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<document id="9DBB843DE4E3CFCC7021EEA71FC802FE" ID-DOI="10.5852/ejt.2024.933.2525" ID-ISSN="2118-9773" ID-Zenodo-Dep="11198874" ID-ZooBank="8F720F2B-BFBC-4CA1-BFF2-A2B8C7C8D3E1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1715771531038" checkinUser="plazi" docAuthor="Stöhr, Sabine" docDate="2024" docId="038987D9FFA3D75D0006C1591CDAF97D" docLanguage="en" docName="EJT.2024.933.1-98.pdf" docOrigin="European Journal of Taxonomy 933" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2525/11335" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophionereis Lutken 1859" docType="treatment" docVersion="3" lastPageNumber="50" masterDocId="FFB0FFA1FF92D76C0253C0121F07FFDA" masterDocTitle="Taxonomic analysis of the genital plates and associated structures in Ophiuroidea (Echinodermata)" masterLastPageNumber="98" masterPageNumber="1" pageNumber="50" updateTime="1715783907690" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="37814759DDD416E0885B98B3A72A7F45">Taxonomic analysis of the genital plates and associated structures in Ophiuroidea (Echinodermata)</mods:title>
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<subSubSection id="C33A6544FFA3D75D0006C1591CD8FEBF" box="[597,991,330,357]" pageId="49" pageNumber="50" type="nomenclature">
<paragraph id="8B9F36CFFFA3D75D0006C1591CD8FEBF" blockId="49.[597,991,330,392]" box="[597,991,330,357]" pageId="49" pageNumber="50">
<heading id="D0D781A3FFA3D75D0006C1591CD8FEBF" box="[597,991,330,357]" centered="true" fontSize="11" level="2" pageId="49" pageNumber="50" reason="2">
Genus
<taxonomicName id="4C204D4CFFA3D75D00F4C1581CD8FEBF" ID-CoL="7PNDM" authority="Lutken, 1859" authorityName="Lutken" authorityYear="1859" box="[679,991,330,357]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="genus">
<emphasis id="B954EADDFFA3D75D00F4C1581C3CFEBF" bold="true" box="[679,827,330,357]" italics="true" pageId="49" pageNumber="50">Ophionereis</emphasis>
Lütken, 1859
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33A6544FFA3D75D00A3C17C1C44FE52" box="[752,835,366,392]" pageId="49" pageNumber="50" type="description">
<paragraph id="8B9F36CFFFA3D75D00A3C17C1C44FE52" blockId="49.[597,991,330,392]" box="[752,835,366,392]" pageId="49" pageNumber="50">
<figureCitation id="131B2A4AFFA3D75D00A3C17C1C44FE52" box="[752,835,366,392]" captionStart="Fig" captionStartId="50.[189,232,1583,1609]" captionTargetBox="[189,1399,265,1534]" captionTargetId="figure-126@50.[189,1399,265,1534]" captionTargetPageId="50" captionText="Fig. 23. Ophionereididae Ljungman, 1867, SEM, unless otherwise noted. AJ. Ophionereis reticulata (Say, 1825).A. Oral aspect, digital image. BC. Micro-CT.B. Oral shield with oGP, inner view.C. abGp, adGB and RS in situ. D. abGP and adGP in situ. E. adPG, abradial aspect. F. abGP. G. Radial shield, inner aspect. H. Oral shield, inner aspect. I. Madreporite, inner aspect. J. Oral genital plates, one with round tubercle/process. KM. Ophionereis porrecta (Lyman, 1860). K. Radial shield, inner aspect. L. adGP, abradial aspect. M. abGP. NP. Ophionereis degeneri (A.H. Clark, 1949). N. Radial shield, inner aspect. O. adGP, abradial aspect. P. abGP. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11198926" httpUri="https://zenodo.org/record/11198926/files/figure.png" pageId="49" pageNumber="50">Fig. 23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33A6544FFA3D75D02EEC1BD1CDAF97D" pageId="49" pageNumber="50" type="type_taxon">
<paragraph id="8B9F36CFFFA3D75D02EEC1BD1E5FFE13" blockId="49.[189,344,431,457]" box="[189,344,431,457]" pageId="49" pageNumber="50">
<heading id="D0D781A3FFA3D75D02EEC1BD1E5FFE13" bold="true" box="[189,344,431,457]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B954EADDFFA3D75D02EEC1BD1E5FFE13" bold="true" box="[189,344,431,457]" pageId="49" pageNumber="50">
<typeStatus id="549B886DFFA3D75D02EEC1BD1FFDFE13" box="[189,250,431,457]" pageId="49" pageNumber="50">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9F36CFFFA3D75D02EEC1CF1D1CFE22" blockId="49.[189,539,477,504]" box="[189,539,477,504]" pageId="49" pageNumber="50">
<taxonomicName id="4C204D4CFFA3D75D02EEC1CF1D10FE22" authority="Say, 1825" authorityName="Say" authorityYear="1825" box="[189,535,477,504]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="reticulata">
<emphasis id="B954EADDFFA3D75D02EEC1CF1E9EFE2D" box="[189,409,477,503]" italics="true" pageId="49" pageNumber="50">Ophiura reticulata</emphasis>
Say, 1825
</taxonomicName>
.
</paragraph>
<paragraph id="8B9F36CFFFA3D75D02EEC20D1E9EFDE3" blockId="49.[189,409,543,569]" box="[189,409,543,569]" pageId="49" pageNumber="50">
<heading id="D0D781A3FFA3D75D02EEC20D1E9EFDE3" bold="true" box="[189,409,543,569]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B954EADDFFA3D75D02EEC20D1E9EFDE3" bold="true" box="[189,409,543,569]" pageId="49" pageNumber="50">Examined species</emphasis>
</heading>
</paragraph>
<paragraph id="8B9F36CFFFA3D75D02EEC25F1B83FDB2" blockId="49.[189,1156,589,616]" box="[189,1156,589,616]" pageId="49" pageNumber="50">
<taxonomicName id="4C204D4CFFA3D75D02EEC25F1EC2FDBD" ID-CoL="49YRC" baseAuthorityName="Say" baseAuthorityYear="1825" box="[189,453,589,615]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="reticulata">
<emphasis id="B954EADDFFA3D75D02EEC25F1EC2FDBD" box="[189,453,589,615]" italics="true" pageId="49" pageNumber="50">Ophionereis reticulata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C204D4CFFA3D75D0387C25C1C07FDBD" ID-CoL="49YRB" authority="Lyman, 1860" authorityName="Lyman" authorityYear="1860" box="[468,768,589,616]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="porrecta">
<emphasis id="B954EADDFFA3D75D0387C25C1D5BFDBD" box="[468,604,590,616]" italics="true" pageId="49" pageNumber="50">O. porrecta</emphasis>
Lyman, 1860
</taxonomicName>
,
<taxonomicName id="4C204D4CFFA3D75D015FC25C1B87FDB2" ID-CoL="6T27M" authority="(A. H. Clark, 1949)" baseAuthorityName="A. H. Clark" baseAuthorityYear="1949" box="[780,1152,589,616]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="degeneri">
<emphasis id="B954EADDFFA3D75D015FC25C1C90FDBD" box="[780,919,589,616]" italics="true" pageId="49" pageNumber="50">O. degeneri</emphasis>
(A.H. Clark, 1949)
</taxonomicName>
.
</paragraph>
<paragraph id="8B9F36CFFFA3D75D02EEC29D1E2FFD73" blockId="49.[189,296,655,681]" box="[189,296,655,681]" pageId="49" pageNumber="50">
<heading id="D0D781A3FFA3D75D02EEC29D1E2FFD73" bold="true" box="[189,296,655,681]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B954EADDFFA3D75D02EEC29D1E2FFD73" bold="true" box="[189,296,655,681]" pageId="49" pageNumber="50">Oral GP</emphasis>
</heading>
</paragraph>
<paragraph id="8B9F36CFFFA3D75D02EEC2AF1A36FCC4" blockId="49.[189,1398,701,798]" pageId="49" pageNumber="50">
Elongated oval, flat, thin, proximal end under OS, pair not meeting (only examined in
<taxonomicName id="4C204D4CFFA3D75D06FDC2AC1A44FD0D" ID-CoL="49YRC" baseAuthorityName="Say" baseAuthorityYear="1825" box="[1198,1347,701,728]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="reticulata">
<emphasis id="B954EADDFFA3D75D06FDC2AC1BC4FD02" box="[1198,1219,702,728]" italics="true" pageId="49" pageNumber="50">O</emphasis>
.
<emphasis id="B954EADDFFA3D75D0681C2AF1A44FD0D" box="[1234,1347,701,727]" italics="true" pageId="49" pageNumber="50">reticulata</emphasis>
</taxonomicName>
, but externally visible in
<taxonomicName id="4C204D4CFFA3D75D03E0C2F31D3AFD21" ID-CoL="6T27M" baseAuthorityName="A. H. Clark" baseAuthorityYear="1949" box="[435,573,737,763]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="degeneri">
<emphasis id="B954EADDFFA3D75D03E0C2F31EC0FD21" box="[435,455,737,763]" italics="true" pageId="49" pageNumber="50">O</emphasis>
.
<emphasis id="B954EADDFFA3D75D0385C2F31D3AFD21" box="[470,573,737,763]" italics="true" pageId="49" pageNumber="50">degeneri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C204D4CFFA3D75D002AC2F31C05FD20" ID-CoL="49YRB" authorityName="Lyman" authorityYear="1860" box="[633,770,737,763]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="porrecta">
<emphasis id="B954EADDFFA3D75D002AC2F31D89FD21" box="[633,654,737,763]" italics="true" pageId="49" pageNumber="50">O</emphasis>
.
<emphasis id="B954EADDFFA3D75D00CDC2F31C05FD20" box="[670,770,737,762]" italics="true" pageId="49" pageNumber="50">porrecta</emphasis>
</taxonomicName>
). On some plates a round lateral process. Beset with small round granules/papillae at abradial edge. Large scales along the genital slit connect to oGP.
</paragraph>
<paragraph id="8B9F36CFFFA3D75D02EEC3571E59FC85" blockId="49.[189,350,837,863]" box="[189,350,837,863]" pageId="49" pageNumber="50">
<heading id="D0D781A3FFA3D75D02EEC3571E59FC85" bold="true" box="[189,350,837,863]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B954EADDFFA3D75D02EEC3571E59FC85" bold="true" box="[189,350,837,863]" pageId="49" pageNumber="50">Adradial GP</emphasis>
</heading>
</paragraph>
<paragraph id="8B9F36CFFFA3D75D02EEC3661D12FB5F" blockId="49.[189,1401,884,1157]" pageId="49" pageNumber="50">
<taxonomicName id="4C204D4CFFA3D75D02EEC3661ECFFC54" ID-CoL="49YRC" baseAuthorityName="Say" baseAuthorityYear="1825" box="[189,456,884,910]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="reticulata">
<emphasis id="B954EADDFFA3D75D02EEC3661ECFFC54" box="[189,456,884,910]" italics="true" pageId="49" pageNumber="50">Ophionereis reticulata</emphasis>
</taxonomicName>
: weakly cleaver-shaped with wavy abradial edge. Proximal half rod-like with longitudinal groove, distal half widened rectangular, protruding abradial edge wavy, articulates with ventral surface to abGP. Abradial surface with small adradial process with coarser stereom, oval hole in distal part. Distal end curved dorsalwards, dorsally rounded, no distinct condyle, but a groove, distal edge concave. Proximal ends of adGP and abGP crossing.
<taxonomicName id="4C204D4CFFA3D75D01D1C4131B87FBC0" ID-CoL="49YRB" authorityName="Lyman" authorityYear="1860" box="[898,1152,1025,1051]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="porrecta">
<emphasis id="B954EADDFFA3D75D01D1C4131B87FBC0" box="[898,1152,1025,1051]" italics="true" pageId="49" pageNumber="50">Ophionereis porrecta</emphasis>
</taxonomicName>
: widened projection smaller, cleaver shape less obvious; distally a condylar process terminating flat.
<taxonomicName id="4C204D4CFFA3D75D0623C4361A68FBE4" ID-CoL="6T27M" baseAuthorityName="A. H. Clark" baseAuthorityYear="1949" box="[1136,1391,1060,1086]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="degeneri">
<emphasis id="B954EADDFFA3D75D0623C4361A68FBE4" box="[1136,1391,1060,1086]" italics="true" pageId="49" pageNumber="50">Ophionereis degeneri</emphasis>
</taxonomicName>
: cleaver shape slightly more pronounced than in
<taxonomicName id="4C204D4CFFA3D75D00A1C4551C7DFBBB" ID-CoL="49YRB" authorityName="Lyman" authorityYear="1860" box="[754,890,1095,1122]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="porrecta">
<emphasis id="B954EADDFFA3D75D00A1C4551C00FBBB" box="[754,775,1095,1121]" italics="true" pageId="49" pageNumber="50">O</emphasis>
.
<emphasis id="B954EADDFFA3D75D0145C45A1C7DFBBB" box="[790,890,1096,1121]" italics="true" pageId="49" pageNumber="50">porrecta</emphasis>
</taxonomicName>
, but not as strong as in
<taxonomicName id="4C204D4CFFA3D75D06C1C4551A20FBBB" ID-CoL="49YRC" baseAuthorityName="Say" baseAuthorityYear="1825" box="[1170,1319,1095,1122]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="reticulata">
<emphasis id="B954EADDFFA3D75D06C1C4551BA0FBBB" box="[1170,1191,1095,1121]" italics="true" pageId="49" pageNumber="50">O</emphasis>
.
<emphasis id="B954EADDFFA3D75D06E5C4551A20FBBB" box="[1206,1319,1095,1121]" italics="true" pageId="49" pageNumber="50">reticulata</emphasis>
</taxonomicName>
; distal condylar process and groove.
</paragraph>
<paragraph id="8B9F36CFFFA3D75D02EEC4B91E59FB1F" blockId="49.[189,350,1195,1221]" box="[189,350,1195,1221]" pageId="49" pageNumber="50">
<heading id="D0D781A3FFA3D75D02EEC4B91E59FB1F" bold="true" box="[189,350,1195,1221]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B954EADDFFA3D75D02EEC4B91E59FB1F" bold="true" box="[189,350,1195,1221]" pageId="49" pageNumber="50">Abradial GP</emphasis>
</heading>
</paragraph>
<paragraph id="8B9F36CFFFA3D75D02EEC4C81E22FA58" blockId="49.[189,1398,1242,1410]" pageId="49" pageNumber="50">
<taxonomicName id="4C204D4CFFA3D75D02EEC4C81ECFFB2E" ID-CoL="49YRC" baseAuthorityName="Say" baseAuthorityYear="1825" box="[189,456,1242,1268]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="reticulata">
<emphasis id="B954EADDFFA3D75D02EEC4C81ECFFB2E" box="[189,456,1242,1268]" italics="true" pageId="49" pageNumber="50">Ophionereis reticulata</emphasis>
</taxonomicName>
: sabre-shaped, as long as adGP, sligthly curved, dorsal edge concave, ventral edge convex, with furrow in abradial side, distal third dorsally flattened concave, articulates to adGP, proximal end crosses adGP.
<taxonomicName id="4C204D4CFFA3D75D0059C5331C02FAE0" ID-CoL="49YRB" authorityName="Lyman" authorityYear="1860" box="[522,773,1313,1339]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="porrecta">
<emphasis id="B954EADDFFA3D75D0059C5331C02FAE0" box="[522,773,1313,1339]" italics="true" pageId="49" pageNumber="50">Ophionereis porrecta</emphasis>
</taxonomicName>
: thin, flat, curved, distally widest with slanting edge, proximally tapered to round point.
<taxonomicName id="4C204D4CFFA3D75D003EC5561C68FA84" ID-CoL="6T27M" baseAuthorityName="A. H. Clark" baseAuthorityYear="1949" box="[621,879,1348,1374]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="degeneri">
<emphasis id="B954EADDFFA3D75D003EC5561C68FA84" box="[621,879,1348,1374]" italics="true" pageId="49" pageNumber="50">Ophionereis degeneri</emphasis>
</taxonomicName>
: similar to
<taxonomicName id="4C204D4CFFA3D75D0651C5561B89FA84" ID-CoL="49YRB" authorityName="Lyman" authorityYear="1860" box="[1026,1166,1348,1374]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="porrecta">
<emphasis id="B954EADDFFA3D75D0651C5561B10FA84" box="[1026,1047,1348,1374]" italics="true" pageId="49" pageNumber="50">O</emphasis>
.
<emphasis id="B954EADDFFA3D75D0679C5571B89FA84" box="[1066,1166,1349,1374]" italics="true" pageId="49" pageNumber="50">porrecta</emphasis>
</taxonomicName>
, but articular edge concave.
</paragraph>
<paragraph id="8B9F36CFFFA3D75D02EEC5BA1E53FA18" blockId="49.[189,340,1448,1474]" box="[189,340,1448,1474]" pageId="49" pageNumber="50">
<heading id="D0D781A3FFA3D75D02EEC5BA1E53FA18" bold="true" box="[189,340,1448,1474]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B954EADDFFA3D75D02EEC5BA1E53FA18" bold="true" box="[189,340,1448,1474]" pageId="49" pageNumber="50">Oral shields</emphasis>
</heading>
</paragraph>
<paragraph id="8B9F36CFFFA3D75D02EEC5C51CAAF9CE" blockId="49.[189,1398,1495,1556]" pageId="49" pageNumber="50">
On inner surface, near distal edge a spur to either side. Madreporite in distal half inflated, with large opening, proximal half thinner (only examined in
<taxonomicName id="4C204D4CFFA3D75D015BC5E81C9BF9CE" ID-CoL="49YRC" baseAuthorityName="Say" baseAuthorityYear="1825" box="[776,924,1530,1556]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="reticulata">
O.
<emphasis id="B954EADDFFA3D75D0178C5E81C9BF9CE" box="[811,924,1530,1556]" italics="true" pageId="49" pageNumber="50">reticulata</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B9F36CFFFA3D75D02EEC6291E67F98F" blockId="49.[189,352,1595,1621]" box="[189,352,1595,1621]" pageId="49" pageNumber="50">
<heading id="D0D781A3FFA3D75D02EEC6291E67F98F" bold="true" box="[189,352,1595,1621]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B954EADDFFA3D75D02EEC6291E67F98F" bold="true" box="[189,352,1595,1621]" pageId="49" pageNumber="50">Radial shield</emphasis>
</heading>
</paragraph>
<paragraph id="8B9F36CFFFA3D75D02EEC6781CDAF97D" blockId="49.[189,1398,1642,1704]" pageId="49" pageNumber="50">
Single condyle and coarser round patch at some distance proximal to condyle in all three examined species, condyle more pronounced in
<taxonomicName id="4C204D4CFFA3D75D002BC69C1C04F97D" ID-CoL="6T27M" baseAuthorityName="A. H. Clark" baseAuthorityYear="1949" box="[632,771,1677,1704]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" kingdom="Animalia" order="Amphilepidida" pageId="49" pageNumber="50" phylum="Echinodermata" rank="species" species="degeneri">
<emphasis id="B954EADDFFA3D75D002BC69C1D8AF972" box="[632,653,1678,1704]" italics="true" pageId="49" pageNumber="50">O</emphasis>
.
<emphasis id="B954EADDFFA3D75D00CFC69F1C04F97D" box="[668,771,1677,1703]" italics="true" pageId="49" pageNumber="50">degeneri</emphasis>
</taxonomicName>
than in the others.
</paragraph>
</subSubSection>
</treatment>
</document>