141 lines
19 KiB
XML
141 lines
19 KiB
XML
<document id="84EED8A25FACE5E810DC5B43963CAE6C" ENCODING="UTF8" ID-GBIF-Dataset="0ad0b492-3b2e-433f-9d58-3b4e05fb5d63" ModsDocID="22171" checkinTime="1249509882759" checkinUser="christiana" docAuthor="Eguchi, K." docDate="2008" docId="9F741699A0F83258BE9465628C951534" docLanguage="en" docName="22171.xml" docOrigin="Zootaxa 1902" docSource="http://hol.osu.edu/reference-full.html?id=22171" docTitle="Pheidole tjibodana Forel" docType="treatment" docVersion="10" lastPageNumber="96" masterDocId="B03BC2DB73BB189B7C160C6A61E2E599" masterDocTitle="A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae)." masterLastPageNumber="118" masterPageNumber="1" pageNumber="93" updateTime="1701311764087" updateUser="plazi">
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<mods:title id="646FFAA4A7F2F22DEFFDBD8FB425AE5B">A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae).</mods:title>
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<mods:namePart id="F7F057E2D246191C13F582921B4943DF">Eguchi, K.</mods:namePart>
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<mods:number id="333318E26236F96F3D656F9518E29ADF">1902</mods:number>
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<mods:identifier id="DEB96892861B31BC3E7DFB1E8E7A06DC" type="HNS-Pub">22171</mods:identifier>
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<treatment id="9F741699A0F83258BE9465628C951534" ID-DOI="http://doi.org/10.5281/zenodo.6235670" ID-GBIF-Taxon="100122336" ID-Zenodo-Dep="6235670" LSID="urn:lsid:plazi:treatment:9F741699A0F83258BE9465628C951534" httpUri="http://treatment.plazi.org/id/9F741699A0F83258BE9465628C951534" lastPageNumber="96" pageNumber="93">
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<subSubSection id="D2F58E1FF600F0C9FD6DD6AFE1022018" type="nomenclature">
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<paragraph id="BEE0532949597A3AC5A55D902B8055FC" pageNumber="93">
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<taxonomicName id="747D49C7A290DEB6FC6EB2850E07D9C7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:175121" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole tjibodana Forel" order="Hymenoptera" pageNumber="93" phylum="Arthropoda" rank="species" species="tjibodana">Pheidole tjibodana Forel</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="0FE1CB5CF397BF58CAD4DB1EFA52F460" type="description">
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<paragraph id="690B6121EDD00703C8826F38312C09C2" pageNumber="93">Figs. 26a-g</paragraph>
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</subSubSection>
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<subSubSection id="839571626AE68D1AA39EF3787C18428B" type="materials_examined">
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<paragraph id="89F7F47006810721B2B2FDEFDE50EEF7" pageNumber="93">
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<taxonomicName id="615D8642C5C8EE561DB319FEBA0DE858" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151448" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole nodgii tjibodana Forel" order="Hymenoptera" pageNumber="93" phylum="Arthropoda" rank="variety" species="nodgii" variety="tjibodana">Pheidole nodgii var. tjibodana Forel</taxonomicName>
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, 1905c: 16. Lectotype: major, "Tjibodas, Java K. Kraepelin leg. 25-28/ III/1904" [Cibodas, Java, Indonesia],
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<collectionCode id="231569D3E9BE3E428B16A065272F75F8" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
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, examined; paralectotypes: 1 minor & 1 queen, same data as lectotype,
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<collectionCode id="088F5327496931357DF5F88C78337A78" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
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, examined.
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</paragraph>
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</subSubSection>
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<subSubSection id="96EE0D12D0D95FC2D254F7F92A7AEEA1" type="reference_group">
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<paragraph id="17344FA701F9E822046D5504BEEEEEA1" pageNumber="93">
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<taxonomicName id="A7FB5C6494368294301861B537EFDEC9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:175121" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole tjibodana Forel" order="Hymenoptera" pageNumber="93" phylum="Arthropoda" rank="species" species="tjibodana">Pheidole tjibodana Forel</taxonomicName>
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. Eguchi 2001a: 29-30 (raised to species, lectotype designation, redescription of major & minor).
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</paragraph>
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<paragraph id="75C3350E9004A7B0F54683DC399F6CB3" pageNumber="93">
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<taxonomicName id="E6DC823E83232DA8AE8D1F59097871E5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="93" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
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sp. eg-161. Yamane et al. 2003: 58.
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</paragraph>
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</subSubSection>
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<subSubSection id="C7D77EAAAACF0187A6CAE7A1EA3DFCE4" type="materials_examined">
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<paragraph id="C5977C1930B2E1573AD7CD67913F05B9" lastPageNumber="94" pageNumber="93">
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Other material examined: Vietnam: Bac Kan: Ba Be N.P., 22°24-25N', 105°37-38', <260 m alt. [Eg04-VN-194, -204, -207, -215]; Quang Ninh: Chua Yen Tu,
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<geoCoordinate id="C1EED713D7B23C4B1457FD88B1EF92EC" direction="north" orientation="latitude" value="21.15">21°09'N</geoCoordinate>
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,
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<geoCoordinate id="CFD06E3EB5939DCCEA90A627DCCC53F9" direction="east" orientation="longitude" value="106.71667">106°43'E</geoCoordinate>
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, 720-845 m alt. [Eg04- VN-024]; Ky Thuong
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<taxonomicNameLabel id="E41DF275D66D4889765F76F42EA4F62D" rank="subSpecies">N.R.</taxonomicNameLabel>
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, 21°09-11'N, 107°06-07'E, 160-570 m alt. [Eg03-VN-175, -215]; Ninh Binh: Cuc Phuong N.P.,
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<geoCoordinate id="87FEA48FE6F1E670863D11B88D21B8FE" direction="north" orientation="latitude" value="20.233334">20°14'N</geoCoordinate>
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,
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<geoCoordinate id="19D055DA428872A05F5CE0016DB406DA" direction="east" orientation="longitude" value="105.6">105°36'E</geoCoordinate>
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[Eg14vi05-05]; Dong Nai: S. Cat Tien N.P., <160 m alt. [Eg04-VN- 584, -644, -651]. Thailand: Nakhonratchasima: Khao Yai N.P., 900-1000 m alt. [TH00-SKY-15]; Chacheongsao
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<pageStartToken id="69ADC0510246C251D8CE031663CD1765" pageNumber="94">:</pageStartToken>
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Lumchangwat Station, Khao Ang Rew Nai W.S. [TH03-SKY-86]; Trang: Khao Chong Waterfall [Eg01- TH-713]. W. Malaysia: Johor: Kota Tingii Fall, 100 m alt. [P.S. Ward #9586-15]. E. Malaysia: Sabah: Tawau Hills Park [Eg96-BOR-031, -039], Guong Rara [Eg97-BOR-565], Poring, 600 m alt. [T. Kikuta's colony: 6XI0106-16-1], Sayap Kinabalu [K. Eguchi's soil sample: S-27]. Indonesia: W. Java: G. Sarak, nr. Bogor [JA97-SKY-12], Halimun [FI98-382], Cibodas [K. Ohkawara's colony: 10/1a (1999)].
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</paragraph>
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</subSubSection>
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<subSubSection id="BE818B77E9AB51AADDCF530EF36FE029" type="description">
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<paragraph id="90B492F84E3C39904413A226650DD8C2" pageNumber="94">Worker measurements & indices: Major (n=5). - HL 0.97-1.14 mm, HW 0.95-1.05 mm, CI 89-100, SL 0.49-0.53 mm, SI 48-52, FL 0.54-0.63 mm, FI 56-61.</paragraph>
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<paragraph id="877043F056F573672336B94697C25775" pageNumber="94">Minor (n=5). - HL 0.51-0.55 mm, HW 0.49-0.55 mm, CI 92-100, SL 0.42-0.47 mm, SI 84-96, FL 0.45-0.51 mm, FI 90-100.</paragraph>
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<paragraph id="856E25EF3082CFC71A37C2BD0EE08B44" pageNumber="94">Worker description</paragraph>
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<paragraph id="D263E4A8135B45273009182CB7CA586C" pageNumber="94">Major. - Head in lateral view not or hardly impressed on vertex, sometimes highly raised on the posterior part of frons; anterior part of frons longitudinally rugose; posterior part of frons, vertex and dorsal and dorsolateral faces of vertexal lobe reticulate; frontal carina well developed horizontally, partly overhanging antennal scrobe; median longitudinal carina of clypeus absent, or sometimes present but very weak; hypostoma with a very large median process in addition to well-developed lateral processes; submedian processes nearly or completely absent; outer surface of mandible (excluding area around the base) smooth, with sparse and very short appressed hairs; antenna with a 3-segmented club; maximal diameter of eye longer or much longer than antennal segment X. Promesonotal dome in dorsal view reticulate, in lateral view at most with an inconspicuous or low mound/process on its posterior slope; humerus moderately produced laterad; the dome at the humeri as broad as or broader than at the bottom; propodeal spine usually well developed as a long horn. Petiole much longer than postpetiole (excluding helcium); postpetiole not massive. First gastral tergite smooth to shagreened largely, or weakly punctured anteriorly and shagreened posteriorly.</paragraph>
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<paragraph id="ADF76F6138A87B763C3CA8AD25289C62" pageNumber="94">Minor. - Dorsum of head well punctured and overlain by weak rugoso-reticulation, or well reticulate with enclosure weakly or dimly punctured; preoccipital carina inconspicuous or very weak dorsally; median part of clypeus rugoso-reticulate, or punctured and overlain by very weak rugoso-reticulation; median longitudinal carina absent, or present as rugula(e); antenna with a 3-segmented club; scape usually exceeding posterior margin of head by less than the length of antennal segment II, but sometimes not exceeding posterior margin; maximal diameter of eye almost as long as or a little longer than antennal segment X (but rarely a little shorther than antennal segment X). Dorsum of promesonotal dome well punctured and overlain by weak rugoso-reticulation, or well reticulate with enclosure weakly or dimly punctured; lateral face of premesonotal dome, mesopleuron, metapleuron and lateral face of propodeum well punctured; premesonotal dome in lateral view lacking a conspicuous mound on its posterior slope; humerus of the dome in dorso-oblique view produced weakly or forming a small tubercle; propodeal spine variable in size and shape. Petiole much longer than postpetiole (excluding helcium); postpetiole not massive.</paragraph>
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</subSubSection>
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<subSubSection id="D95903071B00B9B204B46C243C6F9E38" type="discussion">
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<paragraph id="E46234069FF89A697D7598CE92BCB084" pageNumber="94">
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Recognition: Based on his examination of the type material Eguchi (2001a) distinguished
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<taxonomicName id="4B4A4CC7078B2C36D78C8FFCF60249AA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:175121" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole tjibodana Forel" order="Hymenoptera" pageNumber="94" phylum="Arthropoda" rank="species" species="tjibodana">Pheidole tjibodana</taxonomicName>
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from
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<taxonomicName id="B1F795EBBE5163294BFDB708DF8A17AD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33899" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="3096" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole nodgii Forel" order="Hymenoptera" pageNumber="94" phylum="Arthropoda" rank="species" species="nodgii">P. nodgii Forel</taxonomicName>
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(lectotype (major) & paralectotype minor, "Depok, Java K. Kraepelin leg. 9. III. 1904" [Jakarta, Java, Indonesia],
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<collectionCode id="C9DC5D396F89002DCFD1B179CCD1D463" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
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; 1 paralectotype queen, "Buitenzorg, Java K. Kraepelin leg. 24. II. - 12. III. 1904" [Bogor, Java],
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<collectionCode id="AF6CD03CF400535D1E33B15EBBABDF27" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
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) based on the following differences: posterior part of frons of the major higher in
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<taxonomicName id="CC21FF98FABF7E4AC5A2D2DC46C6E0FD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:175121" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="202" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole tjibodana Forel" order="Hymenoptera" pageNumber="94" phylum="Arthropoda" rank="species" species="tjibodana">P. tjibodana</taxonomicName>
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than in
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<taxonomicName id="8A1CA010889B1B3C2535B7F41F7E0029" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33899" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="3366" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole nodgii Forel" order="Hymenoptera" pageNumber="94" phylum="Arthropoda" rank="species" species="nodgii">P. nodgii</taxonomicName>
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; posterior slope of promesonotal dome of the minor steeper in
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<taxonomicName id="664F5E48AB8B6E6186F8D972E1FF6838" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:175121" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="183" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole tjibodana Forel" order="Hymenoptera" pageNumber="94" phylum="Arthropoda" rank="species" species="tjibodana">P. tjibodana</taxonomicName>
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than in
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<taxonomicName id="2E45DA2FFEDC85F3C457421E80C52B83" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33899" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="3423" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole nodgii Forel" order="Hymenoptera" pageNumber="94" phylum="Arthropoda" rank="species" species="nodgii">P. nodgii</taxonomicName>
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. Intermediate conditions in each of the two "diagnostic characters" were, however, observed in the present examination of nontype material. On the other hand, postpetiole of the minor of
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<taxonomicName id="76799F28DD37EE5221A2A918C467C320" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33899" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="3537" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole nodgii Forel" order="Hymenoptera" pageNumber="94" phylum="Arthropoda" rank="species" species="nodgii">P. nodgii</taxonomicName>
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were always highly raised and somewhat globular in lateral view. I at present treat them as different species, although only two colonies (FI96-253 & FI97-551 from Bogor, W. Java) referable to
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<taxonomicName id="9641ADE05D77DF104DA020689CBDB24A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33899" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="3675" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole nodgii Forel" order="Hymenoptera" pageNumber="94" phylum="Arthropoda" rank="species" species="nodgii">P. nodgii</taxonomicName>
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have been found.
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</paragraph>
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<paragraph id="B64D6CDB775805F2B23ECCCA4052123C" lastPageNumber="96" pageNumber="94">
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This species is well characterized among Indo-Chinese species by the combination of the following features: in the major frontal carina well developed horizontally, partly overhanging antennal scrobe; in the major
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<pageStartToken id="BEBD14B2ECF854E983445BA157360B80" pageNumber="96">median</pageStartToken>
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process well developed but submedian processes absent; in the minor dorsum of head and mesosoma punctured or reticulate with enclosures punctured weakly or dimly; in the major and minor promesonotal dome in lateral view at most with a low or inconspicuous mound on its posterior slope.
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</paragraph>
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</subSubSection>
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<subSubSection id="31C8ED0DD62877F638312C2AAE1681B3" type="multiple">
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<caption id="3FF7EF44EA9BD80C5E1A60E02EB376BC">
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<paragraph id="A2F6A3C2E7DD4FDEEEA64E0D1AA5EA78" pageNumber="95">
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FIGURE 26a-d,
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<taxonomicName id="0BF64F11B028F7A5991BFA97C57B3BD9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:175121" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole tjibodana Forel" order="Hymenoptera" pageNumber="95" phylum="Arthropoda" rank="species" species="tjibodana">Pheidole tjibodana</taxonomicName>
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, major [Eg03-VN-175] - a, head in full-face view; b, head in lateral view; c, mesosoma and waist in dorsal view; d, mesosoma and waist in lateral view.
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</paragraph>
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</caption>
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<caption id="32A8C3A0FF05ACDCAA82A7FF4B265077">
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<paragraph id="D1C56D3BD0CA3AE881DA9694FFBD079B" pageNumber="96">
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FIGURE 26e-g,
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<taxonomicName id="1E306B36D9E049C0BD627AD59592BDB8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:175121" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole tjibodana Forel" order="Hymenoptera" pageNumber="96" phylum="Arthropoda" rank="species" species="tjibodana">Pheidole tjibodana</taxonomicName>
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, minor [Eg03-VN-175] - e, head in full-face view; f, mesosoma and waist in dorsal view; g, mesosoma and waist in lateral view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="85ABCE50AAEB777C8DFB591136A1E55D" type="distribution">
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<paragraph id="C21A4F465EB941E2DD94BC300E4BFA92" lastPageNumber="97" pageNumber="96">
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Distribution & bionomics: Known from the Indo-Chinese and Indo-Malayan subregions. This species is
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<pageStartToken id="8E66131AEF0EA4E335B47CBB006A55B8" pageNumber="97">wide</pageStartToken>
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spread through the Indo-Chinese and Indo-Malayan subregions, and inhabits forest from lowland to hilly areas. Bornean populations usually nest in litter, but Vietnamese populations usually nest in rotting twigs and small wood fragments. Small seeds are sometimes stored inside the nest [Eg96-BOR-031, Eg04-VN-215]. The number of adults per colony is: 8 majors, 71 minors, 1 dealate queen & 3 alate queens [Eg04-VN-204]; 5 majors, 52 minors & 1 dealate queen (with queen pupae) [Eg04-VN-207]; 24 majors, 259 minors, 1 dealate queen & 31 alate queens [Eg04-VN-215].
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |