber="415">Fig. 15G</figureCitation>
): Copulatory tube fused on middle of intersegmental membrane between sterna VII and VIII, approximately
<quantity id="4C879B5CA117FFD86E5DFBDBFBF48624" box="[1047,1129,1118,1141]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="40" pageNumber="415" unit="mm" value="0.8">0.8 mm</quantity>
in length, basally thickened, gradually narrowing toward apex, with a few loops; trunk of conductive tissue not pronounced (or possibly dissolved).
</paragraph>
<paragraph id="8BC036B9A117FFD86910FB3BFAB384E4" blockId="40.[828,1437,190,2069]" pageId="40" pageNumber="415">
<emphasis id="B90BEAABA117FFD86910FB3BFC668684" bold="true" box="[858,1019,1214,1237]" italics="true" pageId="40" pageNumber="415">Measurements</emphasis>
[mm; JJ (n = 10) /
<specimenCount id="9D79FD30A117FFD86E9CFB32FB618687" box="[1238,1276,1207,1238]" pageId="40" pageNumber="415" type="female">♀♀</specimenCount>
(n = 10),
<typeStatus id="54C4881BA117FFD86F2EFB3BFCF786A4" pageId="40" pageNumber="415" type="holotype">holotype</typeStatus>
in parentheses]. Body length 3.50–4.18 (3.75) / 3.50– 4.08; head length (excl. neck) 0.47–0.50 (0.47) / 0.48–0.55; head width across eyes 0.47–0.54 (0.54) / 0.48–0.53; vertex width 0.24–0.27 (0.26) / 0.28–0.30; length of antennal segments I – 0.15–0.18 (0.17) / 0.16–0.19, II – 0.48–0.56 (0.55) / 0.47–0.53, III – 0.32–0.39 (0.36) / 0.32–0.38, and IV – 0.39–0.42 (0.41) / 0.37–0.41; length of labial segments II – 0.12–0.16 (0.12) / 0.13–0.17, III – 0.54–0.62 (0.60) / 0.58–0.65, and IV – 0.30–0.35 (0.33) / 0.32–0.38; anterior pronotal width 0.36–0.41 (0.40) / 0.39–0.45; mesal pronotal length 0.51–0.57 (0.55) / 0.55–0.62; basal pronotal width 0.92–1.13 (1.04) / 0.99–1.17; length of embolial margin 0.91–1.15 (1.03) / 0.97–1.13; length of cuneal margin 0.65–0.80 (0.70) / 0.64–0.78; maximum width across hemelytra 0.96–1.18 (1.07) / 1.05–1.26.
</paragraph>
</subSubSection>
<subSubSection id="C3656532A117FFD86976F93BFB6B8A44" pageId="40" pageNumber="415" type="etymology">
<paragraph id="8BC036B9A117FFD86976F93BFBCF8544" blockId="40.[828,1437,190,2069]" pageId="40" pageNumber="415">
<emphasis id="B90BEAABA117FFD86976F93BFC218484" bold="true" box="[828,956,1726,1749]" pageId="40" pageNumber="415">Etymology.</emphasis>
From Latin,
<taxonomicName id="4C7F4D3AA117FFD86E02F93BFB398484" authorityName="Yamada &amp; Yasunaga" authorityYear="2021" box="[1096,1188,1726,1749]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="40" pageNumber="415" phylum="Arthropoda" rank="species" species="venustus">
<emphasis id="B90BEAABA117FFD86E02F93BFB398484" box="[1096,1188,1726,1749]" italics="true" pageId="40" pageNumber="415">venustus</emphasis>
</taxonomicName>
(= beautiful, graceful), referring to the beautiful and graceful body color of the new species; an adjective.
</paragraph>
<paragraph id="8BC036B9A117FFD86976F89BFB6B8A44" blockId="40.[828,1437,190,2069]" pageId="40" pageNumber="415">
<emphasis id="B90BEAABA117FFD86976F89BFC288564" bold="true" box="[828,949,1822,1845]" pageId="40" pageNumber="415">Bionomics.</emphasis>
<taxonomicName id="4C7F4D3AA117FFD869F0F89BFB148564" authority="Yamada &amp; Yasunaga, 2021" authorityName="Yamada &amp; Yasunaga" authorityYear="2021" box="[954,1161,1822,1845]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="40" pageNumber="415" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="B90BEAABA117FFD869F0F89BFB148564" box="[954,1161,1822,1845]" italics="true" pageId="40" pageNumber="415">Anthocoris venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857D0A117FFD86EC4F89BFB408564" box="[1166,1245,1822,1845]" pageId="40" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
has been frequently collected from
<taxonomicName id="4C7F4D3AA117FFD869A4F8BBFBCF8504" box="[1006,1106,1854,1877]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="40" pageNumber="415" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEAABA117FFD869A4F8BBFBBF8504" box="[1006,1058,1854,1877]" italics="true" pageId="40" pageNumber="415">Salix</emphasis>
spp.
</taxonomicName>
in
<collectingRegion id="49BBF85BA117FFD86E25F8BBFB4B8504" box="[1135,1238,1854,1877]" country="Japan" name="Hokkaido" pageId="40" pageNumber="415">Hokkaido</collectingRegion>
and
<collectingRegion id="49BBF85BA117FFD86F4FF8BBFAC58504" box="[1285,1368,1854,1877]" country="Japan" name="Tochigi" pageId="40" pageNumber="415">Tochigi</collectingRegion>
. It was also collected from
<taxonomicName id="4C7F4D3AA117FFD86E58F8DBFA818524" authority="Blume" authorityName="Blume" box="[1042,1308,1886,1909]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="40" pageNumber="415" phylum="Tracheophyta" rank="species" species="crispula">
<emphasis id="B90BEAABA117FFD86E58F8DBFB578524" box="[1042,1226,1886,1909]" italics="true" pageId="40" pageNumber="415">Quercus crispula</emphasis>
Blume
</taxonomicName>
,
<taxonomicName id="4C7F4D3AA117FFD86F6DF8DBFCC185C4" class="Magnoliopsida" family="Araliaceae" genus="Kalopanax" kingdom="Plantae" order="Apiales" pageId="40" pageNumber="415" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B90BEAABA117FFD86F6DF8DBFA018524" box="[1319,1436,1886,1909]" italics="true" pageId="40" pageNumber="415">Kalopanax</emphasis>
sp.
</taxonomicName>
, and
<taxonomicName id="4C7F4D3AA117FFD869D8F8FBFBAC85C4" box="[914,1073,1918,1941]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="40" pageNumber="415" phylum="Tracheophyta" rank="species" species="japonica">
<emphasis id="B90BEAABA117FFD869D8F8FBFBAC85C4" box="[914,1073,1918,1941]" italics="true" pageId="40" pageNumber="415">Alnus japonica</emphasis>
</taxonomicName>
, and sometimes from the herbaceous plant
<taxonomicName id="4C7F4D3AA117FFD869EAF81BFA0085E4" authority="(F. Schmidt) Ronse Decr." authorityName="Ronse Decr." baseAuthorityName="F. Schmidt" box="[928,1437,1950,1973]" class="Magnoliopsida" family="Polygonaceae" genus="Fallopia" kingdom="Plantae" order="Caryophyllales" pageId="40" pageNumber="415" phylum="Tracheophyta" rank="species" species="sachalinensis">
<emphasis id="B90BEAABA117FFD869EAF81BFB1285E4" box="[928,1167,1950,1973]" italics="true" pageId="40" pageNumber="415">Fallopia sachalinensis</emphasis>
(F. Schmidt) Ronse Decr.
</taxonomicName>
in
<collectingRegion id="49BBF85BA117FFD86910F83BFC5A8584" box="[858,967,1982,2005]" country="Japan" name="Hokkaido" pageId="40" pageNumber="415">Hokkaido</collectingRegion>
. This species was collected from
<taxonomicName id="4C7F4D3AA117FFD86F77F83BFC3E85A4" authorityName="Poppius" authorityYear="1909" class="Magnoliopsida" family="Euphorbiaceae" genus="Mallotus" kingdom="Plantae" order="Malpighiales" pageId="40" pageNumber="415" phylum="Tracheophyta" rank="species" species="japonicus">
<emphasis id="B90BEAABA117FFD86F77F83BFC3E85A4" italics="true" pageId="40" pageNumber="415">Mallotus japonicus</emphasis>
</taxonomicName>
in
<collectingRegion id="49BBF85BA117FFD8698AF85BFBA885A4" box="[960,1077,2014,2037]" country="Japan" name="Tokushima" pageId="40" pageNumber="415">Tokushima</collectingRegion>
and from
<taxonomicName id="4C7F4D3AA117FFD86ED4F85BFA0085A4" authority="Murray" authorityName="Murray" authorityYear="1784" box="[1182,1437,2014,2037]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="40" pageNumber="415" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis id="B90BEAABA117FFD86ED4F85BFADA85A4" box="[1182,1351,2014,2037]" italics="true" pageId="40" pageNumber="415">Quercus serrata</emphasis>
Murray
</taxonomicName>
at Mt. Hikosan,
<collectingRegion id="49BBF85BA117FFD869A2F87BFBD88A44" box="[1000,1093,2046,2069]" country="Japan" name="Fukuoka" pageId="40" pageNumber="415">Fukuoka</collectingRegion>
(TൺKൾඇඈ 1998).
</paragraph>
</subSubSection>
<subSubSection id="C3656532A116FFD96ADDFF3BFDFF8044" pageId="41" pageNumber="416" type="distribution">
<paragraph id="8BC036B9A116FFD96ADDFF3BFDFF8044" blockId="41.[151,760,190,533]" pageId="41" pageNumber="416">
<emphasis id="B90BEAABA116FFD96ADDFF3BFEB08284" bold="true" box="[151,301,190,213]" pageId="41" pageNumber="416">Distribution.</emphasis>
<collectingCountry id="F3687629A116FFD96B7EFF3BFEEB8284" box="[308,374,190,213]" name="Japan" pageId="41" pageNumber="416">Japan</collectingCountry>
:
<collectingRegion id="49BBF85BA116FFD96BCAFF3BFE738284" box="[384,494,190,213]" country="Japan" name="Hokkaido" pageId="41" pageNumber="416">Hokkaido</collectingRegion>
, Rebun-tô Is., Rishiri-tô Is.; Honshu:
<collectingRegion id="49BBF85BA116FFD96B51FF5BFEEC82A4" box="[283,369,222,245]" country="Japan" name="Aomori" pageId="41" pageNumber="416">Aomori</collectingRegion>
,
<collectingRegion id="49BBF85BA116FFD96B30FF5BFE2582A4" box="[378,440,222,245]" country="Japan" name="Iwate" pageId="41" pageNumber="416">Iwate</collectingRegion>
,
<collectingRegion id="49BBF85BA116FFD96B8AFF5BFD8982A4" box="[448,532,222,245]" country="Japan" name="Tochigi" pageId="41" pageNumber="416">Tochigi</collectingRegion>
,
<collectingRegion id="49BBF85BA116FFD96856FF5BFDEF82A4" box="[540,626,222,245]" country="Japan" name="Nagano" pageId="41" pageNumber="416">Nagano</collectingRegion>
,
<collectingRegion id="49BBF85BA116FFD96831FF5BFD3282A4" box="[635,687,222,245]" country="Japan" name="Gifu" pageId="41" pageNumber="416">Gifu</collectingRegion>
,
<collectingRegion id="49BBF85BA116FFD968FCFF5BFD6982A4" box="[694,756,222,245]" country="Japan" name="Aichi" pageId="41" pageNumber="416">Aichi</collectingRegion>
,
<collectingRegion id="49BBF85BA116FFD96ADDFF7BFF528344" box="[151,207,254,277]" country="Japan" name="Nara" pageId="41" pageNumber="416">Nara</collectingRegion>
; Shikoku:
<collectingRegion id="49BBF85BA116FFD96B09FF7BFE238344" box="[323,446,254,277]" country="Japan" name="Tokushima" pageId="41" pageNumber="416">Tokushima</collectingRegion>
,
<collectingRegion id="49BBF85BA116FFD96B83FF7BFD8E8344" box="[457,531,254,277]" country="Japan" name="Ehime" pageId="41" pageNumber="416">Ehime</collectingRegion>
,
<collectingRegion id="49BBF85BA116FFD96854FF7BFDF98344" box="[542,612,254,277]" country="Japan" name="Kochi" pageId="41" pageNumber="416">Kochi</collectingRegion>
; Kyushu:
<collectingRegion id="49BBF85BA116FFD96898FF7BFF418364" country="Japan" name="Fukuoka" pageId="41" pageNumber="416">Fukuoka</collectingRegion>
,
<collectingRegion id="49BBF85BA116FFD96AADFE9BFE848364" box="[231,281,286,309]" country="Japan" name="Oita" pageId="41" pageNumber="416">Oita</collectingRegion>
,
<collectingRegion id="49BBF85BA116FFD96B69FE9BFE178364" box="[291,394,286,309]" country="Japan" name="Miyazaki" pageId="41" pageNumber="416">Miyazaki</collectingRegion>
(
<figureCitation id="13442A3CA116FFD96BD2FE9BFE748364" box="[408,489,286,309]" captionStart="Fig" captionStartId="39.[153,184,2034,2053]" captionTargetBox="[165,1422,635,1952]" captionTargetId="figure-204@39.[151,1437,616,1970]" captionTargetPageId="39" captionText="Fig. 22. Distribution of Anthocoris miyamotoi Hiura, 1959 (circles) and A. venustus sp. nov. (triangles) in Japan. Filled symbols represent records based on specimens examined by us, open symbols represent literature records." figureDoi="http://doi.org/10.5281/zenodo.5821024" httpUri="https://zenodo.org/record/5821024/files/figure.png" pageId="41" pageNumber="416">Fig. 22</figureCitation>
). Compared to the distribution pattern of
<taxonomicName id="4C7F4D3AA116FFD96B05FEBBFE408304" authorityName="Hiura" authorityYear="1959" box="[335,477,318,341]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="416" phylum="Arthropoda" rank="species" species="miyamotoi">
<emphasis id="B90BEAABA116FFD96B05FEBBFE408304" box="[335,477,318,341]" italics="true" pageId="41" pageNumber="416">A. miyamotoi</emphasis>
</taxonomicName>
, the new species is widely distributed throughout
<collectingRegion id="49BBF85BA116FFD96BD8FEDBFE608324" box="[402,509,350,373]" country="Japan" name="Hokkaido" pageId="41" pageNumber="416">Hokkaido</collectingRegion>
and is restricted to the mountainous areas of Honshu, Shikoku, and Kyushu (
<figureCitation id="13442A3CA116FFD96886FEFBFF2B83E4" captionStart="Fig" captionStartId="39.[153,184,2034,2053]" captionTargetBox="[165,1422,635,1952]" captionTargetId="figure-204@39.[151,1437,616,1970]" captionTargetPageId="39" captionText="Fig. 22. Distribution of Anthocoris miyamotoi Hiura, 1959 (circles) and A. venustus sp. nov. (triangles) in Japan. Filled symbols represent records based on specimens examined by us, open symbols represent literature records." figureDoi="http://doi.org/10.5281/zenodo.5821024" httpUri="https://zenodo.org/record/5821024/files/figure.png" pageId="41" pageNumber="416">Fig. 22</figureCitation>
). This species shows an apparently allopatric distribution with
<taxonomicName id="4C7F4D3AA116FFD96AB1FE3BFE158384" authorityName="Hiura" authorityYear="1959" box="[251,392,446,469]" class="Insecta" family="Anthocoridae" genus="Anthocoris" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="416" phylum="Arthropoda" rank="species" species="miyamotoi">
<emphasis id="B90BEAABA116FFD96AB1FE3BFE158384" box="[251,392,446,469]" italics="true" pageId="41" pageNumber="416">A. miyamotoi</emphasis>
</taxonomicName>
. The allopatric distribution in both species might be influenced by differences in life history traits such as diapause and cold tolerance.
</paragraph>
</subSubSection>
</treatment>
</document>
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