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<document id="E99275C3E29BBA232D0847F9CA76C683" ID-DOI="10.11646/zootaxa.3948.2.1" ID-GBIF-Dataset="d0b508ad-9cd9-4f3f-a2fa-edce62cd1900" ID-ISSN="1175-5326" ID-Zenodo-Dep="240293" ID-ZooBank="EC744FBA-7006-4A25-A67D-DC006AD453A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461043511112" checkinUser="plazi" docAuthor="Antić, Dragan, Dražina, Tvrtko, Raða, Tonći, Tomić, Vladimir T. &amp; Makarov, Slobodan E." docDate="2015" docId="03A2AF66FFB9FF9EFF69FE51AE68FBDD" docLanguage="en" docName="zt03948p181.pdf" docOrigin="Zootaxa 3948 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrochaetosoma bertiscea Antic and Makarov, sp. n." docType="treatment" docVersion="8" lastPageNumber="170" masterDocId="FF9BD71EFFB6FF8DFFFEFFE6AB31FFCD" masterDocTitle="Review of the family Anthogonidae (Diplopoda, Chordeumatida), with descriptions of three new species from the Balkan Peninsula" masterLastPageNumber="181" masterPageNumber="151" pageNumber="166" updateTime="1698576207372" updateUser="plazi">
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<mods:title id="B5D005516D16184B1F8E842E1381E1F4">Review of the family Anthogonidae (Diplopoda, Chordeumatida), with descriptions of three new species from the Balkan Peninsula</mods:title>
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<mods:name id="A2A4668290B0F5E003641A6B2CE8C1F4" type="personal">
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<mods:namePart id="D740043A181183664CE35B40E4B00CA2">Antić, Dragan</mods:namePart>
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<mods:namePart id="9FFF6977BA147E1116B9A8C53C8E7DE6">Dražina, Tvrtko</mods:namePart>
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<mods:namePart id="8FD0939D6C6F9AC84154A89D79DBBF2D">Raða, Tonći</mods:namePart>
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<mods:namePart id="7476E955E7CA101EDF7F286DC76044ED">Tomić, Vladimir T.</mods:namePart>
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<mods:namePart id="CF61DEBEDADFC1EB28CE411DE44898FD">Makarov, Slobodan E.</mods:namePart>
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<mods:date id="952823555E6AFD4F88FCE9517476615B">2015</mods:date>
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<mods:number id="1D29CBC47CD60ACC53E8512E1402B321">3948</mods:number>
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<treatment id="03A2AF66FFB9FF9EFF69FE51AE68FBDD" ID-DOI="http://doi.org/10.5281/zenodo.6109319" ID-GBIF-Taxon="119610183" ID-Zenodo-Dep="6109319" LSID="urn:lsid:plazi:treatment:03A2AF66FFB9FF9EFF69FE51AE68FBDD" httpUri="http://treatment.plazi.org/id/03A2AF66FFB9FF9EFF69FE51AE68FBDD" lastPageId="19" lastPageNumber="170" pageId="15" pageNumber="166">
<subSubSection id="C3114DFBFFB9FF82FF69FE51AA24FE3E" pageId="15" pageNumber="166" type="nomenclature">
<paragraph id="8BB41E70FFB9FF82FF69FE51A805FE1F" blockId="15.[151,820,439,499]" box="[151,820,439,466]" pageId="15" pageNumber="166">
<heading id="D0FCA91CFFB9FF82FF69FE51A805FE1F" bold="true" box="[151,820,439,466]" fontSize="11" level="1" pageId="15" pageNumber="166" reason="1">
<emphasis id="B97FC262FFB9FF82FF69FE51A805FE1F" bold="true" box="[151,820,439,466]" pageId="15" pageNumber="166">
<taxonomicName id="4C0B65F3FFB9FF82FF69FE51A9D4FE1F" ID-CoL="849V7" authority="Antic and Makarov" authorityName="Antic and Makarov" box="[151,741,439,466]" class="Diplopoda" family="Anthogonidae" genus="Macrochaetosoma" kingdom="Animalia" order="Chordeumatida" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="bertiscea" status="sp. nov.">
<emphasis id="B97FC262FFB9FF82FF69FE51AAD8FE1C" bold="true" box="[151,489,439,465]" italics="true" pageId="15" pageNumber="166">Macrochaetosoma bertiscea</emphasis>
Antić and Makarov
</taxonomicName>
,
<taxonomicNameLabel id="A24C7F19FFB9FF82FD0CFE5EA805FE1F" box="[754,820,440,466]" pageId="15" pageNumber="166" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB41E70FFB9FF82FF69FE3CAA24FE3E" blockId="15.[151,820,439,499]" box="[151,277,474,499]" pageId="15" pageNumber="166">
<figureCitation id="133002F5FFB9FF82FF69FE3CAA24FE3E" box="[151,277,474,499]" captionStart-0="FIGURES 52 55" captionStart-1="FIGURES 56 61" captionStart-2="FIGURES 62 66" captionStartId-0="15.[151,264,1835,1857]" captionStartId-1="16.[151,264,1835,1857]" captionStartId-2="17.[151,264,1954,1976]" captionTargetBox-0="[151,1436,804,1813]" captionTargetBox-1="[151,1436,193,1813]" captionTargetBox-2="[205,1381,193,1932]" captionTargetId-0="figure@15.[151,1436,804,1814]" captionTargetId-1="figure@16.[151,1436,193,1813]" captionTargetId-2="figure@17.[205,1381,193,1933]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="FIGURES 52 55. Macrochaetosoma bertiscea sp. n., holotype male from the Jama Gorniča Pit, Prokletije Mt. 52: body segment XV, dorsal view; 53: middle area of prozonite XV, dorsal view; 54: cross-section of body segment XV, posterior view; 55: macrochaeta of body segment XV." captionText-1="FIGURES 56 61. Macrochaetosoma bertiscea sp. n., holotype male from the Jama Gorniča Pit, Prokletije Mt., gonopodal block. 56: anterior view; 57: antero-lateral view; 58: antero-lateral and distal view; 59: lateral view; 60: posterior view; 61: postero-lateral and distal view. Designations: a: shield-like coxal process, d: triangular denticles, f: fissure, h: posterior subtriangular horn, m: medial sternal process of posterior gonopods, r: postero-mesal ribs, bb: bilobed basal bulge, b 1: lateral bristles apparatus, b 2: mesal bristles apparatus, cv: coxal vesicle; cip: inner part of lateral coxal process, cop: outer part of lateral coxal process; lcp: lateral coxal process of posterior gonopods." captionText-2="FIGURES 62 66. Macrochaetosoma bertiscea sp. n., holotype male from the Jama Gorniča Pit, Prokletije Mt., gonopodal block. 62: distal view; 63: bristles apparatuses, antero-distal view; 64: lateral bristles apparatus, lateral view; 65: mesal bristles apparatus, posterior view; 66: posterior view." httpUri-0="https://zenodo.org/record/240304/files/figure.png" httpUri-1="https://zenodo.org/record/240305/files/figure.png" httpUri-2="https://zenodo.org/record/240306/files/figure.png" pageId="15" pageNumber="166">Figs 5266</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3114DFBFFB9FF82FF69FDC4AA43FD06" pageId="15" pageNumber="166" type="materials_examined">
<paragraph id="8BB41E70FFB9FF82FF69FDC4AA43FD06" blockId="15.[151,1437,546,751]" pageId="15" pageNumber="166">
<emphasis id="B97FC262FFB9FF82FF69FDC4AAB4FDF6" bold="true" box="[151,389,546,571]" pageId="15" pageNumber="166">Material examined.</emphasis>
<typeStatus id="54B0A0D2FFB9FF82FE6EFDC4AAC9FDF6" box="[400,504,546,571]" pageId="15" pageNumber="166" type="holotype">Holotype</typeStatus>
male: Gorniča Pit, ca
<quantity id="4CF3B395FFB9FF82FCF4FDC5A857FDF7" box="[778,870,547,571]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="166" unit="m" value="2000.0">2000 m</quantity>
, Ćaf Borit, Bjelič Massif, Prokletije Mountains,
<collectingCountry id="F31C5EE0FFB9FF82FF69FDA1AA12FD92" box="[151,291,583,607]" name="Serbia" pageId="15" pageNumber="166">Montenegro</collectingCountry>
,
<geoCoordinate id="EE3F78B7FFB9FF82FED0FDA0AAE9FD92" box="[302,472,582,607]" direction="north" orientation="latitude" pageId="15" pageNumber="166" precision="1" value="42.498974">42°29'56.30&quot;N</geoCoordinate>
,
<geoCoordinate id="EE3F78B7FFB9FF82FE1AFDA0A9BAFD92" box="[484,651,582,607]" direction="east" orientation="longitude" pageId="15" pageNumber="166" precision="1" value="19.88936">19°53'21.70&quot;E</geoCoordinate>
,
<date id="FFB538B0FFB9FF82FD68FDA0A810FD92" box="[662,801,582,607]" pageId="15" pageNumber="166" value="2011-07-25">25.VII.2011</date>
, leg. I. Njunjić (IZB ANG MB 1001). Allotype female (IZB ANG MB 1002) and three
<typeStatus id="54B0A0D2FFB9FF82FDE3FD8DA94FFD4E" box="[541,638,619,643]" pageId="15" pageNumber="166" type="paratype">paratype</typeStatus>
juveniles (IZB ANG MB 1003): same data as for
<typeStatus id="54B0A0D2FFB9FF82FB29FD8CAE0CFD4E" box="[1239,1341,618,643]" pageId="15" pageNumber="166" type="holotype">holotype</typeStatus>
. Further material: Juvenile from the Jama od Karimana Pit, Kučke Planine Mountains,
<date id="FFB538B0FFB9FF82FBDCFD69AF9CFD6A" box="[1058,1197,655,679]" pageId="15" pageNumber="166" value="2010-07-21">21.VII.2010</date>
, leg. I. Njunjić (IZB ANG MBq 1004).
</paragraph>
</subSubSection>
<subSubSection id="C3114DFBFFB9FF82FF39FD30AF3AFD22" box="[199,1035,726,751]" pageId="15" pageNumber="166" type="etymology">
<paragraph id="8BB41E70FFB9FF82FF39FD30AF3AFD22" blockId="15.[151,1437,546,751]" box="[199,1035,726,751]" pageId="15" pageNumber="166">
<emphasis id="B97FC262FFB9FF82FF39FD30AA7FFD22" bold="true" box="[199,334,726,751]" pageId="15" pageNumber="166">Etymology.</emphasis>
After its
<typeStatus id="54B0A0D2FFB9FF82FE46FD31AAD9FD22" box="[440,488,727,751]" pageId="15" pageNumber="166">type</typeStatus>
locality, Prokletije Mountains (Mons Bertiscus).
</paragraph>
</subSubSection>
<caption id="DF744EF8FFB9FF82FF69F8CDA92DF8B3" httpUri="https://zenodo.org/record/240304/files/figure.png" pageId="15" pageNumber="166" targetBox="[151,1436,804,1813]" targetPageId="15">
<paragraph id="8BB41E70FFB9FF82FF69F8CDA92DF8B3" blockId="15.[151,1436,1835,1918]" pageId="15" pageNumber="166">
<emphasis id="B97FC262FFB9FF82FF69F8CDAA66F88C" bold="true" box="[151,343,1835,1857]" pageId="15" pageNumber="166">FIGURES 5255.</emphasis>
<taxonomicName id="4C0B65F3FFB9FF82FE9FF8CDA9B5F88C" box="[353,644,1835,1857]" class="Diplopoda" family="Anthogonidae" genus="Macrochaetosoma" kingdom="Animalia" order="Chordeumatida" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="bertiscea" status="sp. nov.">
<emphasis id="B97FC262FFB9FF82FE9FF8CDA9B5F88C" box="[353,644,1835,1857]" italics="true" pageId="15" pageNumber="166">Macrochaetosoma bertiscea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C7F19FFB9FF82FD70F8CDA9FAF88C" box="[654,715,1835,1857]" pageId="15" pageNumber="166" rank="species">
<emphasis id="B97FC262FFB9FF82FD70F8CDA9FAF88C" bold="true" box="[654,715,1835,1857]" pageId="15" pageNumber="166">sp. n.</emphasis>
</taxonomicNameLabel>
, holotype male from the Jama Gorniča Pit, Prokletije Mt. 52: body segment XV, dorsal view; 53: middle area of prozonite XV, dorsal view; 54: cross-section of body segment XV, posterior view; 55: macrochaeta of body segment XV.
</paragraph>
</caption>
<caption id="DF744EF8FFA6FF9DFF69F8CDA86AF817" httpUri="https://zenodo.org/record/240305/files/figure.png" pageId="16" pageNumber="167" targetBox="[151,1436,193,1813]" targetPageId="16">
<paragraph id="8BB41E70FFA6FF9DFF69F8CDA86AF817" blockId="16.[151,1436,1835,2010]" pageId="16" pageNumber="167">
<emphasis id="B97FC262FFA6FF9DFF69F8CDAA67F88C" bold="true" box="[151,342,1835,1857]" pageId="16" pageNumber="167">FIGURES 5661.</emphasis>
<taxonomicName id="4C0B65F3FFA6FF9DFEA0F8CDA94EF88C" box="[350,639,1835,1857]" class="Diplopoda" family="Anthogonidae" genus="Macrochaetosoma" kingdom="Animalia" order="Chordeumatida" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="bertiscea" status="sp. nov.">
<emphasis id="B97FC262FFA6FF9DFEA0F8CDA94EF88C" box="[350,639,1835,1857]" italics="true" pageId="16" pageNumber="167">Macrochaetosoma bertiscea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C7F19FFA6FF9DFD76F8CDA9F5F88C" box="[648,708,1835,1857]" pageId="16" pageNumber="167" rank="species">
<emphasis id="B97FC262FFA6FF9DFD76F8CDA9F5F88C" bold="true" box="[648,708,1835,1857]" pageId="16" pageNumber="167">sp. n.</emphasis>
</taxonomicNameLabel>
, holotype male from the Jama Gorniča Pit, Prokletije Mt., gonopodal block. 56: anterior view; 57: antero-lateral view; 58: antero-lateral and distal view; 59: lateral view; 60: posterior view; 61: postero-lateral and distal view. Designations:
<emphasis id="B97FC262FFA6FF9DFD7EF88EA9BDF8B3" bold="true" box="[640,652,1896,1918]" pageId="16" pageNumber="167">a</emphasis>
: shield-like coxal process,
<emphasis id="B97FC262FFA6FF9DFC48F88EA8F5F8B3" bold="true" box="[950,964,1896,1918]" pageId="16" pageNumber="167">d</emphasis>
: triangular denticles,
<emphasis id="B97FC262FFA6FF9DFB4FF88EAF88F8B3" bold="true" box="[1201,1209,1896,1918]" pageId="16" pageNumber="167">f</emphasis>
: fissure,
<emphasis id="B97FC262FFA6FF9DFADFF88EAE1EF8B3" bold="true" box="[1313,1327,1896,1918]" pageId="16" pageNumber="167">h</emphasis>
: posterior subtriangular horn,
<emphasis id="B97FC262FFA6FF9DFEA1F861AA45F850" bold="true" box="[351,372,1927,1949]" pageId="16" pageNumber="167">m</emphasis>
: medial sternal process of posterior gonopods,
<emphasis id="B97FC262FFA6FF9DFCA8F861A850F850" bold="true" box="[854,865,1927,1949]" pageId="16" pageNumber="167">r</emphasis>
: postero-mesal ribs,
<emphasis id="B97FC262FFA6FF9DFBC9F861AF62F850" bold="true" box="[1079,1107,1927,1949]" pageId="16" pageNumber="167">bb</emphasis>
: bilobed basal bulge,
<emphasis id="B97FC262FFA6FF9DFACAF861AE7FF850" bold="true" box="[1332,1358,1927,1949]" pageId="16" pageNumber="167">b1</emphasis>
: lateral bristles apparatus,
<emphasis id="B97FC262FFA6FF9DFEA7F843AA45F876" bold="true" box="[345,372,1957,1979]" pageId="16" pageNumber="167">b2</emphasis>
: mesal bristles apparatus,
<emphasis id="B97FC262FFA6FF9DFD77F843A990F876" bold="true" box="[649,673,1957,1979]" pageId="16" pageNumber="167">cv</emphasis>
: coxal vesicle;
<emphasis id="B97FC262FFA6FF9DFCB8F843A857F876" bold="true" box="[838,870,1957,1979]" pageId="16" pageNumber="167">cip</emphasis>
: inner part of lateral coxal process,
<emphasis id="B97FC262FFA6FF9DFB1AF843AE3BF876" bold="true" box="[1252,1290,1957,1979]" pageId="16" pageNumber="167">cop</emphasis>
: outer part of lateral coxal process;
<emphasis id="B97FC262FFA6FF9DFE8DF822AAA2F817" bold="true" box="[371,403,1988,2010]" pageId="16" pageNumber="167">lcp</emphasis>
: lateral coxal process of posterior gonopods.
</paragraph>
</caption>
<caption id="DF744EF8FFA7FF9CFF69F844A950F838" httpUri="https://zenodo.org/record/240306/files/figure.png" pageId="17" pageNumber="168" targetBox="[205,1381,193,1932]" targetPageId="17">
<paragraph id="8BB41E70FFA7FF9CFF69F844A950F838" blockId="17.[151,1436,1954,2037]" pageId="17" pageNumber="168">
<emphasis id="B97FC262FFA7FF9CFF69F844AA67F875" bold="true" box="[151,342,1954,1976]" pageId="17" pageNumber="168">FIGURES 6266.</emphasis>
<taxonomicName id="4C0B65F3FFA7FF9CFEA0F844A94EF875" box="[350,639,1954,1976]" class="Diplopoda" family="Anthogonidae" genus="Macrochaetosoma" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="bertiscea" status="sp. nov.">
<emphasis id="B97FC262FFA7FF9CFEA0F844A94EF875" box="[350,639,1954,1976]" italics="true" pageId="17" pageNumber="168">Macrochaetosoma bertiscea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C7F19FFA7FF9CFD76F844A9F5F875" box="[648,708,1954,1976]" pageId="17" pageNumber="168" rank="species">
<emphasis id="B97FC262FFA7FF9CFD76F844A9F5F875" bold="true" box="[648,708,1954,1976]" pageId="17" pageNumber="168">sp. n.</emphasis>
</taxonomicNameLabel>
, holotype male from the Jama Gorniča Pit, Prokletije Mt., gonopodal block. 62: distal view; 63: bristles apparatuses, antero-distal view; 64: lateral bristles apparatus, lateral view; 65: mesal bristles apparatus, posterior view; 66: posterior view.
</paragraph>
</caption>
<subSubSection id="C3114DFBFFA4FF9FFF39FF71AA06FEF2" pageId="18" pageNumber="169" type="diagnosis">
<paragraph id="8BB41E70FFA4FF9FFF39FF71AA06FEF2" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="169">
<emphasis id="B97FC262FFA4FF9FFF39FF71AA70FF7D" bold="true" box="[199,321,151,176]" pageId="18" pageNumber="169">Diagnosis.</emphasis>
The new taxon differs from the two others described in
<taxonomicName id="4C0B65F3FFA4FF9FFC3FFF71AFA0FF7D" box="[961,1169,151,176]" class="Diplopoda" family="Anthogonidae" genus="Macrochaetosoma" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC262FFA4FF9FFC3FFF71AFA0FF7D" box="[961,1169,151,176]" italics="true" pageId="18" pageNumber="169">Macrochaetosoma</emphasis>
</taxonomicName>
in many characteristics of habitus and gonopodal structure such as: broad, subquadrangular, shield-like anterior coxal processes on the anterior gonopods; bifurcated medial sternal process with bilobed base on the posterior gonopods; and absence of telopodal remnants on posterior gonopods or very unusual promentum in the form of a very low and wide triangle with six setae.
</paragraph>
</subSubSection>
<subSubSection id="C3114DFBFFA4FF9EFF39FEAAAE68FBDD" lastPageId="19" lastPageNumber="170" pageId="18" pageNumber="169" type="description">
<paragraph id="8BB41E70FFA4FF9FFF39FEAAAAC8FE45" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="169">
<emphasis id="B97FC262FFA4FF9FFF39FEAAAA68FEA8" bold="true" box="[199,345,332,357]" pageId="18" pageNumber="169">Description.</emphasis>
Body with 30 segments (including telson), in adults. Juveniles with 28 and 26 segments (including telson), respectively.
</paragraph>
<paragraph id="8BB41E70FFA4FF9FFF39FE73A8EDFE02" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="169">
<emphasis id="B97FC262FFA4FF9FFF39FE73AA43FE61" box="[199,370,405,428]" italics="true" pageId="18" pageNumber="169">Measurements:</emphasis>
<typeStatus id="54B0A0D2FFA4FF9FFE82FE72AAD5FE60" box="[380,484,404,429]" pageId="18" pageNumber="169" type="holotype">Holotype</typeStatus>
male
<quantity id="4CF3B395FFA4FF9FFDD0FE72A9A5FE61" box="[558,660,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="2.73" pageId="18" pageNumber="169" unit="mm" value="27.3">27.3 mm</quantity>
long, vertical diameter of its largest pleurotergite
<quantity id="4CF3B395FFA4FF9FFB27FE72AE05FE61" box="[1241,1332,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="169" unit="mm" value="1.6">1.6 mm</quantity>
; allotype female
<quantity id="4CF3B395FFA4FF9FFF14FE5EAA7FFE02" box="[234,334,440,464]" metricMagnitude="-2" metricUnit="m" metricValue="2.21" pageId="18" pageNumber="169" unit="mm" value="22.1">22.1 mm</quantity>
long, vertical diameter of its largest pleurotergite
<quantity id="4CF3B395FFA4FF9FFC81FE5EA8E9FE02" box="[895,984,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="169" unit="mm" value="1.7">1.7 mm</quantity>
.
</paragraph>
<paragraph id="8BB41E70FFA4FF9FFF39FE3AA990FE38" blockId="18.[151,1437,151,2013]" box="[199,673,476,501]" pageId="18" pageNumber="169">
<emphasis id="B97FC262FFA4FF9FFF39FE3AAA7AFE38" box="[199,331,476,501]" italics="true" pageId="18" pageNumber="169">Coloration:</emphasis>
Pigmentless, yellowish white.
</paragraph>
<paragraph id="8BB41E70FFA4FF9FFF39FE19ABECFCF5" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="169">
<emphasis id="B97FC262FFA4FF9FFF39FE19AA3CFDD5" box="[199,269,511,536]" italics="true" pageId="18" pageNumber="169">Head:</emphasis>
Setose, frontal side in male with slight depression. Labrum (
<typeStatus id="54B0A0D2FFA4FF9FFC2BFE19AF08FDD5" box="[981,1081,511,536]" pageId="18" pageNumber="169" type="holotype">holotype</typeStatus>
♂) with three rounded medial labral teeth, and with 2+2 labral and 8+8 supralabral bacilliform setae. Gnathochilarium (in both sexes) densely setose; promentum in the form of a low and wide triangle with six setae in one row; lingual plates wide, massive, subquadrangular. All setae very long, except setae on basal part of stipites, which are small to medium-sized. Antennae long and thin,
<quantity id="4CF3B395FFA4FF9FFE54FD76AACEFD6A" box="[426,511,656,680]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="18" pageNumber="169" unit="mm" value="4.1">4.1 mm</quantity>
in
<typeStatus id="54B0A0D2FFA4FF9FFDDCFD69A9B5FD65" box="[546,644,655,680]" pageId="18" pageNumber="169" type="holotype">holotype</typeStatus>
male. Length of antennomeres: I (0.16), II (0.37), III (0.91), IV (0.62), V (1.27), VI (0.46), VII (0.28), and VIII (0.03). Length/breadth ratios of antennomeres I-VII: I (1), II (1.9), III (5.3), IV (3.8), V (8.5), VI (3.1), and VII (2.5). Antennomeres II, IV, and VI slightly clavate. Antennomere V curved. Antennomeres II, IV, V, VI, and VII with one, three, one, four and one sensillum (sensilla), respectively. Blind.
</paragraph>
<paragraph id="8BB41E70FFA4FF9FFF39FCA2ABCBFCB2" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="169">
<emphasis id="B97FC262FFA4FF9FFF39FCA2AA13FC90" box="[199,290,836,861]" italics="true" pageId="18" pageNumber="169">Collum:</emphasis>
Narrower than head, with six macrochaetae. Anterior side semicircular, posterior side slightly concave.
</paragraph>
<paragraph id="8BB41E70FFA4FF9FFF39FC6AA926FB95" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="169">
<emphasis id="B97FC262FFA4FF9FFF39FC6AAA48FC69" box="[199,377,908,933]" italics="true" pageId="18" pageNumber="169">Body segments:</emphasis>
Prozonites with hexagonal tiles on the anterior part and with smooth posterior part (
<figureCitation id="133002F5FFA4FF9FFABBFC6AABDAFC05" captionStart="FIGURES 52 55" captionStartId="15.[151,264,1835,1857]" captionTargetBox="[151,1436,804,1813]" captionTargetId="figure@15.[151,1436,804,1814]" captionTargetPageId="15" captionText="FIGURES 52 55. Macrochaetosoma bertiscea sp. n., holotype male from the Jama Gorniča Pit, Prokletije Mt. 52: body segment XV, dorsal view; 53: middle area of prozonite XV, dorsal view; 54: cross-section of body segment XV, posterior view; 55: macrochaeta of body segment XV." httpUri="https://zenodo.org/record/240304/files/figure.png" pageId="18" pageNumber="169">Figs 52 and 53</figureCitation>
). Metazonites wrinkled (
<figureCitation id="133002F5FFA4FF9FFDF0FC49A952FC05" box="[526,611,943,968]" captionStart="FIGURES 52 55" captionStartId="15.[151,264,1835,1857]" captionTargetBox="[151,1436,804,1813]" captionTargetId="figure@15.[151,1436,804,1814]" captionTargetPageId="15" captionText="FIGURES 52 55. Macrochaetosoma bertiscea sp. n., holotype male from the Jama Gorniča Pit, Prokletije Mt. 52: body segment XV, dorsal view; 53: middle area of prozonite XV, dorsal view; 54: cross-section of body segment XV, posterior view; 55: macrochaeta of body segment XV." httpUri="https://zenodo.org/record/240304/files/figure.png" pageId="18" pageNumber="169">Fig. 52</figureCitation>
) with lateral keels in form of swelling (
<figureCitation id="133002F5FFA4FF9FFBBDFC49AFCBFC05" box="[1091,1274,943,968]" captionStart="FIGURES 52 55" captionStartId="15.[151,264,1835,1857]" captionTargetBox="[151,1436,804,1813]" captionTargetId="figure@15.[151,1436,804,1814]" captionTargetPageId="15" captionText="FIGURES 52 55. Macrochaetosoma bertiscea sp. n., holotype male from the Jama Gorniča Pit, Prokletije Mt. 52: body segment XV, dorsal view; 53: middle area of prozonite XV, dorsal view; 54: cross-section of body segment XV, posterior view; 55: macrochaeta of body segment XV." httpUri="https://zenodo.org/record/240304/files/figure.png" pageId="18" pageNumber="169">Figs 52 and 54</figureCitation>
), absent from pleurotergites XXVIII and XXIX. Better developed in male. Limbus absent. Macrochaetae on knobs. In all specimens from
<typeStatus id="54B0A0D2FFA4FF9FFEABFC1EAAB4FBDD" box="[341,389,1016,1040]" pageId="18" pageNumber="169">type</typeStatus>
material, macrochaetae were fallen off. Macrochaetal index CIX (pleurotergite 15) ~ 0.45; median index MIX (pleurotergite 15) ~ 1; paratergal index PIX (pleurotergite 15) ~ 0.35; macrochaetal angle MA (pleurotergite 15) ~160° (
<figureCitation id="133002F5FFA4FF9FFE48FBD9A93BFB95" box="[438,522,1087,1112]" captionStart="FIGURES 52 55" captionStartId="15.[151,264,1835,1857]" captionTargetBox="[151,1436,804,1813]" captionTargetId="figure@15.[151,1436,804,1814]" captionTargetPageId="15" captionText="FIGURES 52 55. Macrochaetosoma bertiscea sp. n., holotype male from the Jama Gorniča Pit, Prokletije Mt. 52: body segment XV, dorsal view; 53: middle area of prozonite XV, dorsal view; 54: cross-section of body segment XV, posterior view; 55: macrochaeta of body segment XV." httpUri="https://zenodo.org/record/240304/files/figure.png" pageId="18" pageNumber="169">Fig. 52</figureCitation>
).
</paragraph>
<paragraph id="8BB41E70FFA4FF9FFF39FB82ABE7FB09" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="169">
<emphasis id="B97FC262FFA4FF9FFF39FB82AA3CFBB0" box="[199,269,1124,1149]" italics="true" pageId="18" pageNumber="169">Telson</emphasis>
: Epiproct with a pair of spinnerets and six setae arranged in two rows (2+2 marginal trichoid setae and 1+1 paramedial bacilliform seta). Hypoproct with two apical trichoid setae. Paraprocts with 3+3 marginal trichoid setae.
</paragraph>
<paragraph id="8BB41E70FFA4FF9FFF39FB29AF71FAC1" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="169">
<emphasis id="B97FC262FFA4FF9FFF39FB29AA69FB25" box="[199,344,1231,1256]" italics="true" pageId="18" pageNumber="169">Walking legs</emphasis>
(both sexes): Elongated and thin. Leg pairs 1 and 2 with tarsal combs, prefemora with several long and robust setae, femora and postfemora with numerous long and robust setae.
</paragraph>
<paragraph id="8BB41E70FFA4FF9FFF39FAF1AA1CF92D" blockId="18.[151,1437,151,2013]" pageId="18" pageNumber="169">
<emphasis id="B97FC262FFA4FF9FFF39FAF1AAD7FAFD" box="[199,486,1303,1328]" italics="true" pageId="18" pageNumber="169">Male sexual characters:</emphasis>
Leg pairs 37 greatly enlarged. Leg pair 3 with posterior excavations on all podomeres; postfemora, tibiae, and tarsi curved caudally and very flattened antero-posteriorly. Leg pair 4 with antero-posteriorly flattened podomeres with excavations; coxae with ventral protuberance, tibiae and tarsi with medio-longitudinal ridge on the oral side. All podomeres very broad and short in leg pairs 3 and 4. Leg pairs 5 and 6 are the most similar to normal walking legs, with robust femora. Coxae of leg pair 7 ventrally densely setose; tarsi broad at base and gradually tapering to last third of length, from there of the same width until claws; last third of tarsi approximately three times narrower than first third (resembling the pedipalpal chela of some pseudoscorpions with closed fingers). Leg pairs 10 and 11 with coxal glands. Leg pair 11 with posterior mesal coxal horns directed toward each other and apically touching. Sternum 10 with medial subtriangular protrusion. Coxae of leg pairs 13 and 15 with well developed, massive mesal process. This structure larger on leg pair 13 than on pair 15. Its anterior part on leg pair 13 is wider and rounded, while the posterior part is acuminating. On leg pair 15, the anterior and posterior parts are the same width, acuminate. Leg pairs 1621 are the same size as the other walking legs.
</paragraph>
<paragraph id="8BB41E70FFA4FF9EFF39F90BAF96FEA8" blockId="18.[151,1437,151,2013]" lastBlockId="19.[151,1437,151,1040]" lastPageId="19" lastPageNumber="170" pageId="18" pageNumber="169">
<emphasis id="B97FC262FFA4FF9FFF39F90BAAAAF8C8" box="[199,411,1772,1797]" italics="true" pageId="18" pageNumber="169">Anterior gonopods</emphasis>
(
<figureCitation id="133002F5FFA4FF9FFE54F90AA91EF8C8" box="[426,559,1772,1797]" captionStart-0="FIGURES 56 61" captionStart-1="FIGURES 62 66" captionStartId-0="16.[151,264,1835,1857]" captionStartId-1="17.[151,264,1954,1976]" captionTargetBox-0="[151,1436,193,1813]" captionTargetBox-1="[205,1381,193,1932]" captionTargetId-0="figure@16.[151,1436,193,1813]" captionTargetId-1="figure@17.[205,1381,193,1933]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURES 56 61. Macrochaetosoma bertiscea sp. n., holotype male from the Jama Gorniča Pit, Prokletije Mt., gonopodal block. 56: anterior view; 57: antero-lateral view; 58: antero-lateral and distal view; 59: lateral view; 60: posterior view; 61: postero-lateral and distal view. Designations: a: shield-like coxal process, d: triangular denticles, f: fissure, h: posterior subtriangular horn, m: medial sternal process of posterior gonopods, r: postero-mesal ribs, bb: bilobed basal bulge, b 1: lateral bristles apparatus, b 2: mesal bristles apparatus, cv: coxal vesicle; cip: inner part of lateral coxal process, cop: outer part of lateral coxal process; lcp: lateral coxal process of posterior gonopods." captionText-1="FIGURES 62 66. Macrochaetosoma bertiscea sp. n., holotype male from the Jama Gorniča Pit, Prokletije Mt., gonopodal block. 62: distal view; 63: bristles apparatuses, antero-distal view; 64: lateral bristles apparatus, lateral view; 65: mesal bristles apparatus, posterior view; 66: posterior view." httpUri-0="https://zenodo.org/record/240305/files/figure.png" httpUri-1="https://zenodo.org/record/240306/files/figure.png" pageId="18" pageNumber="169">Figs 5665</figureCitation>
): Oral side with two parallel-sided, wide, almost subquadrangular, shield-like coxal processes (
<emphasis id="B97FC262FFA4FF9FFEA0F8E9AA5DF8E5" bold="true" box="[350,364,1807,1832]" pageId="18" pageNumber="169">a</emphasis>
) [= Cheirite
<emphasis id="B97FC262FFA4FF9FFDF5F8F7A978F8E5" box="[523,585,1809,1832]" italics="true" pageId="18" pageNumber="169">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA4FF9FFDAAF8E9A83FF8E5" author="Verhoeff" box="[596,782,1807,1832]" pageId="18" pageNumber="169" refString="Verhoeff, K. W. (1942) Eine neue, westbalkanische, cavernicole Familie der Ascospermophora und die fragwurdige Gattung Macrochaetosoma Abs. Zoologischer Anzeiger, 139, 54 - 66." type="journal article" year="1942">Verhoeff (1942)</bibRefCitation>
; telopodites
<emphasis id="B97FC262FFA4FF9FFC5BF8F7A8D2F8E5" box="[933,995,1809,1832]" italics="true" pageId="18" pageNumber="169">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA4FF9FFC10F8F6AFAEF8E5" author="Strasser" box="[1006,1183,1807,1832]" pageId="18" pageNumber="169" refString="Strasser, K. (1962) Diplopoden aus Bulgarien und Jugoslawien. Senckenbergiana biologica, 43, 437 - 470." type="journal article" year="1962">Strasser (1962)</bibRefCitation>
; innerer Coxitfortsatz
<emphasis id="B97FC262FFA4FF9FFF69F8D3ABE4F881" box="[151,213,1845,1868]" italics="true" pageId="18" pageNumber="169">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA4FF9FFF1EF8D2AA91F880" author="Strasser" box="[224,416,1844,1869]" pageId="18" pageNumber="169" refString="Strasser, K. (1971 b) Uber Diplopoden Jugoslawiens. Senckenbergiana biologica, 53, 313 - 345." type="journal article" year="1971" yearSuffix="b">Strasser (1971b)</bibRefCitation>
; or inner/paramedial main branch of the anterior coxal stem
<emphasis id="B97FC262FFA4FF9FFB8DF8D3AF80F881" box="[1139,1201,1845,1868]" italics="true" pageId="18" pageNumber="169">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA4FF9FFB43F8D2AE6BF881" author="Mrsic" box="[1213,1370,1844,1869]" pageId="18" pageNumber="169" refString="Mrsic, N. (1992) Biokoviella mauriesi gen. nov., sp. nov. (Biokoviellidae fam. nov.), Macrochaetosomatinae subf. nov. (Anthogonidae) and superfamily Cleidogonoidea of the western Balkans (Craspedosomidea, Diplopoda). Razprave IV. Razreda SAZU, 33 (3) 51 - 91." type="journal article" year="1992">Mršić (1992)</bibRefCitation>
]. The largest area of the oral sides of these shield-like processes is smooth, except on the top, which is wrinkled. Apically, each shield-like process carrying one outer posterior subtriangular horn (
<emphasis id="B97FC262FFA4FF9FFBAAF89AAF52F858" bold="true" box="[1108,1123,1916,1941]" pageId="18" pageNumber="169">h</emphasis>
). These shields are clearly separated from the rest of the anterior gonopods by a transverse rift. Each lateral side of the anterior gonopods with a very wide, antero-posteriorly directed lateral bristle apparatus (
<emphasis id="B97FC262FFA4FF9FFC60F822A88DF810" bold="true" box="[926,956,1988,2013]" pageId="18" pageNumber="169">b1</emphasis>
) [= äußerer Coxitfortsatz
<emphasis id="B97FC262FFA4FF9FFB06F823AE07F811" box="[1272,1334,1989,2012]" italics="true" pageId="18" pageNumber="169">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA4FF9EFABAF822ABC2FF7D" author="Strasser" lastPageId="19" lastPageNumber="170" pageId="18" pageNumber="169" refString="Strasser, K. (1971 b) Uber Diplopoden Jugoslawiens. Senckenbergiana biologica, 53, 313 - 345." type="journal article" year="1971" yearSuffix="b">Strasser (1971b)</bibRefCitation>
; or outer/lateral branch of the anterior coxal stem
<emphasis id="B97FC262FFA5FF9EFCC1FF7FA84CFF7D" box="[831,893,153,176]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFC79FF71AF12FF7D" author="Mrsic" box="[903,1059,151,176]" pageId="19" pageNumber="170" refString="Mrsic, N. (1992) Biokoviella mauriesi gen. nov., sp. nov. (Biokoviellidae fam. nov.), Macrochaetosomatinae subf. nov. (Anthogonidae) and superfamily Cleidogonoidea of the western Balkans (Craspedosomidea, Diplopoda). Razprave IV. Razreda SAZU, 33 (3) 51 - 91." type="journal article" year="1992">Mršić (1992)</bibRefCitation>
]. Apart from these lateral bristle apparatuses, there are two more, mesal bristle apparatuses (
<emphasis id="B97FC262FFA5FF9EFCABFF5AA843FF18" bold="true" box="[853,882,188,213]" pageId="19" pageNumber="170">b2</emphasis>
), one on each side, with numerous transversoposteriorly directed bristles. The most posterior bristles of these parts reach the cavity of the peculiar paramedial coxal vesicles (
<emphasis id="B97FC262FFA5FF9EFEB0FEE2AA58FED0" bold="true" box="[334,361,260,285]" pageId="19" pageNumber="170">cv</emphasis>
) [= coxite
<emphasis id="B97FC262FFA5FF9EFE0AFEE3A903FED1" box="[500,562,261,284]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFDC0FEE2A9C7FED1" author="Verhoeff" box="[574,758,260,285]" pageId="19" pageNumber="170" refString="Verhoeff, K. W. (1942) Eine neue, westbalkanische, cavernicole Familie der Ascospermophora und die fragwurdige Gattung Macrochaetosoma Abs. Zoologischer Anzeiger, 139, 54 - 66." type="journal article" year="1942">Verhoeff (1942)</bibRefCitation>
and
<bibRefCitation id="EF9A6381FFA5FF9EFCC9FEE2A8D8FED0" author="Strasser" box="[823,1001,260,285]" pageId="19" pageNumber="170" refString="Strasser, K. (1962) Diplopoden aus Bulgarien und Jugoslawien. Senckenbergiana biologica, 43, 437 - 470." type="journal article" year="1962">Strasser (1962)</bibRefCitation>
; hinterer Basalanhang
<emphasis id="B97FC262FFA5FF9EFB07FEE3AE06FED1" box="[1273,1335,261,284]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFABAFEE2ABC2FE8D" author="Strasser" pageId="19" pageNumber="170" refString="Strasser, K. (1971 b) Uber Diplopoden Jugoslawiens. Senckenbergiana biologica, 53, 313 - 345." type="journal article" year="1971" yearSuffix="b">Strasser (1971b)</bibRefCitation>
; or posterior paramedial oval coxal processes
<emphasis id="B97FC262FFA5FF9EFCFDFECFA870FE8D" box="[771,833,297,320]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFCB4FEC1A8D5FE8D" author="Mrsic" box="[842,996,295,320]" pageId="19" pageNumber="170" refString="Mrsic, N. (1992) Biokoviella mauriesi gen. nov., sp. nov. (Biokoviellidae fam. nov.), Macrochaetosomatinae subf. nov. (Anthogonidae) and superfamily Cleidogonoidea of the western Balkans (Craspedosomidea, Diplopoda). Razprave IV. Razreda SAZU, 33 (3) 51 - 91." type="journal article" year="1992">Mršić (1992)</bibRefCitation>
]. These vesicles are the most posterior parts of the anterior gonopods, giving the impression of spiral twisting when viewed distally.
</paragraph>
<paragraph id="8BB41E70FFA5FF9EFF39FE97AA69FC05" blockId="19.[151,1437,151,1040]" pageId="19" pageNumber="170">
<emphasis id="B97FC262FFA5FF9EFF39FE97AA96FE45" box="[199,423,367,392]" italics="true" pageId="19" pageNumber="170">Posterior gonopods</emphasis>
(
<figureCitation id="133002F5FFA5FF9EFE49FE89A906FE45" box="[439,567,367,392]" captionStart="FIGURES 56 61" captionStartId="16.[151,264,1835,1857]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure@16.[151,1436,193,1813]" captionTargetPageId="16" captionText="FIGURES 56 61. Macrochaetosoma bertiscea sp. n., holotype male from the Jama Gorniča Pit, Prokletije Mt., gonopodal block. 56: anterior view; 57: antero-lateral view; 58: antero-lateral and distal view; 59: lateral view; 60: posterior view; 61: postero-lateral and distal view. Designations: a: shield-like coxal process, d: triangular denticles, f: fissure, h: posterior subtriangular horn, m: medial sternal process of posterior gonopods, r: postero-mesal ribs, bb: bilobed basal bulge, b 1: lateral bristles apparatus, b 2: mesal bristles apparatus, cv: coxal vesicle; cip: inner part of lateral coxal process, cop: outer part of lateral coxal process; lcp: lateral coxal process of posterior gonopods." httpUri="https://zenodo.org/record/240305/files/figure.png" pageId="19" pageNumber="170">Figs 5861</figureCitation>
and
<figureCitation id="133002F5FFA5FF9EFD8CFE89A9A3FE45" box="[626,658,367,392]" captionStart="FIGURES 62 66" captionStartId="17.[151,264,1954,1976]" captionTargetBox="[205,1381,193,1932]" captionTargetId="figure@17.[205,1381,193,1933]" captionTargetPageId="17" captionText="FIGURES 62 66. Macrochaetosoma bertiscea sp. n., holotype male from the Jama Gorniča Pit, Prokletije Mt., gonopodal block. 62: distal view; 63: bristles apparatuses, antero-distal view; 64: lateral bristles apparatus, lateral view; 65: mesal bristles apparatus, posterior view; 66: posterior view." httpUri="https://zenodo.org/record/240306/files/figure.png" pageId="19" pageNumber="170">66</figureCitation>
): In Mršićs interpretation of the posterior gonopods, he noted three main structures: a medial sternite process, a paramedial coxal process, and a lateral coxal process. In our view, the posterior gonopods consist of two main structures: a medial sternal process (
<emphasis id="B97FC262FFA5FF9EFC0CFE51AF38FE1D" bold="true" box="[1010,1033,439,464]" pageId="19" pageNumber="170">m</emphasis>
) [= sternalen Mittelfortsatzes
<emphasis id="B97FC262FFA5FF9EFAA0FE5FAEADFE1D" box="[1374,1436,441,464]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFF69FE3AAA65FE39" author="Verhoeff" box="[151,340,476,501]" pageId="19" pageNumber="170" refString="Verhoeff, K. W. (1942) Eine neue, westbalkanische, cavernicole Familie der Ascospermophora und die fragwurdige Gattung Macrochaetosoma Abs. Zoologischer Anzeiger, 139, 54 - 66." type="journal article" year="1942">Verhoeff (1942)</bibRefCitation>
; medianen Sternitfortsatzes/mittlerer Sternitfortsatz
<emphasis id="B97FC262FFA5FF9EFC45FE3BA8C8FE39" box="[955,1017,477,500]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFBF9FE3AAF83FE38" author="Strasser" box="[1031,1202,476,501]" pageId="19" pageNumber="170" refString="Strasser, K. (1962) Diplopoden aus Bulgarien und Jugoslawien. Senckenbergiana biologica, 43, 437 - 470." type="journal article" year="1962">Strasser (1962</bibRefCitation>
,
<bibRefCitation id="EF9A6381FFA5FF9EFB3AFE3AAE3FFE38" author="Strasser" box="[1220,1294,476,501]" pageId="19" pageNumber="170" refString="Strasser, K. (1971 b) Uber Diplopoden Jugoslawiens. Senckenbergiana biologica, 53, 313 - 345." type="journal article" year="1971" yearSuffix="b">1971b</bibRefCitation>
); or medial sternite process
<emphasis id="B97FC262FFA5FF9EFEA4FDE7AAA9FDD5" box="[346,408,513,536]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFE58FE19A977FDD5" author="Mrsic" box="[422,582,511,536]" pageId="19" pageNumber="170" refString="Mrsic, N. (1992) Biokoviella mauriesi gen. nov., sp. nov. (Biokoviellidae fam. nov.), Macrochaetosomatinae subf. nov. (Anthogonidae) and superfamily Cleidogonoidea of the western Balkans (Craspedosomidea, Diplopoda). Razprave IV. Razreda SAZU, 33 (3) 51 - 91." type="journal article" year="1992">Mršić (1992)</bibRefCitation>
]; and lateral coxal process (
<emphasis id="B97FC262FFA5FF9EFC5EFE19A8F5FDD5" bold="true" box="[928,964,511,536]" pageId="19" pageNumber="170">lcp</emphasis>
) on both sides. The medial process is cylindrical, strongly curved posteriorly, and bifurcated distally, with a massive, bilobed basal bulge (
<emphasis id="B97FC262FFA5FF9EFB04FDC2AE28FDF0" bold="true" box="[1274,1305,548,573]" pageId="19" pageNumber="170">bb</emphasis>
). The basal and middle parts are wrinkled.
<emphasis id="B97FC262FFA5FF9EFDFBFDAFA97EFDAD" box="[517,591,585,608]" italics="true" pageId="19" pageNumber="170">In situ</emphasis>
the bifurcated part abuts on the sternal process of leg pair 10. Both
<emphasis id="B97FC262FFA5FF9EFA86FDA1AEADFDAD" bold="true" box="[1400,1436,583,608]" pageId="19" pageNumber="170">lcp</emphasis>
consists of two parts: an inner part (
<emphasis id="B97FC262FFA5FF9EFDBEFD8AA955FD48" bold="true" box="[576,612,620,645]" pageId="19" pageNumber="170">cip</emphasis>
) [= coxale Hörner
<emphasis id="B97FC262FFA5FF9EFCB6FD8BA8B7FD49" box="[840,902,621,644]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFC6EFD8AAF78FD49" author="Verhoeff" box="[912,1097,620,645]" pageId="19" pageNumber="170" refString="Verhoeff, K. W. (1942) Eine neue, westbalkanische, cavernicole Familie der Ascospermophora und die fragwurdige Gattung Macrochaetosoma Abs. Zoologischer Anzeiger, 139, 54 - 66." type="journal article" year="1942">Verhoeff (1942)</bibRefCitation>
; innerer Fortsatz der coxalen Seitenteile/inneren Coxitfortsatz
<emphasis id="B97FC262FFA5FF9EFDF3FD77A97AFD65" box="[525,587,657,680]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFDAAFD76A9C8FD65" author="Strasser" box="[596,761,655,680]" pageId="19" pageNumber="170" refString="Strasser, K. (1962) Diplopoden aus Bulgarien und Jugoslawien. Senckenbergiana biologica, 43, 437 - 470." type="journal article" year="1962">Strasser (1962</bibRefCitation>
,
<bibRefCitation id="EF9A6381FFA5FF9EFCF8FD69A861FD65" author="Strasser" box="[774,848,655,680]" pageId="19" pageNumber="170" refString="Strasser, K. (1971 b) Uber Diplopoden Jugoslawiens. Senckenbergiana biologica, 53, 313 - 345." type="journal article" year="1971" yearSuffix="b">1971b</bibRefCitation>
); or paramedial coxal process
<emphasis id="B97FC262FFA5FF9EFB51FD77AFDCFD65" box="[1199,1261,657,680]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFB08FD69AEA1FD65" author="Mrsic" box="[1270,1424,655,680]" pageId="19" pageNumber="170" refString="Mrsic, N. (1992) Biokoviella mauriesi gen. nov., sp. nov. (Biokoviellidae fam. nov.), Macrochaetosomatinae subf. nov. (Anthogonidae) and superfamily Cleidogonoidea of the western Balkans (Craspedosomidea, Diplopoda). Razprave IV. Razreda SAZU, 33 (3) 51 - 91." type="journal article" year="1992">Mršić (1992)</bibRefCitation>
]; and an outer part (
<emphasis id="B97FC262FFA5FF9EFE97FD52AAA2FD00" bold="true" box="[361,403,692,717]" pageId="19" pageNumber="170">cop</emphasis>
) [= äußerer coxale Lappen
<emphasis id="B97FC262FFA5FF9EFD2EFD53A83FFD01" box="[720,782,693,716]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFCE9FD52A8FFFD01" author="Verhoeff" box="[791,974,692,717]" pageId="19" pageNumber="170" refString="Verhoeff, K. W. (1942) Eine neue, westbalkanische, cavernicole Familie der Ascospermophora und die fragwurdige Gattung Macrochaetosoma Abs. Zoologischer Anzeiger, 139, 54 - 66." type="journal article" year="1942">Verhoeff (1942)</bibRefCitation>
; äußerer Fortsatz der coxalen Seitenteile
<emphasis id="B97FC262FFA5FF9EFF69FD3FABE4FD3D" box="[151,213,729,752]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFF23FD3EAABAFD3D" author="Strasser" box="[221,395,727,752]" pageId="19" pageNumber="170" refString="Strasser, K. (1962) Diplopoden aus Bulgarien und Jugoslawien. Senckenbergiana biologica, 43, 437 - 470." type="journal article" year="1962">Strasser (1962)</bibRefCitation>
; Seitenblatt
<emphasis id="B97FC262FFA5FF9EFDE4FD3FA969FD3D" box="[538,600,729,752]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFD9EFD3EA82CFD3D" author="Strasser" box="[608,797,727,752]" pageId="19" pageNumber="170" refString="Strasser, K. (1971 b) Uber Diplopoden Jugoslawiens. Senckenbergiana biologica, 53, 313 - 345." type="journal article" year="1971" yearSuffix="b">Strasser (1971b)</bibRefCitation>
; or lateral coxal process
<emphasis id="B97FC262FFA5FF9EFBC4FD3FAF49FD3D" box="[1082,1144,729,752]" italics="true" pageId="19" pageNumber="170">sensu</emphasis>
<bibRefCitation id="EF9A6381FFA5FF9EFB7FFD31AE2AFD3D" author="Mrsic" box="[1153,1307,727,752]" pageId="19" pageNumber="170" refString="Mrsic, N. (1992) Biokoviella mauriesi gen. nov., sp. nov. (Biokoviellidae fam. nov.), Macrochaetosomatinae subf. nov. (Anthogonidae) and superfamily Cleidogonoidea of the western Balkans (Craspedosomidea, Diplopoda). Razprave IV. Razreda SAZU, 33 (3) 51 - 91." type="journal article" year="1992">Mršić (1992)</bibRefCitation>
]. The inner parts of the coxal processes are the highest parts of the whole gonopodal block. These structures are curved anteriorly. The first 2/3 from anterior or posterior view are broadened, mainly smooth, while the apical 1/3 is narrower, bearing postero-mesal ribs (
<emphasis id="B97FC262FFA5FF9EFDB0FCA2A96BFC90" bold="true" box="[590,602,836,861]" pageId="19" pageNumber="170">r</emphasis>
) and triangular denticles (
<emphasis id="B97FC262FFA5FF9EFC73FCA2A8ADFC90" bold="true" box="[909,924,836,861]" pageId="19" pageNumber="170">d</emphasis>
). The outer parts of the coxal processes are wide, consisting of two smooth lobes, larger and smaller. The inner and outer parts are divided from each other by a deep gap. There are no traces of telopodal remnants; the border between the inner and outer parts is marked only with a fissure (
<emphasis id="B97FC262FFA5FF9EFEC0FC49AA79FC05" bold="true" box="[318,328,943,968]" pageId="19" pageNumber="170">f</emphasis>
).
</paragraph>
<paragraph id="8BB41E70FFA5FF9EFF39FC32AE68FBDD" blockId="19.[151,1437,151,1040]" pageId="19" pageNumber="170">
<emphasis id="B97FC262FFA5FF9EFF39FC32AA1BFC20" bold="true" box="[199,298,980,1005]" pageId="19" pageNumber="170">Habitat.</emphasis>
Gorniča Pit is one of the deepest pits in
<collectingCountry id="F31C5EE0FFA5FF9EFCFDFC33A8BEFC20" box="[771,911,981,1005]" name="Serbia" pageId="19" pageNumber="170">Montenegro</collectingCountry>
, with a depth of
<quantity id="4CF3B395FFA5FF9EFBADFC32AFADFC21" box="[1107,1180,980,1005]" metricMagnitude="2" metricUnit="m" metricValue="5.16" pageId="19" pageNumber="170" unit="m" value="516.0">516 m</quantity>
and length of
<quantity id="4CF3B395FFA5FF9EFABBFC32AEADFC21" box="[1349,1436,980,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.629" pageId="19" pageNumber="170" unit="m" value="1629.0">1629 m</quantity>
(http://www.speleologija.me/p/najdublje-jame-u-crnoj-gori.html). Unfortunately, no further data is available.
</paragraph>
</subSubSection>
</treatment>
</document>