640 lines
66 KiB
XML
640 lines
66 KiB
XML
<document id="BF59E94D54374CC68D4071EA3AE98462" ID-DOI="10.11646/zootaxa.4006.1.3" ID-GBIF-Dataset="a73584b0-b729-482f-b91f-f0a42bb25591" ID-ISSN="1175-5326" ID-Zenodo-Dep="289264" ID-ZooBank="14516538-9B91-4363-9541-0AF373F9B614" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461202830199" checkinUser="plazi" docAuthor="Thomson, Scott, Amepou, Yolarnie, Anamiato, Jim & Georges, Arthur" docDate="2015" docId="0389878AA812FFE7FF01FF4EFB1BFC5B" docLanguage="en" docName="zt04006p082.pdf" docOrigin="Zootaxa 4006 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Elseya (Hanwarachelys) rhodini Thomson, Amepou, Anamiato & Georges, 2015, sp. nov." docType="treatment" docUuid="AFC2DBEB-C86B-407C-AD5C-2E0D7D130F2F" docUuidSource="ZooBank" docVersion="6" lastPageNumber="73" masterDocId="FFB0FFF2A818FFE9FF96FFD9FFABFFDD" masterDocTitle="A new species and subgenus of Elseya (Testudines: Pleurodira: Chelidae) from New Guinea" masterLastPageNumber="82" masterPageNumber="59" pageNumber="69" updateTime="1698601124389" updateUser="plazi">
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<mods:titleInfo id="62BA6BE42936732081F7FB91FC7C6FDF">
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<mods:title id="90CFEEE56D3B95F1BD3974546A382C30">A new species and subgenus of Elseya (Testudines: Pleurodira: Chelidae) from New Guinea</mods:title>
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</mods:titleInfo>
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<mods:name id="E2A73D5C8E7628C00AA96B5653BA95E0" type="personal">
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<mods:roleTerm id="2926BB98694C76E4B8EEDB6742CF57E4">Author</mods:roleTerm>
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<mods:namePart id="54EE8CC8EE5F24B22C96AA24789D997C">Thomson, Scott</mods:namePart>
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<mods:roleTerm id="7BE352253C987D70130591DBCB2D61DF">Author</mods:roleTerm>
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<mods:namePart id="0628E29A12CC471E390101B8698AD3BF">Amepou, Yolarnie</mods:namePart>
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<mods:roleTerm id="C2583061725077F8249A91A58881475C">Author</mods:roleTerm>
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<mods:namePart id="6FA6CCF6237D22A1092CDE634193A191">Anamiato, Jim</mods:namePart>
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<mods:namePart id="C110EB48FF9A6AA2A6DB302D70012571">Georges, Arthur</mods:namePart>
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<mods:title id="2EF328ED0BF6F373FBF12D19FBF16DDE">Zootaxa</mods:title>
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<mods:date id="A0B2F99B40B93500A935845DD69BD9FA">2015</mods:date>
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<mods:number id="102D7F576783A6481C78D36446687665">4006</mods:number>
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<treatment id="0389878AA812FFE7FF01FF4EFB1BFC5B" ID-DOI="http://doi.org/10.5281/zenodo.6108721" ID-GBIF-Taxon="127692838" ID-Zenodo-Dep="6108721" LSID="urn:lsid:zoobank.org:act:AFC2DBEB-C86B-407C-AD5C-2E0D7D130F2F" httpUri="http://treatment.plazi.org/id/0389878AA812FFE7FF01FF4EFB1BFC5B" lastPageId="14" lastPageNumber="73" pageId="10" pageNumber="69">
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<subSubSection id="C33A6517A812FFE3FF01FF4EFD7BFED1" pageId="10" pageNumber="69" type="nomenclature">
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<paragraph id="8B9F369CA812FFE3FF01FF4EFD2FFF6F" blockId="10.[151,644,151,178]" box="[151,644,151,178]" pageId="10" pageNumber="69">
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<heading id="D0D781F0A812FFE3FF01FF4EFD2FFF6F" bold="true" box="[151,644,151,178]" fontSize="11" level="1" pageId="10" pageNumber="69" reason="1">
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<emphasis id="B954EA8EA812FFE3FF01FF4EFD2FFF6F" bold="true" box="[151,644,151,178]" pageId="10" pageNumber="69">
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<taxonomicName id="4C204D1FA812FFE3FF01FF4EFDB4FF6C" box="[151,543,151,178]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="rhodini" status="sp. nov." subGenus="Hanwarachelys">
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<emphasis id="B954EA8EA812FFE3FF01FF4EFF4CFF6C" bold="true" box="[151,231,151,177]" italics="true" pageId="10" pageNumber="69">Elseya</emphasis>
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(
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<emphasis id="B954EA8EA812FFE3FF6EFF4EFE1EFF6C" bold="true" box="[248,437,151,177]" italics="true" pageId="10" pageNumber="69">Hanwarachelys</emphasis>
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)
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<emphasis id="B954EA8EA812FFE3FE53FF4EFDB4FF6C" bold="true" box="[453,543,151,177]" italics="true" pageId="10" pageNumber="69">rhodini</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26757F5A812FFE3FDB0FF41FD2FFF6F" box="[550,644,152,178]" pageId="10" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
<paragraph id="8B9F369CA812FFE3FF01FF01FDA3FF33" blockId="10.[151,720,216,268]" box="[151,520,216,238]" pageId="10" pageNumber="69">
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Southern New
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<collectingCountry id="F337760CA812FFE3FEB9FF01FED3FF33" box="[303,376,216,238]" name="Guinea-Bissau" pageId="10" pageNumber="69">Guinea</collectingCountry>
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Stream Turtle
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</paragraph>
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<paragraph id="8B9F369CA812FFE3FF01FF2FFD7BFED1" blockId="10.[151,720,216,268]" box="[151,720,246,268]" pageId="10" pageNumber="69">
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2015
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<taxonomicName id="4C204D1FA812FFE3FF46FF2FFECFFED1" box="[208,356,246,268]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="rhodini" status="sp. nov.">
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<emphasis id="B954EA8EA812FFE3FF46FF2FFECFFED1" box="[208,356,246,268]" italics="true" pageId="10" pageNumber="69">Elseya rhodini</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26757F5A812FFE3FEFCFF2EFE10FED1" box="[362,443,247,268]" pageId="10" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
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, Thomson et al., this study.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A6517A812FFE3FF01FEEBFEB7FE06" pageId="10" pageNumber="69" type="materials_examined">
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||
<paragraph id="8B9F369CA812FFE3FF01FEEBFEB7FE06" blockId="10.[151,1437,306,2032]" pageId="10" pageNumber="69">
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||
<emphasis id="B954EA8EA812FFE3FF01FEEBFEA6FE96" bold="true" box="[151,269,306,331]" pageId="10" pageNumber="69">
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||
<typeStatus id="549B883EA812FFE3FF01FEEBFF79FE96" box="[151,210,306,331]" pageId="10" pageNumber="69">Type</typeStatus>
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data
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</emphasis>
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: Holotype—PNGM R25204 (
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<figureCitation id="131B2A19A812FFE3FDC8FEEBFD15FE96" box="[606,702,306,331]" captionStart="FIGURE 8" captionStartId="11.[151,250,1794,1816]" captionTargetBox="[190,1412,215,1757]" captionTargetId="figure@11.[151,1437,195,1770]" captionTargetPageId="11" captionText="FIGURE 8. Dorsal, ventral and lateral views of the holotype PNGM R 25204 of Elseya (Hanwarachelys) rhodini from Wau Creek, Kikori Drainage, Papua New Guinea. Scale: 20 mm." httpUri="https://zenodo.org/record/289270/files/figure.png" pageId="10" pageNumber="69">Figure 8</figureCitation>
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), adult female (carapace length:
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<quantity id="4CD89B79A812FFE3FBBAFEEBFB26FE96" box="[1068,1165,306,331]" metricMagnitude="-1" metricUnit="m" metricValue="2.55" pageId="10" pageNumber="69" unit="mm" value="255.0">255 mm</quantity>
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), collected at Rue Creek (tributary of Wau Creek), Gulf Province,
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<collectingCountry id="F337760CA812FFE3FDF4FE8EFC92FEB2" box="[610,825,342,367]" name="Papua New Guinea" pageId="10" pageNumber="69">Papua New Guinea</collectingCountry>
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(07˚11’ 67.3” S, 144˚37’ 13.8” E) by Yolanie Amepou on
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<date id="FF9E105CA812FFE3FF2BFEA3FE85FE4E" box="[189,302,378,403]" pageId="10" pageNumber="69">6 January</date>
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, 2015 (
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||
<figureCitation id="131B2A19A812FFE3FE11FEA3FE47FE4E" box="[391,492,378,403]" captionStart="FIGURE 9" captionStartId="12.[151,250,1183,1205]" captionTargetBox="[151,1436,195,1159]" captionTargetId="figure@12.[151,1436,195,1159]" captionTargetPageId="12" captionText="FIGURE 9. Habitat of Elseya rhodini Wau Creek, Sirebi River, Kikori drainage (07 ˚ 11 ’ 67.3 ” S, 144 ˚ 37 ’ 13.8 ” E)." httpUri="https://zenodo.org/record/289271/files/figure.png" pageId="10" pageNumber="69">Figure 9</figureCitation>
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).
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<typeStatus id="549B883EA812FFE3FD92FEA2FDC0FE4E" box="[516,619,379,403]" pageId="10" pageNumber="69" type="paratype">Paratype</typeStatus>
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:
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<collectionCode id="ED31AE59A812FFE3FDEFFEA2FD61FE4E" box="[633,714,379,403]" pageId="10" pageNumber="69">PNGM</collectionCode>
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||
R25203, adult male (carapace length:
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||
<quantity id="4CD89B79A812FFE3FB06FEA3FB5FFE4E" box="[1168,1268,378,403]" metricMagnitude="-1" metricUnit="m" metricValue="2.05" pageId="10" pageNumber="69" unit="mm" value="205.0">205 mm</quantity>
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||
), herein set as Allotype, collection data as per
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<typeStatus id="549B883EA812FFE3FE6DFE47FDCCFE6A" box="[507,615,414,439]" pageId="10" pageNumber="69" type="holotype">Holotype</typeStatus>
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||
. For basic measurements of these
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<typeStatus id="549B883EA812FFE3FC70FE46FB8AFE6A" box="[998,1057,415,439]" pageId="10" pageNumber="69">types</typeStatus>
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and others from the genus
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<taxonomicName id="4C204D1FA812FFE3FAC7FE47FA37FE6B" box="[1361,1436,414,438]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="genus">
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<emphasis id="B954EA8EA812FFE3FAC7FE47FA37FE6B" box="[1361,1436,414,438]" italics="true" pageId="10" pageNumber="69">Elseya</emphasis>
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</taxonomicName>
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see Table 1.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33A6517A812FFE3FF51FE3FFBB4FD52" pageId="10" pageNumber="69" type="etymology">
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<paragraph id="8B9F369CA812FFE3FF51FE3FFBB4FD52" blockId="10.[151,1437,306,2032]" pageId="10" pageNumber="69">
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<emphasis id="B954EA8EA812FFE3FF51FE3FFEE3FE22" bold="true" box="[199,328,486,511]" pageId="10" pageNumber="69">Etymology</emphasis>
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—Named in honor of Anders G. J. Rhodin; Chairman Emeritus of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group; Founder and Director, Chelonian Research Foundation. He also researched the turtles of
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<collectingCountry id="F337760CA812FFE3FE92FDF6FE75FD9A" box="[260,478,558,583]" name="Papua New Guinea" pageId="10" pageNumber="69">Papua New Guinea</collectingCountry>
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describing two species,
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<taxonomicName id="4C204D1FA812FFE3FD65FDF7FAC7FD9A" authority="Rhodin and Mittermeier, 1976" authorityName="Rhodin and Mittermeier" authorityYear="1976" box="[755,1388,558,583]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="parkeri" subGenus="M.">
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<emphasis id="B954EA8EA812FFE3FD65FDF7FCC3FD9B" box="[755,872,558,582]" italics="true" pageId="10" pageNumber="69">Chelodina</emphasis>
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(
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<emphasis id="B954EA8EA812FFE3FCEFFDF6FC3BFD9B" box="[889,912,559,582]" italics="true" pageId="10" pageNumber="69">M</emphasis>
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||
.)
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<emphasis id="B954EA8EA812FFE3FC3FFDF7FC50FD9B" box="[937,1019,558,582]" italics="true" pageId="10" pageNumber="69">parkeri</emphasis>
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(
|
||
<bibRefCitation id="EFB14B6DA812FFE3FB9CFDF7FAC8FD9A" author="Rhodin" box="[1034,1379,558,583]" pageId="10" pageNumber="69" refString="Rhodin, A. G. J. & Mittermeier, R. A. (1976) Chelodina parkeri, a new species of chelid turtle from New Guinea, with a discussion of Chelodina siebenrocki Werner, 1901. Bulletin of the Museum of Comparative Zoology, 147 (11), 465 - 488." type="journal article" year="1976">Rhodin and Mittermeier, 1976</bibRefCitation>
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)
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</taxonomicName>
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and
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<taxonomicName id="4C204D1FA812FFE3FF01FD8AFDDCFDB6" authority="Rhodin, 1994" authorityName="Rhodin" authorityYear="1994" box="[151,631,594,619]" class="Reptilia" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="pritchardi" subGenus="C.">
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<emphasis id="B954EA8EA812FFE3FF01FD8AFEA7FDB6" box="[151,268,595,619]" italics="true" pageId="10" pageNumber="69">Chelodina</emphasis>
|
||
(
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||
<emphasis id="B954EA8EA812FFE3FE89FD8AFE99FDB7" box="[287,306,595,618]" italics="true" pageId="10" pageNumber="69">C</emphasis>
|
||
.)
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<emphasis id="B954EA8EA812FFE3FEDAFD8AFE17FDB6" box="[332,444,595,619]" italics="true" pageId="10" pageNumber="69">pritchardi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB14B6DA812FFE3FE58FD8BFDC4FDB6" author="Rhodin" box="[462,623,594,619]" pageId="10" pageNumber="69" refString="Rhodin, A. G. J. (1994) Chelid turtles of the Australasian Archipelago: I. A new species of Chelodina from southeastern Papua New Guinea. Breviora, 497, 1 - 36." type="journal article" year="1994">Rhodin, 1994</bibRefCitation>
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)
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</taxonomicName>
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. He was also one of the first to collect this new species (
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<bibRefCitation id="EFB14B6DA812FFE3FA83FD8BFE98FD52" author="Rhodin" pageId="10" pageNumber="69" refString="Rhodin, A. G. J. & Rhodin, S. D. (1977) Iakttagelser fran en herpetologisk samlingsresa till Papua New Guinea. [Observations from a herpetological collecting trip to Papua New Guinea]. Snoken - National Swedish Herpetological Association, 7 (2 / 3), 65 - 72. [in Swedish]" type="journal article" year="1977">Rhodin and Rhodin, 1977</bibRefCitation>
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), and the first to recognize it as new (
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||
<bibRefCitation id="EFB14B6DA812FFE3FD40FDAFFBB8FD52" author="Rhodin" box="[726,1043,630,655]" pageId="10" pageNumber="69" refString="Rhodin, A. G. J. & Genorupa, V. R. (2000) Conservation status of freshwater turtles in Papua New Guinea. Chelonian Research Monographs, 2, 129 - 136." type="journal article" year="2000">Rhodin and Genorupa 2000</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A6517A812FFE3FF51FD43FAE4FD0A" pageId="10" pageNumber="69" type="distribution">
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<paragraph id="8B9F369CA812FFE3FF51FD43FAE4FD0A" blockId="10.[151,1437,306,2032]" pageId="10" pageNumber="69">
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<emphasis id="B954EA8EA812FFE3FF51FD43FEF2FD6E" bold="true" box="[199,345,666,691]" pageId="10" pageNumber="69">Distribution</emphasis>
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—Streams draining into rivers in New
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<collectingCountry id="F337760CA812FFE3FC9EFD43FCF7FD6E" box="[776,860,666,691]" name="Guinea-Bissau" pageId="10" pageNumber="69">Guinea</collectingCountry>
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, south of the central ranges, from the Sitekwa River region of
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<collectingCountry id="F337760CA812FFE3FE92FD66FEE0FD0B" box="[260,331,703,726]" name="Papua New Guinea" pageId="10" pageNumber="69">Papua</collectingCountry>
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,
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<collectingCountry id="F337760CA812FFE3FEC0FD67FE6DFD0A" box="[342,454,702,727]" name="Indonesia" pageId="10" pageNumber="69">Indonesia</collectingCountry>
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, to the Kikori-Purari rivers region of
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<collectingCountry id="F337760CA812FFE3FCFEFD66FBEFFD0A" box="[872,1092,702,727]" name="Papua New Guinea" pageId="10" pageNumber="69">Papua New Guinea</collectingCountry>
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. Aru Islands. (
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<figureCitation id="131B2A19A812FFE3FB70FD67FAEFFD0A" box="[1254,1348,702,727]" captionStart="FIGURE 1" captionStartId="1.[151,250,1799,1821]" captionTargetBox="[151,1436,968,1774]" captionTargetId="figure@1.[151,1436,968,1774]" captionTargetPageId="1" captionText="FIGURE 1. Map of New Guinea showing the distribution of Elseya novaeguineae (Birds Head), E. schultzei (north of Central Ranges) and E. rhodini (south of Central Ranges). The locations of the holotypes for each species are shown as red dots; the type locality for the neotype of Elseya branderhorsti is shown as a green square." httpUri="https://zenodo.org/record/289265/files/figure.png" pageId="10" pageNumber="69">Figure 1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A6517A812FFE3FF51FD3BFDC5FB2E" pageId="10" pageNumber="69" type="diagnosis">
|
||
<paragraph id="8B9F369CA812FFE3FF51FD3BFDC5FB2E" blockId="10.[151,1437,306,2032]" pageId="10" pageNumber="69">
|
||
<emphasis id="B954EA8EA812FFE3FF51FD3BFE91FD26" bold="true" box="[199,314,738,763]" pageId="10" pageNumber="69">Diagnosis</emphasis>
|
||
–Distinguished by gold speckling usually present in the sclera of the eye, which partially obscures the iris; base color green (
|
||
<figureCitation id="131B2A19A812FFE3FE23FCDFFD8DFCC2" box="[437,550,774,799]" captionStart="FIGURE 10" captionStartId="12.[151,250,1887,1909]" captionTargetBox="[340,1246,1260,1862]" captionTargetId="figure@12.[340,1247,1260,1862]" captionTargetPageId="12" captionText="FIGURE 10. Lateral view of the head of Elseya rhodini. Wild caught individual (UC <Aus> AA 42045)." httpUri="https://zenodo.org/record/289272/files/figure.png" pageId="10" pageNumber="69">Figure 10</figureCitation>
|
||
). In
|
||
<emphasis id="B954EA8EA812FFE3FDC1FCDFFB86FCC3" box="[599,1069,774,798]" italics="true" pageId="10" pageNumber="69">
|
||
<taxonomicName id="4C204D1FA812FFE3FDC1FCDFFCEFFCC3" box="[599,836,774,798]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="novaeguineae">Elseya novaeguineae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D1FA812FFE3FCECFCDFFB86FCC3" box="[890,1069,774,798]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="schultzei">Elseya schultzei</taxonomicName>
|
||
</emphasis>
|
||
the sclera is bright green and the iris is starkly contrasted, though the base colors of the sclera in the later is a duller and plainer green. The head shield in
|
||
<taxonomicName id="4C204D1FA812FFE3FF6BFC96FEC5FCBB" box="[253,366,846,870]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA812FFE3FF6BFC96FEA5FCBB" box="[253,270,847,870]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FE8AFC97FEC5FCBB" box="[284,366,846,870]" italics="true" pageId="10" pageNumber="69">rhodini</emphasis>
|
||
</taxonomicName>
|
||
is reduced when compared to other species of this subgenus. The lateral edge of the head shield at its midpoint between the eye and the tympanum is above the level of the eye, below it in the other species. The ventral extension of the head shield down the parietal arch is also reduced, typically still reaching the tympanum, but narrower throughout its length. Posterior to the eye in
|
||
<taxonomicName id="4C204D1FA812FFE3FCDBFC62FBA6FC0F" box="[845,1037,955,978]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="novaeguineae">
|
||
<emphasis id="B954EA8EA812FFE3FCDBFC62FCF5FC0F" box="[845,862,955,978]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FCE7FC62FBA6FC0F" box="[881,1037,955,978]" italics="true" pageId="10" pageNumber="69">novaeguineae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D1FA812FFE3FBDAFC62FB7AFC0E" box="[1100,1233,955,979]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="schultzei">
|
||
<emphasis id="B954EA8EA812FFE3FBDAFC62FBF6FC0F" box="[1100,1117,955,978]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FBF9FC62FB7AFC0E" box="[1135,1233,955,979]" italics="true" pageId="10" pageNumber="69">schultzei</emphasis>
|
||
</taxonomicName>
|
||
is a variable but generally present extension of the head shield that extends down behind the eye, in some specimens to nearly the midpoint of the eye. This is absent in
|
||
<taxonomicName id="4C204D1FA812FFE3FDAAFBDAFD06FBC6" box="[572,685,1027,1051]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA812FFE3FDAAFBDAFDE6FBC7" box="[572,589,1027,1050]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FDCDFBDAFD06FBC6" box="[603,685,1027,1051]" italics="true" pageId="10" pageNumber="69">rhodini</emphasis>
|
||
</taxonomicName>
|
||
. On the carapace the spots on the scutes of
|
||
<taxonomicName id="4C204D1FA812FFE3FB02FBDAFAAEFBC6" box="[1172,1285,1027,1051]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA812FFE3FB02FBDAFB0EFBC7" box="[1172,1189,1027,1050]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FB25FBDAFAAEFBC6" box="[1203,1285,1027,1051]" italics="true" pageId="10" pageNumber="69">rhodini</emphasis>
|
||
</taxonomicName>
|
||
are visible in all growth stages, whereas in the other two species they are often faded or absent in adults. The cervical is generally present in the members of this group however it is narrower (approximately 30% of length) in
|
||
<taxonomicName id="4C204D1FA812FFE3FB25FB92FA8FFBBE" box="[1203,1316,1099,1123]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA812FFE3FB25FB92FB6FFBBF" box="[1203,1220,1099,1122]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FB44FB92FA8FFBBE" box="[1234,1316,1099,1123]" italics="true" pageId="10" pageNumber="69">rhodini</emphasis>
|
||
</taxonomicName>
|
||
whereas it is usually 50% of length in the other two species. The lateral sulcus of the first vertebral scute in both
|
||
<taxonomicName id="4C204D1FA812FFE3FA8CFBB6FA37FB5B" box="[1306,1436,1134,1158]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="schultzei">
|
||
<emphasis id="B954EA8EA812FFE3FA8CFBB6FA80FB5B" box="[1306,1323,1135,1158]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FAACFBB7FA37FB5B" box="[1338,1436,1134,1158]" italics="true" pageId="10" pageNumber="69">schultzei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D1FA812FFE3FF50FB4AFE2AFB77" box="[198,385,1171,1194]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="novaeguineae">
|
||
<emphasis id="B954EA8EA812FFE3FF50FB4AFF7CFB77" box="[198,215,1171,1194]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FF73FB4AFE2AFB77" box="[229,385,1171,1194]" italics="true" pageId="10" pageNumber="69">novaeguineae</emphasis>
|
||
</taxonomicName>
|
||
contacts the center of the second marginal scute, whereas in
|
||
<taxonomicName id="4C204D1FA812FFE3FBBDFB4AFB37FB76" box="[1067,1180,1171,1195]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA812FFE3FBBDFB4AFB97FB77" box="[1067,1084,1171,1194]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FBDCFB4AFB37FB76" box="[1098,1180,1171,1195]" italics="true" pageId="10" pageNumber="69">rhodini</emphasis>
|
||
</taxonomicName>
|
||
it contacts the anterior ¼ of the second marginal. The skull of
|
||
<taxonomicName id="4C204D1FA812FFE3FDDAFB6EFD15FB13" box="[588,702,1206,1230]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA812FFE3FDDAFB6EFDF6FB13" box="[588,605,1207,1230]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FDFAFB6FFD15FB13" box="[620,702,1206,1230]" italics="true" pageId="10" pageNumber="69">rhodini</emphasis>
|
||
</taxonomicName>
|
||
can be diagnosed by the contact of the vomer with the pterygoids posteriorly and the premaxillae anteriorly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6517A812FFE7FF51FB27FB1BFC5B" lastPageId="14" lastPageNumber="73" pageId="10" pageNumber="69" type="description">
|
||
<paragraph id="8B9F369CA812FFE3FF51FB27FEFAFACA" blockId="10.[151,1437,306,2032]" box="[199,337,1278,1303]" pageId="10" pageNumber="69">
|
||
<emphasis id="B954EA8EA812FFE3FF51FB27FEFAFACA" bold="true" box="[199,337,1278,1303]" pageId="10" pageNumber="69">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F369CA812FFE3FF51FAFBFE12FAE6" blockId="10.[151,1437,306,2032]" box="[199,441,1314,1339]" pageId="10" pageNumber="69">External Morphology</paragraph>
|
||
<paragraph id="8B9F369CA812FFE3FF51FA93FD90F95A" blockId="10.[151,1437,306,2032]" pageId="10" pageNumber="69">
|
||
<emphasis id="B954EA8EA812FFE3FF51FA93FE9EFABC" box="[199,309,1354,1377]" italics="true" pageId="10" pageNumber="69">Carapace</emphasis>
|
||
—The carapace is broadly oval slightly flattened at the anterior from the 2nd marginal. The first vertebral scute is approximately the same width as the second. The outer sulci of the first vertebral tend to intersect the 2nd marginal close to the sulcus between the 1st and 2nd marginal, differing from
|
||
<taxonomicName id="4C204D1FA812FFE3FBD3FA4CFAAAFA71" box="[1093,1281,1429,1452]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="novaeguineae">
|
||
<emphasis id="B954EA8EA812FFE3FBD3FA4CFBFDFA71" box="[1093,1110,1429,1452]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FBF3FA4CFAAAFA71" box="[1125,1281,1429,1452]" italics="true" pageId="10" pageNumber="69">novaeguineae</emphasis>
|
||
</taxonomicName>
|
||
where it is in the center of the 2nd marginal. There is some flaring of the marginal over the hind legs, though it tends to be more pronounced in
|
||
<taxonomicName id="4C204D1FA812FFE3FEDEFA39FDA3FA2A" box="[328,520,1504,1527]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="novaeguineae">
|
||
<emphasis id="B954EA8EA812FFE3FEDEFA39FEF2FA2A" box="[328,345,1504,1527]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FEFAFA39FDA3FA2A" box="[364,520,1504,1527]" italics="true" pageId="10" pageNumber="69">novaeguineae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D1FA812FFE3FDD1FA39FD66FA2A" box="[583,717,1503,1527]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="schultzei">
|
||
<emphasis id="B954EA8EA812FFE3FDD1FA39FDF3FA2A" box="[583,600,1504,1527]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FDFDFA06FD66FA2A" box="[619,717,1503,1527]" italics="true" pageId="10" pageNumber="69">schultzei</emphasis>
|
||
</taxonomicName>
|
||
than is apparent in similarly sized individuals of
|
||
<taxonomicName id="4C204D1FA812FFE3FA89FA39FA3EFA2A" box="[1311,1429,1503,1527]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA812FFE3FA89FA39FA9BFA2A" box="[1311,1328,1504,1527]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FAD5FA06FA3EFA2A" box="[1347,1429,1503,1527]" italics="true" pageId="10" pageNumber="69">rhodini</emphasis>
|
||
</taxonomicName>
|
||
. Cervical scute is present for all three taxa, though it is usually relatively narrower in
|
||
<taxonomicName id="4C204D1FA812FFE3FBF2F9DDFB7CF9C6" box="[1124,1239,1539,1563]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA812FFE3FBF2F9DDFBDEF9C6" box="[1124,1141,1540,1563]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FB13F9DAFB7CF9C6" box="[1157,1239,1539,1563]" italics="true" pageId="10" pageNumber="69">rhodini</emphasis>
|
||
</taxonomicName>
|
||
than in the other two species of the subgenus. The carapace is brown in color for all three species, spots may be present at the center of each major scute, these persist into adult hood in
|
||
<taxonomicName id="4C204D1FA812FFE3FD6FF995FCC6F9BE" box="[761,877,1611,1635]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA812FFE3FD6FF995FCA1F9BE" box="[761,778,1612,1635]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FC8DF992FCC6F9BE" box="[795,877,1611,1635]" italics="true" pageId="10" pageNumber="69">rhodini</emphasis>
|
||
</taxonomicName>
|
||
, are faded if present in
|
||
<taxonomicName id="4C204D1FA812FFE3FB11F995FAA0F9BE" box="[1159,1291,1611,1635]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="schultzei">
|
||
<emphasis id="B954EA8EA812FFE3FB11F995FB33F9BE" box="[1159,1176,1612,1635]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FB3FF992FAA0F9BE" box="[1193,1291,1611,1635]" italics="true" pageId="10" pageNumber="69">schultzei</emphasis>
|
||
</taxonomicName>
|
||
and may be absent altogether in
|
||
<taxonomicName id="4C204D1FA812FFE3FEEFF9A9FD9FF95A" box="[377,564,1648,1671]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="novaeguineae">
|
||
<emphasis id="B954EA8EA812FFE3FEEFF9A9FE21F95A" box="[377,394,1648,1671]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FE0EF9A9FD9FF95A" box="[408,564,1648,1671]" italics="true" pageId="10" pageNumber="69">novaeguineae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F369CA812FFE3FF51F94AFA36F8C5" blockId="10.[151,1437,306,2032]" pageId="10" pageNumber="69">
|
||
<emphasis id="B954EA8EA812FFE3FF51F94AFE8DF976" box="[199,294,1683,1707]" italics="true" pageId="10" pageNumber="69">Plastron</emphasis>
|
||
—The plastra of all three species are very similar, being rectangular in general shape but widest at the posterior of the anterior lobe and with lateral flanging in older individuals just anterior to the rear legs on the posterior lobe. The plastral formula is generally pec>fem>abd>ana>int>hum with the gulars excluded from the midline by the intergular. The plastron is cream to yellow in color, very occasionally there is a slight pink suffusion.
|
||
</paragraph>
|
||
<paragraph id="8B9F369CA812FFE3FF51F8FAFAC6F82D" blockId="10.[151,1437,306,2032]" pageId="10" pageNumber="69">
|
||
<emphasis id="B954EA8EA812FFE3FF51F8FAFE0DF8E6" box="[199,422,1827,1851]" italics="true" pageId="10" pageNumber="69">Head and soft parts</emphasis>
|
||
—The color of the sclera of
|
||
<taxonomicName id="4C204D1FA812FFE3FD72F8FDFCFEF8E6" box="[740,853,1827,1851]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA812FFE3FD72F8FDFD5EF8E6" box="[740,757,1828,1851]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FC95F8FAFCFEF8E6" box="[771,853,1827,1851]" italics="true" pageId="10" pageNumber="69">rhodini</emphasis>
|
||
</taxonomicName>
|
||
is gold flecked green, 2.5
|
||
<collectingCountry id="F337760CA812FFE3FBEFF8FAFB25F8E1" box="[1145,1166,1827,1852]" name="Italy" pageId="10" pageNumber="69">Y</collectingCountry>
|
||
7/8 to 2.5
|
||
<collectingCountry id="F337760CA812FFE3FB6FF8FAFAA5F8E1" box="[1273,1294,1827,1852]" name="Italy" pageId="10" pageNumber="69">Y</collectingCountry>
|
||
8/8 (
|
||
<bibRefCitation id="EFB14B6DA812FFE3FAA8F8FAFF7AF882" author="Munsell" pageId="10" pageNumber="69" refString="Munsell, A. H. (Munsell Color) (2000) Munsell Soil Color Charts, Revised washable edition. Gretag Macbeth, New York, 10 pp." type="book" year="2000">Munsell 2000</bibRefCitation>
|
||
) with the iris gold and relatively indistinct. Compared with the color of the sclera of
|
||
<taxonomicName id="4C204D1FA812FFE3FB15F891FA95F882" box="[1155,1342,1864,1887]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="novaeguineae">
|
||
<emphasis id="B954EA8EA812FFE3FB15F891FB3FF882" box="[1155,1172,1864,1887]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FB34F891FA95F882" box="[1186,1342,1864,1887]" italics="true" pageId="10" pageNumber="69">novaeguineae</emphasis>
|
||
</taxonomicName>
|
||
is green with no flecks, 5y7/3 to 5y8/3 (
|
||
<bibRefCitation id="EFB14B6DA812FFE3FE60F8B2FD3FF85E" author="Munsell" box="[502,660,1899,1924]" pageId="10" pageNumber="69" refString="Munsell, A. H. (Munsell Color) (2000) Munsell Soil Color Charts, Revised washable edition. Gretag Macbeth, New York, 10 pp." type="book" year="2000">Munsell 2000</bibRefCitation>
|
||
) with a very distinct gold iris, and in
|
||
<taxonomicName id="4C204D1FA812FFE3FBAEF8B5FB12F85E" box="[1080,1209,1899,1923]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="schultzei">
|
||
<emphasis id="B954EA8EA812FFE3FBAEF8B5FBE2F85E" box="[1080,1097,1900,1923]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FBC1F8B2FB12F85E" box="[1111,1209,1899,1923]" italics="true" pageId="10" pageNumber="69">schultzei</emphasis>
|
||
</taxonomicName>
|
||
it is also green with no flecks, 5y7/6 to 5y8/6 (
|
||
<bibRefCitation id="EFB14B6DA812FFE3FE54F856FDCAF87A" author="Munsell" box="[450,609,1935,1960]" pageId="10" pageNumber="69" refString="Munsell, A. H. (Munsell Color) (2000) Munsell Soil Color Charts, Revised washable edition. Gretag Macbeth, New York, 10 pp." type="book" year="2000">Munsell 2000</bibRefCitation>
|
||
) again with a very distinct gold iris. The dorsal surface of the head of
|
||
<taxonomicName id="4C204D1FA812FFE3FA12F849FF42F816" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA812FFE3FA12F849FA3EF87A" box="[1412,1429,1936,1959]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FF01F86AFF42F816" box="[151,233,1971,1995]" italics="true" pageId="10" pageNumber="69">rhodini</emphasis>
|
||
</taxonomicName>
|
||
is variegated in color between yellow brown to dark brown, a similar pattern is seen in
|
||
<taxonomicName id="4C204D1FA812FFE3FA99F86DFA3EF816" box="[1295,1429,1971,1995]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="schultzei">
|
||
<emphasis id="B954EA8EA812FFE3FA99F86DFA8BF816" box="[1295,1312,1972,1995]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FAA5F86AFA3EF816" box="[1331,1429,1971,1995]" italics="true" pageId="10" pageNumber="69">schultzei</emphasis>
|
||
</taxonomicName>
|
||
, however
|
||
<taxonomicName id="4C204D1FA812FFE3FF68F801FE12F832" box="[254,441,2008,2031]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="69" phylum="Chordata" rank="species" species="novaeguineae">
|
||
<emphasis id="B954EA8EA812FFE3FF68F801FEA4F832" box="[254,271,2008,2031]" italics="true" pageId="10" pageNumber="69">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA812FFE3FE8BF801FE12F832" box="[285,441,2008,2031]" italics="true" pageId="10" pageNumber="69">novaeguineae</emphasis>
|
||
</taxonomicName>
|
||
tends to be far less variegated without the striking patterns seen in the other species.
|
||
</paragraph>
|
||
<caption id="DF5F6614A813FFE2FF01F8DBFD5EF8E6" httpUri="https://zenodo.org/record/289270/files/figure.png" pageId="11" pageNumber="70" targetBox="[190,1412,215,1757]" targetPageId="11">
|
||
<paragraph id="8B9F369CA813FFE2FF01F8DBFD5EF8E6" blockId="11.[151,1436,1794,1851]" pageId="11" pageNumber="70">
|
||
<emphasis id="B954EA8EA813FFE2FF01F8DBFEBFF8CA" bold="true" box="[151,276,1794,1816]" pageId="11" pageNumber="70">FIGURE 8.</emphasis>
|
||
Dorsal, ventral and lateral views of the holotype PNGM R25204 of
|
||
<taxonomicName id="4C204D1FA813FFE2FC4FF8DBFA86F8C5" box="[985,1325,1794,1816]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="70" phylum="Chordata" rank="species" species="rhodini" subGenus="Hanwarachelys">
|
||
<emphasis id="B954EA8EA813FFE2FC4FF8DBFA86F8C5" box="[985,1325,1794,1816]" italics="true" pageId="11" pageNumber="70">Elseya (Hanwarachelys) rhodini</emphasis>
|
||
</taxonomicName>
|
||
from Wau Creek, Kikori Drainage, Papua New Guinea. Scale: 20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F369CA813FFE2FF51F8B5FF76F82D" blockId="11.[151,1436,1899,2032]" pageId="11" pageNumber="70">
|
||
<emphasis id="B954EA8EA813FFE2FF51F8B5FDB1F859" box="[199,538,1900,1924]" italics="true" pageId="11" pageNumber="70">Size and Sexual Dimorphism</emphasis>
|
||
—This species attains a size of approximately
|
||
<quantity id="4CD89B79A813FFE2FBDEF8B5FB35F859" box="[1096,1182,1900,1924]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" pageId="11" pageNumber="70" unit="mm" value="240.0">240mm</quantity>
|
||
carapace length with females being slightly larger than males. This is similar for all members of the subgenus
|
||
<emphasis id="B954EA8EA813FFE2FB39F849FAF4F875" box="[1199,1375,1936,1960]" italics="true" pageId="11" pageNumber="70">Hanwarachelys</emphasis>
|
||
. The species attains a smaller adult size than
|
||
<taxonomicName id="4C204D1FA813FFE2FDFFF86DFCF3F811" box="[617,856,1972,1996]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="70" phylum="Chordata" rank="species" species="branderhorsti">
|
||
<emphasis id="B954EA8EA813FFE2FDFFF86DFCF3F811" box="[617,856,1972,1996]" italics="true" pageId="11" pageNumber="70">Elseya branderhorsti</emphasis>
|
||
</taxonomicName>
|
||
with which it is sympatric throughout parts of its range.
|
||
</paragraph>
|
||
<caption id="DF5F6614A814FFE5FF01FB46FA8DFB68" box="[151,1318,1183,1205]" httpUri="https://zenodo.org/record/289271/files/figure.png" pageId="12" pageNumber="71" targetBox="[151,1436,195,1159]" targetPageId="12">
|
||
<paragraph id="8B9F369CA814FFE5FF01FB46FA8DFB68" blockId="12.[151,1318,1183,1205]" box="[151,1318,1183,1205]" pageId="12" pageNumber="71">
|
||
<emphasis id="B954EA8EA814FFE5FF01FB46FEB9FB69" bold="true" box="[151,274,1183,1205]" pageId="12" pageNumber="71">FIGURE 9.</emphasis>
|
||
Habitat of
|
||
<taxonomicName id="4C204D1FA814FFE5FE13FB46FCCEFB68" authority="Wau Creek, Sirebi River, Kikori" authorityName="Wau Creek, Sirebi River, Kikori" box="[389,869,1183,1205]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="71" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA814FFE5FE13FB46FDB2FB68" box="[389,537,1183,1205]" italics="true" pageId="12" pageNumber="71">Elseya rhodini</emphasis>
|
||
Wau Creek, Sirebi River, Kikori
|
||
</taxonomicName>
|
||
drainage (07˚11’ 67.3” S, 144˚37’ 13.8” E).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F6614A814FFE5FF01F886FB1BF8A8" box="[151,1200,1887,1909]" httpUri="https://zenodo.org/record/289272/files/figure.png" pageId="12" pageNumber="71" targetBox="[340,1246,1260,1862]" targetPageId="12">
|
||
<paragraph id="8B9F369CA814FFE5FF01F886FB1BF8A8" blockId="12.[151,1200,1887,1909]" box="[151,1200,1887,1909]" pageId="12" pageNumber="71">
|
||
<emphasis id="B954EA8EA814FFE5FF01F886FEB4F8A9" bold="true" box="[151,287,1887,1909]" pageId="12" pageNumber="71">FIGURE 10.</emphasis>
|
||
Lateral view of the head of
|
||
<taxonomicName id="4C204D1FA814FFE5FDABF886FD7AF8A8" box="[573,721,1887,1909]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="71" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA814FFE5FDABF886FD7AF8A8" box="[573,721,1887,1909]" italics="true" pageId="12" pageNumber="71">Elseya rhodini</emphasis>
|
||
</taxonomicName>
|
||
. Wild caught individual (UC<Aus>AA42045).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F369CA815FFE4FF51FF4EFE96FF6D" blockId="13.[151,1436,151,357]" box="[199,317,151,176]" pageId="13" pageNumber="72">
|
||
<emphasis id="B954EA8EA815FFE4FF51FF4EFE96FF6D" bold="true" box="[199,317,151,176]" pageId="13" pageNumber="72">Osteology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9F369CA815FFE4FF51FF65FE97FEB9" blockId="13.[151,1436,151,357]" pageId="13" pageNumber="72">
|
||
<emphasis id="B954EA8EA815FFE4FF51FF65FF54FF09" box="[199,255,188,212]" italics="true" pageId="13" pageNumber="72">Skull</emphasis>
|
||
(
|
||
<figureCitation id="131B2A19A815FFE4FE99FF65FE25FF08" box="[271,398,188,213]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStartId-0="13.[151,246,912,934]" captionStartId-1="13.[151,250,1944,1966]" captionStartId-2="14.[151,250,1891,1913]" captionTargetBox-0="[360,1226,433,869]" captionTargetBox-1="[441,1139,1031,1891]" captionTargetBox-2="[473,1116,972,1857]" captionTargetId-0="figure@13.[340,1248,411,888]" captionTargetId-1="figure@13.[431,1156,1010,1917]" captionTargetId-2="figure@14.[458,1128,957,1866]" captionTargetPageId-0="13" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURE 11. Lateral views of the skulls of A. Elseya (Hanwarachelys) rhodini, B. Elseya (Hanwarachelys) novaeguineae, C. Elseya (Elseya) dentata, and D. Elseya (Pelocomastes) albagula. po—postorbital, sq—squamosal, q—quadrate, j—jugal, mx—maxilla." captionText-1="FIGURE 12. Ventral views of the skulls of A. Elseya (Hanwarachelys) rhodini, B. Elseya (Hanwarachelys) novaeguineae, C. Elseya (Elseya) dentata, and D. Elseya (Pelocomastes) albagula. pm—premaxilla, mx—maxilla, pal—palatine, vo—vomer, bs—basisphenoid, bo—basooccipital. Also labelled are the alveolar and lingual ridges." captionText-2="FIGURE 13. Dorsal view of the skulls of A. Elseya (Hanwarachelys) rhodini, B. Elseya (Hanwarachelys) novaeguineae, C. Elseya (Elseya) dentata, and D. Elseya (Pelocomastes) albagula. fr—frontal, po—postorbital, pa—parietal, sosupraoccipital." httpUri-0="https://zenodo.org/record/289273/files/figure.png" httpUri-1="https://zenodo.org/record/289274/files/figure.png" httpUri-2="https://zenodo.org/record/289275/files/figure.png" pageId="13" pageNumber="72">Figs. 11–13</figureCitation>
|
||
)—Dermal roofing elements of the skull are similar in all species of
|
||
<taxonomicName id="4C204D1FA815FFE4FB0FFF65FB4FFF09" box="[1177,1252,188,212]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="72" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA815FFE4FB0FFF65FB4FFF09" box="[1177,1252,188,212]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
</taxonomicName>
|
||
, the frontals are paired and the anterior process separates the nasals almost in their entirety. Parietals are large and postero-laterally suture to the squamosal to form the parietal arch. The arch is narrow in the
|
||
<taxonomicName id="4C204D1FA815FFE4FC78FEDDFB35FEC1" box="[1006,1182,260,284]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="72" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA815FFE4FC78FEDDFB92FEC1" box="[1006,1081,260,284]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA815FFE4FBDCFEDDFB3EFEC1" box="[1098,1173,260,284]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
much wider in
|
||
<taxonomicName id="4C204D1FA815FFE4FAC7FEDDFEF0FE9D" authority="Hanwarachelys" authorityName="Hanwarachelys" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="72" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA815FFE4FAC7FEDDFA37FEC1" box="[1361,1436,260,284]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA815FFE4FF36FEF1FEFBFE9D" box="[160,336,296,320]" italics="true" pageId="13" pageNumber="72">Hanwarachelys</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
. This feature also determines the degree of lateral extension of the head shield as it requires bone for its support.
|
||
</paragraph>
|
||
<caption id="DF5F6614A815FFE4FF01FC49FAD8FC14" httpUri="https://zenodo.org/record/289273/files/figure.png" pageId="13" pageNumber="72" targetBox="[360,1226,433,869]" targetPageId="13">
|
||
<paragraph id="8B9F369CA815FFE4FF01FC49FAD8FC14" blockId="13.[151,1436,912,969]" pageId="13" pageNumber="72">
|
||
<emphasis id="B954EA8EA815FFE4FF01FC49FEB2FC78" bold="true" box="[151,281,912,934]" pageId="13" pageNumber="72">FIGURE 11.</emphasis>
|
||
Lateral views of the skulls of A.
|
||
<taxonomicName id="4C204D1FA815FFE4FDCEFC49FC39FC7B" box="[600,914,912,934]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="72" phylum="Chordata" rank="species" species="rhodini" subGenus="Hanwarachelys">
|
||
<emphasis id="B954EA8EA815FFE4FDCEFC49FD32FC7B" box="[600,665,912,934]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA815FFE4FD30FC49FC95FC7B" box="[678,830,912,934]" italics="true" pageId="13" pageNumber="72">Hanwarachelys</emphasis>
|
||
)
|
||
<emphasis id="B954EA8EA815FFE4FCDAFC49FC39FC7B" box="[844,914,912,934]" italics="true" pageId="13" pageNumber="72">rhodini</emphasis>
|
||
</taxonomicName>
|
||
, B.
|
||
<taxonomicName id="4C204D1FA815FFE4FC2CFC49FA98FC7B" box="[954,1331,912,934]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="72" phylum="Chordata" rank="species" species="novaeguineae" subGenus="Hanwarachelys">
|
||
<emphasis id="B954EA8EA815FFE4FC2CFC49FC50FC7B" box="[954,1019,912,934]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA815FFE4FB9EFC49FB0BFC7B" box="[1032,1184,912,934]" italics="true" pageId="13" pageNumber="72">Hanwarachelys</emphasis>
|
||
)
|
||
<emphasis id="B954EA8EA815FFE4FB3BFC48FA98FC7B" box="[1197,1331,913,934]" italics="true" pageId="13" pageNumber="72">novaeguineae</emphasis>
|
||
</taxonomicName>
|
||
, C.
|
||
<taxonomicName id="4C204D1FA815FFE4FACAFC49FE9EFC14" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="72" phylum="Chordata" rank="species" species="dentata" subGenus="Elseya">
|
||
<emphasis id="B954EA8EA815FFE4FACAFC49FA37FC7B" box="[1372,1436,912,934]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA815FFE4FF09FC6AFF74FC14" box="[159,223,947,969]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
)
|
||
<emphasis id="B954EA8EA815FFE4FF7BFC6AFE9EFC14" box="[237,309,947,969]" italics="true" pageId="13" pageNumber="72">dentata</emphasis>
|
||
</taxonomicName>
|
||
, and D.
|
||
<taxonomicName id="4C204D1FA815FFE4FE11FC6AFD15FC14" box="[391,702,947,969]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="72" phylum="Chordata" rank="species" species="albagula" subGenus="Pelocomastes">
|
||
<emphasis id="B954EA8EA815FFE4FE11FC6AFE63FC14" box="[391,456,947,969]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA815FFE4FE43FC6AFDF1FC14" box="[469,602,947,969]" italics="true" pageId="13" pageNumber="72">Pelocomastes</emphasis>
|
||
)
|
||
<emphasis id="B954EA8EA815FFE4FDFEFC6AFD15FC14" box="[616,702,947,969]" italics="true" pageId="13" pageNumber="72">albagula</emphasis>
|
||
</taxonomicName>
|
||
. po—postorbital, sq—squamosal, q—quadrate, j—jugal, mx—maxilla.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F6614A815FFE4FF01F841FBA3F836" httpUri="https://zenodo.org/record/289274/files/figure.png" pageId="13" pageNumber="72" targetBox="[441,1139,1031,1891]" targetPageId="13">
|
||
<paragraph id="8B9F369CA815FFE4FF01F841FBA3F836" blockId="13.[151,1436,1944,2027]" pageId="13" pageNumber="72">
|
||
<emphasis id="B954EA8EA815FFE4FF01F841FEB4F870" bold="true" box="[151,287,1944,1966]" pageId="13" pageNumber="72">FIGURE 12.</emphasis>
|
||
Ventral views of the skulls of A.
|
||
<taxonomicName id="4C204D1FA815FFE4FDE3F841FC6BF873" box="[629,960,1944,1966]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="72" phylum="Chordata" rank="species" species="rhodini" subGenus="Hanwarachelys">
|
||
<emphasis id="B954EA8EA815FFE4FDE3F841FD12F873" box="[629,697,1944,1966]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA815FFE4FD5EF841FCCCF873" box="[712,871,1944,1966]" italics="true" pageId="13" pageNumber="72">Hanwarachelys</emphasis>
|
||
)
|
||
<emphasis id="B954EA8EA815FFE4FCE0F841FC6BF873" box="[886,960,1944,1966]" italics="true" pageId="13" pageNumber="72">rhodini</emphasis>
|
||
</taxonomicName>
|
||
, B.
|
||
<taxonomicName id="4C204D1FA815FFE4FC7DF841FADCF873" box="[1003,1399,1944,1966]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="72" phylum="Chordata" rank="species" species="novaeguineae" subGenus="Hanwarachelys">
|
||
<emphasis id="B954EA8EA815FFE4FC7DF841FB84F873" box="[1003,1071,1944,1966]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA815FFE4FBABF841FB77F873" box="[1085,1244,1944,1966]" italics="true" pageId="13" pageNumber="72">Hanwarachelys</emphasis>
|
||
)
|
||
<emphasis id="B954EA8EA815FFE4FB7DF840FADCF873" box="[1259,1399,1945,1966]" italics="true" pageId="13" pageNumber="72">novaeguineae</emphasis>
|
||
</taxonomicName>
|
||
, C.
|
||
<taxonomicName id="4C204D1FA815FFE4FF01F86FFE27F811" box="[151,396,1974,1996]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="72" phylum="Chordata" rank="species" species="dentata" subGenus="Elseya">
|
||
<emphasis id="B954EA8EA815FFE4FF01F86FFF70F811" box="[151,219,1974,1996]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA815FFE4FF7DF86FFE84F811" box="[235,303,1974,1996]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
)
|
||
<emphasis id="B954EA8EA815FFE4FED6F86FFE27F811" box="[320,396,1974,1996]" italics="true" pageId="13" pageNumber="72">dentata</emphasis>
|
||
</taxonomicName>
|
||
, and D.
|
||
<taxonomicName id="4C204D1FA815FFE4FE7FF86FFC9FF811" box="[489,820,1974,1996]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="72" phylum="Chordata" rank="species" species="albagula" subGenus="Pelocomastes">
|
||
<emphasis id="B954EA8EA815FFE4FE7FF86FFD86F811" box="[489,557,1974,1996]" italics="true" pageId="13" pageNumber="72">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA815FFE4FDABF86FFD63F811" box="[573,712,1974,1996]" italics="true" pageId="13" pageNumber="72">Pelocomastes</emphasis>
|
||
)
|
||
<emphasis id="B954EA8EA815FFE4FD4CF86FFC9FF811" box="[730,820,1974,1996]" italics="true" pageId="13" pageNumber="72">albagula</emphasis>
|
||
</taxonomicName>
|
||
. pm—premaxilla, mx—maxilla, pal—palatine, vo—vomer, bs—basisphenoid, bo—basooccipital. Also labelled are the alveolar and lingual ridges.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F369CA816FFE7FF51FF41FBAEFE12" blockId="14.[151,1436,151,902]" pageId="14" pageNumber="73">
|
||
On the palatal surface there are significant differences at the species level. The vomer of
|
||
<taxonomicName id="4C204D1FA816FFE7FB3EFF40FAB2FF6D" box="[1192,1305,152,176]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA816FFE7FB3EFF40FB12FF6D" box="[1192,1209,153,176]" italics="true" pageId="14" pageNumber="73">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA816FFE7FB51FF41FAB2FF6D" box="[1223,1305,152,176]" italics="true" pageId="14" pageNumber="73">rhodini</emphasis>
|
||
</taxonomicName>
|
||
completely divides the maxillae anteriorly to meet the pre-maxillae and posteriorly the palatines to meet the pterygoids. In
|
||
<taxonomicName id="4C204D1FA816FFE7FA12FF64FE98FF2A" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="species" species="novaeguineae">
|
||
<emphasis id="B954EA8EA816FFE7FA12FF64FA3EFF09" box="[1412,1429,189,212]" italics="true" pageId="14" pageNumber="73">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA816FFE7FF01FF39FE98FF2A" box="[151,307,224,247]" italics="true" pageId="14" pageNumber="73">novaeguineae</emphasis>
|
||
</taxonomicName>
|
||
both the maxillae and palatines meet at the midline. In the description of
|
||
<taxonomicName id="4C204D1FA816FFE7FB0CFF39FF50FEC1" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="species" species="albagula" subGenus="Pelocomastes">
|
||
<emphasis id="B954EA8EA816FFE7FB0CFF39FB4EFF25" box="[1178,1253,224,248]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA816FFE7FB6FFF39FA38FF25" box="[1273,1427,224,248]" italics="true" pageId="14" pageNumber="73">Pelocomastes</emphasis>
|
||
)
|
||
<emphasis id="B954EA8EA816FFE7FF01FEDDFF50FEC1" box="[151,251,260,284]" italics="true" pageId="14" pageNumber="73">albagula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFB14B6DA816FFE7FE9DFEDDFE57FEC0" author="Thomson" box="[267,508,260,285]" pageId="14" pageNumber="73" refString="Thomson, S., Georges, A. & Limpus, C. J. (2006) A new species of freshwater turtle in the genus Elseya (Testudines: Chelidae) from central coastal Queensland, Australia. Chelonian Conservation and Biology, 5, 74 - 86. http: // dx. doi. org / 10.2744 / 1071 - 8443 (2006) 5 [74: ANSOFT] 2.0. CO; 2" type="journal article" year="2006">Thomson et al, 2006</bibRefCitation>
|
||
) the form of the alveolar ridge was further defined and we follow this here. The lingual ridge (sensu
|
||
<bibRefCitation id="EFB14B6DA816FFE7FEEAFEFEFDCCFE9D" author="Thomson" box="[380,615,295,320]" pageId="14" pageNumber="73" refString="Thomson, S., Georges, A. & Limpus, C. J. (2006) A new species of freshwater turtle in the genus Elseya (Testudines: Chelidae) from central coastal Queensland, Australia. Chelonian Conservation and Biology, 5, 74 - 86. http: // dx. doi. org / 10.2744 / 1071 - 8443 (2006) 5 [74: ANSOFT] 2.0. CO; 2" type="journal article" year="2006">Thomson et al, 2006</bibRefCitation>
|
||
) is completely absent in all members of both
|
||
<taxonomicName id="4C204D1FA816FFE7FBFCFEF1FAB2FE9D" box="[1130,1305,296,320]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA816FFE7FBFCFEF1FB1EFE9D" box="[1130,1205,296,320]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA816FFE7FB50FEF1FABAFE9D" box="[1222,1297,296,320]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C204D1FA816FFE7FAC7FEF1FEF2FEB9" authority="Hanwarachelys" authorityName="Hanwarachelys" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA816FFE7FAC7FEF1FA37FE9D" box="[1361,1436,296,320]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA816FFE7FF36FE95FEFBFEB9" box="[160,336,332,356]" italics="true" pageId="14" pageNumber="73">Hanwarachelys</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
its presence being a synapomorphy for the
|
||
<taxonomicName id="4C204D1FA816FFE7FCD5FE95FBE8FEB9" authority="(Pelocomastes)" baseAuthorityName="Pelocomastes" box="[835,1091,332,356]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA816FFE7FCD5FE95FC25FEB9" box="[835,910,332,356]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA816FFE7FC08FE95FB93FEB9" box="[926,1080,332,356]" italics="true" pageId="14" pageNumber="73">Pelocomastes</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
. However the alveolar ridge is present in members of the
|
||
<taxonomicName id="4C204D1FA816FFE7FE29FEA9FDC6FE55" box="[447,621,368,392]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA816FFE7FE29FEA9FDA1FE55" box="[447,522,368,392]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA816FFE7FD8CFEA9FDCEFE55" box="[538,613,368,392]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
and is only partially present in the members of
|
||
<taxonomicName id="4C204D1FA816FFE7FB17FEA9FA3CFE55" authority="Hanwarachelys" authorityName="Hanwarachelys" box="[1153,1431,368,392]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA816FFE7FB17FEA9FB67FE55" box="[1153,1228,368,392]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA816FFE7FB4AFEA9FA27FE55" box="[1244,1420,368,392]" italics="true" pageId="14" pageNumber="73">Hanwarachelys</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C204D1FA816FFE7FF21FE4CFE81FE71" box="[183,298,404,428]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="species" species="rhodini">
|
||
<emphasis id="B954EA8EA816FFE7FF21FE4CFF63FE71" box="[183,200,405,428]" italics="true" pageId="14" pageNumber="73">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA816FFE7FF4EFE4DFE81FE71" box="[216,298,404,428]" italics="true" pageId="14" pageNumber="73">rhodini</emphasis>
|
||
</taxonomicName>
|
||
there is no obvious alveolar ridge on the ramphotheca of the upper jaw whereas there is a reduced, though present ridge consisting of a row of raised nodules in
|
||
<taxonomicName id="4C204D1FA816FFE7FCD2FE60FC54FE12" box="[836,1023,440,464]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="species" species="novaeguineae">
|
||
<emphasis id="B954EA8EA816FFE7FCD2FE60FCFEFE0D" box="[836,853,441,464]" italics="true" pageId="14" pageNumber="73">E</emphasis>
|
||
.
|
||
<emphasis id="B954EA8EA816FFE7FCF5FE61FC54FE12" box="[867,1023,440,463]" italics="true" pageId="14" pageNumber="73">novaeguineae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F369CA816FFE7FF51FE05FDE5FDC5" blockId="14.[151,1436,151,902]" pageId="14" pageNumber="73">
|
||
Cervicals—The articulation formula is the same in all three species and follows the typical chelid condition of (2(3(4(5)6)7(8) (sensu
|
||
<bibRefCitation id="EFB14B6DA816FFE7FE0FFE26FDEAFDC5" author="Williams" box="[409,577,511,536]" pageId="14" pageNumber="73" refString="Williams, E. E. (1950) Variation in the cervical articulation in recent and fossil turtles. Bulletin of the American Museum of Natural History, 94, 509 - 561." type="journal article" year="1950">Williams 1950</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F369CA816FFE7FF51FDFDFBA2FCC7" blockId="14.[151,1436,151,902]" pageId="14" pageNumber="73">
|
||
Shell—The carapace of members of the
|
||
<taxonomicName id="4C204D1FA816FFE7FD04FDFDFCE9FDE1" box="[658,834,548,572]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA816FFE7FD04FDFDFD76FDE1" box="[658,733,548,572]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA816FFE7FD78FDFDFC92FDE1" box="[750,825,548,572]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
differ from members of the
|
||
<taxonomicName id="4C204D1FA816FFE7FB11FDFDFA37FDE1" authority="Hanwarachelys" authorityName="Hanwarachelys" box="[1159,1436,548,572]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA816FFE7FB11FDFDFB79FDE1" box="[1159,1234,548,572]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA816FFE7FB75FDFDFA39FDE1" box="[1251,1426,548,572]" italics="true" pageId="14" pageNumber="73">Hanwarachelys</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
in the structure of the bridge strut suture, the former having a medial constriction, present in all three species, whereas this is absent in all members of the latter. Further the sutural scar of the ileum in the
|
||
<emphasis id="B954EA8EA816FFE7FB6BFDB4FAA5FD59" box="[1277,1294,621,644]" italics="true" pageId="14" pageNumber="73">E</emphasis>
|
||
. (
|
||
<taxonomicName id="4C204D1FA816FFE7FABCFDB5FADEFD59" box="[1322,1397,620,644]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA816FFE7FABCFDB5FADEFD59" box="[1322,1397,620,644]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
</taxonomicName>
|
||
) is separated from the anterior suture of the 8th pleural with this scar occupying the posterior 2/3 of the 8th pleural and continuing onto the pygal. In
|
||
<emphasis id="B954EA8EA816FFE7FE61FD63FDA3FD0C" box="[503,520,698,721]" italics="true" pageId="14" pageNumber="73">E</emphasis>
|
||
. (
|
||
<emphasis id="B954EA8EA816FFE7FDB5FD60FD78FD0C" box="[547,723,697,721]" italics="true" pageId="14" pageNumber="73">Hanwarachelys</emphasis>
|
||
) this scar is adjacent to the anterior suture of the 8th pleural occupying the full width and also continuing on to the pygal (Figure 2). All members of the
|
||
<emphasis id="B954EA8EA816FFE7FB2FFD07FB61FD28" box="[1209,1226,734,757]" italics="true" pageId="14" pageNumber="73">E</emphasis>
|
||
. (
|
||
<emphasis id="B954EA8EA816FFE7FB75FD04FA39FD28" box="[1251,1426,733,757]" italics="true" pageId="14" pageNumber="73">Hanwarachelys</emphasis>
|
||
) normally possess a cervical scute, absent in
|
||
<emphasis id="B954EA8EA816FFE7FD12FCDBFD3EFCC4" box="[644,661,770,793]" italics="true" pageId="14" pageNumber="73">E</emphasis>
|
||
. (
|
||
<taxonomicName id="4C204D1FA816FFE7FD3AFCD8FD5CFCC4" box="[684,759,769,793]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA816FFE7FD3AFCD8FD5CFCC4" box="[684,759,769,793]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<emphasis id="B954EA8EA816FFE7FCA0FCDBFCECFCC4" box="[822,839,770,793]" italics="true" pageId="14" pageNumber="73">E</emphasis>
|
||
. (
|
||
<taxonomicName id="4C204D1FA816FFE7FCC9FCD8FC52FCC4" box="[863,1017,769,793]" class="Reptilia" family="Chelidae" genus="Pelocomastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA816FFE7FCC9FCD8FC52FCC4" box="[863,1017,769,793]" italics="true" pageId="14" pageNumber="73">Pelocomastes</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F369CA816FFE7FF51FCFCFB1BFC5B" blockId="14.[151,1436,151,902]" pageId="14" pageNumber="73">
|
||
All three species of
|
||
<taxonomicName id="4C204D1FA816FFE7FE27FCFCFD12FCE3" authority="Hawarachelys" authorityName="Hawarachelys" box="[433,697,805,830]" class="Reptilia" family="Chelidae" genus="Elseya" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EA8EA816FFE7FE27FCFCFE57FCE0" box="[433,508,805,829]" italics="true" pageId="14" pageNumber="73">Elseya</emphasis>
|
||
(
|
||
<emphasis id="B954EA8EA816FFE7FD98FCFCFD1BFCE0" box="[526,688,805,829]" italics="true" pageId="14" pageNumber="73">Hawarachelys</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
lack exposed neural bones (sensu
|
||
<bibRefCitation id="EFB14B6DA816FFE7FBDAFCFCFAD4FCE3" author="Thomson" box="[1100,1407,805,830]" pageId="14" pageNumber="73" refString="Thomson, S. A. & Georges, A. (1996) Neural bones in chelid turtles. Chelonian Conservation and Biology, 2, 82 - 86." type="journal article" year="1996">Thomson & Georges 1996</bibRefCitation>
|
||
) a feature common to the entire genus. There are slight differences in the general morphology of the shell between the three species within
|
||
<emphasis id="B954EA8EA816FFE7FEEAFCB7FE26FC58" box="[380,397,878,901]" italics="true" pageId="14" pageNumber="73">E</emphasis>
|
||
. (
|
||
<emphasis id="B954EA8EA816FFE7FE32FCB4FDFFFC58" box="[420,596,877,901]" italics="true" pageId="14" pageNumber="73">Hanwarachelys</emphasis>
|
||
), but subject to considerable within species variation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |