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<document id="06561683A0673270A5F774BFA17AABE1" ID-DOI="10.26879/841" ID-ISSN="1094-8074" ID-Zenodo-Dep="11062827" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1714011148597" checkinUser="felipe" docAuthor="Wang, Hongshan &amp; Dilcher, David L." docDate="2018" docId="03838A22FF89AA3FFEF3F9BEFE94300C" docLanguage="en" docName="PalaeontolElectron.21.3.34A.pdf" docOrigin="Palaeontologia Electronica (34 A) 21 (3)" docSource="http://dx.doi.org/10.26879/841" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.4:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="4" docTitle="Sapindopsis retallackii Wang &amp; Dilcher 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="28" masterDocId="FFBAF25AFF93AA24FFC4FFEEFFCB377D" masterDocTitle="Early Cretaceous angiosperm leaves from the Dakota Formation, Hoisington III locality, Kansas, USA" masterLastPageNumber="49" masterPageNumber="1" pageNumber="27" updateTime="1715622024402" updateUser="carolina" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="FA21E75AD9A96AF49986AEC814AC1C71">Early Cretaceous angiosperm leaves from the Dakota Formation, Hoisington III locality, Kansas, USA</mods:title>
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<mods:namePart id="485AE7F340021DA1E86C36EE2A1B79E6">Wang, Hongshan</mods:namePart>
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<mods:namePart id="790DFBD8E19E377511EEAECFA28C8E39">Dilcher, David L.</mods:namePart>
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<mods:title id="A9118B3F69314F420E1E5FD5DDE10041">Palaeontologia Electronica</mods:title>
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<mods:date id="969376660297629E78C563B9A41880EB">2018</mods:date>
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<mods:title id="E6C6C75B7D12A428BC2E0471C65486A4">34 A</mods:title>
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<mods:number id="D6114F5520896F8110B1D2512217A932">2018-09-30</mods:number>
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<mods:identifier id="3F118AFCFB095AB7D5DF796C9DB83DF0" type="DOI">10.26879/841</mods:identifier>
<mods:identifier id="97C142C178FDB46E923999D32ABE23BB" type="ISSN">1094-8074</mods:identifier>
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<treatment id="03838A22FF89AA3FFEF3F9BEFE94300C" LSID="urn:lsid:plazi:treatment:03838A22FF89AA3FFEF3F9BEFE94300C" httpUri="http://treatment.plazi.org/id/03838A22FF89AA3FFEF3F9BEFE94300C" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<subSubSection id="C33068BFFF89AA3EFEF3F9BEFD693117" box="[311,674,1616,1642]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8B953B34FF89AA3EFEF3F9BEFD693117" blockId="26.[311,674,1616,1674]" box="[311,674,1616,1642]" pageId="26" pageNumber="27">
<heading id="D0DD8C58FF89AA3EFEF3F9BEFD693117" box="[311,674,1616,1642]" centered="true" fontSize="10" level="2" pageId="26" pageNumber="27" reason="7">
<taxonomicName id="4C2A40B7FF89AA3EFEF3F9BEFDF53117" authority="Wang &amp; Dilcher, 2018" authorityName="Wang &amp; Dilcher" authorityYear="2018" box="[311,574,1616,1642]" class="Magnoliopsida" family="Sapindaceae" genus="Sapindopsis" kingdom="Plantae" order="Sapindales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="retallackii" status="sp. nov.">
<emphasis id="B95EE726FF89AA3EFEF3F9BEFDF53117" box="[311,574,1616,1642]" italics="true" pageId="26" pageNumber="27">Sapindopsis retallackii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5A5DFF89AA3EFD81F9BFFD693117" box="[581,674,1617,1642]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33068BFFF89AA3EFE70F99FFDEE31F7" box="[436,549,1649,1674]" pageId="26" pageNumber="27" type="description">
<paragraph id="8B953B34FF89AA3EFE70F99FFDEE31F7" blockId="26.[311,674,1616,1674]" box="[436,549,1649,1674]" pageId="26" pageNumber="27">
<figureCitation id="131127B1FF89AA3EFE70F99FFDEE31F7" box="[436,549,1649,1674]" captionStart="FIGURE 17" captionStartId="27.[197,288,1298,1321]" captionTargetBox="[394,1239,165,1258]" captionTargetId="figure-204@27.[394,1239,165,1259]" captionTargetPageId="27" captionText="FIGURE 17. Sapindopsis retallackii sp. nov. 1, UF15706-3153, showing a leaflet with entire margin. Note the petiolule at the base (indicated by arrow). Scale bar equals 5 mm. 2, Enlargement of Figure 17.1 to show numerous thin secondary and intersecondary veins. Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11062861" httpUri="https://zenodo.org/record/11062861/files/figure.png" pageId="26" pageNumber="27">Figure 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33068BFFF89AA3EFF04F975FC0935E2" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8B953B34FF89AA3EFF04F975FC0935E2" blockId="26.[192,793,1691,1908]" lastBlockId="26.[840,1441,166,1887]" pageId="26" pageNumber="27">
<emphasis id="B95EE726FF89AA3EFF04F975FE7931C9" bold="true" box="[192,434,1691,1717]" pageId="26" pageNumber="27">Specific diagnosis.</emphasis>
Leaf compound, with three or four leaflets; most commonly three leaflets closely arranged on the distal portion of the rachis, occasionally with one leaflet attached at some distance proximally on rachis; rachis thin. Leaflet lorate, always curved abaxially; apex attenuate; base normal acute; lateral leaflets sessile and terminal leaflets petiolulate; margin entire, usually revolute. Primary venation pinnate; primary vein stout; course straight (medial leaflet) or recurved (lateral leaflets). Secondary venation brochidodromous; secondary veins thin relative to primary vein; opposite to subopposite, decurrent; angle of divergence acute, joining superadjacent secondary veins to form a loop enclosing an intercostal area, with two series of loops (tertiary and quaternary in order) in excostal region; exmedial branches of secondary veins common, forming tertiary veins. Intersecondary veins common. Tertiary vein moderate; orthogonal reticulate but tending to be percurrent, retroflexed. Quaternary veins random reticulate, forming incompletely closed meshes. Veinlets simple, linear.
</paragraph>
</subSubSection>
<subSubSection id="C33068BFFF89AA3EFC8CFD48FA6B3162" pageId="26" pageNumber="27" type="description">
<paragraph id="8B953B34FF89AA3EFC8CFD48FA6B3162" blockId="26.[840,1441,166,1887]" pageId="26" pageNumber="27">
<emphasis id="B95EE726FF89AA3EFC8CFD48FC2935C2" bold="true" box="[840,994,678,703]" pageId="26" pageNumber="27">Description.</emphasis>
Leaflet lamina base symmetrical. Form lanceolate,
<collectionCode id="ED3BA3F1FF89AA3EFBDBFD28FBE735A2" box="[1055,1068,710,735]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="26" pageNumber="27" type="Herbarium">L</collectionCode>
/
<collectionCode id="ED3BA3F1FF89AA3EFBF0FD28FB8535A2" box="[1076,1102,710,735]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="26" pageNumber="27" type="Herbarium">W</collectionCode>
&gt;5,
<quantity id="4CD296D1FF89AA3EFB48FD28FB1235A2" box="[1164,1241,710,735]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="27" unit="cm" value="10.0">10 cm</quantity>
long (estimated length) and
<quantity id="4CD296D1FF89AA3EFC1CFD08FBE13582" box="[984,1066,742,767]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="26" pageNumber="27" unit="cm" value="1.8">1.8 cm</quantity>
wide. Apex missing. Base normal, acute; petiolule
<quantity id="4CD296D1FF89AA3EFB8FFCE8FB413462" box="[1099,1162,774,799]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="27" unit="cm" value="1.0">1 cm</quantity>
long and
<quantity id="4CD296D1FF89AA3EFACAFCE8FA9E3462" box="[1294,1365,774,799]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="27" unit="mm" value="1.0">1 mm</quantity>
wide. Margin entire. Petiole normal, short,
<quantity id="4CD296D1FF89AA3EFAC9FCC8FAA93442" box="[1293,1378,806,831]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="27" unit="cm" value="0.8">0.8 cm</quantity>
long and
<quantity id="4CD296D1FF89AA3EFC45FCA8FC153422" box="[897,990,838,863]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="mm" value="0.5">0.5 mm</quantity>
wide. Primary venation pinnate; primary vein stout, slightly curved, up to
<quantity id="4CD296D1FF89AA3EFACCFC88FAAA3402" box="[1288,1377,870,895]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="mm" value="0.5">0.5 mm</quantity>
wide at the widest portion of lamina. Secondary venation festooned brochidodromous; secondary vein fine relative to primary vein; secondary veins diverging from primary vein at narrow acute angles, with lowest one to two pairs more acute than pairs above; secondary veins slightly recurved after diverging from primary vein, then extending a distance to about one fourth to one half of half lamina before joining superadjacent secondary or intersecondary veins to
<taxonomicName id="4C2A40B7FF89AA3EFC71FB48FBD433C2" box="[949,1055,1190,1215]" form="two" pageId="26" pageNumber="27" rank="form">form two</taxonomicName>
series of loops in the excostal region. Intersecondary veins common, one to three pairs per intercostal region; intersecondary veins simple or occasionally forking at variable distance after diverging from primary vein; intersecondary veins almost the same width as secondary veins. Tertiary veins hair-like; diverging from primary veins, exmedial (lower) side of secondary veins or intersecondary veins at moderate acute to narrow acute angles, predominately exmedially ramified, orientation parallel to secondary or intersecondary veins, connected by cross veins of the same order.
</paragraph>
</subSubSection>
<subSubSection id="C33068BFFF89AA3EFC8CF9C8FB8E31C2" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8B953B34FF89AA3EFC8CF9C8FADE3142" blockId="26.[840,1441,166,1887]" box="[840,1301,1574,1599]" pageId="26" pageNumber="27">
<emphasis id="B95EE726FF89AA3EFC8CF9C8FB233142" bold="true" box="[840,1256,1574,1599]" pageId="26" pageNumber="27">Number of specimens examined.</emphasis>
30.
</paragraph>
<paragraph id="8B953B34FF89AA3EFC8CF9A8FAC73122" blockId="26.[840,1441,166,1887]" box="[840,1292,1606,1631]" pageId="26" pageNumber="27">
<materialsCitation id="3B423169FF89AA3EFC8CF9A8FAC73122" box="[840,1292,1606,1631]" collectionCode="UF" pageId="26" pageNumber="27" specimenCode="UF15706-3153" specimenCount="1" typeStatus="holotype">
<emphasis id="B95EE726FF89AA3EFC8CF9A8FC0A3122" bold="true" box="[840,961,1606,1631]" pageId="26" pageNumber="27">
<typeStatus id="54918596FF89AA3EFC8CF9A8FC763122" box="[840,957,1606,1631]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DB8C934FFF89AA3EFC0DF9A8FBB13122" box="[969,1146,1606,1631]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="26" pageNumber="27" type="Museum">UF15706-3153</specimenCode>
(
<figureCitation id="131127B1FF89AA3EFB4DF9A8FACB3122" box="[1161,1280,1606,1631]" captionStart="FIGURE 17" captionStartId="27.[197,288,1298,1321]" captionTargetBox="[394,1239,165,1258]" captionTargetId="figure-204@27.[394,1239,165,1259]" captionTargetPageId="27" captionText="FIGURE 17. Sapindopsis retallackii sp. nov. 1, UF15706-3153, showing a leaflet with entire margin. Note the petiolule at the base (indicated by arrow). Scale bar equals 5 mm. 2, Enlargement of Figure 17.1 to show numerous thin secondary and intersecondary veins. Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11062861" httpUri="https://zenodo.org/record/11062861/files/figure.png" pageId="26" pageNumber="27">Figure 17</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8B953B34FF89AA3EFC8CF988FB8E31C2" blockId="26.[840,1441,166,1887]" pageId="26" pageNumber="27">
<emphasis id="B95EE726FF89AA3EFC8CF988FBDE3102" bold="true" box="[840,1045,1638,1664]" pageId="26" pageNumber="27">Species epithet.</emphasis>
In recognition of Greg Retallack and his contribution to the Dakota Formation geology and paleobotany.
</paragraph>
</subSubSection>
<subSubSection id="C33068BFFF89AA3EFC8CF928FA9731A2" box="[840,1372,1734,1760]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="8B953B34FF89AA3EFC8CF928FA9731A2" blockId="26.[840,1441,166,1887]" box="[840,1372,1734,1760]" pageId="26" pageNumber="27">
<emphasis id="B95EE726FF89AA3EFC8CF928FC39319D" bold="true" box="[840,1010,1734,1760]" pageId="26" pageNumber="27">Occurrences.</emphasis>
Hoisington III locality,
<collectingRegion id="49EEF5D6FF89AA3EFB3FF928FA9331A2" box="[1275,1368,1734,1759]" country="United States of America" name="Kansas" pageId="26" pageNumber="27">Kansas</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C33068BFFF89AA3FFC8CF908FE94300C" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8B953B34FF89AA3FFC8CF908FD1D32AC" blockId="26.[840,1441,166,1887]" lastBlockId="27.[192,793,1432,1905]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<emphasis id="B95EE726FF89AA3EFC8CF908FC0B3182" bold="true" box="[840,960,1766,1791]" pageId="26" pageNumber="27">Remarks.</emphasis>
Only
<specimenCount id="9D2CF0BDFF89AA3EFBCEF908FB7D3182" box="[1034,1206,1766,1791]" count="1" pageId="26" pageNumber="27" type="generic">one specimen</specimenCount>
, representing a terminal leaflet of a compound leaf, is observed from the Hoisington III locality. The venation pattern matches that of the compound leaves from the Springfield locality and Pleasant Dale locality,
<collectingRegion id="49EEF5D6FF88AA3FFF04FA56FEF932AC" box="[192,306,1464,1489]" country="United States of America" name="Nebraska" pageId="27" pageNumber="28">Nebraska</collectingRegion>
(Wang and Dilcher, in preparation).
</paragraph>
<caption id="DF556BBCFF88AA3FFF01FAFCFCE13219" ID-DOI="http://doi.org/10.5281/zenodo.11062861" ID-Zenodo-Dep="11062861" httpUri="https://zenodo.org/record/11062861/files/figure.png" pageId="27" pageNumber="28" startId="27.[197,288,1298,1321]" targetBox="[394,1239,165,1258]" targetPageId="27" targetType="figure">
<paragraph id="8B953B34FF88AA3FFF01FAFCFCE13219" blockId="27.[197,1435,1298,1380]" pageId="27" pageNumber="28">
<emphasis id="B95EE726FF88AA3FFF01FAFCFE8C3254" bold="true" box="[197,327,1298,1321]" pageId="27" pageNumber="28">FIGURE 17.</emphasis>
<taxonomicName id="4C2A40B7FF88AA3FFE8AFAFCFDF03254" authority="Wang &amp; Dilcher, 2018" authorityName="Wang &amp; Dilcher" authorityYear="2018" box="[334,571,1298,1321]" class="Magnoliopsida" family="Sapindaceae" genus="Sapindopsis" kingdom="Plantae" order="Sapindales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="retallackii" status="sp. nov.">
<emphasis id="B95EE726FF88AA3FFE8AFAFCFDF03254" box="[334,571,1298,1321]" italics="true" pageId="27" pageNumber="28">Sapindopsis retallackii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5A5DFF88AA3FFD84FAFCFD5F3254" box="[576,660,1298,1321]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B95EE726FF88AA3FFD5CFAFCFD6E3254" bold="true" box="[664,677,1298,1321]" pageId="27" pageNumber="28">1</emphasis>
, UF15706-3153, showing a leaflet with entire margin. Note the petiolule at the base (indicated by arrow). Scale bar equals 5 mm.
<emphasis id="B95EE726FF88AA3FFCE8FADEFCF2323A" bold="true" box="[812,825,1328,1351]" pageId="27" pageNumber="28">2</emphasis>
, Enlargement of Figure 17.1 to show numerous thin secondary and intersecondary veins. Scale bar equals 1 mm.
</paragraph>
</caption>
<paragraph id="8B953B34FF88AA3FFF3EFA36FE94300C" blockId="27.[192,793,1432,1905]" pageId="27" pageNumber="28">
<taxonomicName id="4C2A40B7FF88AA3FFF3EFA36FDC8328C" authorityName="Wang &amp; Dilcher" authorityYear="2018" box="[250,515,1496,1521]" class="Magnoliopsida" family="Sapindaceae" genus="Sapindopsis" kingdom="Plantae" order="Sapindales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="retallackii">
<emphasis id="B95EE726FF88AA3FFF3EFA36FDC8328C" box="[250,515,1496,1521]" italics="true" pageId="27" pageNumber="28">Sapindopsis retallackii</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C2A40B7FF88AA3FFD5BFA36FEEE316C" class="Magnoliopsida" family="Sapindaceae" genus="Sapindopsis" kingdom="Plantae" order="Sapindales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="undefined-A">
<emphasis id="B95EE726FF88AA3FFD5BFA36FF2B316C" italics="true" pageId="27" pageNumber="28">Sapindopsis</emphasis>
sp. A
</taxonomicName>
(Huang and Dilcher, 1994) in having three leaflets but they differ in that
<taxonomicName id="4C2A40B7FF88AA3FFDD4F9F6FD54314C" authorityName="Wang &amp; Dilcher" authorityYear="2018" box="[528,671,1560,1585]" class="Magnoliopsida" family="Sapindaceae" genus="Sapindopsis" kingdom="Plantae" order="Sapindales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="retallackii">
<emphasis id="B95EE726FF88AA3FFDD4F9F6FD54314C" box="[528,671,1560,1585]" italics="true" pageId="27" pageNumber="28">S. retallackii</emphasis>
</taxonomicName>
occasionally has four lorate leaflets, and brochidodromous secondary venation.
<taxonomicName id="4C2A40B7FF88AA3FFE73F9B6FD74310C" authorityName="Wang &amp; Dilcher" authorityYear="2018" box="[439,703,1624,1649]" class="Magnoliopsida" family="Sapindaceae" genus="Sapindopsis" kingdom="Plantae" order="Sapindales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="retallackii">
<emphasis id="B95EE726FF88AA3FFE73F9B6FD74310C" box="[439,703,1624,1649]" italics="true" pageId="27" pageNumber="28">Sapindopsis retallackii</emphasis>
</taxonomicName>
can be distinguished from other
<taxonomicName id="4C2A40B7FF88AA3FFE2CF996FD1131EC" box="[488,730,1656,1681]" pageId="27" pageNumber="28">
<emphasis id="B95EE726FF88AA3FFE2CF996FDB231EC" box="[488,633,1656,1681]" italics="true" pageId="27" pageNumber="28">Sapindopsis</emphasis>
species
</taxonomicName>
(see previous section) by the following suite of characters: compound leaf with petiolulate terminal leaflets, sessile lateral leaflets, lorate leaflet shape, brochidodromous secondary venation, the presence of exmedial branches of secondary veins that branch into tertiary veins, and common intersecondary veins.
</paragraph>
</subSubSection>
</treatment>
</document>
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