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<document id="30563B9B2BC005E66A3EB7053F48348D" ID-DOI="10.5281/zenodo.206579" ID-GBIF-Dataset="d02a1a04-922c-443d-b60d-8f30075211a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="206579" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460228489759" checkinUser="plazi" docAuthor="Okanishi, Masanori &amp; Fujita, Toshihiko" docDate="2011" docId="03901004FFC0FFABFF1948A2A881ABC0" docLanguage="en" docName="zt02751p039.pdf" docOrigin="Zootaxa 2751" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Asteroporpa (Asteromoana) muricatopatella Okanishi &amp; Fujita, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="33" masterDocId="FFA9687CFFC3FFA3FF8E4835AA22AA71" masterDocTitle="Two new species of the subgenus Asteroporpa (Asteromoana) (Ophiuroidea: Euryalida: Gorgonocephalidae) from Japan" masterLastPageNumber="39" masterPageNumber="25" pageNumber="28" updateTime="1698238769680" updateUser="plazi">
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<mods:title id="C1718BB211AA3F26BEE2887BB0AF1B4F">Two new species of the subgenus Asteroporpa (Asteromoana) (Ophiuroidea: Euryalida: Gorgonocephalidae) from Japan</mods:title>
</mods:titleInfo>
<mods:name id="53A5A1CCB01A74728E3994457EE9F7A0" type="personal">
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<mods:namePart id="32D2A4214CC0D003F929C2093AA1F653">Okanishi, Masanori</mods:namePart>
</mods:name>
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<mods:namePart id="5C174EECADE8DE6B1807390319EBCEED">Fujita, Toshihiko</mods:namePart>
</mods:name>
<mods:typeOfResource id="DAA165EA29511455E3D05F8D820AD151">text</mods:typeOfResource>
<mods:relatedItem id="D4150C399C3E66F9BCCB0FAC5BB5A60A" type="host">
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<mods:title id="F796F5EFA421679B8F387C562F70331B">Zootaxa</mods:title>
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<mods:part id="C94F4D3BD48AA814D91FC4E015553DF3">
<mods:date id="5FE39F9AB4D209194826236C2DE3554B">2011</mods:date>
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<mods:number id="056077B820F0A5263DF7DC1E110226AA">2751</mods:number>
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<mods:identifier id="CA59A250B07B7CF925336D232AC21E60" type="DOI">10.5281/zenodo.206579</mods:identifier>
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<treatment id="03901004FFC0FFABFF1948A2A881ABC0" ID-DOI="http://doi.org/10.5281/zenodo.6191645" ID-GBIF-Taxon="119386958" ID-Zenodo-Dep="6191645" LSID="urn:lsid:plazi:treatment:03901004FFC0FFABFF1948A2A881ABC0" httpUri="http://treatment.plazi.org/id/03901004FFC0FFABFF1948A2A881ABC0" lastPageId="8" lastPageNumber="33" pageId="3" pageNumber="28">
<subSubSection id="C323F299FFC0FFA0FF1948A2AB29AA86" pageId="3" pageNumber="28" type="nomenclature">
<paragraph id="8B86A112FFC0FFA0FF1948A2A933AAC0" blockId="3.[151,785,151,247]" box="[151,785,151,177]" pageId="3" pageNumber="28">
<heading id="D0CE167EFFC0FFA0FF1948A2A933AAC0" bold="true" box="[151,785,151,177]" fontSize="11" level="1" pageId="3" pageNumber="28" reason="1">
<emphasis id="B94D7D00FFC0FFA0FF1948A2A933AAC0" bold="true" box="[151,785,151,177]" pageId="3" pageNumber="28">
<taxonomicName id="4C39DA91FFC0FFA0FF1948A2A88EAAC0" ID-CoL="90641ae2-8162-472a-a403-d78bb9e52493" box="[151,684,151,177]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="3" pageNumber="28" phylum="Echinodermata" rank="species" species="muricatopatella" status="sp. nov." subGenus="Asteromoana">
<emphasis id="B94D7D00FFC0FFA0FF1948A2AB09AAC0" bold="true" box="[151,299,151,177]" italics="true" pageId="3" pageNumber="28">Asteroporpa</emphasis>
(
<emphasis id="B94D7D00FFC0FFA0FEB248A2ABFFAAC0" bold="true" box="[316,477,151,177]" italics="true" pageId="3" pageNumber="28">Asteromoana</emphasis>
)
<emphasis id="B94D7D00FFC0FFA0FE6048A2A88EAAC0" bold="true" box="[494,684,151,177]" italics="true" pageId="3" pageNumber="28">muricatopatella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EC07BFFC0FFA0FD3A48A2A933AAC0" box="[692,785,151,177]" pageId="3" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B86A112FFC0FFA0FF19488EAB29AA86" blockId="3.[151,785,151,247]" pageId="3" pageNumber="28">
[New Japanese name: Toge-moana-moduru] (
<figureCitation id="1302BD97FFC0FFA0FF1148EAAB21AA86" box="[159,259,223,247]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="4.[151,250,1928,1951]" captionStartId-1="5.[151,250,1922,1945]" captionStartId-2="6.[151,250,758,781]" captionStartId-3="7.[151,250,1265,1288]" captionTargetBox-0="[154,1430,193,1905]" captionTargetBox-1="[228,1355,181,1892]" captionTargetBox-2="[167,1419,187,738]" captionTargetBox-3="[154,1436,193,1234]" captionTargetId-0="figure@4.[151,1436,119,1907]" captionTargetId-1="figure@5.[228,1362,178,1895]" captionTargetId-2="figure@6.[167,1424,140,741]" captionTargetId-3="figure@7.[154,1439,131,1242]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 2. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral side; B, oral side." captionText-1="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." captionText-2="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." captionText-3="FIGURE 5. Asteroporpa (Asteromoana) muricatopatella sp. nov., one paratype (NSMT E- 5619 - B). SEM photographs of internal ossicles. A B, a hooklet-bearing plate (A) and a hooklet (B) at the aboral periphery of the disc; C D, cone-shaped dermal ossicles at the aboral periphery of the disc, external (C) and lateral (D) views; E, a hooklet at the middle portion of the arm; F G, plate-shaped dermal ossicles at the aboral basal portion of the arm, external (F) and lateral (G) views; H I, plate-shaped dermal ossicles at the aboral distal portion of the arm, external (H) and lateral (I) views; J K, arm spines from basal portion of the arm (J) and distal portion of the arm (K) with an arc indicating reticular structure. Arrows indicate the orientation (D, G, I): ext, external side; int, internal side." httpUri-0="https://zenodo.org/record/206581/files/figure.png" httpUri-1="https://zenodo.org/record/206582/files/figure.png" httpUri-2="https://zenodo.org/record/206583/files/figure.png" httpUri-3="https://zenodo.org/record/206584/files/figure.png" pageId="3" pageNumber="28">Figs 25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C323F299FFC0FFA0FF19491AAFA9AB8E" pageId="3" pageNumber="28" type="materials_examined">
<paragraph id="8B86A112FFC0FFA0FF19491AAFA9AB8E" blockId="3.[151,1437,302,1698]" pageId="3" pageNumber="28">
<emphasis id="B94D7D00FFC0FFA0FF19491AAB72AB36" bold="true" box="[151,336,302,327]" pageId="3" pageNumber="28">
<typeStatus id="54821FB0FFC0FFA0FF19491AAAF2AB36" box="[151,208,303,327]" pageId="3" pageNumber="28">Type</typeStatus>
materials.
</emphasis>
<collectionCode id="ED2839D7FFC0FFA0FED2491BAB88AB37" box="[348,426,302,326]" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="3" pageNumber="28">NSMT</collectionCode>
<collectionCode id="ED2839D7FFC0FFA0FE3C491BABEFAB37" box="[434,461,302,326]" pageId="3" pageNumber="28">E-</collectionCode>
6548,
<typeStatus id="54821FB0FFC0FFA0FD9B491BA858AB37" box="[533,634,302,326]" pageId="3" pageNumber="28" type="holotype">holotype</typeStatus>
, collected by fishing boat
<emphasis id="B94D7D00FFC0FFA0FC23491AAE3DAB37" box="[941,1055,303,326]" italics="true" pageId="3" pageNumber="28">Mio-maru</emphasis>
, off Kume-jima Island, Okinawa Prefecture, southwestern
<collectingCountry id="F32EE182FFC0FFA0FE374961ABDEAB1A" box="[441,508,340,363]" name="Japan" pageId="3" pageNumber="28">Japan</collectingCountry>
, 26˚14.116N, 126˚50.120E,
<quantity id="4CC10CF7FFC0FFA0FCD84966A982AB1A" box="[854,928,339,363]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="28" unit="m" value="410.0">410 m</quantity>
deep,
<date id="FF8787D2FFC0FFA0FC7E4966AEE0AB1A" box="[1008,1218,339,363]" pageId="3" pageNumber="28" value="2010-02-20">20 February 2010</date>
.
<collectionCode id="ED2839D7FFC0FFA0FB414966AF3FAB1A" box="[1231,1309,339,363]" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="3" pageNumber="28">NSMT</collectionCode>
<collectionCode id="ED2839D7FFC0FFA0FAA94966AF60AB1A" box="[1319,1346,339,363]" pageId="3" pageNumber="28">E-</collectionCode>
5619-B, one
<typeStatus id="54821FB0FFC0FFA0FF40494CAB13ABE0" box="[206,305,377,401]" pageId="3" pageNumber="28" type="paratype">paratype</typeStatus>
, collected with ROV
<emphasis id="B94D7D00FFC0FFA0FDCC494CA8BBABE0" box="[578,665,377,401]" italics="true" pageId="3" pageNumber="28">Hakuyo</emphasis>
of a manned submersible
<emphasis id="B94D7D00FFC0FFA0FC68494CAE55ABE0" box="[998,1143,377,401]" italics="true" pageId="3" pageNumber="28">Shinyu-maru</emphasis>
, off Yaku-shima Island, Kagoshima Prefecture, southwestern
<collectingCountry id="F32EE182FFC0FFA0FDB349AAA8A2ABC7" box="[573,640,415,438]" name="Japan" pageId="3" pageNumber="28">Japan</collectingCountry>
, 30˚49.5N, 130˚48.52E,
<quantity id="4CC10CF7FFC0FFA0FC2549ABA9D6ABC7" box="[939,1012,414,438]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="28" unit="m" value="140.0">140 m</quantity>
deep,
<date id="FF8787D2FFC0FFA0FBCE49ABAF02ABC7" box="[1088,1312,414,438]" pageId="3" pageNumber="28" value="2007-09-26">26 September 2007</date>
.
<collectionCode id="ED2839D7FFC0FFA0FAA249ABAF58ABC7" box="[1324,1402,414,438]" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="3" pageNumber="28">NSMT</collectionCode>
<collectionCode id="ED2839D7FFC0FFA0FA0C49AAAFBEABC7" box="[1410,1436,415,438]" pageId="3" pageNumber="28">E-</collectionCode>
6547, two
<typeStatus id="54821FB0FFC0FFA0FE8149F6AB5CABAA" box="[271,382,451,475]" pageId="3" pageNumber="28" type="paratype">paratypes</typeStatus>
, collected off Kagoshima Prefecture or off Okinawa Prefecture, southwestern
<collectingCountry id="F32EE182FFC0FFA0FB6149F6AF10ABAB" box="[1263,1330,451,474]" name="Japan" pageId="3" pageNumber="28">Japan</collectingCountry>
, approximately
<quantity id="4CC10CF7FFC0FFA0FF6449D2AB4BAB8E" box="[234,361,487,511]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="28" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200300 m</quantity>
(no further data are available and this site is not shown in
<figureCitation id="1302BD97FFC0FFA0FC7649D2AE1AAB8E" box="[1016,1080,487,511]" captionStart="FIGURE 1" captionStartId="1.[151,250,1801,1824]" captionTargetBox="[174,1418,327,1773]" captionTargetId="figure@1.[166,1422,321,1782]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of Asteroporpa (Asteromoana) muricatopatella sp. nov. and A. (Asterom.) koyoae sp. nov." httpUri="https://zenodo.org/record/206580/files/figure.png" pageId="3" pageNumber="28">Fig. 1</figureCitation>
),
<date id="FF8787D2FFC0FFA0FBDF49D2AF0BAB8E" box="[1105,1321,487,511]" pageId="3" pageNumber="28" value="2007-11-16">16 November 2007</date>
(
<figureCitation id="1302BD97FFC0FFA0FAB649D2AF58AB8E" box="[1336,1402,487,511]" captionStart="FIGURE 1" captionStartId="1.[151,250,1801,1824]" captionTargetBox="[174,1418,327,1773]" captionTargetId="figure@1.[166,1422,321,1782]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of Asteroporpa (Asteromoana) muricatopatella sp. nov. and A. (Asterom.) koyoae sp. nov." httpUri="https://zenodo.org/record/206580/files/figure.png" pageId="3" pageNumber="28">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C323F299FFC0FFA0FF494A39AB27AF99" pageId="3" pageNumber="28" type="description">
<paragraph id="8B86A112FFC0FFA0FF494A39AE87A854" blockId="3.[151,1437,302,1698]" box="[199,1189,524,549]" pageId="3" pageNumber="28">
<emphasis id="B94D7D00FFC0FFA0FF494A39ABC6A854" bold="true" box="[199,484,524,549]" pageId="3" pageNumber="28">
Description of
<typeStatus id="54821FB0FFC0FFA0FEF84A38ABFDA854" box="[374,479,525,549]" pageId="3" pageNumber="28" type="holotype">holotype</typeStatus>
.
</emphasis>
Size is
<quantity id="4CC10CF7FFC0FFA0FDB24A38A8B0A854" box="[572,658,525,549]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="28" unit="mm" value="4.8">4.8 mm</quantity>
in disc diameter,
<quantity id="4CC10CF7FFC0FFA0FCD74A38A985A854" box="[857,935,525,549]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="28" unit="mm" value="35.0">35 mm</quantity>
in arm length (
<figureCitation id="1302BD97FFC0FFA0FBDD4A38AEBAA854" box="[1107,1176,525,549]" captionStart="FIGURE 2" captionStartId="4.[151,250,1928,1951]" captionTargetBox="[154,1430,193,1905]" captionTargetId="figure@4.[151,1436,119,1907]" captionTargetPageId="4" captionText="FIGURE 2. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral side; B, oral side." httpUri="https://zenodo.org/record/206581/files/figure.png" pageId="3" pageNumber="28">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8B86A112FFC0FFA0FF4B4A06AF0BA9E6" blockId="3.[151,1437,302,1698]" pageId="3" pageNumber="28">
<emphasis id="B94D7D00FFC0FFA0FF4B4A06AADDA83B" box="[197,255,563,586]" italics="true" pageId="3" pageNumber="28">Disc.</emphasis>
Disc is five-lobed in shape with notched interradial edges (
<figureCitation id="1302BD97FFC0FFA0FC354A07AE25A83B" box="[955,1031,562,586]" captionStart="FIGURE 2" captionStartId="4.[151,250,1928,1951]" captionTargetBox="[154,1430,193,1905]" captionTargetId="figure@4.[151,1436,119,1907]" captionTargetPageId="4" captionText="FIGURE 2. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral side; B, oral side." httpUri="https://zenodo.org/record/206581/files/figure.png" pageId="3" pageNumber="28">Figs 2</figureCitation>
,
<figureCitation id="1302BD97FFC0FFA0FB984A07AE01A83B" box="[1046,1059,562,586]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">3</figureCitation>
AB). On the aboral side, radial shields and their surrounds are tumid (
<figureCitation id="1302BD97FFC0FFA0FDC84A62A8AAA81E" box="[582,648,599,623]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">Fig. 3</figureCitation>
A). There are no conspicuous ridges of rows of raised hooklet-bearing plates and the hooklet-bearing plates are scattered on periphery of each radial shield (
<figureCitation id="1302BD97FFC0FFA0FBD44A49AEBFA8E5" box="[1114,1181,636,660]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">Fig. 3</figureCitation>
E). Each hooklet-bearing plate possesses eight to eleven tubercle-shaped articulations for hooklets. Each hooklet has two inner teeth. Except for the hooklet-bearing plates, the aboral disc is covered by cone-shaped dermal ossicles with one or two terminal projections which are approximately one-fourth the length of the height of the cone-shaped dermal ossicles (
<figureCitation id="1302BD97FFC0FFA0FF5C4B25AB37A959" box="[210,277,784,808]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">Fig. 3</figureCitation>
D). The cone-shaped dermal ossicles are approximately 150230 µm in length and 270 µm in height at the periphery of the disc (
<figureCitation id="1302BD97FFC0FFA0FE394B00ABDAA93C" box="[439,504,821,845]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">Fig. 3</figureCitation>
D), and approximately 130210 µm in length and 240 µm in height at center of the disc (
<figureCitation id="1302BD97FFC0FFA0FF5B4B6FAB3BA903" box="[213,281,858,882]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">Fig. 3</figureCitation>
C). Radial shields are completely concealed by dermal ossicles, oval shape, approximately
<quantity id="4CC10CF7FFC0FFA0FAA94B6FAF5CA903" box="[1319,1406,858,882]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="28" unit="mm" value="4.5">4.5 mm</quantity>
in length, 3.0 mm in width distally,
<quantity id="4CC10CF7FFC0FFA0FD874B4AA87DA9E6" box="[521,607,895,919]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="28" unit="mm" value="1.5">1.5 mm</quantity>
in width proximally, and do not reach the disc center (
<figureCitation id="1302BD97FFC0FFA0FB4C4B4AAF27A9E6" box="[1218,1285,895,919]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">Fig. 3</figureCitation>
A).
</paragraph>
<paragraph id="8B86A112FFC0FFA0FF4B4B91A8B1AF7B" blockId="3.[151,1437,302,1698]" pageId="3" pageNumber="28">
The oral surface of the disc is covered both by flat, polygonal plate-shaped dermal ossicles and by cone-shaped dermal ossicles with mostly one or rarely two terminal projections similar to the aboral disc (
<figureCitation id="1302BD97FFC0FFA0FB344BFCAEDFA990" box="[1210,1277,969,993]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">Fig. 3</figureCitation>
G). The coneshaped dermal ossicles are scattered only on the disc periphery, not on the oral plates (
<figureCitation id="1302BD97FFC0FFA0FB1E4BDAAEF6AE76" box="[1168,1236,1007,1031]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">Fig. 3</figureCitation>
GH). The plateshaped dermal ossicles are approximately 140190 µm in length and 50 µm in height on periphery of the disc, and approximately 6080 µm in length and 20 µm in height on oral plates (
<figureCitation id="1302BD97FFC0FFA0FC344C0CA9D9AE20" box="[954,1019,1081,1105]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">Fig. 3</figureCitation>
GH). The cone-shaped dermal ossicles are approximately 150190 µm in length and 190 µm in height (
<figureCitation id="1302BD97FFC0FFA0FC394C6BA9D9AE07" box="[951,1019,1118,1142]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">Fig. 3</figureCitation>
G). Oral shields, adoral shields, oral plates and ventral arm plates are completely concealed by dermal ossicles (
<figureCitation id="1302BD97FFC0FFA0FC524CB6AE3DAEEA" box="[988,1055,1155,1179]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">Fig. 3</figureCitation>
B, G). Uniformly acute and spineformed teeth are situated on the jaws (
<figureCitation id="1302BD97FFC0FFA0FDDF4C9DA8B7AEB1" box="[593,661,1192,1216]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">Fig. 3</figureCitation>
H). Teeth length varies with their position of the jaw. On the top of jaws, they are approximately 190 µm in length, and toward the basal part of the jaws, the teeth gradually decrease in length to approximately 135 µm (
<figureCitation id="1302BD97FFC0FFA0FDA24CC7A84DAF7B" box="[556,623,1266,1290]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="3" pageNumber="28">Fig. 3</figureCitation>
H).
</paragraph>
<paragraph id="8B86A112FFC0FFA0FF194D22AB27AF99" blockId="3.[151,1437,302,1698]" pageId="3" pageNumber="28">
The lateral interradial surface of the disc is slightly inclined to the oral side and covered only by cone-shaped dermal ossicles with mostly one or rarely two terminal projections which are approximately one-third length of the height of the cone-shaped dermal ossicles (
<figureCitation id="1302BD97FFC0FFA0FDF34D54A8E3AF08" box="[637,705,1377,1401]" captionStart="FIGURE 4" captionStartId="6.[151,250,758,781]" captionTargetBox="[167,1419,187,738]" captionTargetId="figure@6.[167,1424,140,741]" captionTargetPageId="6" captionText="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="3" pageNumber="28">Fig. 4</figureCitation>
A). The cone-shaped dermal ossicles are approximately 150190 µm in length and 240 µm in height (
<figureCitation id="1302BD97FFC0FFA0FDA94DB3A848AFEF" box="[551,618,1414,1438]" captionStart="FIGURE 4" captionStartId="6.[151,250,758,781]" captionTargetBox="[167,1419,187,738]" captionTargetId="figure@6.[167,1424,140,741]" captionTargetPageId="6" captionText="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="3" pageNumber="28">Fig. 4</figureCitation>
A). Two genital slits (
<quantity id="4CC10CF7FFC0FFA0FCD44DB3A992AFEF" box="[858,944,1414,1438]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="28" unit="mm" value="0.3">0.3 mm</quantity>
long and
<quantity id="4CC10CF7FFC0FFA0FB934DB3AEA2AFEF" box="[1053,1152,1414,1438]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="28" unit="mm" value="0.05">0.05 mm</quantity>
wide) are present in each interradius (
<figureCitation id="1302BD97FFC0FFA0FE914D9EAB47AFB2" box="[287,357,1451,1475]" captionStart="FIGURE 4" captionStartId="6.[151,250,758,781]" captionTargetBox="[167,1419,187,738]" captionTargetId="figure@6.[167,1424,140,741]" captionTargetPageId="6" captionText="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="3" pageNumber="28">Fig. 4</figureCitation>
A). One circular madreporite is situated on oral interradius, approximately 440 µm in diameter (
<figureCitation id="1302BD97FFC0FFA0FF114DE5AAC6AF99" box="[159,228,1488,1512]" captionStart="FIGURE 4" captionStartId="6.[151,250,758,781]" captionTargetBox="[167,1419,187,738]" captionTargetId="figure@6.[167,1424,140,741]" captionTargetPageId="6" captionText="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="3" pageNumber="28">Fig. 4</figureCitation>
B).
</paragraph>
</subSubSection>
<subSubSection id="C323F299FFC0FFA5FF494DC0A83BADC0" lastPageId="6" lastPageNumber="31" pageId="3" pageNumber="28" type="diagnosis">
<paragraph id="8B86A112FFC0FFA0FF494DC0AEA8AC42" blockId="3.[151,1437,302,1698]" pageId="3" pageNumber="28">
<emphasis id="B94D7D00FFC0FFA0FF494DC0AB63AC7F" bold="true" box="[199,321,1525,1550]" pageId="3" pageNumber="28">Diagnosis.</emphasis>
Dermal ossicles are cone-shaped with terminal projections on the aboral disc; plate-shaped on oral plates; cone-shaped with relatively short terminal projections at the lateral interradial disc.
</paragraph>
<paragraph id="8B86A112FFC0FFA0FF4B4E75ABEAACD3" blockId="3.[151,1437,302,1698]" pageId="3" pageNumber="28">
<emphasis id="B94D7D00FFC0FFA0FF4B4E75AB2AAC29" box="[197,264,1600,1624]" italics="true" pageId="3" pageNumber="28">Arms.</emphasis>
Arms are simple, five in number, have no abrupt gap in thickness. The basal portion of the arm is
<quantity id="4CC10CF7FFC0FFA0FAF74E0AAAE1AC0C" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="28" unit="mm" value="2.2">2.2 mm</quantity>
wide and
<quantity id="4CC10CF7FFC0FFA0FEB64E50ABADAC0C" box="[312,399,1637,1661]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="28" unit="mm" value="2.1">2.1 mm</quantity>
high, with an arched aboral surface and flattened oral surface. Arms taper gradually toward the arm tip (
<figureCitation id="1302BD97FFC0FFA0FE914EBFAB4AACD3" box="[287,360,1674,1698]" captionStart="FIGURE 2" captionStartId="4.[151,250,1928,1951]" captionTargetBox="[154,1430,193,1905]" captionTargetId="figure@4.[151,1436,119,1907]" captionTargetPageId="4" captionText="FIGURE 2. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral side; B, oral side." httpUri="https://zenodo.org/record/206581/files/figure.png" pageId="3" pageNumber="28">Figs 2</figureCitation>
,
<figureCitation id="1302BD97FFC0FFA0FEFA4EBFABA1ACD3" box="[372,387,1674,1698]" captionStart="FIGURE 4" captionStartId="6.[151,250,758,781]" captionTargetBox="[167,1419,187,738]" captionTargetId="figure@6.[167,1424,140,741]" captionTargetPageId="6" captionText="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="3" pageNumber="28">4</figureCitation>
CG).
</paragraph>
<caption id="DF46F19AFFC7FFA7FF194FBDAF4BADEF" box="[151,1385,1928,1951]" httpUri="https://zenodo.org/record/206581/files/figure.png" pageId="4" pageNumber="29" targetBox="[154,1430,193,1905]" targetPageId="4">
<paragraph id="8B86A112FFC7FFA7FF194FBDAF4BADEF" blockId="4.[151,1385,1928,1951]" box="[151,1385,1928,1951]" pageId="4" pageNumber="29">
<emphasis id="B94D7D00FFC7FFA7FF194FBDAB31ADEC" bold="true" box="[151,275,1928,1951]" pageId="4" pageNumber="29">FIGURE 2.</emphasis>
<taxonomicName id="4C39DA91FFC7FFA7FE974FBCA8FEADEF" box="[281,732,1928,1950]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="4" pageNumber="29" phylum="Echinodermata" rank="species" species="muricatopatella" status="sp. nov." subGenus="Asteromoana">
<emphasis id="B94D7D00FFC7FFA7FE974FBCABB5ADEF" box="[281,407,1929,1950]" italics="true" pageId="4" pageNumber="29">Asteroporpa</emphasis>
(
<emphasis id="B94D7D00FFC7FFA7FE2B4FBCA809ADEF" box="[421,555,1929,1950]" italics="true" pageId="4" pageNumber="29">Asteromoana</emphasis>
)
<emphasis id="B94D7D00FFC7FFA7FDB44FBDA8FEADEF" box="[570,732,1928,1950]" italics="true" pageId="4" pageNumber="29">muricatopatella</emphasis>
</taxonomicName>
<emphasis id="B94D7D00FFC7FFA7FD6C4FBCA910ADEF" bold="true" box="[738,818,1929,1950]" pageId="4" pageNumber="29">
<taxonomicNameLabel id="A27EC07BFFC7FFA7FD6C4FBCA910ADEF" box="[738,818,1929,1950]" pageId="4" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (NSMT E-6548). A, aboral side; B, oral side.
</paragraph>
</caption>
<caption id="DF46F19AFFC6FFA6FF194FB7AEB6AD84" httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="5" pageNumber="30" targetBox="[228,1355,181,1892]" targetPageId="5">
<paragraph id="8B86A112FFC6FFA6FF194FB7AEB6AD84" blockId="5.[151,1436,1922,2037]" pageId="5" pageNumber="30">
<emphasis id="B94D7D00FFC6FFA6FF194FB7AB31ADE9" bold="true" box="[151,275,1922,1945]" pageId="5" pageNumber="30">FIGURE 3.</emphasis>
<taxonomicName id="4C39DA91FFC6FFA6FE944FB6A8FCADE8" box="[282,734,1923,1945]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="5" pageNumber="30" phylum="Echinodermata" rank="species" species="muricatopatella" status="sp. nov." subGenus="Asteromoana">
<emphasis id="B94D7D00FFC6FFA6FE944FB6ABBAADE9" box="[282,408,1923,1944]" italics="true" pageId="5" pageNumber="30">Asteroporpa</emphasis>
(
<emphasis id="B94D7D00FFC6FFA6FE284FB6A80EADE9" box="[422,556,1923,1944]" italics="true" pageId="5" pageNumber="30">Asteromoana</emphasis>
)
<emphasis id="B94D7D00FFC6FFA6FDB24FB6A8FCADE8" box="[572,734,1923,1945]" italics="true" pageId="5" pageNumber="30">muricatopatella</emphasis>
</taxonomicName>
<emphasis id="B94D7D00FFC6FFA6FD6A4FB6A917ADE9" bold="true" box="[740,821,1923,1944]" pageId="5" pageNumber="30">
<taxonomicNameLabel id="A27EC07BFFC6FFA6FD6A4FB6A917ADE9" box="[740,821,1923,1944]" pageId="5" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (NSMT E-6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws.
</paragraph>
</caption>
<caption id="DF46F19AFFC5FFA5FF194AC3AEFEA918" httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="6" pageNumber="31" targetBox="[167,1419,187,738]" targetPageId="6">
<paragraph id="8B86A112FFC5FFA5FF194AC3AEFEA918" blockId="6.[151,1436,758,873]" pageId="6" pageNumber="31">
<emphasis id="B94D7D00FFC5FFA5FF194AC3AB30A97D" bold="true" box="[151,274,758,781]" pageId="6" pageNumber="31">FIGURE 4.</emphasis>
<taxonomicName id="4C39DA91FFC5FFA5FE974ACDA8FEA97C" box="[281,732,759,781]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="6" pageNumber="31" phylum="Echinodermata" rank="species" species="muricatopatella" status="sp. nov." subGenus="Asteromoana">
<emphasis id="B94D7D00FFC5FFA5FE974ACDABB5A97C" box="[281,407,760,781]" italics="true" pageId="6" pageNumber="31">Asteroporpa</emphasis>
(
<emphasis id="B94D7D00FFC5FFA5FE2B4ACDA809A97C" box="[421,555,760,781]" italics="true" pageId="6" pageNumber="31">Asteromoana</emphasis>
)
<emphasis id="B94D7D00FFC5FFA5FDB44AC2A8FEA97C" box="[570,732,759,781]" italics="true" pageId="6" pageNumber="31">muricatopatella</emphasis>
</taxonomicName>
<emphasis id="B94D7D00FFC5FFA5FD6C4AC2A910A97D" bold="true" box="[738,818,759,780]" pageId="6" pageNumber="31">
<taxonomicNameLabel id="A27EC07BFFC5FFA5FD6C4AC2A910A97D" box="[738,818,759,780]" pageId="6" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (NSMT E-6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G).
</paragraph>
</caption>
<paragraph id="8B86A112FFC5FFA5FF4B4BA6A852AFDF" blockId="6.[151,1436,915,1969]" pageId="6" pageNumber="31">
On the aboral and lateral surface, each arm segment is surrounded by single annular transverse row of hookletbearing plates (
<figureCitation id="1302BD97FFC5FFA5FECC4B82ABA5A9BE" box="[322,391,951,975]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="6" pageNumber="31">Figs 3</figureCitation>
F, 4C, E, G). Each hooklet bears two inner teeth. On the basal one or two arm segments, each hooklet-bearing plate is separated by plate-shaped dermal ossicles. On third and subsequent distal segments, those plates are fully in contact with each other (
<figureCitation id="1302BD97FFC5FFA5FDFD4C34A89BAE68" box="[627,697,1025,1049]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1945]" captionTargetBox="[228,1355,181,1892]" captionTargetId="figure@5.[228,1362,178,1895]" captionTargetPageId="5" captionText="FIGURE 3. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral disc and basal portion of the arm; B, oral disc and basal portion of the arm; C, aboral central part of the disc; D, aboral periphery of the disc; E, aboral periphery of the disc, a hooklet-bearing plate is indicated by an arrow; F, aboral basal portion of the arm, hooklet-bearing plates are indicated by arrows; G, oral disc, a cone-shaped dermal ossicle is indicated by an arrow; H, jaws." httpUri="https://zenodo.org/record/206582/files/figure.png" pageId="6" pageNumber="31">Figs 3</figureCitation>
F, 4C, E, G). Except for hooklet-bearing plates, the aboral and lateral surface of the basal arm is completely covered by domed and polygonal plate-shaped dermal ossicles of approximately 150220 µm in length and 200 µm in height (
<figureCitation id="1302BD97FFC5FFA5FCB64C7EA95EAE12" box="[824,892,1099,1123]" captionStart="FIGURE 4" captionStartId="6.[151,250,758,781]" captionTargetBox="[167,1419,187,738]" captionTargetId="figure@6.[167,1424,140,741]" captionTargetPageId="6" captionText="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="6" pageNumber="31">Fig. 4</figureCitation>
C). The oral surface is covered by flat and polygonal plate-shaped dermal ossicles, similar to those on the oral disc of approximately 80160 µm in length and 60 µm in height (
<figureCitation id="1302BD97FFC5FFA5FEB44CA1ABA2AEDD" box="[314,384,1172,1196]" captionStart="FIGURE 4" captionStartId="6.[151,250,758,781]" captionTargetBox="[167,1419,187,738]" captionTargetId="figure@6.[167,1424,140,741]" captionTargetPageId="6" captionText="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="6" pageNumber="31">Fig. 4</figureCitation>
D). In the middle portion of the arm, the aboral and lateral surface is covered by domed and round granule-shaped dermal ossicles of approximately 120150 µm in length and 130 µm in height (
<figureCitation id="1302BD97FFC5FFA5FABB4C8CAF5EAEA0" box="[1333,1404,1209,1233]" captionStart="FIGURE 4" captionStartId="6.[151,250,758,781]" captionTargetBox="[167,1419,187,738]" captionTargetId="figure@6.[167,1424,140,741]" captionTargetPageId="6" captionText="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="6" pageNumber="31">Fig. 4</figureCitation>
E). There are also some smaller, round, relatively flat and round granule-shaped dermal ossicles of approximately 100 120 µm in length and 110 µm in height. In the distal portion of the arm, the aboral and lateral surface is covered by flat and round granule-shaped dermal ossicles of approximately 80120 µm in length and 100 µm in height (
<figureCitation id="1302BD97FFC5FFA5FAE14D12AA84AF15" captionStart="FIGURE 4" captionStartId="6.[151,250,758,781]" captionTargetBox="[167,1419,187,738]" captionTargetId="figure@6.[167,1424,140,741]" captionTargetPageId="6" captionText="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="6" pageNumber="31">Fig. 4</figureCitation>
G). The oral surface is covered by flat and round plate-shaped dermal ossicles of approximately 6080 µm in length and 30 µm in height (
<figureCitation id="1302BD97FFC5FFA5FE584D44A838AFF8" box="[470,538,1393,1417]" captionStart="FIGURE 4" captionStartId="6.[151,250,758,781]" captionTargetBox="[167,1419,187,738]" captionTargetId="figure@6.[167,1424,140,741]" captionTargetPageId="6" captionText="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="6" pageNumber="31">Fig. 4</figureCitation>
F). Throughout the arm, lateral arm plates and ventral arm plates are completely concealed by dermal ossicles (
<figureCitation id="1302BD97FFC5FFA5FE624DA3A80DAFDF" box="[492,559,1430,1454]" captionStart="FIGURE 4" captionStartId="6.[151,250,758,781]" captionTargetBox="[167,1419,187,738]" captionTargetId="figure@6.[167,1424,140,741]" captionTargetPageId="6" captionText="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="6" pageNumber="31">Fig. 4</figureCitation>
D, F).
</paragraph>
<paragraph id="8B86A112FFC5FFA5FF4B4D8EA903AC88" blockId="6.[151,1436,915,1969]" pageId="6" pageNumber="31">
First tentacle pores have no arm spine; second has four, third has five or six, and from fifth pores to the pores at middle arm segments there are six arm spines (
<figureCitation id="1302BD97FFC5FFA5FD354DEAA920AF86" box="[699,770,1503,1527]" captionStart="FIGURE 4" captionStartId="6.[151,250,758,781]" captionTargetBox="[167,1419,187,738]" captionTargetId="figure@6.[167,1424,140,741]" captionTargetPageId="6" captionText="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="6" pageNumber="31">Fig. 4</figureCitation>
D). Subsequently, the number of the arm spines decreases gradually to one toward the arm tip. Throughout the arm, all arm spines are approximately half the length of the corresponding arm segment, and are covered by thin integument (
<figureCitation id="1302BD97FFC5FFA5FCFC4E1CA997AC30" box="[882,949,1577,1601]" captionStart="FIGURE 4" captionStartId="6.[151,250,758,781]" captionTargetBox="[167,1419,187,738]" captionTargetId="figure@6.[167,1424,140,741]" captionTargetPageId="6" captionText="FIGURE 4. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, lateral interradius of the disc; B, a madreporite; C, aboral view of basal portion of the arm; D, oral view of basal portion of the arm; E, lateral view of middle portion of the arm; F, oral view of distal portion of the arm; G, aboral view of distal portion of the arm. Abbreviations: AS, arm spine; GS, genital slit; M, madreporite. Arrows indicate transverse rows of hooklet-bearing plates (C, E, G)." httpUri="https://zenodo.org/record/206583/files/figure.png" pageId="6" pageNumber="31">Fig. 4</figureCitation>
D, F). Arm spines in the basal one-fourth of the arm are ovoid, carrying two or three terminal projections. The inner most arm spine has three and the others have two terminal projections. In the middle portion, from one-fourth to three-fourths of the arm length, the arm spines have two terminal projections. In the distal quarter of the arm, arm spines are transformed into hooks. Those hook-shaped spines have one inner tooth, and they can be distinguished from hooklets on aboral and lateral surface of the arm by the reticular structure on their basal portion.
</paragraph>
<paragraph id="8B86A112FFC5FFA5FF4B4F33A83BADC0" blockId="6.[151,1436,915,1969]" pageId="6" pageNumber="31">
<emphasis id="B94D7D00FFC5FFA5FF4B4F33AB29AD6F" box="[197,267,1798,1822]" italics="true" pageId="6" pageNumber="31">Color.</emphasis>
Aboral side of the body is basically gray. Distal half of each radial shield are white and the disc center is dark brown (
<figureCitation id="1302BD97FFC5FFA5FEA94F1EAB49AD32" box="[295,363,1835,1859]" captionStart="FIGURE 2" captionStartId="4.[151,250,1928,1951]" captionTargetBox="[154,1430,193,1905]" captionTargetId="figure@4.[151,1436,119,1907]" captionTargetPageId="4" captionText="FIGURE 2. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral side; B, oral side." httpUri="https://zenodo.org/record/206581/files/figure.png" pageId="6" pageNumber="31">Fig. 2</figureCitation>
A). Between the proximal part of paired radial shields, one yellow spot is present (
<figureCitation id="1302BD97FFC5FFA5FA824F1EAF6DAD32" box="[1292,1359,1835,1859]" captionStart="FIGURE 2" captionStartId="4.[151,250,1928,1951]" captionTargetBox="[154,1430,193,1905]" captionTargetId="figure@4.[151,1436,119,1907]" captionTargetPageId="4" captionText="FIGURE 2. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral side; B, oral side." httpUri="https://zenodo.org/record/206581/files/figure.png" pageId="6" pageNumber="31">Fig. 2</figureCitation>
A). On basal aboral median line of three arms, there are two yellow spots. Lateral interradial disc is also basically gray. In the central part of each lateral oral interradial disc, one larger yellow spot is present (
<figureCitation id="1302BD97FFC5FFA5FBDD4F41AEBAADFD" box="[1107,1176,1908,1932]" captionStart="FIGURE 2" captionStartId="4.[151,250,1928,1951]" captionTargetBox="[154,1430,193,1905]" captionTargetId="figure@4.[151,1436,119,1907]" captionTargetPageId="4" captionText="FIGURE 2. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral side; B, oral side." httpUri="https://zenodo.org/record/206581/files/figure.png" pageId="6" pageNumber="31">Fig. 2</figureCitation>
B). The oral side of the body is uniformly white (
<figureCitation id="1302BD97FFC5FFA5FE3A4FACABDAADC0" box="[436,504,1945,1969]" captionStart="FIGURE 2" captionStartId="4.[151,250,1928,1951]" captionTargetBox="[154,1430,193,1905]" captionTargetId="figure@4.[151,1436,119,1907]" captionTargetPageId="4" captionText="FIGURE 2. Asteroporpa (Asteromoana) muricatopatella sp. nov., holotype (NSMT E- 6548). A, aboral side; B, oral side." httpUri="https://zenodo.org/record/206581/files/figure.png" pageId="6" pageNumber="31">Fig. 2</figureCitation>
B).
</paragraph>
</subSubSection>
<caption id="DF46F19AFFC4FFA4FF194CC4ABD9AFCE" httpUri="https://zenodo.org/record/206584/files/figure.png" pageId="7" pageNumber="32" targetBox="[154,1436,193,1234]" targetPageId="7">
<paragraph id="8B86A112FFC4FFA4FF194CC4ABD9AFCE" blockId="7.[151,1436,1265,1471]" pageId="7" pageNumber="32">
<emphasis id="B94D7D00FFC4FFA4FF194CC4AB34AF76" bold="true" box="[151,278,1265,1288]" pageId="7" pageNumber="32">FIGURE 5.</emphasis>
<taxonomicName id="4C39DA91FFC4FFA4FEAF4CC7A8CEAF76" box="[289,748,1265,1287]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Asteroporpa" kingdom="Animalia" order="Euryalida" pageId="7" pageNumber="32" phylum="Echinodermata" rank="species" species="muricatopatella" status="sp. nov." subGenus="Asteromoana">
<emphasis id="B94D7D00FFC4FFA4FEAF4CC7ABBDAF76" box="[289,415,1266,1287]" italics="true" pageId="7" pageNumber="32">Asteroporpa</emphasis>
(
<emphasis id="B94D7D00FFC4FFA4FE3F4CC7A815AF76" box="[433,567,1266,1287]" italics="true" pageId="7" pageNumber="32">Asteromoana</emphasis>
)
<emphasis id="B94D7D00FFC4FFA4FDC44CC4A8CEAF76" box="[586,748,1265,1287]" italics="true" pageId="7" pageNumber="32">muricatopatella</emphasis>
</taxonomicName>
<emphasis id="B94D7D00FFC4FFA4FD784CC7A968AF76" bold="true" box="[758,842,1266,1287]" pageId="7" pageNumber="32">
<taxonomicNameLabel id="A27EC07BFFC4FFA4FD784CC7A968AF76" box="[758,842,1266,1287]" pageId="7" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, one paratype (NSMT E-5619-B). SEM photographs of internal ossicles. AB, a hooklet-bearing plate (A) and a hooklet (B) at the aboral periphery of the disc; CD, cone-shaped dermal ossicles at the aboral periphery of the disc, external (C) and lateral (D) views; E, a hooklet at the middle portion of the arm; FG, plate-shaped dermal ossicles at the aboral basal portion of the arm, external (F) and lateral (G) views; HI, plate-shaped dermal ossicles at the aboral distal portion of the arm, external (H) and lateral (I) views; JK, arm spines from basal portion of the arm (J) and distal portion of the arm (K) with an arc indicating reticular structure. Arrows indicate the orientation (D, G, I): ext, external side; int, internal side.
</paragraph>
</caption>
<subSubSection id="C323F299FFC4FFABFF4B4DDCA96EAA88" lastPageId="8" lastPageNumber="33" pageId="7" pageNumber="32" type="description">
<paragraph id="8B86A112FFC4FFA4FF4B4DDCAF7AAC70" blockId="7.[151,1437,1513,1979]" box="[197,1368,1513,1538]" pageId="7" pageNumber="32">
<emphasis id="B94D7D00FFC4FFA4FF4B4DDCA856AC73" bold="true" box="[197,628,1513,1538]" pageId="7" pageNumber="32">
Ossicle morphology of the
<typeStatus id="54821FB0FFC4FFA4FD8C4DDFA84DAC73" box="[514,623,1514,1538]" pageId="7" pageNumber="32" type="paratype">paratype</typeStatus>
.
</emphasis>
E-5619-B: Size is
<quantity id="4CC10CF7FFC4FFA4FCC44DDCA9BDAC70" box="[842,927,1513,1537]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="7" pageNumber="32" unit="mm" value="9.1">9.1 mm</quantity>
in disc diameter,
<quantity id="4CC10CF7FFC4FFA4FBE84DDCAE97AC70" box="[1126,1205,1513,1537]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="32" unit="mm" value="45.0">45 mm</quantity>
in arm length.
</paragraph>
<paragraph id="8B86A112FFC4FFA4FF4B4E3BAB49AD3C" blockId="7.[151,1437,1513,1979]" pageId="7" pageNumber="32">
Hooklet-bearing plates on radial shield possess eight to eleven tubercle-shaped articulations for hooklets (
<figureCitation id="1302BD97FFC4FFA4FAE14E3BAA87AC3A" captionStart="FIGURE 5" captionStartId="7.[151,250,1265,1288]" captionTargetBox="[154,1436,193,1234]" captionTargetId="figure@7.[154,1439,131,1242]" captionTargetPageId="7" captionText="FIGURE 5. Asteroporpa (Asteromoana) muricatopatella sp. nov., one paratype (NSMT E- 5619 - B). SEM photographs of internal ossicles. A B, a hooklet-bearing plate (A) and a hooklet (B) at the aboral periphery of the disc; C D, cone-shaped dermal ossicles at the aboral periphery of the disc, external (C) and lateral (D) views; E, a hooklet at the middle portion of the arm; F G, plate-shaped dermal ossicles at the aboral basal portion of the arm, external (F) and lateral (G) views; H I, plate-shaped dermal ossicles at the aboral distal portion of the arm, external (H) and lateral (I) views; J K, arm spines from basal portion of the arm (J) and distal portion of the arm (K) with an arc indicating reticular structure. Arrows indicate the orientation (D, G, I): ext, external side; int, internal side." httpUri="https://zenodo.org/record/206584/files/figure.png" pageId="7" pageNumber="32">Fig. 5</figureCitation>
A). The tubercle-shaped articulations form two or three parallel rows (
<figureCitation id="1302BD97FFC4FFA4FC394E06A9D8AC3A" box="[951,1018,1587,1611]" captionStart="FIGURE 5" captionStartId="7.[151,250,1265,1288]" captionTargetBox="[154,1436,193,1234]" captionTargetId="figure@7.[154,1439,131,1242]" captionTargetPageId="7" captionText="FIGURE 5. Asteroporpa (Asteromoana) muricatopatella sp. nov., one paratype (NSMT E- 5619 - B). SEM photographs of internal ossicles. A B, a hooklet-bearing plate (A) and a hooklet (B) at the aboral periphery of the disc; C D, cone-shaped dermal ossicles at the aboral periphery of the disc, external (C) and lateral (D) views; E, a hooklet at the middle portion of the arm; F G, plate-shaped dermal ossicles at the aboral basal portion of the arm, external (F) and lateral (G) views; H I, plate-shaped dermal ossicles at the aboral distal portion of the arm, external (H) and lateral (I) views; J K, arm spines from basal portion of the arm (J) and distal portion of the arm (K) with an arc indicating reticular structure. Arrows indicate the orientation (D, G, I): ext, external side; int, internal side." httpUri="https://zenodo.org/record/206584/files/figure.png" pageId="7" pageNumber="32">Fig. 5</figureCitation>
A). The hooklets bear two inner teeth (
<figureCitation id="1302BD97FFC4FFA4FF114E6DAAC1AC01" box="[159,227,1624,1648]" captionStart="FIGURE 5" captionStartId="7.[151,250,1265,1288]" captionTargetBox="[154,1436,193,1234]" captionTargetId="figure@7.[154,1439,131,1242]" captionTargetPageId="7" captionText="FIGURE 5. Asteroporpa (Asteromoana) muricatopatella sp. nov., one paratype (NSMT E- 5619 - B). SEM photographs of internal ossicles. A B, a hooklet-bearing plate (A) and a hooklet (B) at the aboral periphery of the disc; C D, cone-shaped dermal ossicles at the aboral periphery of the disc, external (C) and lateral (D) views; E, a hooklet at the middle portion of the arm; F G, plate-shaped dermal ossicles at the aboral basal portion of the arm, external (F) and lateral (G) views; H I, plate-shaped dermal ossicles at the aboral distal portion of the arm, external (H) and lateral (I) views; J K, arm spines from basal portion of the arm (J) and distal portion of the arm (K) with an arc indicating reticular structure. Arrows indicate the orientation (D, G, I): ext, external side; int, internal side." httpUri="https://zenodo.org/record/206584/files/figure.png" pageId="7" pageNumber="32">Fig. 5</figureCitation>
B). The hooklets on middle portion of the arm also bear two inner teeth (
<figureCitation id="1302BD97FFC4FFA4FB814E6DAE73AC01" box="[1039,1105,1624,1648]" captionStart="FIGURE 5" captionStartId="7.[151,250,1265,1288]" captionTargetBox="[154,1436,193,1234]" captionTargetId="figure@7.[154,1439,131,1242]" captionTargetPageId="7" captionText="FIGURE 5. Asteroporpa (Asteromoana) muricatopatella sp. nov., one paratype (NSMT E- 5619 - B). SEM photographs of internal ossicles. A B, a hooklet-bearing plate (A) and a hooklet (B) at the aboral periphery of the disc; C D, cone-shaped dermal ossicles at the aboral periphery of the disc, external (C) and lateral (D) views; E, a hooklet at the middle portion of the arm; F G, plate-shaped dermal ossicles at the aboral basal portion of the arm, external (F) and lateral (G) views; H I, plate-shaped dermal ossicles at the aboral distal portion of the arm, external (H) and lateral (I) views; J K, arm spines from basal portion of the arm (J) and distal portion of the arm (K) with an arc indicating reticular structure. Arrows indicate the orientation (D, G, I): ext, external side; int, internal side." httpUri="https://zenodo.org/record/206584/files/figure.png" pageId="7" pageNumber="32">Fig. 5</figureCitation>
E). Dermal ossicles on aboral periphery of the disc are cone-shaped, approximately 180 µm in length and 200 µm in height, with two or three terminal projections (
<figureCitation id="1302BD97FFC4FFA4FEE24E94AB8DACC8" box="[364,431,1697,1721]" captionStart="FIGURE 5" captionStartId="7.[151,250,1265,1288]" captionTargetBox="[154,1436,193,1234]" captionTargetId="figure@7.[154,1439,131,1242]" captionTargetPageId="7" captionText="FIGURE 5. Asteroporpa (Asteromoana) muricatopatella sp. nov., one paratype (NSMT E- 5619 - B). SEM photographs of internal ossicles. A B, a hooklet-bearing plate (A) and a hooklet (B) at the aboral periphery of the disc; C D, cone-shaped dermal ossicles at the aboral periphery of the disc, external (C) and lateral (D) views; E, a hooklet at the middle portion of the arm; F G, plate-shaped dermal ossicles at the aboral basal portion of the arm, external (F) and lateral (G) views; H I, plate-shaped dermal ossicles at the aboral distal portion of the arm, external (H) and lateral (I) views; J K, arm spines from basal portion of the arm (J) and distal portion of the arm (K) with an arc indicating reticular structure. Arrows indicate the orientation (D, G, I): ext, external side; int, internal side." httpUri="https://zenodo.org/record/206584/files/figure.png" pageId="7" pageNumber="32">Fig. 5</figureCitation>
CD). The length of the terminal projections are approximately one-fourth the height of the cone-shaped dermal ossicles (
<figureCitation id="1302BD97FFC4FFA4FD9B4EF3A87AACAF" box="[533,600,1734,1758]" captionStart="FIGURE 5" captionStartId="7.[151,250,1265,1288]" captionTargetBox="[154,1436,193,1234]" captionTargetId="figure@7.[154,1439,131,1242]" captionTargetPageId="7" captionText="FIGURE 5. Asteroporpa (Asteromoana) muricatopatella sp. nov., one paratype (NSMT E- 5619 - B). SEM photographs of internal ossicles. A B, a hooklet-bearing plate (A) and a hooklet (B) at the aboral periphery of the disc; C D, cone-shaped dermal ossicles at the aboral periphery of the disc, external (C) and lateral (D) views; E, a hooklet at the middle portion of the arm; F G, plate-shaped dermal ossicles at the aboral basal portion of the arm, external (F) and lateral (G) views; H I, plate-shaped dermal ossicles at the aboral distal portion of the arm, external (H) and lateral (I) views; J K, arm spines from basal portion of the arm (J) and distal portion of the arm (K) with an arc indicating reticular structure. Arrows indicate the orientation (D, G, I): ext, external side; int, internal side." httpUri="https://zenodo.org/record/206584/files/figure.png" pageId="7" pageNumber="32">Fig. 5</figureCitation>
CD). Dermal ossicles on the aboral basal portion of the arm are domed, round and plate-shaped, approximately 220 µm in length and 200 µm in height (
<figureCitation id="1302BD97FFC4FFA4FBBD4EDEAE55AD72" box="[1075,1143,1771,1795]" captionStart="FIGURE 5" captionStartId="7.[151,250,1265,1288]" captionTargetBox="[154,1436,193,1234]" captionTargetId="figure@7.[154,1439,131,1242]" captionTargetPageId="7" captionText="FIGURE 5. Asteroporpa (Asteromoana) muricatopatella sp. nov., one paratype (NSMT E- 5619 - B). SEM photographs of internal ossicles. A B, a hooklet-bearing plate (A) and a hooklet (B) at the aboral periphery of the disc; C D, cone-shaped dermal ossicles at the aboral periphery of the disc, external (C) and lateral (D) views; E, a hooklet at the middle portion of the arm; F G, plate-shaped dermal ossicles at the aboral basal portion of the arm, external (F) and lateral (G) views; H I, plate-shaped dermal ossicles at the aboral distal portion of the arm, external (H) and lateral (I) views; J K, arm spines from basal portion of the arm (J) and distal portion of the arm (K) with an arc indicating reticular structure. Arrows indicate the orientation (D, G, I): ext, external side; int, internal side." httpUri="https://zenodo.org/record/206584/files/figure.png" pageId="7" pageNumber="32">Fig. 5</figureCitation>
FG). Dermal ossicles on aboral distal portion of the arm are flat, round and granule-shaped, approximately 70 µm in length and 30 µm in height (
<figureCitation id="1302BD97FFC4FFA4FF624F00AB0FAD3C" box="[236,301,1845,1869]" captionStart="FIGURE 5" captionStartId="7.[151,250,1265,1288]" captionTargetBox="[154,1436,193,1234]" captionTargetId="figure@7.[154,1439,131,1242]" captionTargetPageId="7" captionText="FIGURE 5. Asteroporpa (Asteromoana) muricatopatella sp. nov., one paratype (NSMT E- 5619 - B). SEM photographs of internal ossicles. A B, a hooklet-bearing plate (A) and a hooklet (B) at the aboral periphery of the disc; C D, cone-shaped dermal ossicles at the aboral periphery of the disc, external (C) and lateral (D) views; E, a hooklet at the middle portion of the arm; F G, plate-shaped dermal ossicles at the aboral basal portion of the arm, external (F) and lateral (G) views; H I, plate-shaped dermal ossicles at the aboral distal portion of the arm, external (H) and lateral (I) views; J K, arm spines from basal portion of the arm (J) and distal portion of the arm (K) with an arc indicating reticular structure. Arrows indicate the orientation (D, G, I): ext, external side; int, internal side." httpUri="https://zenodo.org/record/206584/files/figure.png" pageId="7" pageNumber="32">Fig. 5</figureCitation>
HI).
</paragraph>
<paragraph id="8B86A112FFC4FFA4FF4B4F6CA99FADCA" blockId="7.[151,1437,1513,1979]" pageId="7" pageNumber="32">
The inner most arm spines on the basal one-fourth of the arm are ovoid, carrying two or three terminal projections on the tip (
<figureCitation id="1302BD97FFC4FFA4FEDB4F4BABBBADE7" box="[341,409,1918,1942]" captionStart="FIGURE 5" captionStartId="7.[151,250,1265,1288]" captionTargetBox="[154,1436,193,1234]" captionTargetId="figure@7.[154,1439,131,1242]" captionTargetPageId="7" captionText="FIGURE 5. Asteroporpa (Asteromoana) muricatopatella sp. nov., one paratype (NSMT E- 5619 - B). SEM photographs of internal ossicles. A B, a hooklet-bearing plate (A) and a hooklet (B) at the aboral periphery of the disc; C D, cone-shaped dermal ossicles at the aboral periphery of the disc, external (C) and lateral (D) views; E, a hooklet at the middle portion of the arm; F G, plate-shaped dermal ossicles at the aboral basal portion of the arm, external (F) and lateral (G) views; H I, plate-shaped dermal ossicles at the aboral distal portion of the arm, external (H) and lateral (I) views; J K, arm spines from basal portion of the arm (J) and distal portion of the arm (K) with an arc indicating reticular structure. Arrows indicate the orientation (D, G, I): ext, external side; int, internal side." httpUri="https://zenodo.org/record/206584/files/figure.png" pageId="7" pageNumber="32">Fig. 5</figureCitation>
J) Inner most arm spines on the distal quarter of the arm are hook-shaped, with one inner teeth along inner edge and reticular structure on basal portion (
<figureCitation id="1302BD97FFC4FFA4FCD84F96A9BAADCA" box="[854,920,1955,1979]" captionStart="FIGURE 5" captionStartId="7.[151,250,1265,1288]" captionTargetBox="[154,1436,193,1234]" captionTargetId="figure@7.[154,1439,131,1242]" captionTargetPageId="7" captionText="FIGURE 5. Asteroporpa (Asteromoana) muricatopatella sp. nov., one paratype (NSMT E- 5619 - B). SEM photographs of internal ossicles. A B, a hooklet-bearing plate (A) and a hooklet (B) at the aboral periphery of the disc; C D, cone-shaped dermal ossicles at the aboral periphery of the disc, external (C) and lateral (D) views; E, a hooklet at the middle portion of the arm; F G, plate-shaped dermal ossicles at the aboral basal portion of the arm, external (F) and lateral (G) views; H I, plate-shaped dermal ossicles at the aboral distal portion of the arm, external (H) and lateral (I) views; J K, arm spines from basal portion of the arm (J) and distal portion of the arm (K) with an arc indicating reticular structure. Arrows indicate the orientation (D, G, I): ext, external side; int, internal side." httpUri="https://zenodo.org/record/206584/files/figure.png" pageId="7" pageNumber="32">Fig. 5</figureCitation>
K).
</paragraph>
<paragraph id="8B86A112FFCBFFABFF4B48A2A96EAA88" blockId="8.[151,1436,151,433]" pageId="8" pageNumber="33">
<emphasis id="B94D7D00FFCBFFABFF4B48A2AB1BAAC1" bold="true" box="[197,313,151,176]" pageId="8" pageNumber="33">Variation.</emphasis>
Some color variations can be observed in four specimens. The periphery of the aboral disc and aboral surface of the arms are gray for the
<typeStatus id="54821FB0FFCBFFABFDCB4889A888AAA5" box="[581,682,188,212]" pageId="8" pageNumber="33" type="holotype">holotype</typeStatus>
, dark red for one
<typeStatus id="54821FB0FFCBFFABFCE24888A9EEAAA4" box="[876,972,189,213]" pageId="8" pageNumber="33" type="paratype">paratype</typeStatus>
(NSMT E-5619-B), and the entire bodies of the other two
<typeStatus id="54821FB0FFCBFFABFEDE48D7AB99AA8B" box="[336,443,226,250]" pageId="8" pageNumber="33" type="paratype">paratypes</typeStatus>
(NSMT E-6547) are creamy white.
</paragraph>
</subSubSection>
<subSubSection id="C323F299FFCBFFABFF4B4930A805AB32" pageId="8" pageNumber="33" type="distribution">
<paragraph id="8B86A112FFCBFFABFF4B4930A805AB32" blockId="8.[151,1436,151,433]" pageId="8" pageNumber="33">
<emphasis id="B94D7D00FFCBFFABFF4B4930AB75AB6F" bold="true" box="[197,343,261,286]" pageId="8" pageNumber="33">Distribution</emphasis>
.
<emphasis id="B94D7D00FFCBFFABFEE94930ABE3AB6F" bold="true" box="[359,449,261,286]" pageId="8" pageNumber="33">
<collectingCountry id="F32EE182FFCBFFABFEE94930ABE3AB6F" box="[359,449,261,286]" name="Japan" pageId="8" pageNumber="33">JAPAN</collectingCountry>
</emphasis>
: off Yaku-shima Island,
<quantity id="4CC10CF7FFCBFFABFD504933A905AB6F" box="[734,807,262,286]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="33" unit="m" value="140.0">140 m</quantity>
(present study), off Kume-jima Island,
<quantity id="4CC10CF7FFCBFFABFB7E4933AF1BAB6F" box="[1264,1337,262,286]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="8" pageNumber="33" unit="m" value="410.0">410 m</quantity>
(present study), southwestern
<collectingCountry id="F32EE182FFCBFFABFE0B4919ABE7AB32" box="[389,453,300,323]" name="Japan" pageId="8" pageNumber="33">Japan</collectingCountry>
(
<figureCitation id="1302BD97FFCBFFABFE5A491EA834AB32" box="[468,534,299,323]" captionStart="FIGURE 1" captionStartId="1.[151,250,1801,1824]" captionTargetBox="[174,1418,327,1773]" captionTargetId="figure@1.[166,1422,321,1782]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of Asteroporpa (Asteromoana) muricatopatella sp. nov. and A. (Asterom.) koyoae sp. nov." httpUri="https://zenodo.org/record/206580/files/figure.png" pageId="8" pageNumber="33">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C323F299FFCBFFABFF4B4965A881ABC0" pageId="8" pageNumber="33" type="etymology">
<paragraph id="8B86A112FFCBFFABFF4B4965A881ABC0" blockId="8.[151,1436,151,433]" pageId="8" pageNumber="33">
<emphasis id="B94D7D00FFCBFFABFF4B4965AB66AB19" bold="true" box="[197,324,336,360]" pageId="8" pageNumber="33">Etymology</emphasis>
. The specific name is a feminine noun in apposition formed as a compound of Latin words,
<emphasis id="B94D7D00FFCBFFABFACD4964AA9AABFC" italics="true" pageId="8" pageNumber="33">muricatus</emphasis>
(adjective, meaning “covered by spines”) and
<emphasis id="B94D7D00FFCBFFABFD594940A907ABFC" box="[727,805,373,397]" italics="true" pageId="8" pageNumber="33">patella</emphasis>
(feminine noun, meaning “disc”), referring to the disc covered by spiny cone-shaped dermal ossicles.
</paragraph>
</subSubSection>
</treatment>
</document>