213 lines
23 KiB
XML
213 lines
23 KiB
XML
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<mods:title id="0466F1B3E41DACD67F30AFCC0C713BF8">A taxonomic revision of the genus Hexarhopalus (Coleoptera, Tenebrionidae Cnodalonini) from China</mods:title>
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<mods:namePart id="8EAD20DDA57BF5D7645CB8D63FB48D6C">Jiang, Ri-Xin</mods:namePart>
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<mods:affiliation id="CFCEC653D0102E58678F85824D53EB26">College of Marine Science and Biological Engineering, Qingdao University of Science and Technology, 53 Zhengzhou Road, Qingdao, 266042, P. R. China. & maoshuwuyouzhi @ 163. com; https: // orcid. org / 0000 - 0002 - 5339 - 853 X & 3581505520 @ qq. com; https: // orcid. org / 0000 - 0001 - 7919 - 8432 & shuowang @ qust. edu. cn; https: // orcid. org / 0000 - 0003 - 0412 - 3799</mods:affiliation>
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<mods:namePart id="9DC1D3D772DAC24AE88FD62BF548C20A">Bai, Xing-Long</mods:namePart>
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<mods:affiliation id="78424A8C1C1D3179F0085C47528AD7A6">College of Life Science, Institute of Life Science and Green Development, Hebei University, Baoding, Hebei, 071002, P. R. China Corresponding author. & bxlngg @ 163. com; https: // orcid. org / 0000 - 0002 - 0448 - 8272 & gdren @ hbu. edu. cn; https: // orcid. org / 0000 - 0001 - 5808 - 9122</mods:affiliation>
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<mods:namePart id="F44163B8ECEAD93EA09F79650D52DB75">Ren, Guo-Dong</mods:namePart>
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<mods:affiliation id="429BF67B896838185522734AFE1A5FCC">College of Life Science, Institute of Life Science and Green Development, Hebei University, Baoding, Hebei, 071002, P. R. China Corresponding author. & bxlngg @ 163. com; https: // orcid. org / 0000 - 0002 - 0448 - 8272 & gdren @ hbu. edu. cn; https: // orcid. org / 0000 - 0001 - 5808 - 9122</mods:affiliation>
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<mods:namePart id="F385D61839B9DE3F0A28BE141A139759">Yang, Hui-Yu</mods:namePart>
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<mods:affiliation id="8BBA1541B53C228BCE2987430962215F">College of Marine Science and Biological Engineering, Qingdao University of Science and Technology, 53 Zhengzhou Road, Qingdao, 266042, P. R. China. & maoshuwuyouzhi @ 163. com; https: // orcid. org / 0000 - 0002 - 5339 - 853 X & 3581505520 @ qq. com; https: // orcid. org / 0000 - 0001 - 7919 - 8432 & shuowang @ qust. edu. cn; https: // orcid. org / 0000 - 0003 - 0412 - 3799</mods:affiliation>
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<mods:namePart id="7156CB1B7E38A9CDFF5144EA1E14FEE7">Wang, Shuo</mods:namePart>
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<mods:affiliation id="B1CD2ECDA141A52C2FEEDC1DCA2EF270">College of Marine Science and Biological Engineering, Qingdao University of Science and Technology, 53 Zhengzhou Road, Qingdao, 266042, P. R. China. & maoshuwuyouzhi @ 163. com; https: // orcid. org / 0000 - 0002 - 5339 - 853 X & 3581505520 @ qq. com; https: // orcid. org / 0000 - 0001 - 7919 - 8432 & shuowang @ qust. edu. cn; https: // orcid. org / 0000 - 0003 - 0412 - 3799 & shuowang @ qust. edu. cn; https: // orcid. org / 0000 - 0003 - 0412 - 3799</mods:affiliation>
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<mods:title id="858E51EE138AE045E6AADFD26F75294F">Zootaxa</mods:title>
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<mods:part id="BDFD22D57A3A0B83F2396DBFD68730FA">
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<mods:date id="5B23B8EF622909CB7F4130DFECD336D6">2020</mods:date>
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<mods:number id="A400F773DEB328AB7705DFD47F1868E3">2020-07-31</mods:number>
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<mods:number id="E8DB955B72B617189017E5B2CE74054D">4821</mods:number>
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<treatment id="038382170D48703A2FBD4674FE4DFF6A" ID-DOI="http://doi.org/10.5281/zenodo.4441054" ID-GBIF-Taxon="176580102" ID-Zenodo-Dep="4441054" LSID="urn:lsid:plazi:treatment:038382170D48703A2FBD4674FE4DFF6A" httpUri="http://treatment.plazi.org/id/038382170D48703A2FBD4674FE4DFF6A" lastPageId="10" lastPageNumber="286" pageId="9" pageNumber="286">
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<subSubSection id="C330608A0D4870392FBD4674FCA6F983" box="[151,797,1570,1597]" pageId="9" pageNumber="286" type="nomenclature">
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<paragraph id="8B9533010D4870392FBD4674FCA6F983" blockId="9.[151,797,1570,1632]" box="[151,797,1570,1597]" pageId="9" pageNumber="286">
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<heading id="D0DD846D0D4870392FBD4674FCA6F983" bold="true" box="[151,797,1570,1597]" fontSize="11" level="1" pageId="9" pageNumber="286" reason="1">
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<taxonomicName id="4C2A48820D4870392FBD4674FCA6F983" authority="(Pic, 1922)" baseAuthorityName="Pic" baseAuthorityYear="1922" box="[151,797,1570,1597]" class="Insecta" family="Tenebrionidae" genus="Hexarhopalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="286" phylum="Arthropoda" rank="species" species="attenuatus" subGenus="Hexarhopalus">
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<emphasis id="B95EEF130D4870392FBD4674FCA6F983" bold="true" box="[151,797,1570,1597]" pageId="9" pageNumber="286">
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<emphasis id="B95EEF130D4870392FBD4674FD29F982" bold="true" box="[151,658,1570,1597]" italics="true" pageId="9" pageNumber="286">Hexarhopalus (Hexarhopalus) attenuatus</emphasis>
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||
(Pic, 1922)
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||
</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C330608A0D4870392FBD4610FEC9F9DF" box="[151,370,1606,1632]" pageId="9" pageNumber="286" type="description">
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<paragraph id="8B9533010D4870392FBD4610FEC9F9DF" blockId="9.[151,797,1570,1632]" box="[151,370,1606,1632]" pageId="9" pageNumber="286">
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<heading id="D0DD846D0D4870392FBD4610FEC9F9DF" bold="true" box="[151,370,1606,1632]" fontSize="11" level="1" pageId="9" pageNumber="286" reason="1">
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<emphasis id="B95EEF130D4870392FBD4610FEC9F9DF" bold="true" box="[151,370,1606,1632]" pageId="9" pageNumber="286">
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(
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<figureCitation id="13112F840D4870392FB54610FF4EF9DF" box="[159,245,1606,1632]" captionStart="FIGURE 20" captionStartId="22.[151,250,1872,1896]" captionTargetBox="[210,1371,351,1840]" captionTargetId="figure-58@22.[201,1386,345,1848]" captionTargetPageId="22" captionText="FIGURE 20. Types of Chinese Hexarhopalus species.A. Hexarhopalus (H.) mangshanicus, Ren & xu, 2011, holotype, male; B. Hexarhopalus (L.) bisinuatus, Ren & xu, 2011, holotype, male; C. Hexarhopalus (L.) xui, Ren & xu, 2011, holotype, female; D. Hexarhopalus (H.) attenuatus (Pic, 1922), paralectotype; E. Same, in lateral view (20A–C deposited in MHBU; 20D deposited in HNHM; figs 20D–E were taken by Tamás Németh)." figureDoi="http://doi.org/10.5281/zenodo.4440980" httpUri="https://zenodo.org/record/4440980/files/figure.png" pageId="9" pageNumber="286">Figs 20</figureCitation>
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D–E, 21A)
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C330608A0D4870392FBD46DDFC26F95F" pageId="9" pageNumber="286" type="reference_group">
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<paragraph id="8B9533010D4870392FBD46DDFB3BF97D" blockId="9.[151,1426,1675,1760]" pageId="9" pageNumber="286">
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<treatmentCitationGroup id="AB3A142F0D4870392FBD46DDFB3BF97D" pageId="9" pageNumber="286">
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<treatmentCitation id="0A8B15100D4870392FBD46DDFDA4F91C" author="Pic, M." box="[151,543,1675,1699]" page="24" pageId="9" pageNumber="286" year="1922">
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<taxonomicName id="4C2A48820D4870392FBD46DDFDA4F91C" authority="Pic, 1922 a: 24" authorityName="Pic" authorityPageNumber="24" authorityYear="1922" box="[151,543,1675,1699]" class="Insecta" family="Tenebrionidae" genus="Leprocaulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Phasmida" pageId="9" pageNumber="286" phylum="Arthropoda" rank="species" species="attenuatus">
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<emphasis id="B95EEF130D4870392FBD46DDFE3EF91C" box="[151,389,1675,1699]" italics="true" pageId="9" pageNumber="286">Leprocaulus attenuatus</emphasis>
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<bibRefCitation id="EFBB4EF00D4870392EA146DDFDA4F91C" author="Pic, M." box="[395,543,1675,1699]" pageId="9" pageNumber="286" pagination="504 - 505" refId="ref10941" refString="Pic, M. (1922 a) Sur deux Leprocaulus (Coleopteres, Heteromeres). Bulletin du Museum National d'Histoire Naturelle Paris, 28, 504 - 505." type="journal article" year="1922">Pic, 1922a: 24</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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;
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||
<treatmentCitation id="0A8B15100D4870392D0046DDFD51F91C" author="Kaszab, Z." box="[554,746,1675,1699]" page="177" pageId="9" pageNumber="286" year="1983">
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<bibRefCitation id="EFBB4EF00D4870392D0046DDFD51F91C" author="Kaszab, Z." box="[554,746,1675,1699]" pageId="9" pageNumber="286" pagination="177 - 183" refId="ref10828" refString="Kaszab, Z. (1983) Uber die Gattung Leprocaulus Fairmaire, 1896 (Coleoptera, Tenebrionidae). Annales Historico-Naturales Musei Nationalis Hungarici, 75, 177 - 183." type="journal article" year="1983">Kaszab, 1983: 177</bibRefCitation>
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</treatmentCitation>
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;
|
||
<treatmentCitation id="0A8B15100D4870392DDF46DDFC76F91C" author="Lobl, I. & Merkl, O. & Ando, K. & Bouchard, P. & Lillig, M. & Masumoto, K. & Schawaller, W." box="[757,973,1675,1699]" page="343" pageId="9" pageNumber="286" year="2008">
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<bibRefCitation id="EFBB4EF00D4870392DDF46DDFC76F91C" author="Lobl, I. & Merkl, O. & Ando, K. & Bouchard, P. & Lillig, M. & Masumoto, K. & Schawaller, W." box="[757,973,1675,1699]" pageId="9" pageNumber="286" pagination="1802" refId="ref10860" refString="Lobl, I., Merkl, O., Ando, K., Bouchard, P., Lillig, M., Masumoto, K. & Schawaller, W. (2008) Family Tenebrionidae Latreille, 1802. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 5. Tenebrionoidea. Apollo Books, Stenstrup, pp. 105 - 352." type="book chapter" year="2008">
|
||
Löbl
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||
<emphasis id="B95EEF130D4870392C0146DAFCE5F91C" box="[811,862,1675,1699]" italics="true" pageId="9" pageNumber="286">et al.</emphasis>
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2008: 343
|
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</bibRefCitation>
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</treatmentCitation>
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.
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||
<typeStatus id="54918DA30D4870392CFC46DAFBB3F91B" box="[982,1032,1676,1700]" pageId="9" pageNumber="286">Type</typeStatus>
|
||
locality.
|
||
<collectingCountry id="F33D73910D4870392B4C46DDFB1DF91C" box="[1126,1190,1675,1699]" name="China" pageId="9" pageNumber="286">China</collectingCountry>
|
||
:
|
||
<collectingRegion id="49EEFDE30D4870392B8546DAFB47F91C" box="[1199,1276,1676,1699]" country="China" name="Yunnan" pageId="9" pageNumber="286">Yunnan</collectingRegion>
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||
(Pe Yen Tsin).
|
||
<taxonomicName id="4C2A48820D4870392FBD46FCFD78F97D" authority="Pic, 1922 a" authorityName="Pic" authorityYear="1922" box="[151,707,1706,1730]" class="Insecta" family="Tenebrionidae" genus="Leprocaulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Phasmida" pageId="9" pageNumber="286" phylum="Arthropoda" rank="species" species="attenuatus" subGenus="Pseudocoelophus">
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||
<emphasis id="B95EEF130D4870392FBD46FCFEAEF97D" box="[151,277,1706,1730]" italics="true" pageId="9" pageNumber="286">Leprocaulus</emphasis>
|
||
(
|
||
<emphasis id="B95EEF130D4870392E0E46FCFE6EF97D" box="[292,469,1706,1730]" italics="true" pageId="9" pageNumber="286">Pseudocoelophus</emphasis>
|
||
)
|
||
<emphasis id="B95EEF130D4870392ECE46FCFDF5F97E" box="[484,590,1706,1729]" italics="true" pageId="9" pageNumber="286">attenuatus</emphasis>
|
||
<bibRefCitation id="EFBB4EF00D4870392D7E46FCFD78F97D" author="Pic, M." box="[596,707,1706,1730]" pageId="9" pageNumber="286" pagination="504 - 505" refId="ref10941" refString="Pic, M. (1922 a) Sur deux Leprocaulus (Coleopteres, Heteromeres). Bulletin du Museum National d'Histoire Naturelle Paris, 28, 504 - 505." type="journal article" year="1922">Pic, 1922a</bibRefCitation>
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</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0A8B15100D4870392DE646FCFC32F97D" author="Kaszab, Z." box="[716,905,1706,1730]" page="179" pageId="9" pageNumber="286" year="1983">
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<bibRefCitation id="EFBB4EF00D4870392DE646FCFC32F97D" author="Kaszab, Z." box="[716,905,1706,1730]" pageId="9" pageNumber="286" pagination="177 - 183" refId="ref10828" refString="Kaszab, Z. (1983) Uber die Gattung Leprocaulus Fairmaire, 1896 (Coleoptera, Tenebrionidae). Annales Historico-Naturales Musei Nationalis Hungarici, 75, 177 - 183." type="journal article" year="1983">Kaszab, 1983: 179</bibRefCitation>
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</treatmentCitation>
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||
(with photo of habitus).
|
||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B9533010D4870392FBD469EFC26F95F" blockId="9.[151,1426,1675,1760]" box="[151,925,1736,1760]" pageId="9" pageNumber="286">
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<treatmentCitationGroup id="AB3A142F0D4870392FBD469EFC26F95F" box="[151,925,1736,1760]" pageId="9" pageNumber="286">
|
||
<taxonomicName id="4C2A48820D4870392FBD469EFDD2F95F" authority="(Pic, 1922 a)" baseAuthorityName="Pic" baseAuthorityYear="1922" box="[151,617,1736,1760]" class="Insecta" family="Tenebrionidae" genus="Hexarhopalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="286" phylum="Arthropoda" rank="species" species="attenuatus">
|
||
<emphasis id="B95EEF130D4870392FBD469EFE9DF95F" box="[151,294,1736,1760]" italics="true" pageId="9" pageNumber="286">Hexarhopalus</emphasis>
|
||
(
|
||
<emphasis id="B95EEF130D4870392E1F469FFED0F95F" box="[309,363,1737,1760]" italics="true" pageId="9" pageNumber="286">s. str.</emphasis>
|
||
)
|
||
<emphasis id="B95EEF130D4870392E53469FFE58F95F" box="[377,483,1737,1760]" italics="true" pageId="9" pageNumber="286">attenuatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBB4EF00D4870392EDA469EFDDAF95F" author="Pic, M." box="[496,609,1736,1760]" pageId="9" pageNumber="286" pagination="504 - 505" refId="ref10941" refString="Pic, M. (1922 a) Sur deux Leprocaulus (Coleopteres, Heteromeres). Bulletin du Museum National d'Histoire Naturelle Paris, 28, 504 - 505." type="journal article" year="1922">Pic, 1922a</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0A8B15100D4870392D58469EFC23F95F" author="Becvar, S. & Purchart, L." box="[626,920,1736,1760]" page="40" pageId="9" pageNumber="286" year="2008">
|
||
<bibRefCitation id="EFBB4EF00D4870392D58469EFC23F95F" author="Becvar, S. & Purchart, L." box="[626,920,1736,1760]" pageId="9" pageNumber="286" pagination="35 - 70" refId="ref10437" refString="Becvar, S. & Purchart, L. (2008) Revision of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera: Tenebrionidae: Cnodaloninae), with description of Malaysphena gen. nov. Annales Zoologici, 58, 35 - 70. https: // doi. org / 10.3161 / 067.058.0102" type="journal article" year="2008">Bečvář & Purchart, 2008: 40</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C330608A0D48703A2FBD4759FE4DFF6A" lastPageId="10" lastPageNumber="287" pageId="9" pageNumber="286" type="discussion">
|
||
<paragraph id="8B9533010D48703A2FBD4759FE4DFF6A" blockId="9.[151,1437,1806,2013]" lastBlockId="10.[151,1436,151,213]" lastPageId="10" lastPageNumber="287" pageId="9" pageNumber="286">
|
||
<emphasis id="B95EEF130D4870392FBD4759FEBBF896" bold="true" box="[151,256,1807,1833]" pageId="9" pageNumber="286">Remark.</emphasis>
|
||
The
|
||
<typeStatus id="54918DA30D4870392E174759FED6F896" box="[317,365,1807,1833]" pageId="9" pageNumber="286">type</typeStatus>
|
||
locality ‘Pe Yen Tsin’ is now called Shiyang Town (
|
||
<emphasis id="B95EEF130D4870392CE04746FBA5F898" bold="true" box="[970,1054,1808,1831]" pageId="9" pageNumber="286">Ƃŧẘ</emphasis>
|
||
) of Dayao County (
|
||
<emphasis id="B95EEF130D4870392A2D4746FAE0F898" bold="true" box="[1287,1371,1808,1831]" pageId="9" pageNumber="286">Ęďss</emphasis>
|
||
), and formerly called Baiyanjing (
|
||
<emphasis id="B95EEF130D4870392ECA4762FD88F8F4" bold="true" box="[480,563,1844,1867]" pageId="9" pageNumber="286">àü#</emphasis>
|
||
) of Yanfeng County (
|
||
<emphasis id="B95EEF130D4870392C1D4762FC30F8F4" bold="true" box="[823,907,1844,1867]" pageId="9" pageNumber="286">üƚss</emphasis>
|
||
).
|
||
<bibRefCitation id="EFBB4EF00D4870392C8F4765FBC0F8F2" author="Ren, G. - D. & Xu, J. - S." box="[933,1147,1842,1869]" pageId="9" pageNumber="286" pagination="23 - 34" refId="ref11188" refString="Ren, G. - D. & Xu, J. - S. (2011) The genus Hexarhopalus Fairmaire, 1891 in China, with description of three new species (Coleoptera, Tenebrionidae, Cnodalonini), Acta Zoologica Academiae Scientiarum Hungaricae, 57, 23 - 34." type="journal article" year="2011">Ren and Xu, 2011</bibRefCitation>
|
||
identified the specimens from Ailao Mountain (
|
||
<collectingRegion id="49EEFDE30D4870392EBD4701FDE0F8CE" box="[407,603,1878,1905]" country="China" name="Yunnan" pageId="9" pageNumber="286">Yunnan Province</collectingRegion>
|
||
) as
|
||
<taxonomicName id="4C2A48820D4870392DA14700FC67F8CF" baseAuthorityName="Pic" baseAuthorityYear="1922" box="[651,988,1878,1905]" class="Insecta" family="Tenebrionidae" genus="Hexarhopalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="286" phylum="Arthropoda" rank="species" species="attenuatus" subGenus="Hexarhopalus">
|
||
<emphasis id="B95EEF130D4870392DA14700FC92F8CF" box="[651,809,1878,1904]" italics="true" pageId="9" pageNumber="286">Hexarhopalus</emphasis>
|
||
(
|
||
<emphasis id="B95EEF130D4870392C104701FCEDF8CF" box="[826,854,1879,1904]" italics="true" pageId="9" pageNumber="286">H.</emphasis>
|
||
)
|
||
<emphasis id="B95EEF130D4870392C4D4701FC67F8CF" box="[871,988,1879,1904]" italics="true" pageId="9" pageNumber="286">attenuatus</emphasis>
|
||
</taxonomicName>
|
||
. However, we examined the specimens from the same locality, and the different characters suggest that specimens from Ailao Mountain is a new species. Although they are similar in general appearance, and both possess three longitudinal grooves on pronotum. But they can be easily separated from each other, the new species with shoulders fully developed (shouldersless in
|
||
<taxonomicName id="4C2A48820D4B703A2FBD40C1FE5CFF0F" baseAuthorityName="Pic" baseAuthorityYear="1922" box="[151,487,151,177]" class="Insecta" family="Tenebrionidae" genus="Hexarhopalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="attenuatus" subGenus="Hexarhopalus">
|
||
<emphasis id="B95EEF130D4B703A2FBD40C1FE8EFF0E" box="[151,309,151,177]" italics="true" pageId="10" pageNumber="287">Hexarhopalus</emphasis>
|
||
(
|
||
<emphasis id="B95EEF130D4B703A2E6C40C1FED9FF0F" box="[326,354,151,176]" italics="true" pageId="10" pageNumber="287">H.</emphasis>
|
||
)
|
||
<emphasis id="B95EEF130D4B703A2E5840C1FE5CFF0F" box="[370,487,151,176]" italics="true" pageId="10" pageNumber="287">attenuatus</emphasis>
|
||
</taxonomicName>
|
||
), and except the lateral one, other two grooves of pronotum are much weak (deep in
|
||
<taxonomicName id="4C2A48820D4B703A2FBD40EDFE5DFF6B" baseAuthorityName="Pic" baseAuthorityYear="1922" box="[151,486,187,213]" class="Insecta" family="Tenebrionidae" genus="Hexarhopalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="attenuatus" subGenus="Hexarhopalus">
|
||
<emphasis id="B95EEF130D4B703A2FBD40EDFE8EFF6A" box="[151,309,187,213]" italics="true" pageId="10" pageNumber="287">Hexarhopalus</emphasis>
|
||
(
|
||
<emphasis id="B95EEF130D4B703A2E6C40EDFEDAFF6B" box="[326,353,187,212]" italics="true" pageId="10" pageNumber="287">H.</emphasis>
|
||
)
|
||
<emphasis id="B95EEF130D4B703A2E5B40EDFE5DFF6B" box="[369,486,187,212]" italics="true" pageId="10" pageNumber="287">attenuatus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |