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<document id="5450BE54B7AB9B1B9E1BFABFAE6197BF" ID="10.11646/zootaxa.4258.6.3SLASH10954" ID-CLB-Dataset="33117" ID-DOI="10.11646/zootaxa.4258.6.3" ID-GBIF-Dataset="777f9d1d-53e8-4a40-ae71-3e93569f5eb4" ID-ISSN="1175-5326" ID-Zenodo-Dep="571027" ID-ZooBank="D8121CF9-5F38-45B1-AB19-CAD03A95CC0B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493797901851" checkinUser="plazi" docAuthor="Pui, Yong Min, Karin, Benjamin R., Bauer, Aaron M. &amp; Das, Indraneil" docDate="2017" docId="03A7FF78FF899960FF556B487D9DFE52" docLanguage="en" docName="zootaxa.4258.6.3SLASH10954.pdf" docOrigin="Zootaxa 4258 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tropidophorus sebi Pui, Karin, Bauer &amp; Das, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="547" masterDocId="FF9E8700FF8B9968FFC268557C28FF9D" masterDocTitle="A new species of Tropidophorus Duméril &amp; Bibron, 1839 (Squamata: Sauria: Scincidae) from Sarawak, East Malaysia (Borneo)" masterLastPageNumber="550" masterPageNumber="539" pageNumber="541" updateTime="1698439227195" updateUser="ExternalLinkService">
<mods:mods id="235119FA4D6677BD8B29F074EEE470EF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B39DEE8AACD6C3E88F4A690DA8BF7E1A">A new species of Tropidophorus Duméril &amp; Bibron, 1839 (Squamata: Sauria: Scincidae) from Sarawak, East Malaysia (Borneo)</mods:title>
</mods:titleInfo>
<mods:name id="D1C031F1E33A8FFD7D7E050233EF52AC" type="personal">
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<mods:namePart id="F179790E046181C46484C18DCDE7EE3B">Pui, Yong Min</mods:namePart>
</mods:name>
<mods:name id="3FD88B39B5FA87816F83533AC9F3FA14" type="personal">
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<mods:namePart id="59E2B33F847069C978B2112AF863F930">Karin, Benjamin R.</mods:namePart>
</mods:name>
<mods:name id="DA17685519960AB3853E235390BD1FBB" type="personal">
<mods:role id="C3E412A2FFBF06BB1115464C09F62B90">
<mods:roleTerm id="8484783F50461822CD272A4CB3487B0B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ACFFFA4D2A7F2DBE59A63EFCF0E6D933">Bauer, Aaron M.</mods:namePart>
</mods:name>
<mods:name id="D6BA9865D33BC89E7FC7CA0A5DFB52D1" type="personal">
<mods:role id="7BF872B52D8FB536B64F39750DA0CCD3">
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<mods:namePart id="56B1F94B4E89DEA61886154F3A93A75E">Das, Indraneil</mods:namePart>
</mods:name>
<mods:typeOfResource id="C0319D07A75E1BE80EC483161A9C252E">text</mods:typeOfResource>
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<mods:title id="94CBD87C418836176124DB95D79EA0A8">Zootaxa</mods:title>
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<mods:part id="6F90B8E4867A656AC0039E52B9B0558D">
<mods:date id="398928A14C69D2AF86981BD1A2D74E1D">2017</mods:date>
<mods:detail id="C0D225D8D6F304DF483B7EA06DF5696A" type="volume">
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<mods:identifier id="FAF6F408E993134D26BEA7CCF236994C" type="CLB-Dataset">33117</mods:identifier>
<mods:identifier id="BEED6CA131A514C3011F39A2885A3F2D" type="DOI">10.11646/zootaxa.4258.6.3</mods:identifier>
<mods:identifier id="5EE91F3B06E3E77232656A45F8CCB1E9" type="GBIF-Dataset">777f9d1d-53e8-4a40-ae71-3e93569f5eb4</mods:identifier>
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<treatment id="03A7FF78FF899960FF556B487D9DFE52" ID-DOI="http://doi.org/10.5281/zenodo.6049585" ID-GBIF-Taxon="128835047" ID-Zenodo-Dep="6049585" LSID="urn:lsid:plazi:treatment:03A7FF78FF899960FF556B487D9DFE52" httpUri="http://treatment.plazi.org/id/03A7FF78FF899960FF556B487D9DFE52" lastPageId="8" lastPageNumber="547" pageId="2" pageNumber="541">
<subSubSection id="C3141DE5FF89996AFF556B487D3AFCC5" pageId="2" pageNumber="541" type="nomenclature">
<paragraph id="8BB14E6EFF89996AFF556B487DCEFCAA" blockId="2.[151,486,797,856]" box="[151,486,797,823]" pageId="2" pageNumber="541">
<heading id="D0F9F902FF89996AFF556B487DCEFCAA" bold="true" box="[151,486,797,823]" fontSize="11" level="1" pageId="2" pageNumber="541" reason="1">
<emphasis id="B97A927CFF89996AFF556B487DCEFCAA" bold="true" box="[151,486,797,823]" pageId="2" pageNumber="541">
<taxonomicName id="4C0E35EDFF89996AFF556B487DA9FCAA" box="[151,385,797,823]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="541" phylum="Chordata" rank="species" species="sebi" status="sp. nov.">
<emphasis id="B97A927CFF89996AFF556B487DA9FCAA" bold="true" box="[151,385,797,823]" italics="true" pageId="2" pageNumber="541">Tropidophorus sebi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2492F07FF89996AFE4A6B487DCEFCAA" box="[392,486,797,823]" pageId="2" pageNumber="541" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB14E6EFF89996AFF556B6A7D3AFCC5" blockId="2.[151,486,797,856]" box="[151,274,831,856]" pageId="2" pageNumber="541">
(
<figureCitation id="133552EBFF89996AFF5D6B6A7D20FCC5" box="[159,264,831,856]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1322,1344]" captionStartId-1="4.[151,250,1061,1083]" captionTargetBox-0="[151,1436,589,1301]" captionTargetBox-1="[344,1240,197,1032]" captionTargetId-0="figure@3.[151,1436,589,1301]" captionTargetId-1="figure@4.[340,1247,193,1040]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Female holotype of Tropidophorus sebi sp. nov. (UNIMAS P 1167) in life. (SVL = 83.8 mm)." captionText-1="FIGURE 2. Female holotype of Tropidophorus sebi sp. nov. (UNIMAS P 1167), showing head in a. lateral; b. ventral; and c. dorsal views (scale markers = 10 mm); and d. pes and e. manus. (scale markers = 1 mm)." httpUri-0="https://zenodo.org/record/571028/files/figure.png" httpUri-1="https://zenodo.org/record/571029/files/figure.png" pageId="2" pageNumber="541">Figs. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3141DE5FF89996AFF556BD278CEFBAD" pageId="2" pageNumber="541" type="materials_examined">
<paragraph id="8BB14E6EFF89996AFF556BD27817FB91" blockId="2.[151,1437,903,2009]" pageId="2" pageNumber="541">
<materialsCitation id="3B664433FF89996AFF556BD27817FB91" ID-GBIF-Occurrence="1499986382" collectingDate="2015-05-19" collectionCode="UNIMAS" collectorName="Pui Yong Min &amp; Mohamad Paisal bin Wahab &amp; Adult &amp; The &amp; Fig." country="Malaysia" county="Kapit district" elevation="117" location="Museum" municipality="upper Baleh" pageId="2" pageNumber="541" specimenCount="2" stateProvince="Sarawak" typeStatus="holotype">
<typeStatus id="54B5F0CCFF89996AFF556BD27D2CFC3D" box="[151,260,903,928]" pageId="2" pageNumber="541" type="holotype">
<emphasis id="B97A927CFF89996AFF556BD27D2CFC3D" bold="true" box="[151,260,903,928]" pageId="2" pageNumber="541">Holotype</emphasis>
</typeStatus>
.
<location id="8ED118B5FF89996AFED46BDD7D50FC02" LSID="urn:lsid:plazi:treatment:03A7FF78FF899960FF556B487D9DFE52:8ED118B5FF89996AFED46BDD7D50FC02" box="[278,376,904,927]" country="Malaysia" county="Kapit district" municipality="upper Baleh" name="Museum" pageId="2" pageNumber="541" stateProvince="Sarawak">Museum</location>
of the
<location id="8ED118B5FF89996AFE106BD27E06FC3D" LSID="urn:lsid:plazi:treatment:03A7FF78FF899960FF556B487D9DFE52:8ED118B5FF89996AFE106BD27E06FC3D" box="[466,558,903,928]" country="Malaysia" county="Kapit district" municipality="upper Baleh" name="Institute" pageId="2" pageNumber="541" stateProvince="Sarawak">Institute</location>
of
<location id="8ED118B5FF89996AFD996BD27ECEFC3D" LSID="urn:lsid:plazi:treatment:03A7FF78FF899960FF556B487D9DFE52:8ED118B5FF89996AFD996BD27ECEFC3D" box="[603,742,903,928]" country="Malaysia" county="Kapit district" municipality="upper Baleh" name="Biodiversity" pageId="2" pageNumber="541" stateProvince="Sarawak">Biodiversity</location>
and
<location id="8ED118B5FF89996AFCE66BD27845FC3D" LSID="urn:lsid:plazi:treatment:03A7FF78FF899960FF556B487D9DFE52:8ED118B5FF89996AFCE66BD27845FC3D" box="[804,1133,903,928]" country="Malaysia" county="Kapit district" municipality="upper Baleh" name="Environmental Conservation" pageId="2" pageNumber="541" stateProvince="Sarawak">Environmental Conservation</location>
,
<collectionCode id="ED1FD6ABFF89996AFBBE6BDD78C2FC3D" box="[1148,1258,904,928]" country="Malaysia" httpUri="http://grbio.org/cool/z1cp-qx8g" name="Universiti Malaysia Sarawak" pageId="2" pageNumber="541">UNIMAS</collectionCode>
P1167, from a small stream located at the
<location id="8ED118B5FF89996AFE0E6BF97ED4FC59" LSID="urn:lsid:plazi:treatment:03A7FF78FF899960FF556B487D9DFE52:8ED118B5FF89996AFE0E6BF97ED4FC59" box="[460,764,940,965]" country="Malaysia" county="Kapit district" municipality="upper Baleh" name="First Count Logging Camp" pageId="2" pageNumber="541" stateProvince="Sarawak">First Count Logging Camp</location>
(01.35.644°N; 113.47.377°E; datum WGS84),
<location id="8ED118B5FF89996AFAD66BF97978FC58" LSID="urn:lsid:plazi:treatment:03A7FF78FF899960FF556B487D9DFE52:8ED118B5FF89996AFAD66BF97978FC58" box="[1300,1360,940,965]" country="Malaysia" county="Kapit district" municipality="upper Baleh" name="Putai" pageId="2" pageNumber="541" stateProvince="Sarawak">Putai</location>
,
<collectingMunicipality id="6BD5D414FF89996AFA9F6BF87CF3FC75" pageId="2" pageNumber="541">upper Baleh</collectingMunicipality>
,
<collectingCounty id="62D036E2FF89996AFF2B6B9A7DA9FC75" box="[233,385,975,1000]" pageId="2" pageNumber="541">Kapit district</collectingCounty>
,
<collectingRegion id="49CA808CFF89996AFE4C6B9A7DD9FC75" box="[398,497,975,1000]" country="Malaysia" name="Sarawak" pageId="2" pageNumber="541">Sarawak</collectingRegion>
, East
<collectingCountry id="F3190EFEFF89996AFDFB6B9A7E88FC75" box="[569,672,975,1000]" name="Malaysia" pageId="2" pageNumber="541">Malaysia</collectingCountry>
(Borneo), altitude
<elevation id="0023A95DFF89996AFCBE6B857FECFC7A" box="[892,964,976,1000]" metricMagnitude="2" metricUnit="m" metricValue="1.17" pageId="2" pageNumber="541" unit="m" value="117.0">
<quantity id="4CF6E38BFF89996AFCBE6B857FECFC7A" box="[892,964,976,1000]" metricMagnitude="2" metricUnit="m" metricValue="1.17" pageId="2" pageNumber="541" unit="m" value="117.0">117 m</quantity>
</elevation>
ASL, coll.
<collectorName id="26FB2BB8FF89996AFB8F6B9A78D8FC75" box="[1101,1264,975,1000]" pageId="2" pageNumber="541">Pui Yong Min</collectorName>
and
<collectorName id="26FB2BB8FF89996AFAE86B9A7D74FB90" pageId="2" pageNumber="541">Mohamad Paisal bin Wahab</collectorName>
,
<collectingDate id="EFF49146FF89996AFEA46BA17E28FB90" box="[358,512,1012,1037]" pageId="2" pageNumber="541" value="2015-05-19">
<date id="FFB068AEFF89996AFEA46BA17E28FB90" box="[358,512,1012,1037]" pageId="2" pageNumber="541" value="2015-05-19">19 May 2015</date>
</collectingDate>
.
<collectorName id="26FB2BB8FF89996AFDC86BA17E62FB90" box="[522,586,1012,1037]" pageId="2" pageNumber="541">Adult</collectorName>
female.
<collectorName id="26FB2BB8FF89996AFD696BA17EFFFB90" box="[683,727,1012,1037]" pageId="2" pageNumber="541">The</collectorName>
<typeStatus id="54B5F0CCFF89996AFD1F6BA07F25FB90" box="[733,781,1013,1037]" pageId="2" pageNumber="541">type</typeStatus>
locality is shown in
<figureCitation id="133552EBFF89996AFC346BA17812FB91" box="[1014,1082,1012,1037]" captionStart="FIGURE 3" captionStartId="5.[151,250,1351,1373]" captionTargetBox="[303,1284,265,1329]" captionTargetId="figure@5.[302,1284,265,1330]" captionTargetPageId="5" captionText="FIGURE 3. Map showing type locality of Tropidophorus sebi sp. nov., at Putai, Upper Sungei Baleh, Kapit District, Sarawak, Malaysia (Borneo). Inset: Map of the Indo-Pacific, showing the location of Borneo." httpUri="https://zenodo.org/record/571030/files/figure.png" pageId="2" pageNumber="541">
<collectorName id="26FB2BB8FF89996AFC346BA1780BFB90" box="[1014,1059,1012,1037]" pageId="2" pageNumber="541">Fig.</collectorName>
3
</figureCitation>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB14E6EFF89996AFF056C4278CEFBAD" blockId="2.[151,1437,903,2009]" box="[199,1254,1047,1072]" pageId="2" pageNumber="541">
<bibRef id="52AEDA98FF89996AFF056C4278CEFBAD" box="[199,1254,1047,1072]" pageId="2" pageNumber="541" type="book">
<materialsCitation id="3B664433FF89996AFF056C4278CEFBAD" ID-GBIF-Occurrence="1499986381" box="[199,1254,1047,1072]" collectingDate="2015-05-20" collectionCode="UNIMAS" collectorName="Pui Yong Min &amp; Mohamad Paisal bin Wahab &amp; Adult &amp; The &amp; Fig." country="Malaysia" county="Kapit district" elevation="117" location="Museum" municipality="upper Baleh" pageId="2" pageNumber="541" specimenCount="1" stateProvince="Sarawak" typeStatus="paratype">
<typeStatus id="54B5F0CCFF89996AFF056C427D1CFBAD" box="[199,308,1047,1072]" pageId="2" pageNumber="541" type="paratype">
<emphasis id="B97A927CFF89996AFF056C427D1CFBAD" bold="true" box="[199,308,1047,1072]" pageId="2" pageNumber="541">Paratype</emphasis>
</typeStatus>
.
<author id="53B3A68BFF89996AFE806C4D7DEFFBAD" box="[322,455,1047,1072]" fns="I" npo="LF" pageId="2" pageNumber="541">
<collectionCode id="ED1FD6ABFF89996AFE806C4D7D98FBAD" box="[322,432,1048,1072]" country="Malaysia" httpUri="http://grbio.org/cool/z1cp-qx8g" name="Universiti Malaysia Sarawak" pageId="2" pageNumber="541">UNIMAS</collectionCode>
P
</author>
1166, same locality as holotype. Collected
<collectingDate id="EFF49146FF89996AFC6A6C4D7868FBAD" box="[936,1088,1047,1072]" pageId="2" pageNumber="541" value="2015-05-20">
20 May
<year id="FFA60F3DFF89996AFBC76C427868FBAD" box="[1029,1088,1047,1072]" pageId="2" pageNumber="541">2015</year>
</collectingDate>
.
<journalOrPublisher id="51CF44A2FF89996AFB896C4278CAFBAD" box="[1099,1250,1047,1072]" jopScore="0.050000004" pageId="2" pageNumber="541">Adult female</journalOrPublisher>
.
</materialsCitation>
</bibRef>
</paragraph>
</subSubSection>
<subSubSection id="C3141DE5FF89996AFF056C6978D3FB79" pageId="2" pageNumber="541" type="diagnosis">
<paragraph id="8BB14E6EFF89996AFF056C6978D3FB79" blockId="2.[151,1437,903,2009]" pageId="2" pageNumber="541">
<emphasis id="B97A927CFF89996AFF056C697D12FBC8" bold="true" box="[199,314,1084,1109]" pageId="2" pageNumber="541">Diagnosis</emphasis>
. A large-sized (SVL
<quantity id="4CF6E38BFF89996AFDEA6C697EA4FBC9" box="[552,652,1084,1108]" metricMagnitude="-2" metricUnit="m" metricValue="8.379999999999999" pageId="2" pageNumber="541" unit="mm" value="83.8">83.8 mm</quantity>
and
<quantity id="4CF6E38BFF89996AFD036C697F0DFBC9" box="[705,805,1084,1109]" metricMagnitude="-2" metricUnit="m" metricValue="8.59" pageId="2" pageNumber="541" unit="mm" value="85.9">85.9 mm</quantity>
in the two specimens known, both adult females); upper head shields, dorsal and lateral scales smooth; parietal scales in two pairs; presuboculars three; supraciliaries eight, first largest; supraoculars four, fourth contacting orbit; supralabials seven (supralabial five contacting orbit); infralabials four, second longest; postmental undivided; longitudinal scale rows 58; ventrals 53; transverse scale rows at midbody 34; subcaudals 98; preanals enlarged, single; and subdigital lamellae on Toe
<date id="FFB068AEFF89996AFB766C9878D3FB79" box="[1204,1275,1228,1252]" pageId="2" pageNumber="541">IV 19.</date>
</paragraph>
</subSubSection>
<subSubSection id="C3141DE5FF89996AFF056CBA7D5AFAE8" pageId="2" pageNumber="541" type="etymology">
<paragraph id="8BB14E6EFF89996AFF056CBA7D5AFAE8" blockId="2.[151,1437,903,2009]" pageId="2" pageNumber="541">
<emphasis id="B97A927CFF89996AFF056CBA7D6EFA95" bold="true" box="[199,326,1263,1288]" pageId="2" pageNumber="541">Etymology</emphasis>
. The species name,
<taxonomicName id="4C0E35EDFF89996AFDFA6CBA7E4DFA95" box="[568,613,1263,1288]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="541" phylum="Chordata" rank="species" species="sebi">
<emphasis id="B97A927CFF89996AFDFA6CBA7E4DFA95" box="[568,613,1263,1288]" italics="true" pageId="2" pageNumber="541">sebi</emphasis>
</taxonomicName>
derives from the acronym SEB, for
<collectingRegion id="49CA808CFF89996AFBE86CBA78A2FA95" box="[1066,1162,1263,1288]" country="Malaysia" name="Sarawak" pageId="2" pageNumber="541">
<collectingCountry id="F3190EFEFF89996AFBE86CBA78A2FA95" box="[1066,1162,1263,1288]" name="Malaysia" pageId="2" pageNumber="541">Sarawak</collectingCountry>
</collectingRegion>
Energy Berhad, with a genitive ending. We are pleased to name the new species after this organisation in acknowledgement of support received for our long-term research of the herpetofauna of the Upper Baleh region. Suggested common name: Baleh Water Skink.
</paragraph>
</subSubSection>
<subSubSection id="C3141DE5FF899960FF056D2A7D9DFE52" lastPageId="8" lastPageNumber="547" pageId="2" pageNumber="541" type="description">
<paragraph id="8BB14E6EFF89996BFF056D2A7D43FF65" blockId="2.[151,1437,903,2009]" lastBlockId="3.[151,1436,151,536]" lastPageId="3" lastPageNumber="542" pageId="2" pageNumber="541">
<emphasis id="B97A927CFF89996AFF056D2A7DC2FA05" bold="true" box="[199,490,1407,1432]" pageId="2" pageNumber="541">
Description of
<typeStatus id="54B5F0CCFF89996AFEBE6D2A7DCDFA05" box="[380,485,1407,1432]" pageId="2" pageNumber="541" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult female. Large size, SVL
<quantity id="4CF6E38BFF89996AFC9D6DD57FE0FA0A" box="[863,968,1408,1432]" metricMagnitude="-2" metricUnit="m" metricValue="8.379999999999999" pageId="2" pageNumber="541" unit="mm" value="83.8">83.8 mm</quantity>
, TL
<quantity id="4CF6E38BFF89996AFBC06DD57852FA0A" box="[1026,1146,1408,1432]" metricMagnitude="-1" metricUnit="m" metricValue="1.214" pageId="2" pageNumber="541" unit="mm" value="121.4">121.4 mm</quantity>
; snout acute (IN:IO ratio 0.37), projecting beyond lower jaws; nostril laterally oriented; oval, situated closer to snout-tip than to orbit; eye nostril distance
<quantity id="4CF6E38BFF89996AFE856D9D7D85FA42" box="[327,429,1480,1504]" metricMagnitude="-3" metricUnit="m" metricValue="4.83" pageId="2" pageNumber="541" unit="mm" value="4.83">4.83 mm</quantity>
, eyesnout distance
<quantity id="4CF6E38BFF89996AFD506D927EDEFA42" box="[658,758,1479,1504]" metricMagnitude="-3" metricUnit="m" metricValue="6.58" pageId="2" pageNumber="541" unit="mm" value="6.58">6.58 mm</quantity>
(E-N:E-S ratio 0.73); head long, much longer than wide, HL
<quantity id="4CF6E38BFF89996AFF556DB97D24F999" box="[151,268,1516,1541]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="2" pageNumber="541" unit="mm" value="13.6">13.60 mm</quantity>
, HW
<quantity id="4CF6E38BFF89996AFE936DB97DEDF999" box="[337,453,1516,1540]" metricMagnitude="-2" metricUnit="m" metricValue="1.248" pageId="2" pageNumber="541" unit="mm" value="12.48">12.48 mm</quantity>
(HL:HW ratio 1.09); head shape slightly flattened, HD
<quantity id="4CF6E38BFF89996AFB8A6DB97886F999" box="[1096,1198,1516,1540]" metricMagnitude="-3" metricUnit="m" metricValue="8.91" pageId="2" pageNumber="541" unit="mm" value="8.91">8.91 mm</quantity>
(HL:HD ratio 1.53); upper head shields smooth; supranasals absent; frontonasal trapezoidal, as long as wide; prefrontals trapezoidal, in narrow contact medially; frontal elongated, trapezoidal, wider anteriorly; frontoparietals joined; interparietal single, with small transparent spot; parietal scales in two pairs; large posterior parietals, in contact posteriorly; small anterior parietal, in contact with frontoparietal and supraocular IV; nuchal scales in two pairs; nostrils on nasal located closer to anterior loreal; postnasal absent; supranasals absent; loreals two, anterior lower than posterior; preoculars two; presuboculars three, anterior large, second in contact with third and fourth supralabials, and third in contact with fourth and fifth supralabials; supraciliaries eight, first largest; supraoculars four, fourth contacting orbit; postsupraocular present; two postocular; pretemporal single; six postsuboculars, lower one contact with fifth and sixth supralabials; six primary temporals, lower one in contact with sixth and seventh supralabials; five secondary temporals; seven supralabials (supralabial five contacting orbit); four infralabials, second longest; mental smaller than rostral; rostral broad, projecting onto snout; posterior border of rostral straight; postmental undivided; genials in three pairs, first and second pair in broad contact, and third pair separated by three scales; auricular opening scaleless, ovoid and smaller than orbit of eye, its location indicated by a shallow depression; eyes relatively small; pupil discernable in preserved specimen; moveable eyelids; upper palpebrals 14; lower palpebrals 14; tongue short; undivided anteriorly, tip obtuse, not pointed; teeth relatively small and somewhat pointed.
</paragraph>
<paragraph id="8BB14E6EFF88996BFF0569517E9EFE4D" blockId="3.[151,1436,151,536]" pageId="3" pageNumber="542">
Body slender, BW
<quantity id="4CF6E38BFF88996BFE6069517E3DFE81" box="[418,533,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.252" pageId="3" pageNumber="542" unit="mm" value="12.52">12.52 mm</quantity>
(BW:SVL ratio 0.15); head distinct from neck and body; 58 longitudinal scale rows from parietal to above level of anterior margin of hind limb; dorsal and lateral scales smooth; ventrals 53, counted from first postgular to last scale before preanals; transverse scale rows at midbody 34; subcaudals 98; preanals enlarged, single; tail laterally compressed, relatively long, longer than snout-vent length (TL:SVL ratio 1.45); tail tip acute; tail base wider than rest of tail; tail gradually tapering to a point; median row of subcaudals enlarged. Visceral fat bodies absent in abdomen.
</paragraph>
<paragraph id="8BB14E6EFF88996BFF0569897901FD85" blockId="3.[151,1436,151,536]" pageId="3" pageNumber="542">
Limbs short and pentadactyl, digits short and clawed; lamellae smooth, enlarged; adpressed limbs touching; lamellae under Finger
<date id="FFB068AEFF88996BFE566A557D9FFD85" box="[404,439,512,536]" pageId="3" pageNumber="542">I-8</date>
;
<date id="FFB068AEFF88996BFE016A557DD6FD85" box="[451,510,512,536]" pageId="3" pageNumber="542" value="2012-02">II-12</date>
;
<date id="FFB068AEFF88996BFDC86A557E65FD85" box="[522,589,512,536]" pageId="3" pageNumber="542" value="2011-03">III-11</date>
;
<date id="FFB068AEFF88996BFD9B6A557EA6FD85" box="[601,654,512,536]" pageId="3" pageNumber="542">IV-9</date>
;
<date id="FFB068AEFF88996BFD5869AA7EEBFD85" box="[666,707,511,536]" pageId="3" pageNumber="542">V-5</date>
and lamellae under Toe
<date id="FFB068AEFF88996BFC146A557FD2FD8A" box="[982,1018,512,535]" pageId="3" pageNumber="542">I-7</date>
;
<date id="FFB068AEFF88996BFBC46A557868FD85" box="[1030,1088,512,536]" pageId="3" pageNumber="542" value="2011-02">II-11</date>
;
<date id="FFB068AEFF88996BFB8E69AA78B8FD85" box="[1100,1168,511,536]" pageId="3" pageNumber="542" value="2015-03">III-15</date>
;
<date id="FFB068AEFF88996BFB5E6A5578F7FD85" box="[1180,1247,512,536]" pageId="3" pageNumber="542" value="1919-04">IV-19</date>
;
<date id="FFB068AEFF88996BFB296A557901FD85" box="[1259,1321,512,536]" pageId="3" pageNumber="542">V-12.</date>
</paragraph>
<caption id="DF711EE6FF88996BFF556D7F78E4FADD" box="[151,1228,1322,1344]" httpUri="https://zenodo.org/record/571028/files/figure.png" pageId="3" pageNumber="542" targetBox="[151,1436,589,1301]" targetPageId="3">
<paragraph id="8BB14E6EFF88996BFF556D7F78E4FADD" blockId="3.[151,1228,1322,1344]" box="[151,1228,1322,1344]" pageId="3" pageNumber="542">
<emphasis id="B97A927CFF88996BFF556D7F7D3BFAA2" bold="true" box="[151,275,1322,1344]" pageId="3" pageNumber="542">FIGURE 1.</emphasis>
Female holotype of
<taxonomicName id="4C0E35EDFF88996BFE266D7F7E81FADD" box="[484,681,1322,1344]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="542" phylum="Chordata" rank="species" species="sebi" status="sp. nov.">
<emphasis id="B97A927CFF88996BFE266D7F7E81FADD" box="[484,681,1322,1344]" italics="true" pageId="3" pageNumber="542">Tropidophorus sebi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2492F07FF88996BFD6D6D7F7ED7FADD" box="[687,767,1322,1344]" pageId="3" pageNumber="542" rank="species">
<emphasis id="B97A927CFF88996BFD6D6D7F7ED2FAA2" bold="true" box="[687,762,1322,1343]" pageId="3" pageNumber="542">sp. nov</emphasis>
.
</taxonomicNameLabel>
(UNIMAS P1167) in life. (SVL = 83.8 mm).
</paragraph>
</caption>
<paragraph id="8BB14E6EFF88996BFF056D327FE4F891" blockId="3.[151,1436,1383,2020]" pageId="3" pageNumber="542">
<emphasis id="B97A927CFF88996BFF056D327D82FA1D" bold="true" box="[199,426,1383,1408]" pageId="3" pageNumber="542">Colouration in life.</emphasis>
Dorsum chocolate brown, with dark greyish-brown transverse bars on trunk and tail; bars on nape fused to form collar-like pattern; blackish-grey postocular band, starting from posterior corner of orbit and broadening to axillary region, and narrowing thereafter along the flanks of torsi; forehead unpatterned chocolate brown; temporal region grey with cream-coloured flecks; axilla and lower flanks with scattered yellow flecks; labials brown with greyish-black bars; dorsal surface of tail greyish-brown with darker areas forming incomplete bands; dorsal surface of fore- and hindlimbs chocolate brown with darker variegation; digits brown with darker bands; gular region pale grey, the chin with dark grey variegation, forming about 10 lines that extend to the pectoral region; upper two-thirds of pectoral region pale yellow with indistinct grey stripes; lower third orange, each scale edged with yellow; cloacal region deep orange; lowest edge of anal scale dark grey; tail base grey with large, orange-coloured scales; rest of tail dark grey; lower parts of forearm unpatterned pale grey; hindlimbs unpatterned pale orange; manus and pes dark grey; pupil rounded, black, with a narrow, orange ring; iris grey with an orange cast; oral cavity pale cream-coloured; tongue dark grey, not pale apically.
</paragraph>
<paragraph id="8BB14E6EFF88996BFF056F427F04F8E5" blockId="3.[151,1436,1383,2020]" pageId="3" pageNumber="542">
<emphasis id="B97A927CFF88996BFF056F42783EF8AD" bold="true" box="[199,1046,1815,1840]" pageId="3" pageNumber="542">
Measurements (in millimeters;
<typeStatus id="54B5F0CCFF88996BFDFF6F427E8CF8AD" box="[573,676,1815,1840]" pageId="3" pageNumber="542" type="holotype">holotype</typeStatus>
with
<typeStatus id="54B5F0CCFF88996BFD256F427F7CF8AD" box="[743,852,1815,1840]" pageId="3" pageNumber="542" type="paratype">paratype</typeStatus>
, in parentheses)
</emphasis>
. SVL 83.8 (85.9); HL 13.6 (14.7); HW 12.5 (13.6); HD 8.9 (9.8); BW 12.5 (13.5); TBL 11.3 (11.9); ED 4.1 (4.3); IN 3.1 (3.2); E
<geoCoordinate id="EE3A28A9FF88996BFB7C6F6978DDF8C9" box="[1214,1269,1851,1876]" direction="south" orientation="latitude" pageId="3" pageNumber="542" precision="5555" value="-6.9">S 6.9</geoCoordinate>
(6.9); E
<geoCoordinate id="EE3A28A9FF88996BFA9C6F6979B4F8C9" box="[1374,1436,1852,1876]" direction="north" orientation="latitude" pageId="3" pageNumber="542" precision="5555" value="4.8">N 4.8</geoCoordinate>
(5.3); N
<geoCoordinate id="EE3A28A9FF88996BFF3D6F357D1FF8E5" box="[255,311,1888,1912]" direction="south" orientation="latitude" pageId="3" pageNumber="542" precision="5555" value="-1.7">S 1.7</geoCoordinate>
(1.7); AG 36.7 (41.5) and TL 121.4 (87.2).
</paragraph>
<paragraph id="8BB14E6EFF88996BFF056FD67EDDF879" blockId="3.[151,1436,1383,2020]" pageId="3" pageNumber="542">
<emphasis id="B97A927CFF88996BFF056FD67F5AF801" bold="true" box="[199,882,1923,1948]" pageId="3" pageNumber="542">Squamation (holotype with paratype in parentheses).</emphasis>
Transverse scale rows at midbody 34 (33); longitudinal scale rows 58 (57); ventral scale rows 53 (53); supralabials 7 (7); infralabials 4 (4); subcaudals 98 (22 +; regenerated tail) and lamellae under toe
<date id="FFB068AEFF88996BFDBA6F997E91F879" box="[632,697,1996,2020]" pageId="3" pageNumber="542" value="1919-04">IV 19</date>
(19).
</paragraph>
<caption id="DF711EE6FF8F996CFF556C707834FBC7" httpUri="https://zenodo.org/record/571029/files/figure.png" pageId="4" pageNumber="543" targetBox="[344,1240,197,1032]" targetPageId="4">
<paragraph id="8BB14E6EFF8F996CFF556C707834FBC7" blockId="4.[151,1436,1061,1114]" pageId="4" pageNumber="543">
<emphasis id="B97A927CFF8F996CFF556C707D3BFBA7" bold="true" box="[151,275,1061,1083]" pageId="4" pageNumber="543">FIGURE 2.</emphasis>
Female holotype of
<taxonomicName id="4C0E35EDFF8F996CFE2B6C707E87FBA6" box="[489,687,1061,1083]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="species" species="sebi" status="sp. nov.">
<emphasis id="B97A927CFF8F996CFE2B6C707E87FBA6" box="[489,687,1061,1083]" italics="true" pageId="4" pageNumber="543">Tropidophorus sebi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2492F07FF8F996CFD746C737F20FBA6" box="[694,776,1062,1083]" pageId="4" pageNumber="543" rank="species">
<emphasis id="B97A927CFF8F996CFD746C737F2AFBA6" bold="true" box="[694,770,1062,1083]" pageId="4" pageNumber="543">sp. nov</emphasis>
.
</taxonomicNameLabel>
(UNIMAS P1167), showing head in a. lateral; b. ventral; and c. dorsal views (scale markers = 10 mm); and d. pes and e. manus. (scale markers = 1 mm).
</paragraph>
</caption>
<paragraph id="8BB14E6EFF8F996CFF056CD47EB8FB7F" blockId="4.[151,1437,1153,2006]" pageId="4" pageNumber="543">
<emphasis id="B97A927CFF8F996CFF056CD47E41FB07" bold="true" box="[199,617,1153,1178]" pageId="4" pageNumber="543">Ecological Notes and Distribution.</emphasis>
The
<typeStatus id="54B5F0CCFF8F996CFD6A6CD77EF0FB07" box="[680,728,1154,1178]" pageId="4" pageNumber="543">type</typeStatus>
series was collected from narrow crevices of rocky banks of small streams at the headwaters of Sungei Baleh. Currently, the new species is only known from its
<typeStatus id="54B5F0CCFF8F996CFAC86CF37912FB23" box="[1290,1338,1190,1214]" pageId="4" pageNumber="543">type</typeStatus>
locality, Putai, Upper Baleh, central
<collectingRegion id="49CA808CFF8F996CFE0C6C9C7E06FB7F" box="[462,558,1225,1250]" country="Malaysia" name="Sarawak" pageId="4" pageNumber="543">
<collectingCountry id="F3190EFEFF8F996CFE0C6C9C7E06FB7F" box="[462,558,1225,1250]" name="Malaysia" pageId="4" pageNumber="543">Sarawak</collectingCountry>
</collectingRegion>
(
<figureCitation id="133552EBFF8F996CFDFC6C9C7EABFB7F" box="[574,643,1225,1250]" captionStart="FIGURE 4" captionStartId="6.[151,250,1999,2021]" captionTargetBox="[204,1375,203,1959]" captionTargetId="figure@6.[204,1375,203,1959]" captionTargetPageId="6" captionText="FIGURE 4. Habitat of Tropidophorus sebi sp. nov. at Putai, Upper Sungei Baleh, Sarawak, Malaysia (Borneo)." httpUri="https://zenodo.org/record/571031/files/figure.png" pageId="4" pageNumber="543">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BB14E6EFF8F996CFF056CB87E01F99F" blockId="4.[151,1437,1153,2006]" pageId="4" pageNumber="543">
<emphasis id="B97A927CFF8F996CFF056CB87E22FA9B" bold="true" box="[199,522,1261,1286]" pageId="4" pageNumber="543">Phylogenetic relationships.</emphasis>
The ML and BI topologies were identical with respect to relationships within
<taxonomicName id="4C0E35EDFF8F996CFF556D447D14FAB7" box="[151,316,1297,1322]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="genus">
<emphasis id="B97A927CFF8F996CFF556D447D14FAB7" box="[151,316,1297,1322]" italics="true" pageId="4" pageNumber="543">Tropidophorus</emphasis>
</taxonomicName>
. Phylogenetic analysis of mitochondrial l2s and 16s provide strong support for
<taxonomicName id="4C0E35EDFF8F996CFB346D447CDBFAD3" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="species" species="beccarii">
<emphasis id="B97A927CFF8F996CFB346D447CDBFAD3" italics="true" pageId="4" pageNumber="543">Tropidophorus beccarii</emphasis>
</taxonomicName>
as the closest relative of the new species (see
<figureCitation id="133552EBFF8F996CFCCB6D607F67FAD3" box="[777,847,1333,1358]" captionStart="FIGURE 5" captionStartId="7.[151,250,1696,1718]" captionTargetBox="[151,1435,196,1666]" captionTargetId="figure@7.[151,1436,193,1675]" captionTargetPageId="7" captionText="FIGURE 5. Concatenated ML phylogram of the mitochondrial 12 s and 16 s genes for Tropidophorus and close relatives with ML bootstraps (left number) and BI posterior probabilities (right number) shown at the nodes. Nodes with ML bootstrap ± 70 and BI posterior probability ± 0.95 are denoted with a black dot. The new species is indicated in bold." httpUri="https://zenodo.org/record/571032/files/figure.png" pageId="4" pageNumber="543">Fig. 5</figureCitation>
). This sister relationship suggests a single origin of smooth dorsal scales among
<taxonomicName id="4C0E35EDFF8F996CFE266D0C7EA1FAEF" box="[484,649,1369,1394]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="genus">
<emphasis id="B97A927CFF8F996CFE266D0C7EA1FAEF" box="[484,649,1369,1394]" italics="true" pageId="4" pageNumber="543">Tropidophorus</emphasis>
</taxonomicName>
on Borneo (see Comparison), assuming an ancestral state of keeled dorsal scales for the genus. The phylogenetic results presented here are generally concordant with results of
<bibRefCitation id="EF9F339FFF8F996CFA936D287D02FA27" pageId="4" pageNumber="543" refString="Honda, M., Ota, H., Murphy, R. W. &amp; Hikida, T. (2006) Phylogeny and biogeography of water skinks of the genus Tropidophorus (Reptilia: Scincidae): a molecular approach. Zoologica Scripta, 35, 85 - 95. https: // doi. org / 10.1111 / j. 1463 - 6409.2005.00215. x" type="journal article">
Honda
<emphasis id="B97A927CFF8F996CFF556DF77CFBFA27" box="[151,211,1441,1466]" italics="true" pageId="4" pageNumber="543">et al.</emphasis>
(2006)
</bibRefCitation>
, though we recover much weaker support for the paraphyly of
<taxonomicName id="4C0E35EDFF8F996CFBD16DF47890FA27" box="[1043,1208,1441,1466]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="genus">
<emphasis id="B97A927CFF8F996CFBD16DF47890FA27" box="[1043,1208,1441,1466]" italics="true" pageId="4" pageNumber="543">Tropidophorus</emphasis>
</taxonomicName>
with respect to the
<taxonomicName id="4C0E35EDFF8F996CFF556D907D64FA43" box="[151,332,1477,1502]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="genus">
<emphasis id="B97A927CFF8F996CFF556D907D64FA43" box="[151,332,1477,1502]" italics="true" pageId="4" pageNumber="543">Sphenomorphus</emphasis>
</taxonomicName>
group (ML bootstrap 36 vs. 73). The ML and BI topologies were identical with respect to relationships within
<emphasis id="B97A927CFF8F996CFEBE6DBC7E01F99F" box="[380,553,1513,1538]" italics="true" pageId="4" pageNumber="543">
<taxonomicName id="4C0E35EDFF8F996CFEBE6DBC7E0DF99F" box="[380,549,1513,1538]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="genus">Tropidophorus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BB14E6EFF8F996CFF056E5B7F3EF928" blockId="4.[151,1437,1153,2006]" pageId="4" pageNumber="543">
Raw pairwise sequence divergence between
<taxonomicName id="4C0E35EDFF8F996CFD796E5B7F1AF9BB" box="[699,818,1549,1574]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="species" species="beccarii">
<emphasis id="B97A927CFF8F996CFD796E5B7F1AF9BB" box="[699,818,1549,1574]" italics="true" pageId="4" pageNumber="543">T. beccarii</emphasis>
</taxonomicName>
and the new species is 6.87.0% for 12s and 5.45.6% for 16s. Genetic divergence between the two
<typeStatus id="54B5F0CCFF8F996CFD506E677EEAF9D7" box="[658,706,1586,1610]" pageId="4" pageNumber="543">type</typeStatus>
specimens of the new species is low, with raw distances of 0.2% and 0.3% for 12s and 16s, respectively. However, this measure of genetic diversity within the new species may be an underestimate as the two
<typeStatus id="54B5F0CCFF8F996CFE396E2F7E03F90F" box="[507,555,1658,1682]" pageId="4" pageNumber="543">type</typeStatus>
specimens were collected from the same locality, and increased genetic divergence would likely exist across a larger spatial area.
</paragraph>
<paragraph id="8BB14E6EFF8F996DFF056E947EA2FF49" blockId="4.[151,1437,1153,2006]" lastBlockId="5.[151,1436,151,213]" lastPageId="5" lastPageNumber="544" pageId="4" pageNumber="543">
<emphasis id="B97A927CFF8F996CFF056E947D4AF947" bold="true" box="[199,354,1729,1754]" pageId="4" pageNumber="543">Comparison.</emphasis>
We initially compare the new species with Bornean congeners here.
<taxonomicName id="4C0E35EDFF8F996CFBA16E947914F947" box="[1123,1340,1729,1754]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="species" species="sebi" status="sp. nov.">
<emphasis id="B97A927CFF8F996CFBA16E947914F947" box="[1123,1340,1729,1754]" italics="true" pageId="4" pageNumber="543">Tropidophorus sebi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2492F07FF8F996CFA816E9479B4F944" box="[1347,1436,1729,1754]" pageId="4" pageNumber="543" rank="species">
<emphasis id="B97A927CFF8F996CFA816E9479BDF947" bold="true" box="[1347,1429,1729,1754]" pageId="4" pageNumber="543">sp. nov</emphasis>
.
</taxonomicNameLabel>
differs from
<taxonomicName id="4C0E35EDFF8F996CFEE46EB37DBFF963" box="[294,407,1765,1790]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="species" species="brookei">
<emphasis id="B97A927CFF8F996CFEE46EB37DBFF963" box="[294,407,1765,1790]" italics="true" pageId="4" pageNumber="543">T. brookei</emphasis>
</taxonomicName>
(distribution: widespread on Borneo),
<taxonomicName id="4C0E35EDFF8F996CFC8D6EB37F96F960" box="[847,958,1766,1789]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="species" species="iniquus">
<emphasis id="B97A927CFF8F996CFC8D6EB37F96F960" box="[847,958,1766,1789]" italics="true" pageId="4" pageNumber="543">T. iniquus</emphasis>
</taxonomicName>
(distribution: upper Sungei Kajan, central Kalimantan, Indonesia),
<taxonomicName id="4C0E35EDFF8F996CFE6C6F5F7E1AF8BC" box="[430,562,1802,1825]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="species" species="micropus">
<emphasis id="B97A927CFF8F996CFE6C6F5F7E1AF8BC" box="[430,562,1802,1825]" italics="true" pageId="4" pageNumber="543">T. micropus</emphasis>
</taxonomicName>
(distribution: Long Bloe in upper Sungei Mahakkam, Kalimantan, Indonesia and Putai, Sarawak, Malaysia) and
<taxonomicName id="4C0E35EDFF8F996CFDE66F7B7E85F8DB" box="[548,685,1837,1862]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="species" species="perplexus">
<emphasis id="B97A927CFF8F996CFDE66F7B7E85F8DB" box="[548,685,1837,1862]" italics="true" pageId="4" pageNumber="543">T. perplexus</emphasis>
</taxonomicName>
(distribution: Sungei Tinjar, Sarawak, Malaysia) in having smooth dorsal scales (versus keeled in these four species). The new species is further distinguished from another two smooth dorsal scales congeners,
<taxonomicName id="4C0E35EDFF8F996CFDD46F237EBAF813" box="[534,658,1909,1934]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="species" species="beccarii">
<emphasis id="B97A927CFF8F996CFDD46F237EBAF813" box="[534,658,1909,1934]" italics="true" pageId="4" pageNumber="543">T. beccarii</emphasis>
</taxonomicName>
(widespread on Borneo) and
<taxonomicName id="4C0E35EDFF8F996CFC366F2378BCF813" box="[1012,1172,1909,1934]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="species" species="mocquardii">
<emphasis id="B97A927CFF8F996CFC366F2378BCF813" box="[1012,1172,1909,1934]" italics="true" pageId="4" pageNumber="543">T. mocquardii</emphasis>
</taxonomicName>
(distribution: northern Sabah, Malaysia; treated as a synonym of
<taxonomicName id="4C0E35EDFF8F996CFDB46FCF7EC7F82F" box="[630,751,1945,1970]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="species" species="beccarii">
<emphasis id="B97A927CFF8F996CFDB46FCF7EC7F82F" box="[630,751,1945,1970]" italics="true" pageId="4" pageNumber="543">T. beccarii</emphasis>
</taxonomicName>
by some authors but considered valid here; see below) from Borneo, in showing a higher count of midbody scale rows (3334) versus lower in
<taxonomicName id="4C0E35EDFF8F996CFB8B6FEB78E9F84B" box="[1097,1217,1981,2006]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="543" phylum="Chordata" rank="species" species="beccarii">
<emphasis id="B97A927CFF8F996CFB8B6FEB78E9F84B" box="[1097,1217,1981,2006]" italics="true" pageId="4" pageNumber="543">T. beccarii</emphasis>
</taxonomicName>
(2830), in having four supraoculars versus five supraoculars, and in lacking a thick, dark, dorsolateral stripe along flanks, that are with light transverse bands in
<taxonomicName id="4C0E35EDFF8E996DFE2568E87EABFF48" box="[487,643,188,213]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="mocquardii">
<emphasis id="B97A927CFF8E996DFE2568E87EABFF48" box="[487,643,188,213]" italics="true" pageId="5" pageNumber="544">T. mocquardii</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF711EE6FF8E996DFF556D127FCEFAE1" httpUri="https://zenodo.org/record/571030/files/figure.png" pageId="5" pageNumber="544" targetBox="[303,1284,265,1329]" targetPageId="5">
<paragraph id="8BB14E6EFF8E996DFF556D127FCEFAE1" blockId="5.[151,1436,1351,1404]" pageId="5" pageNumber="544">
<emphasis id="B97A927CFF8E996DFF556D127D3BFAC1" bold="true" box="[151,275,1351,1373]" pageId="5" pageNumber="544">FIGURE 3.</emphasis>
Map showing type locality of
<taxonomicName id="4C0E35EDFF8E996DFD886D127F27FAC0" box="[586,783,1351,1373]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="sebi" status="sp. nov.">
<emphasis id="B97A927CFF8E996DFD886D127F27FAC0" box="[586,783,1351,1373]" italics="true" pageId="5" pageNumber="544">Tropidophorus sebi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2492F07FF8E996DFCD76D1D7F4EFAC0" box="[789,870,1352,1373]" pageId="5" pageNumber="544" rank="species">
<emphasis id="B97A927CFF8E996DFCD76D1D7F48FAC0" bold="true" box="[789,864,1352,1373]" pageId="5" pageNumber="544">sp. nov</emphasis>
.
</taxonomicNameLabel>
, at Putai, Upper Sungei Baleh, Kapit District, Sarawak, Malaysia (Borneo). Inset: Map of the Indo-Pacific, showing the location of Borneo.
</paragraph>
</caption>
<paragraph id="8BB14E6EFF8E996FFF056DF67F9FF877" blockId="5.[151,1437,1443,2008]" lastBlockId="7.[151,1437,1821,2026]" lastPageId="7" lastPageNumber="546" pageId="5" pageNumber="544">
<materialsCitation id="3B664433FF8E996DFF056DF67FE4F9B5" ID-GBIF-Occurrence="1499986384" country="Bangladesh" location="Sulawesi" pageId="5" pageNumber="544" specimenCount="1" stateProvince="Sulawesi">
The presence of smooth dorsal scales distinguishes the new species from the following extra-Bornean congeners that possess keeled dorsal scales:
<taxonomicName id="4C0E35EDFF8E996DFD0C6D9D7879FA7D" authority="Annandale, 1912" authorityName="Annandale" authorityYear="1912" box="[718,1105,1479,1504]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="assamensis">
<emphasis id="B97A927CFF8E996DFD0C6D9D7F5BFA42" box="[718,883,1480,1503]" italics="true" pageId="5" pageNumber="544">T. assamensis</emphasis>
Annandale, 1912
</taxonomicName>
(distribution: north-eastern
<collectingCountry id="F3190EFEFF8E996DFF556DBE7D32F999" box="[151,282,1515,1540]" name="Bangladesh" pageId="5" pageNumber="544">Bangladesh</collectingCountry>
and north-eastern
<collectingCountry id="F3190EFEFF8E996DFE286DBE7E0EF999" box="[490,550,1515,1540]" name="India" pageId="5" pageNumber="544">India</collectingCountry>
, including Mizoram and Assam states);
<taxonomicName id="4C0E35EDFF8E996DFC2B6DB879B4F999" authority="Hikida, Riyanto &amp; Ota, 2003" authorityName="Hikida, Riyanto &amp; Ota" authorityYear="2003" box="[1001,1436,1515,1540]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="baconi">
<emphasis id="B97A927CFF8E996DFC2B6DB87878F999" box="[1001,1104,1515,1540]" italics="true" pageId="5" pageNumber="544">T. baconi</emphasis>
Hikida, Riyanto &amp; Ota, 2003
</taxonomicName>
(distribution: Patunuang Natural Reserve, southern
<collectingRegion id="49CA808CFF8E996DFD1F6E5A7F6DF9B5" box="[733,837,1551,1576]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="544">Sulawesi</collectingRegion>
,
<collectingCountry id="F3190EFEFF8E996DFC906E5A7FECF9B5" box="[850,964,1551,1576]" name="Indonesia" pageId="5" pageNumber="544">Indonesia</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="3B664433FF8E996DFC1A6E457872F9D1" ID-GBIF-Occurrence="1499986387" country="Thailand" location="Mount Ba Vi" municipality="northern Vietnam" pageId="5" pageNumber="544" specimenCount="1" stateProvince="Ha Tay Province">
<taxonomicName id="4C0E35EDFF8E996DFC1A6E45792AF9B5" authority="Bourret, 1939" authorityName="Bourret" authorityYear="1939" box="[984,1282,1551,1576]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="baviensis">
<emphasis id="B97A927CFF8E996DFC1A6E457874F9B5" box="[984,1116,1551,1576]" italics="true" pageId="5" pageNumber="544">T. baviensis</emphasis>
Bourret, 1939
</taxonomicName>
(distribution:
<location id="8ED118B5FF8E996DFF556E617D19F9D1" LSID="urn:lsid:plazi:treatment:03A7FF78FF899960FF556B487D9DFE52:8ED118B5FF8E996DFF556E617D19F9D1" box="[151,305,1587,1612]" country="Thailand" municipality="northern Vietnam" name="Mount Ba Vi" pageId="5" pageNumber="544" stateProvince="Ha Tay Province">Mount Ba Vi</location>
,
<collectingRegion id="49CA808CFF8E996DFEFD6E617E2DF9D1" box="[319,517,1587,1612]" inferredBySuffix="true" pageId="5" pageNumber="544">Ha Tay Province</collectingRegion>
,
<collectingMunicipality id="6BD5D414FF8E996DFDD06E667EF2F9D1" box="[530,730,1587,1612]" pageId="5" pageNumber="544">northern Vietnam</collectingMunicipality>
and possibly, adjacent
<collectingCountry id="F3190EFEFF8E996DFC2B6E66787AF9D1" box="[1001,1106,1587,1612]" name="Thailand" pageId="5" pageNumber="544">Thailand</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="3B664433FF8E996DFBA56E607FDDF9ED" ID-GBIF-Occurrence="1499986390" country="China, Myanmar, Thailand" location="Yunnan" pageId="5" pageNumber="544" specimenCount="1" stateProvince="Yunnan">
<taxonomicName id="4C0E35EDFF8E996DFBA56E6079B4F9D1" authority="Blyth, 1853" authorityName="Blyth" authorityYear="1853" box="[1127,1436,1587,1612]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="B97A927CFF8E996DFBA56E6078D1F9D1" box="[1127,1273,1587,1612]" italics="true" pageId="5" pageNumber="544">T. berdmorei</emphasis>
(Blyth, 1853)
</taxonomicName>
(distribution:
<collectingRegion id="49CA808CFF8E996DFEF16E0D7DA0F9F2" box="[307,392,1624,1647]" country="China" name="Yunnan" pageId="5" pageNumber="544">Yunnan</collectingRegion>
in southern
<collectingCountry id="F3190EFEFF8E996DFDD96E027F66F9ED" box="[539,846,1623,1648]" isEnumeration="true" pageId="5" pageNumber="544">China, Myanmar, Thailand</collectingCountry>
and
<collectingCountry id="F3190EFEFF8E996DFC4B6E027FC5F9ED" box="[905,1005,1623,1648]" name="Vietnam" pageId="5" pageNumber="544">Vietnam</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="3B664433FF8E996DFBC06E0D7F46F909" ID-GBIF-Occurrence="1499986392" country="Thailand" location="southern Vietnam" pageId="5" pageNumber="544" specimenCount="1">
<taxonomicName id="4C0E35EDFF8E996DFBC06E0D7CE7F909" authority="Dumeril &amp; Bibron, 1839" authorityName="Dumeril &amp; Bibron" authorityYear="1839" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="cocincinensis">
<emphasis id="B97A927CFF8E996DFBC06E0D789FF9F2" box="[1026,1207,1624,1647]" italics="true" pageId="5" pageNumber="544">T. cocincinensis</emphasis>
<bibRefCitation id="EF9F339FFF8E996DFB036E027CE7F909" author="Dumeril" pageId="5" pageNumber="544" refString="Dumeril, A. - M. - C. &amp; Bibron, G. (1839) Erpetologie Generale ou Histoire Naturelle complete des Reptiles. Fol. 5, contenant l'histoire de quatre-vingt-trois genres et de deux cent sept especes des trois dernieres familles de l'ordre des Sauriens, savoir: les Lacertiliens, les Chalcidiens et les Scincoidiens. Ouvrage illustre de planches. Librarie Encyclopedique de Roret, Paris, viii + 854 pp., errata, 4 folding tables. [pls. 37, 39, 39 bis, 41 bis, 49, 51 - 54 and 56 - 58]" type="book chapter" year="1839">Duméril &amp; Bibron, 1839</bibRefCitation>
</taxonomicName>
(distribution:
<collectingCountry id="F3190EFEFF8E996DFEB66E2E7DF2F909" box="[372,474,1659,1684]" name="Thailand" pageId="5" pageNumber="544">Thailand</collectingCountry>
,
<location id="8ED118B5FF8E996DFE2B6E2E7E9AF909" LSID="urn:lsid:plazi:treatment:03A7FF78FF899960FF556B487D9DFE52:8ED118B5FF8E996DFE2B6E2E7E9AF909" box="[489,690,1659,1684]" country="Thailand" name="southern Vietnam" pageId="5" pageNumber="544">southern Vietnam</location>
and
<collectingCountry id="F3190EFEFF8E996DFD2C6E2E7F4EF909" box="[750,870,1659,1684]" name="Cambodia" pageId="5" pageNumber="544">Cambodia</collectingCountry>
)
</materialsCitation>
;
<taxonomicName id="4C0E35EDFF8E996DFCB96E287898F909" authority="Bacon, 1980" authorityName="Bacon" authorityYear="1980" box="[891,1200,1659,1684]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="davaoensis">
<emphasis id="B97A927CFF8E996DFCB96E28783EF909" box="[891,1046,1659,1684]" italics="true" pageId="5" pageNumber="544">T. davaoensis</emphasis>
Bacon, 1980
</taxonomicName>
(distribution: southcentral Mindanao, Philippines);
<materialsCitation id="3B664433FF8E996DFDD46EF57DF5F941" ID-GBIF-Occurrence="1499986394" country="Philippines" location="Leyte" pageId="5" pageNumber="544" specimenCount="1" stateProvince="Leyte">
<taxonomicName id="4C0E35EDFF8E996DFDD46EF57F02F925" authority="Gunther, 1861" authorityName="Gunther" authorityYear="1861" box="[534,810,1695,1720]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="grayi">
<emphasis id="B97A927CFF8E996DFDD46EF57E5AF92A" box="[534,626,1696,1719]" italics="true" pageId="5" pageNumber="544">T. grayi</emphasis>
Günther, 1861
</taxonomicName>
(distribution: Panay, Luzon, Polillo,
<collectingRegion id="49CA808CFF8E996DFB2C6EF57919F925" box="[1262,1329,1696,1720]" country="Philippines" name="Southern Leyte" pageId="5" pageNumber="544">Leyte</collectingRegion>
, Negros, Mastabe,
<collectingRegion id="49CA808CFF8E996DFEC46E967D6CF941" box="[262,324,1731,1756]" country="Philippines" name="Cebu" pageId="5" pageNumber="544">Cebu</collectingRegion>
,
<collectingCountry id="F3190EFEFF8E996DFE906E967DFDF941" box="[338,469,1731,1756]" name="Philippines" pageId="5" pageNumber="544">Philippines</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="3B664433FF8E996DFE286E907922F941" ID-GBIF-Occurrence="1499986388" box="[490,1290,1731,1756]" country="China" location="Guangxi" pageId="5" pageNumber="544" specimenCount="1" stateProvince="Guangxi">
<taxonomicName id="4C0E35EDFF8E996DFE286E907F36F941" authority="Wen, 1992" authorityName="Wen" authorityYear="1992" box="[490,798,1732,1756]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="guangxiensis">
<emphasis id="B97A927CFF8E996DFE286E907EB2F941" box="[490,666,1733,1756]" italics="true" pageId="5" pageNumber="544">T. guangxiensis</emphasis>
<bibRefCitation id="EF9F339FFF8E996DFD666E917F36F941" author="Wen" box="[676,798,1732,1756]" pageId="5" pageNumber="544" refString="Wen, Y. (1992) A new species of the genus Tropidophorus (Reptilia: Lacertilia) from Guangxi Zhuang Autonomous Region, China. Asiatic Herpetological Research, 4, 18 - 22." type="journal article" year="1992">Wen, 1992</bibRefCitation>
</taxonomicName>
(distribution:
<collectingRegion id="49CA808CFF8E996DFC066E96780CF941" box="[964,1060,1731,1756]" country="China" name="Guangxi" pageId="5" pageNumber="544">Guangxi</collectingRegion>
and
<collectingRegion id="49CA808CFF8E996DFB9D6E917884F946" box="[1119,1196,1732,1755]" country="China" name="Hunan" pageId="5" pageNumber="544">Hunan</collectingRegion>
,
<collectingCountry id="F3190EFEFF8E996DFB786E96792AF941" box="[1210,1282,1731,1756]" name="China" pageId="5" pageNumber="544">China</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="3B664433FF8E996DFAD56E907DDBF8D5" ID-GBIF-Occurrence="1499986385" country="China" location="Bac Kan" pageId="5" pageNumber="544" specimenCount="1" stateProvince="Bac Kan">
<taxonomicName id="4C0E35EDFF8E996DFAD56E907D37F89D" authority="Smith, 1923" authorityName="Smith" authorityYear="1923" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="hainanus">
<emphasis id="B97A927CFF8E996DFAD56E9079B4F941" box="[1303,1436,1731,1756]" italics="true" pageId="5" pageNumber="544">T. hainanus</emphasis>
<bibRefCitation id="EF9F339FFF8E996DFF556EB27D37F89D" author="Smith" box="[151,287,1767,1792]" pageId="5" pageNumber="544" refString="Smith, M. A. (1923) A review of the lizards of the genus Tropidophorus on the Asiatic mainland. Proceedings of the Zoological Society of London, 1923, 775 - 781." type="proceedings paper" year="1923">Smith, 1923</bibRefCitation>
</taxonomicName>
(distribution: Hainan, Guangxi, Jiangxi, Hunan, Guangdong and Guizhou in
<collectingCountry id="F3190EFEFF8E996DFB426EB278EEF89D" box="[1152,1222,1767,1792]" name="China" pageId="5" pageNumber="544">China</collectingCountry>
, and
<collectingRegion id="49CA808CFF8E996DFAC26EBD794CF962" box="[1280,1380,1768,1791]" country="Vietnam" name="Bac Kan" pageId="5" pageNumber="544">Bac Kan</collectingRegion>
,
<collectingRegion id="49CA808CFF8E996DFAAD6EBD7CFDF8B9" country="Vietnam" name="Cao Bang" pageId="5" pageNumber="544">Cao Bang</collectingRegion>
,
<collectingRegion id="49CA808CFF8E996DFF236F5E7D6DF8B9" box="[225,325,1803,1828]" country="Vietnam" name="Dak Lak" pageId="5" pageNumber="544">Dak Lak</collectingRegion>
, Ha tay,
<collectingRegion id="49CA808CFF8E996DFE686F5E7E01F8B9" box="[426,553,1803,1828]" country="Vietnam" name="Hai Duong" pageId="5" pageNumber="544">Hai Duong</collectingRegion>
,
<collectingRegion id="49CA808CFF8E996DFDF76F597E8CF8B9" box="[565,676,1803,1828]" country="Vietnam" name="Hoa Binh" pageId="5" pageNumber="544">Hoa Binh</collectingRegion>
,
<collectingRegion id="49CA808CFF8E996DFD736F597F08F8BE" box="[689,800,1804,1827]" country="Vietnam" name="Kon Tum" pageId="5" pageNumber="544">Kon Tum</collectingRegion>
,
<collectingRegion id="49CA808CFF8E996DFCEE6F5E7FB9F8B9" box="[812,913,1803,1828]" country="Vietnam" name="Lai Chau" pageId="5" pageNumber="544">Lai chau</collectingRegion>
,
<collectingRegion id="49CA808CFF8E996DFC5F6F5E7839F8B9" box="[925,1041,1803,1828]" country="Vietnam" name="Ninh Binh" pageId="5" pageNumber="544">Ninh binh</collectingRegion>
,
<collectingRegion id="49CA808CFF8E996DFBDF6F5E78A9F8B9" box="[1053,1153,1803,1828]" country="Vietnam" name="Phu Tho" pageId="5" pageNumber="544">Phu Tho</collectingRegion>
,
<collectingRegion id="49CA808CFF8E996DFB4F6F597932F8B9" box="[1165,1306,1803,1828]" country="Vietnam" name="Quang Ninh" pageId="5" pageNumber="544">Quang Ninh</collectingRegion>
,
<collectingRegion id="49CA808CFF8E996DFAE46F5E79B4F8B9" box="[1318,1436,1803,1828]" country="Vietnam" name="Vinh Phuc" pageId="5" pageNumber="544">Vinh Phuc</collectingRegion>
and
<collectingRegion id="49CA808CFF8E996DFF126F657D67F8D5" box="[208,335,1839,1864]" country="Vietnam" name="Nam Dinh" pageId="5" pageNumber="544">Nam Dinh</collectingRegion>
in
<collectingCountry id="F3190EFEFF8E996DFE446F7A7DC2F8D5" box="[390,490,1839,1864]" name="Vietnam" pageId="5" pageNumber="544">Vietnam</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="3B664433FF8E996DFDC56F65785EF80D" ID-GBIF-Occurrence="1499986383" country="Thailand" location="Nong Kai Province" municipality="eastern Thailand" pageId="5" pageNumber="544" specimenCount="1" stateProvince="Chaiyaphum Province">
<taxonomicName id="4C0E35EDFF8E996DFDC56F6579B4F8D5" authority="Chuaynkern, Nabhitabhata, Inthara, Kamsook &amp; Somsri, 2005" authorityName="Chuaynkern, Nabhitabhata, Inthara, Kamsook &amp; Somsri" authorityYear="2005" box="[519,1436,1839,1864]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="hangnam">
<emphasis id="B97A927CFF8E996DFDC56F657EBCF8D5" box="[519,660,1839,1864]" italics="true" pageId="5" pageNumber="544">T. hangnam</emphasis>
<bibRefCitation id="EF9F339FFF8E996DFD676F7A79B4F8D5" author="Chuaynkern" box="[677,1436,1839,1864]" pageId="5" pageNumber="544" refString="Chuaynkern, Y., Nabhitabhata, J., Inthara, C., Kamsook, M. &amp; Somsri, K. (2005) A new species of the water skink Tropidophorus (Reptilia: Squamata: Scincidae) from northeastern Thailand. Thailand Natural History Museum Journal, 1, 165 - 176." type="journal article" year="2005">Chuaynkern, Nabhitabhata, Inthara, Kamsook &amp; Somsri, 2005</bibRefCitation>
</taxonomicName>
(distribution:
<collectingRegion id="49CA808CFF8E996DFEF16F067E1CF8F1" box="[307,564,1875,1900]" country="Thailand" name="Chaiyaphum" pageId="5" pageNumber="544">Chaiyaphum Province</collectingRegion>
, north-easten
<collectingCountry id="F3190EFEFF8E996DFD146F067F13F8F1" box="[726,827,1875,1900]" name="Thailand" pageId="5" pageNumber="544">Thailand</collectingCountry>
),
<taxonomicName id="4C0E35EDFF8E996DFC966F007CE7F80D" authority="Hikida, Orlov, Nabhitabhata &amp; Ota, 2002" authorityName="Hikida, Orlov, Nabhitabhata &amp; Ota" authorityYear="2002" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="latiscutatus">
<emphasis id="B97A927CFF8E996DFC966F007FDCF8F1" box="[852,1012,1875,1900]" italics="true" pageId="5" pageNumber="544">T. latiscutatus</emphasis>
<bibRefCitation id="EF9F339FFF8E996DFC3C6F067CE7F80D" author="Hikida" pageId="5" pageNumber="544" refString="Hikida, T., Orlov, N. L., Nabhitabhata, J. &amp; Ota, H. (2002) Three new depressed-bodied water skinks of the genus Tropidophorus (Lacertilia: Scincidae) from Thailand and Vietnam. Current Herpetology, 21, 9 - 23. https: // doi. org / 10.5358 / hsj. 21.9" type="journal article" year="2002">Hikida, Orlov, Nabhitabhata &amp; Ota, 2002</bibRefCitation>
</taxonomicName>
(distribution: Phu Wua Wildlife Sanctuary,
<location id="8ED118B5FF8E996DFD066F2D7F8AF80D" LSID="urn:lsid:plazi:treatment:03A7FF78FF899960FF556B487D9DFE52:8ED118B5FF8E996DFD066F2D7F8AF80D" box="[708,930,1911,1936]" country="Thailand" municipality="eastern Thailand" name="Nong Kai Province" pageId="5" pageNumber="544" stateProvince="Chaiyaphum Province">
Nong
<collectingRegion id="49CA808CFF8E996DFCC86F227F8AF80D" box="[778,930,1911,1936]" inferredBySuffix="true" pageId="5" pageNumber="544">Kai Province</collectingRegion>
</location>
,
<collectingMunicipality id="6BD5D414FF8E996DFC6C6F2D7847F80D" box="[942,1135,1911,1936]" pageId="5" pageNumber="544">eastern Thailand</collectingMunicipality>
)
</materialsCitation>
;
<materialsCitation id="3B664433FF8E996DFB406F2D78B1F829" ID-GBIF-Occurrence="1499986386" country="Thailand" location="Roi Et Province" pageId="5" pageNumber="544" specimenCount="1" stateProvince="Roi Et Province">
<taxonomicName id="4C0E35EDFF8E996DFB406F2D7D95F829" authority="Hikida, Orlov, Nabhitabhata &amp; Ota, 2002" authorityName="Hikida, Orlov, Nabhitabhata &amp; Ota" authorityYear="2002" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="matsuii">
<emphasis id="B97A927CFF8E996DFB406F2D78DAF812" box="[1154,1266,1912,1935]" italics="true" pageId="5" pageNumber="544">T. matsuii</emphasis>
<bibRefCitation id="EF9F339FFF8E996DFB386F227D95F829" author="Hikida" pageId="5" pageNumber="544" refString="Hikida, T., Orlov, N. L., Nabhitabhata, J. &amp; Ota, H. (2002) Three new depressed-bodied water skinks of the genus Tropidophorus (Lacertilia: Scincidae) from Thailand and Vietnam. Current Herpetology, 21, 9 - 23. https: // doi. org / 10.5358 / hsj. 21.9" type="journal article" year="2002">Hikida, Orlov, Nabhitabhata &amp; Ota, 2002</bibRefCitation>
</taxonomicName>
(distribution: Phu Pa Namtip,
<collectingRegion id="49CA808CFF8E996DFCD16FCE7FE0F829" box="[787,968,1947,1972]" country="Thailand" name="Roi Et" pageId="5" pageNumber="544">
Roi
<collectingRegion id="49CA808CFF8E996DFC806FC97FE0F829" box="[834,968,1947,1972]" inferredBySuffix="true" pageId="5" pageNumber="544">Et Province</collectingRegion>
</collectingRegion>
, eastern
<collectingCountry id="F3190EFEFF8E996DFBEA6FCE78B9F829" box="[1064,1169,1947,1972]" name="Thailand" pageId="5" pageNumber="544">Thailand</collectingCountry>
)
</materialsCitation>
;
<taxonomicName id="4C0E35EDFF8E996DFB666FC87CE7F845" authority="Gunther, 1861" authorityName="Gunther" authorityYear="1861" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B97A927CFF8E996DFB666FC8791BF829" box="[1188,1331,1947,1972]" italics="true" pageId="5" pageNumber="544">T. microlepis</emphasis>
Günther, 1861
</taxonomicName>
(distribution: southern Thailand, Vietnam, Laos and Cambodia);
<materialsCitation id="3B664433FF8E996FFC6F6F957E81F8AB" ID-GBIF-Occurrence="1499986391" country="Philippines" lastPageId="7" lastPageNumber="546" location="Basilan" pageId="5" pageNumber="544" specimenCount="1" stateProvince="Basilan">
<taxonomicName id="4C0E35EDFF8E996DFC6F6F9578D5F845" authority="Stejneger, 1908" authorityName="Stejneger" authorityYear="1908" box="[941,1277,1983,2008]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="544" phylum="Chordata" rank="species" species="misaminius">
<emphasis id="B97A927CFF8E996DFC6F6F95786FF84A" box="[941,1095,1984,2007]" italics="true" pageId="5" pageNumber="544">T. misaminius</emphasis>
Stejneger, 1908
</taxonomicName>
(distribution:
<collectingRegion id="49CA808CFF8C996FFF556F487CC6F8AB" box="[151,238,1821,1846]" country="Philippines" name="Basilan" pageId="7" pageNumber="546">Basilan</collectingRegion>
,
<collectingRegion id="49CA808CFF8C996FFF3B6F487D4FF8AB" box="[249,359,1821,1846]" country="Philippines" name="Camiguin" pageId="7" pageNumber="546">Camiguin</collectingRegion>
and Mindanao,
<collectingCountry id="F3190EFEFF8C996FFDDC6F487E89F8AB" box="[542,673,1821,1846]" name="Philippines" pageId="7" pageNumber="546">Philippines</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="3B664433FF8C996FFD766F4B7E72F8C7" ID-GBIF-Occurrence="1499986389" country="Vietnam" location="Cao Bang Province" pageId="7" pageNumber="546" specimenCount="1" stateProvince="Cao Bang Province">
<taxonomicName id="4C0E35EDFF8C996FFD766F4B792BF8AB" authority="Hikida, Orlov, Nabhitabhata &amp; Ota, 2002" authorityName="Hikida, Orlov, Nabhitabhata &amp; Ota" authorityYear="2002" box="[692,1283,1821,1846]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="546" phylum="Chordata" rank="species" species="murphyi">
<emphasis id="B97A927CFF8C996FFD766F4B7F04F8AB" box="[692,812,1821,1846]" italics="true" pageId="7" pageNumber="546">T. murphyi</emphasis>
<bibRefCitation id="EF9F339FFF8C996FFCF16F48792BF8AB" author="Hikida" box="[819,1283,1821,1846]" pageId="7" pageNumber="546" refString="Hikida, T., Orlov, N. L., Nabhitabhata, J. &amp; Ota, H. (2002) Three new depressed-bodied water skinks of the genus Tropidophorus (Lacertilia: Scincidae) from Thailand and Vietnam. Current Herpetology, 21, 9 - 23. https: // doi. org / 10.5358 / hsj. 21.9" type="journal article" year="2002">Hikida, Orlov, Nabhitabhata &amp; Ota, 2002</bibRefCitation>
</taxonomicName>
(distribution:
<collectingRegion id="49CA808CFF8C996FFF556F177D51F8C7" box="[151,377,1857,1882]" country="Vietnam" name="Cao Bang" pageId="7" pageNumber="546">
<collectingRegion id="49CA808CFF8C996FFF556F177D20F8C7" box="[151,264,1858,1882]" inferredBySuffix="true" pageId="7" pageNumber="546">Cao Bang</collectingRegion>
Province
</collectingRegion>
, northern
<collectingCountry id="F3190EFEFF8C996FFE2C6F147E7AF8C7" box="[494,594,1857,1882]" name="Vietnam" pageId="7" pageNumber="546">Vietnam</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="3B664433FF8C996FFDA56F177F29F8E3" ID-GBIF-Occurrence="1499986393" country="Vietnam" location="Quang Binh Province" pageId="7" pageNumber="546" specimenCount="1" stateProvince="Quang Binh Province">
<taxonomicName id="4C0E35EDFF8C996FFDA56F177813F8C7" authority="Ziegler, Thanh &amp; Thanh, 2005" authorityName="Ziegler, Thanh &amp; Thanh" authorityYear="2005" box="[615,1083,1857,1882]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="546" phylum="Chordata" rank="species" species="noggei">
<emphasis id="B97A927CFF8C996FFDA56F177EFAF8C4" box="[615,722,1858,1881]" italics="true" pageId="7" pageNumber="546">T. noggei</emphasis>
<bibRefCitation id="EF9F339FFF8C996FFD196F147813F8C7" author="Ziegler" box="[731,1083,1857,1882]" pageId="7" pageNumber="546" refString="Ziegler, T., Thanh, V. N. &amp; Thanh, B. N. (2005) A new water skink of the genus Tropidophorus from the Phong Nha - Ke Bang National Park, central Vietnam (Squamata: Sauria: Scincidae). Salamandra, 41, 137 - 146." type="journal article" year="2005">Ziegler, Thanh &amp; Thanh, 2005</bibRefCitation>
</taxonomicName>
(distribution: Phong Nha - Ke Bang National Park,
<collectingRegion id="49CA808CFF8C996FFE496F337EAEF8E3" box="[395,646,1893,1918]" country="Vietnam" name="Quang Binh" pageId="7" pageNumber="546">
<collectingRegion id="49CA808CFF8C996FFE496F337E3DF8E3" box="[395,533,1893,1918]" inferredBySuffix="true" pageId="7" pageNumber="546">Quang Binh</collectingRegion>
Province
</collectingRegion>
,
<collectingCountry id="F3190EFEFF8C996FFD576F307ED1F8E3" box="[661,761,1893,1918]" name="Vietnam" pageId="7" pageNumber="546">Vietnam</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="3B664433FF8C996FFCCC6F337EECF83F" ID-GBIF-Occurrence="1499986395" country="Philippines" location="Mataling River Falls" pageId="7" pageNumber="546" specimenCount="1" stateProvince="Cotabato Province">
<taxonomicName id="4C0E35EDFF8C996FFCCC6F337860F8E3" authority="Stejneger, 1910" authorityName="Stejneger" authorityYear="1910" box="[782,1096,1893,1918]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="546" phylum="Chordata" rank="species" species="partelloi">
<emphasis id="B97A927CFF8C996FFCCC6F337FA4F8E3" box="[782,908,1893,1918]" italics="true" pageId="7" pageNumber="546">T. partelloi</emphasis>
Stejneger, 1910
</taxonomicName>
(distribution:
<location id="8ED118B5FF8C996FFB2C6F307CF8F83F" LSID="urn:lsid:plazi:treatment:03A7FF78FF899960FF556B487D9DFE52:8ED118B5FF8C996FFB2C6F307CF8F83F" country="Philippines" name="Mataling River Falls" pageId="7" pageNumber="546" stateProvince="Cotabato Province">Mataling River Falls</location>
,
<collectingRegion id="49CA808CFF8C996FFF196FDC7D86F83F" box="[219,430,1929,1954]" country="Philippines" name="South Cotabato" pageId="7" pageNumber="546">Cotabato Province</collectingRegion>
, Mindanao,
<collectingCountry id="F3190EFEFF8C996FFDFA6FDC7E93F83F" box="[568,699,1929,1954]" name="Philippines" pageId="7" pageNumber="546">Philippines</collectingCountry>
)
</materialsCitation>
;
<taxonomicName id="4C0E35EDFF8C996FFD0C6FDF7FCCF83F" authority="Smith, 1919" authorityName="Smith" authorityYear="1919" box="[718,996,1929,1954]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="546" phylum="Chordata" rank="species" species="robinsoni">
<emphasis id="B97A927CFF8C996FFD0C6FDF7F7CF83F" box="[718,852,1929,1954]" italics="true" pageId="7" pageNumber="546">T. robinsoni</emphasis>
<bibRefCitation id="EF9F339FFF8C996FFC996FDC7FCCF83F" author="Smith" box="[859,996,1929,1954]" pageId="7" pageNumber="546" refString="Smith, M. A. (1919) The lizards of the genus Tropidophorus in Siam, with descriptions of two new species. Journal of the Natural History Society of Siam, 3, 223 - 228." type="journal article" year="1919">Smith, 1919</bibRefCitation>
</taxonomicName>
(distribution: Myanmar and Thailand);
<materialsCitation id="3B664433FF8C996FFF556FFB7F9BF877" ID-GBIF-Occurrence="1499986380" country="Hong Kong" location="Guangxi, Guangdong" pageId="7" pageNumber="546" specimenCount="1" stateProvince="Guangxi, Guangdong">
<taxonomicName id="4C0E35EDFF8C996FFF556FFB7D83F85B" authority="Boettger, 1886" authorityName="Boettger" authorityYear="1886" box="[151,427,1965,1990]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="546" phylum="Chordata" rank="species" species="sinicus">
<emphasis id="B97A927CFF8C996FFF556FFB7D28F858" box="[151,256,1966,1989]" italics="true" pageId="7" pageNumber="546">T. sinicus</emphasis>
Boettger, 1886
</taxonomicName>
(distribution:
<collectingRegion id="49CA808CFF8C996FFD896FF87F16F85B" box="[587,830,1965,1990]" isEnumeration="true" pageId="7" pageNumber="546">Guangxi, Guangdong</collectingRegion>
,
<collectingCountry id="F3190EFEFF8C996FFC8B6FFB7FE4F85B" box="[841,972,1966,1990]" name="Hong Kong" pageId="7" pageNumber="546">Hong Kong</collectingCountry>
in
<collectingCountry id="F3190EFEFF8C996FFC2D6FF8781AF85B" box="[1007,1074,1965,1990]" name="China" pageId="7" pageNumber="546">China</collectingCountry>
and
<collectingCountry id="F3190EFEFF8C996FFBA56FF878E0F85B" box="[1127,1224,1965,1990]" name="Vietnam" pageId="7" pageNumber="546">Vietnam</collectingCountry>
) and
<taxonomicName id="4C0E35EDFF8C996FFAC76FFB7CE7F877" authority="Smith, 1919" authorityName="Smith" authorityYear="1919" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="546" phylum="Chordata" rank="species" species="thai">
<emphasis id="B97A927CFF8C996FFAC76FFB7962F85B" box="[1285,1354,1965,1990]" italics="true" pageId="7" pageNumber="546">T. thai</emphasis>
<bibRefCitation id="EF9F339FFF8C996FFA906FF87CE7F877" author="Smith" pageId="7" pageNumber="546" refString="Smith, M. A. (1919) The lizards of the genus Tropidophorus in Siam, with descriptions of two new species. Journal of the Natural History Society of Siam, 3, 223 - 228." type="journal article" year="1919">Smith, 1919</bibRefCitation>
</taxonomicName>
(distribution:
<collectingCountry id="F3190EFEFF8C996FFEAD6F877DF4F877" box="[367,476,2002,2026]" name="Myanmar" pageId="7" pageNumber="546">Myanmar</collectingCountry>
and northern and north-eastern
<collectingCountry id="F3190EFEFF8C996FFC806F847F83F877" box="[834,939,2001,2026]" name="Thailand" pageId="7" pageNumber="546">Thailand</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<caption id="DF711EE6FF8D996EFF556F9A792BF878" box="[151,1283,1999,2021]" httpUri="https://zenodo.org/record/571031/files/figure.png" pageId="6" pageNumber="545" targetBox="[204,1375,203,1959]" targetPageId="6">
<paragraph id="8BB14E6EFF8D996EFF556F9A792BF878" blockId="6.[151,1283,1999,2021]" box="[151,1283,1999,2021]" pageId="6" pageNumber="545">
<emphasis id="B97A927CFF8D996EFF556F9A7D3AF879" bold="true" box="[151,274,1999,2021]" pageId="6" pageNumber="545">FIGURE 4.</emphasis>
Habitat of
<taxonomicName id="4C0E35EDFF8D996EFE476F9A7E62F878" box="[389,586,1999,2021]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="545" phylum="Chordata" rank="species" species="sebi" status="sp. nov.">
<emphasis id="B97A927CFF8D996EFE476F9A7E62F878" box="[389,586,1999,2021]" italics="true" pageId="6" pageNumber="545">Tropidophorus sebi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2492F07FF8D996EFD926F9A7E88F878" box="[592,672,1999,2021]" pageId="6" pageNumber="545" rank="species">
<emphasis id="B97A927CFF8D996EFD926F9A7EB2F879" bold="true" box="[592,666,1999,2020]" pageId="6" pageNumber="545">sp. nov</emphasis>
.
</taxonomicNameLabel>
at Putai, Upper Sungei Baleh, Sarawak, Malaysia (Borneo).
</paragraph>
</caption>
<caption id="DF711EE6FF8C996FFF556EF578B7F96E" httpUri="https://zenodo.org/record/571032/files/figure.png" pageId="7" pageNumber="546" targetBox="[151,1435,196,1666]" targetPageId="7">
<paragraph id="8BB14E6EFF8C996FFF556EF578B7F96E" blockId="7.[151,1435,1696,1779]" pageId="7" pageNumber="546">
<emphasis id="B97A927CFF8C996FFF556EF57D3BF928" bold="true" box="[151,275,1696,1718]" pageId="7" pageNumber="546">FIGURE 5.</emphasis>
Concatenated ML phylogram of the mitochondrial 12s and 16s genes for
<taxonomicName id="4C0E35EDFF8C996FFBCF6EF5788BF92B" box="[1037,1187,1696,1718]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="546" phylum="Chordata" rank="genus">
<emphasis id="B97A927CFF8C996FFBCF6EF5788BF92B" box="[1037,1187,1696,1718]" italics="true" pageId="7" pageNumber="546">Tropidophorus</emphasis>
</taxonomicName>
and close relatives with ML bootstraps (left number) and BI posterior probabilities (right number) shown at the nodes. Nodes with ML bootstrap ± 70 and BI posterior probability ± 0.95 are denoted with a black dot. The new species is indicated in bold.
</paragraph>
</caption>
<paragraph id="8BB14E6EFF839960FF0568CD7849FE15" blockId="8.[151,1436,151,464]" pageId="8" pageNumber="547">
Two extra-Bornean congeners possess smooth scales:
<taxonomicName id="4C0E35EDFF839960FCE468CC7CE7FF49" authority="Nguyen, Nguyen, Schmitz, Orlov &amp; Ziegler, 2010" authorityName="Nguyen, Nguyen, Schmitz, Orlov &amp; Ziegler" authorityYear="2010" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="547" phylum="Chordata" rank="species" species="boehmei">
<emphasis id="B97A927CFF839960FCE468CC7F89FF2D" box="[806,929,151,176]" italics="true" pageId="8" pageNumber="547">T. boehmei</emphasis>
<bibRefCitation id="EF9F339FFF839960FC6A68CD7CE7FF49" author="Nguyen" pageId="8" pageNumber="547" refString="Nguyen, T. Q., Nguyen, T. T., Schmitz, A., Orlov, N. L. &amp; Ziegler, T. (2010 b) A new species of the genus Tropidophorus Dumeril &amp; Bibron, 1839 (Squamata: Sauria: Scincidae) from Vietnam. Zootaxa, 2439, 53 - 68." type="journal article" year="2010">Nguyen, Nguyen, Schmitz, Orlov &amp; Ziegler, 2010</bibRefCitation>
</taxonomicName>
(distribution: Hoang Lien Mountain,
<collectingRegion id="49CA808CFF839960FD5968E87F59FF48" box="[667,881,188,213]" country="Vietnam" name="Lao Cai" pageId="8" pageNumber="547">Lao Cai Province</collectingRegion>
, northern
<collectingCountry id="F3190EFEFF839960FC3268E97879FF48" box="[1008,1105,188,213]" name="Vietnam" pageId="8" pageNumber="547">Vietnam</collectingCountry>
) and
<taxonomicName id="4C0E35EDFF839960FB5F68E879B4FF49" authority="Smith 1923" authorityName="Smith" authorityYear="1923" box="[1181,1436,188,213]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="547" phylum="Chordata" rank="species" species="laotus">
<emphasis id="B97A927CFF839960FB5F68E8792BFF48" box="[1181,1283,188,213]" italics="true" pageId="8" pageNumber="547">T. laotus</emphasis>
<bibRefCitation id="EF9F339FFF839960FAD168E979B4FF49" author="Smith" box="[1299,1436,188,213]" pageId="8" pageNumber="547" refString="Smith, M. A. (1923) A review of the lizards of the genus Tropidophorus on the Asiatic mainland. Proceedings of the Zoological Society of London, 1923, 775 - 781." type="proceedings paper" year="1923">Smith 1923</bibRefCitation>
</taxonomicName>
(distribution: Pak Lai district in
<collectingCountry id="F3190EFEFF839960FDD568B57E65FF6A" box="[535,589,224,247]" name="Laos" pageId="8" pageNumber="547">Laos</collectingCountry>
and
<collectingCountry id="F3190EFEFF839960FD4C688A7EDFFF65" box="[654,759,223,248]" name="Thailand" pageId="8" pageNumber="547">Thailand</collectingCountry>
).
<taxonomicName id="4C0E35EDFF839960FCCC688A7836FF65" box="[782,1054,223,248]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="547" phylum="Chordata" rank="species" species="boehmei">
<emphasis id="B97A927CFF839960FCCC688A7836FF65" box="[782,1054,223,248]" italics="true" pageId="8" pageNumber="547">Tropidophorus boehmei</emphasis>
</taxonomicName>
differs from the new species in showing tiny, reduced prefrontals vs. trapezoidal prefrontals that are in contact, supralabials six vs. seven, longitudinal scale rows 6069 vs. 58, and transverse scale rows at midbody 3032 vs.
<quantity id="4CF6E38BFF839960FB52697D78E5FEDD" box="[1168,1229,295,320]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="8" pageNumber="547" unit="in" value="34.0">34 in</quantity>
the new Bornean species.
<taxonomicName id="4C0E35EDFF839960FF3869197DC6FEF8" box="[250,494,332,357]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="547" phylum="Chordata" rank="species" species="laotus">
<emphasis id="B97A927CFF839960FF3869197DC6FEF8" box="[250,494,332,357]" italics="true" pageId="8" pageNumber="547">Tropidophorus laotus</emphasis>
</taxonomicName>
differs from the new species in showing a frontonasal that is divided vs. fused, supralabials six vs. seven, and anterior loreal divided vs. undivided in the new species.
</paragraph>
<paragraph id="8BB14E6EFF839960FF0569C17D9DFE52" blockId="8.[151,1436,151,464]" pageId="8" pageNumber="547">
Finally, to aid field identification, we present below a dichotomous identification key to the Bornean species of the genus
<taxonomicName id="4C0E35EDFF839960FECA69E27D85FE4D" box="[264,429,439,464]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="547" phylum="Chordata" rank="genus">
<emphasis id="B97A927CFF839960FECA69E27D85FE4D" box="[264,429,439,464]" italics="true" pageId="8" pageNumber="547">Tropidophorus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>