2177 lines
253 KiB
XML
2177 lines
253 KiB
XML
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<mods:title id="D35338999EB42745BAB789410C4BC295">The nomenclatural status of “ Anoplophallus maculatus ” Cope, 1895, its consequences on the systematics of Lycodon subcinctus auctorum, and the description of a new species (Squamata: Colubridae)</mods:title>
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<mods:namePart id="FA00B41F46E17B9E6DEDC07DC92528B8">Nguyen, Tan Van</mods:namePart>
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<mods:affiliation id="86C43A4D4F386D899E550F22D2AA6224">Institute for Research and Training in Medicine, Biology and Pharmacy, Duy Tan University, Da Nang, 550000, Vietnam & College of Medicine and Pharmacy, Duy Tan University, 120 Hoang Minh Thao, Lien Chieu, Da Nang, 550000, Vietnam</mods:affiliation>
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<mods:namePart id="120FC51556D5C9F39D1591DC8A16DDA0">Lee, Justin L.</mods:namePart>
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<mods:affiliation id="5B763DF39C1D0A6A4B32012470B957CE">Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109 USA</mods:affiliation>
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<mods:namePart id="E60CBD83E4B059EEDA86D786C61C8795">Pauwels, Olivier S. G.</mods:namePart>
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<mods:affiliation id="BD3B4B819A1527E0BE46D0EC59D61F58">Royal Belgian Institute of Natural Sciences, Rue Vautier 29, B- 1000 Brussels, Belgium</mods:affiliation>
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<mods:namePart id="28E06F7E4E37CA633FCB48547C83CB62">Kennedy-Gold, Stevie R.</mods:namePart>
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<mods:affiliation id="4157ECA7F6A261DE122EE9B378FA623E">Department of Herpetology, Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA</mods:affiliation>
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<mods:namePart id="F61649AC175C5FC276C506B0E732FAAB">Poyarkov, Nikolay A.</mods:namePart>
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<mods:affiliation id="45010C0806FBB883170AE7CF540D2E32">Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- 1, Moscow 119991, Russia & Joint Vietnam - Russia Tropical Science and Technology Research Centre, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="90CAF95A9F4991FCC1AF42A8FCEE1061">David, Patrick</mods:namePart>
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<treatment id="D76B87C72705321CB1C6FB14FA6CFBC1" ID-DOI="http://doi.org/10.5281/zenodo.13921457" ID-Zenodo-Dep="13921457" LSID="urn:lsid:plazi:treatment:D76B87C72705321CB1C6FB14FA6CFBC1" httpUri="http://treatment.plazi.org/id/D76B87C72705321CB1C6FB14FA6CFBC1" lastPageId="21" lastPageNumber="508" pageId="7" pageNumber="494">
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<heading id="043581BD2705320EB1C6FB14FDFFFB5D" bold="true" box="[151,532,1194,1221]" fontSize="11" level="1" pageId="7" pageNumber="494" reason="1">
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<taxonomicName id="98C24D522705320EB1C6FB14FE5BFB5D" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[151,432,1194,1221]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="494" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
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<emphasis id="6DB6EAC32705320EB1C6FB14FE5BFB5D" bold="true" box="[151,432,1194,1221]" italics="true" pageId="7" pageNumber="494">Lycodon neomaculatus</emphasis>
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<taxonomicNameLabel id="768557B82705320EB0E6FB15FDFFFB5D" box="[439,532,1195,1221]" pageId="7" pageNumber="494" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="17D8655A2705320EB1C6FB4CFDC7FA94" box="[151,556,1266,1293]" pageId="7" pageNumber="494" type="vernacular_names">
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<paragraph id="5F7D36D12705320EB1C6FB4CFDC7FA94" blockId="7.[151,931,1266,1329]" box="[151,556,1266,1293]" pageId="7" pageNumber="494">
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<heading id="043581BD2705320EB1C6FB4CFDC7FA94" bold="true" box="[151,556,1266,1293]" fontSize="11" level="1" pageId="7" pageNumber="494" reason="1">
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<emphasis id="6DB6EAC32705320EB1C6FB4CFDC7FA94" bold="true" box="[151,556,1266,1293]" pageId="7" pageNumber="494">Indochinese Banded Wolf Snake</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="17D8655A2705320EB1C6FAA9FC48FAA9" box="[151,931,1302,1329]" pageId="7" pageNumber="494" type="description">
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<paragraph id="5F7D36D12705320EB1C6FAA9FC48FAA9" blockId="7.[151,931,1266,1329]" box="[151,931,1302,1329]" pageId="7" pageNumber="494">
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(
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<tableCitation id="1240036A2705320EB1CEFAA9FF1FFAA9" box="[159,244,1303,1329]" captionStart="TABLE 1" captionStartId="16.[152,244,151,175]" captionTargetPageId="16" captionText="TABLE 1. Comparison of morphological characters of Lycodon neomaculatus sp. nov. with L. subcinctus and L. sealei. Diagnostic differences between the new species and L. subcinctus + L. sealei are marked in bold." httpUri="http://table.plazi.org/id/0BBD665927123219B1C9FF29FB48FF4B" pageId="7" pageNumber="494" tableUuid="0BBD665927123219B1C9FF29FB48FF4B">Table 1</tableCitation>
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;
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<figureCitation id="C7F92A542705320EB053FAA9FEA5FAA9" box="[258,334,1303,1329]" captionStart="FIGURE 1" captionStartId="8.[151,250,1495,1519]" captionTargetBox="[151,1436,181,1471]" captionTargetId="figure-29@8.[151,1436,181,1471]" captionTargetPageId="8" captionText="FIGURE 1. Colubrid snake species involved in the case of Anoplophallus maculatus Cope, 1895.—USNM 7339, previously considered the holotype of “Lycodon maculatus” reported by Liu et al. (2023): A—General dorsal view; B—General ventral view; C—Dorsalview ofthe head;D—Lateralview of thehead,leftside; E—Livespecimen of Leptodeira maculata (= Megalops maculatus) from Minatitlán, Colima, Mexico; F—Live specimen of Lycodon subannulatus from Penang Island, Malaysia, of which Nymphophidium maculatum is a junior synonym. Images not to scale. Photos by T. Hsu (A–D), C. Grünwald (E), J. L. Lee (F)." figureDoi="http://doi.org/10.5281/zenodo.13916221" httpUri="https://zenodo.org/record/13916221/files/figure.png" pageId="7" pageNumber="494">Figs. 1</figureCitation>
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,
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||
<figureCitation id="C7F92A542705320EB00DFAA9FE80FAA9" box="[348,363,1303,1329]" captionStart="FIGURE 3" captionStartId="10.[151,250,1790,1814]" captionTargetBox="[153,1423,463,1754]" captionTargetId="graphics-175@10.[153,1423,463,1742]" captionTargetPageId="10" captionText="FIGURE 3. Bayesian phylogenetic tree of Lycodon inferred from the mitochondrial cytb sequences. Numbers before slashes indicate Bayesian posterior probabilities and numbers after slashes indicate ML bootstrap supports. Nodes filled in with a black circle indicate strong support by both the ML and BI analyses, whereas nodes filled in with a white circle indicate strong support only in the BI analysis. Inset photo of Lycodon neomaculatus sp. nov. taken by A.M. Bragin." figureDoi="http://doi.org/10.5281/zenodo.13916225" httpUri="https://zenodo.org/record/13916225/files/figure.png" pageId="7" pageNumber="494">3</figureCitation>
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,
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<figureCitation id="C7F92A542705320EB029FAA8FE63FAA9" box="[376,392,1302,1329]" captionStart="FIGURE 5" captionStartId="13.[151,250,1495,1519]" captionTargetBox="[151,1436,181,1471]" captionTargetId="figure-18@13.[151,1436,181,1471]" captionTargetPageId="13" captionText="FIGURE 5. Lycodon neomaculatus sp. nov. in preservative—MCZ R-175968 (holotype, adult male) A—General dorsal view; B—General ventral view; C—Lateral view of the head, right side; D—Lateral view of the head, left side; E—Dorsal view of the head; F—Ventral view of the tail with the everted hemipenes. Images not to scale. Photos by M. Gage and J. Martinez." figureDoi="http://doi.org/10.5281/zenodo.13916229" httpUri="https://zenodo.org/record/13916229/files/figure.png" pageId="7" pageNumber="494">5</figureCitation>
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,
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||
<figureCitation id="C7F92A542705320EB0C5FAA9FE4FFAA9" box="[404,420,1303,1329]" captionStart="FIGURE 6" captionStartId="14.[151,250,1495,1519]" captionTargetBox="[151,1436,181,1471]" captionTargetId="figure-29@14.[151,1436,181,1471]" captionTargetPageId="14" captionText="FIGURE 6. Lycodon neomaculatus sp. nov. in preservative—specimen CAS 235846 (paratype, adult male) A—General dorsal view; B—General ventral view; C—dorsal view of the head; D—Ventral view of the head; E—Lateral view of the head, right side; F—Ventral view of the tail with the everted hemipenes. Photos by J.L. Lee." figureDoi="http://doi.org/10.5281/zenodo.13916233" httpUri="https://zenodo.org/record/13916233/files/figure.png" pageId="7" pageNumber="494">6</figureCitation>
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,
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||
<figureCitation id="C7F92A542705320EB0E1FAA9FE55FAA9" box="[432,446,1303,1329]" captionStart="FIGURE 7" captionStartId="15.[151,250,1323,1347]" captionTargetBox="[397,1191,669,1299]" captionTargetId="figure-260@15.[397,1191,669,1299]" captionTargetPageId="15" captionText="FIGURE 7. Differences in dorsal body band coloration in preservative between Lycodon neomaculatus sp. nov. (A, B) and L. subcinctus (C, D). A—FMNH 180149 from Nakhon Ratchasima, Thailand; B—CAS 229729 from Tanintharyi, Myanmar; C—FMNH 229855 from Selangor, Malaysia; D—FMNH 183772 from Selangor, Malaysia. Images not to scale. Photos by J.L. Lee." figureDoi="http://doi.org/10.5281/zenodo.13916235" httpUri="https://zenodo.org/record/13916235/files/figure.png" pageId="7" pageNumber="494">7</figureCitation>
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||
(A, B), 8, 12; Appendix
|
||
<tableCitation id="1240036A2705320EB384FAA9FCAFFAA9" box="[725,836,1303,1329]" captionStart="TABLE 1" captionStartId="16.[152,244,151,175]" captionTargetPageId="16" captionText="TABLE 1. Comparison of morphological characters of Lycodon neomaculatus sp. nov. with L. subcinctus and L. sealei. Diagnostic differences between the new species and L. subcinctus + L. sealei are marked in bold." httpUri="http://table.plazi.org/id/0BBD665927123219B1C9FF29FB48FF4B" pageId="7" pageNumber="494" tableUuid="0BBD665927123219B1C9FF29FB48FF4B">Tables S1</tableCitation>
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||
, S2, S6)
|
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</paragraph>
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</subSubSection>
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<subSubSection id="17D8655A2705320EB1C6FAC2FB45F8BB" pageId="7" pageNumber="494" type="reference_group">
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<paragraph id="5F7D36D12705320EB1C6FAC2FEDDFA0E" blockId="7.[151,310,1404,1430]" box="[151,310,1404,1430]" pageId="7" pageNumber="494">
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<heading id="043581BD2705320EB1C6FAC2FEDDFA0E" bold="true" box="[151,310,1404,1430]" fontSize="11" level="1" pageId="7" pageNumber="494" reason="1">
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<emphasis id="6DB6EAC32705320EB1C6FAC2FEDDFA0E" bold="true" box="[151,310,1404,1430]" pageId="7" pageNumber="494">Chresonymy</emphasis>
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</heading>
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</paragraph>
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<paragraph id="5F7D36D12705320EB1C6FA04FAC4F9D5" blockId="7.[151,1437,1466,1828]" pageId="7" pageNumber="494">
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<taxonomicName id="98C24D522705320EB1C6FA04FEB4FA4A" baseAuthorityName="F. Boie" baseAuthorityYear="1827" box="[151,351,1466,1490]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="494" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC32705320EB1C6FA04FEB4FA4A" box="[151,351,1466,1490]" italics="true" pageId="7" pageNumber="494">Lycodon subcinctus</emphasis>
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</taxonomicName>
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||
(
|
||
<emphasis id="6DB6EAC32705320EB03FFA05FE7FFA4A" box="[366,404,1467,1490]" italics="true" pageId="7" pageNumber="494">non</emphasis>
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||
<taxonomicName id="98C24D522705320EB0CAFA04FD33FA4A" authority="Boie, 1827" authorityName="Boie" authorityYear="1827" box="[411,728,1466,1490]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="494" phylum="Chordata" rank="species" species="subcinctus">
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||
<emphasis id="6DB6EAC32705320EB0CAFA04FD88FA4A" box="[411,611,1466,1490]" italics="true" pageId="7" pageNumber="494">Lycodon subcinctus</emphasis>
|
||
<bibRefCitation id="3B534B202705320EB33BFA04FD33FA4A" author="Boie, F." box="[618,728,1466,1490]" pageId="7" pageNumber="494" pagination="508 - 566" refId="ref18299" refString="Boie, F. (1827) Bemerkungen uber Merrem's Versuch eines Systems der Amphibien, 1. Lieferung: Ophidier. Isis von Oken, 20 (6), 508 - 566." type="journal article" year="1827">Boie, 1827</bibRefCitation>
|
||
</taxonomicName>
|
||
)—
|
||
<bibRefCitation id="3B534B202705320EB3ABFA04FC53FA4A" author="Smith, M. A." box="[762,952,1466,1490]" pageId="7" pageNumber="494" pagination="195 - 212" refId="ref22878" refString="Smith, M. A. (1923) On a collection of reptiles and batrachians from the island of Hainan. Journal of the Natural History Society of Siam, 6, 195 - 212." type="journal article" year="1923">Smith (1923: 202)</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B202705320EB292FA04FB5EFA4A" author="Stejneger, L. H." box="[963,1205,1466,1490]" pageId="7" pageNumber="494" pagination="1 - 115" refId="ref23013" refString="Stejneger, L. H. (1925) Chinese amphibians and reptiles in the United States National Museum. Proceedings of the United States National Museum, 66 (25), 1 - 115. https: // doi. org / 10.5479 / si. 00963801.66 - 2562.1" type="journal article" year="1925">Stejneger (1925: 90–91</bibRefCitation>
|
||
, in part);
|
||
<bibRefCitation id="3B534B202705320EB446FA04FF04FA68" author="Smith, M. A." pageId="7" pageNumber="494" refId="ref22915" refString="Smith, M. A. (1943) The fauna of British India, Ceylon and Burma, including the whole of the Indo-chinese subregion. Reptilia and Amphibia. Vol. III. Serpentes. Secretary of State for India, London, xii + 583 pp., folding map." type="book" year="1943">Smith (1943: 258</bibRefCitation>
|
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, in part);
|
||
<bibRefCitation id="3B534B202705320EB01CFA67FDEFFA68" author="Deuve, J." box="[333,516,1496,1521]" pageId="7" pageNumber="494" pagination="1 - 252" refId="ref19096" refString="Deuve, J. (1970) Serpents du Laos. Memoires O. R. S. T. O. M., Paris, (39), 1 - 252, 18 figs., 32 pls." type="journal article" year="1970">Deuve (1970: 126</bibRefCitation>
|
||
, in part);
|
||
<bibRefCitation id="3B534B202705320EB333FA66FC9FFA69" author="Zhao, E. M. & Huang, M. H. & Zong, Y. & Jiang, Y. M. & Huang, Q. Y. & Zhao, H. & Ma, J. F. & Zheng, J. & Huang, Z. J. & Wei, G. & Yang, D. Q. & Li, D. J." box="[610,884,1496,1521]" pageId="7" pageNumber="494" refId="ref24466" refString="Zhao, E. M., Huang, M. H., Zong, Y., Jiang, Y. M., Huang, Q. Y., Zhao, H., Ma, J. F., Zheng, J., Huang, Z. J., Wei, G., Yang, D. Q. & Li, D. J. (1998) Fauna Sinica Reptilia. Vol. 3. Squamata: Serpentes. Science Press, Beijing, 522 pp. [in Chinese]" type="book" year="1998">
|
||
Zhao
|
||
<emphasis id="6DB6EAC32705320EB3CAFA67FD2DFA68" box="[667,710,1496,1520]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (1998: 183–184
|
||
</bibRefCitation>
|
||
, in part);
|
||
<bibRefCitation id="3B534B202705320EB280FA66FB91FA69" author="Zhao, E. M." box="[977,1146,1496,1521]" pageId="7" pageNumber="494" refId="ref24423" refString="Zhao, E. M. (2006) Snakes of China. Vols. I - II. Anhui Science and Technology Publishing House, Hefei, 390 + 287 pp., 194 col. pls. [in Chinese]" type="book" year="2006">Zhao (2006: 218</bibRefCitation>
|
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, in part);
|
||
<bibRefCitation id="3B534B202705320EB586FA67FF00F997" author="Wogan, G. O. U. & Vindum, J. V. & Wilkinson, J. A. & Koo, M. S. & Slowinski, J. B. & Win, H. & Thin, T. & Kyi, S. W. & Oo, S. L. & Lwin, K. S. & Shein, A. K." pageId="7" pageNumber="494" pagination="83 - 96" refId="ref23772" refString="Wogan, G. O. U., Vindum, J. V., Wilkinson, J. A., Koo, M. S., Slowinski, J. B., Win, H., Thin, T., Kyi, S. W., Oo, S. L., Lwin, K. S. & Shein, A. K. (2008) New country records and range extensions for Myanmar amphibians and reptiles. Hamadryad, 33 (1), 83 - 96." type="journal article" year="2008">
|
||
Wogan
|
||
<emphasis id="6DB6EAC32705320EB475FA67FAA4FA68" box="[1316,1359,1496,1520]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2008: 87)
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B202705320EB1A6FA46FE11F997" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[247,506,1527,1552]" pageId="7" pageNumber="494" refId="ref21584" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2009) Herpetofauna of Vietnam. Edition Chimaira, Frankfurt am Main, 768 pp." type="book" year="2009">
|
||
Nguyen
|
||
<emphasis id="6DB6EAC32705320EB01FFA46FE90F997" box="[334,379,1527,1551]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2009: 321
|
||
</bibRefCitation>
|
||
; in part);
|
||
<bibRefCitation id="3B534B202705320EB30CFA49FCB6F997" author="Geissler, P. & Nguyen, T. Q. & Poyarkov, N. A. & Bohme, W." box="[605,861,1527,1551]" pageId="7" pageNumber="494" pagination="9 - 16" refId="ref19433" refString="Geissler, P., Nguyen, T. Q., Poyarkov, N. A. & Bohme, W. (2011) New records of snakes from Cat Tien National Park Dong Nai and Lam Dong provinces southern Vietnam. Bonn Zoological Bulletin, 60 (1), 9 - 16." type="journal article" year="2011">
|
||
Geissler
|
||
<emphasis id="6DB6EAC32705320EB3E6FA46FD0FF997" box="[695,740,1527,1551]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2011: 13)
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B202705320EB239FA49FBFCF997" author="Siler, C. D. & Oliveros, C. H. & Santanen, A. & Brown, R. M." box="[872,1047,1527,1551]" pageId="7" pageNumber="494" pagination="262 - 277" refId="ref22814" refString="Siler, C. D., Oliveros, C. H., Santanen, A. & Brown, R. M. (2013) Multilocus phylogeny reveals unexpected diversification patterns in Asian Wolf Snakes (genus Lycodon). Zoologica Scripta, 42, 262 - 277. https: // doi. org / 10.1111 / zsc. 12007" type="journal article" year="2013">
|
||
Siler
|
||
<emphasis id="6DB6EAC32705320EB2CEFA46FC27F997" box="[927,972,1527,1551]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2013
|
||
</bibRefCitation>
|
||
: in part);
|
||
<bibRefCitation id="3B534B202705320EB528FA49FA96F997" author="Wallach, V. & Williams, K. L. & Boundy, J." box="[1145,1405,1527,1551]" pageId="7" pageNumber="494" refId="ref23535" refString="Wallach, V., Williams, K. L. & Boundy, J. (2014) Snakes of the wworld. A catalogue of living and extinct species. CRC Press, Boca Raton, Florida, xxvii + 1209 pp." type="book" year="2014">
|
||
Wallach
|
||
<emphasis id="6DB6EAC32705320EB583FA46FAEBF997" box="[1234,1280,1527,1551]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2014: 396
|
||
</bibRefCitation>
|
||
, in part); Chan-ard
|
||
<emphasis id="6DB6EAC32705320EB039F9A9FE7FF9B6" box="[360,404,1558,1582]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2015: in part);
|
||
<bibRefCitation id="3B534B202705320EB36DF9A8FC68F9B6" author="Vassilieva, A. B. & Galoyan, E. A. & Poyarkov, N. A. & Geissler P." box="[572,899,1558,1582]" pageId="7" pageNumber="494" refId="ref23332" refString="Vassilieva, A. B., Galoyan, E. A., Poyarkov, N. A. & Geissler P. (2016) A photographic field guide to the amphibians and reptiles of the lowland monsoon forests of southern Vietnam. Edition Chimaira, Frankfurt am Main, 324 pp." type="book" year="2016">
|
||
Vassilieva
|
||
<emphasis id="6DB6EAC32705320EB3F8F9A9FD3EF9B6" box="[681,725,1558,1582]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2016: 140–141
|
||
</bibRefCitation>
|
||
, in part);
|
||
<bibRefCitation id="3B534B202705320EB2B3F9A8FB56F9B6" author="Wang, J. & Qi, S. & Lyu, Z. T. & Zeng, Z. C. & Wang, Y. Y." box="[994,1213,1558,1582]" pageId="7" pageNumber="494" pagination="85 - 108" refId="ref23580" refString="Wang, J., Qi, S., Lyu, Z. T., Zeng, Z. C. & Wang, Y. Y. (2020) A new species of the genus Lycodon (Serpentes, Colubridae) from Guangxi, China. Zookeys, 954, 85 - 108. https: // doi. org / 10.3897 / zookeys. 954.53432" type="journal article" year="2020">
|
||
Wang
|
||
<emphasis id="6DB6EAC32705320EB573F9A9FBA5F9B6" box="[1058,1102,1558,1582]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2020: 93
|
||
</bibRefCitation>
|
||
, 108);
|
||
<bibRefCitation id="3B534B202705320EB453F9A8FEC8F9D5" author="Le, D. T. & Luong, A. M. & Pham, C. T. & Pham, T. Q. & Nguyen, S. L. H. & Ziegler, T. & Nguyen, T. Q." pageId="7" pageNumber="494" pagination="201 - 219" refId="ref20602" refString="Le, D. T., Luong, A. M., Pham, C. T., Pham, T. Q., Nguyen, S. L. H., Ziegler, T. & Nguyen, T. Q. (2021) New records and an updated checklist of amphibians and snakes from Tuyen Quang Province, Vietnam. Bonn zoological Bulletin, 70 (1), 201 - 219. https: // doi. org / 10.20363 / BZB- 2021.70.1.201" type="journal article" year="2021">
|
||
Le
|
||
<emphasis id="6DB6EAC32705320EB473F9A9FAA5F9B6" box="[1314,1358,1558,1582]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2021: 210–201
|
||
</bibRefCitation>
|
||
, in part);
|
||
<bibRefCitation id="3B534B202705320EB0D5F98AFD4EF9D4" author="David, P. & Teynie, A. & Vogel, G." box="[388,677,1588,1613]" pageId="7" pageNumber="494" refId="ref19064" refString="David, P., Teynie, A. & Vogel, G. (2023) The Snakes of Laos. Edition Chimaira, Frankfurt am Main, 960 pp." type="book" year="2023">
|
||
David
|
||
<emphasis id="6DB6EAC32705320EB099F98BFE1EF9D4" box="[456,501,1588,1612]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2023: 328–335
|
||
</bibRefCitation>
|
||
, in part);
|
||
<bibRefCitation id="3B534B202705320EB257F98BFC59F9D5" author="Uetz, P. & Freed, P. & Aguilar, R. & Hosek, J." box="[774,946,1588,1613]" pageId="7" pageNumber="494" refId="ref23225" refString="Uetz, P., Freed, P., Aguilar, R. & Hosek, J. (2024) The Reptile Database. The Reptile Database. Available from: http: // www. reptile-database. org / (accessed 15 January 2024)" type="url" year="2024">
|
||
Uetz
|
||
<emphasis id="6DB6EAC32705320EB26DF98BFC82F9D4" box="[828,873,1588,1612]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2024
|
||
</bibRefCitation>
|
||
, page “
|
||
<taxonomicName id="98C24D522705320EB2AFF98AFB2DF9D4" authorityName="Boie" authorityYear="1827" box="[1022,1222,1588,1612]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="494" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC32705320EB2AFF98AFB2DF9D4" box="[1022,1222,1588,1612]" italics="true" pageId="7" pageNumber="494">Lycodon subcinctus</emphasis>
|
||
</taxonomicName>
|
||
”, in part).
|
||
</paragraph>
|
||
<paragraph id="5F7D36D12705320EB1A6F9EDFCDBF9F3" blockId="7.[151,1437,1466,1828]" box="[247,816,1619,1643]" pageId="7" pageNumber="494">
|
||
<taxonomicName id="98C24D522705320EB1A6F9EDFE5CF9F3" authorityName="F. Boie" authorityYear="1827" box="[247,439,1619,1643]" class="Reptilia" family="Colubridae" genus="Ophites" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="494" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC32705320EB1A6F9EDFE5CF9F3" box="[247,439,1619,1643]" italics="true" pageId="7" pageNumber="494">Ophites subcinctus</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="3B534B202705320EB080F9EDFD3FF9F3" author="Zhao, E. M. & Adler, K." box="[465,724,1619,1643]" pageId="7" pageNumber="494" refId="ref24383" refString="Zhao, E. M. & Adler, K. (1993) Herpetology of China. Contributions to Herpetology. No. 10. Society for the Study of Amphibians and Reptiles, Oxford, 522 pp." type="book" year="1993">Zhao & Adler (1993: 247</bibRefCitation>
|
||
, in part).
|
||
</paragraph>
|
||
<paragraph id="5F7D36D12705320EB1A6F9CCFCB5F912" blockId="7.[151,1437,1466,1828]" box="[247,862,1650,1674]" pageId="7" pageNumber="494">
|
||
<taxonomicName id="98C24D522705320EB1A6F9CCFDDBF912" box="[247,560,1650,1674]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="494" phylum="Chordata" rank="subSpecies" species="subcinctus" subSpecies="subcinctus">
|
||
<emphasis id="6DB6EAC32705320EB1A6F9CCFDDBF912" box="[247,560,1650,1674]" italics="true" pageId="7" pageNumber="494">Lycodon subcinctus subcinctus</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="3B534B202705320EB31BF9CCFCE9F912" author="Taylor, E. H." box="[586,770,1650,1674]" pageId="7" pageNumber="494" pagination="609 - 1096" refId="ref23192" refString="Taylor, E. H. (1965) The serpents of Thailand and adjacent waters. The University of Kansas Science Bulletin, 45 (9), 609 - 1096." type="journal article" year="1965">Taylor (1965: 739</bibRefCitation>
|
||
, in part).
|
||
</paragraph>
|
||
<paragraph id="5F7D36D12705320EB1A6F92EFC58F931" blockId="7.[151,1437,1466,1828]" box="[247,947,1680,1705]" pageId="7" pageNumber="494">
|
||
<taxonomicName id="98C24D522705320EB1A6F92EFDC6F930" authorityName="Boie" authorityYear="1827" box="[247,557,1680,1705]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="494" phylum="Chordata" rank="species" species="subcinctus" subGenus="Lycodon">
|
||
<emphasis id="6DB6EAC32705320EB1A6F92EFEA6F930" box="[247,333,1680,1704]" italics="true" pageId="7" pageNumber="494">Lycodon</emphasis>
|
||
(
|
||
<emphasis id="6DB6EAC32705320EB00DF92EFE59F930" box="[348,434,1680,1704]" italics="true" pageId="7" pageNumber="494">Lycodon</emphasis>
|
||
)
|
||
<emphasis id="6DB6EAC32705320EB093F92EFDC6F930" box="[450,557,1680,1704]" italics="true" pageId="7" pageNumber="494">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="3B534B202705320EB317F92EFCBCF931" author="Poyarkov, N. A. & Nguyen, T. V. & Popov, E. S. & Geissler, P. & Pawangkhanant, P. & Neang, T. & Suwannapoom, C. & Ananjeva, N. B. & Orlov, N. L." box="[582,855,1680,1705]" pageId="7" pageNumber="494" pagination="255 - 476" refId="ref22080" refString="Poyarkov, N. A., Nguyen, T. V., Popov, E. S., Geissler, P., Pawangkhanant, P., Neang, T., Suwannapoom, C., Ananjeva, N. B. & Orlov, N. L. (2023) Recent progress in taxonomic studies, biogeographic analysis, and revised checklist of Reptiles in Indochina. Russian Journal of Herpetology, 30 (5), 255 - 476. https: // doi. org / 10.30906 / 1026 - 2296 - 2023 - 30 - 5 - 255 - 476" type="journal article" year="2023">
|
||
Poyarkov
|
||
<emphasis id="6DB6EAC32705320EB3FCF92FFD31F930" box="[685,730,1680,1704]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2023: 341
|
||
</bibRefCitation>
|
||
, in part).
|
||
</paragraph>
|
||
<paragraph id="5F7D36D12705320EB1A6F911FCFBF95F" blockId="7.[151,1437,1466,1828]" box="[247,784,1711,1735]" pageId="7" pageNumber="494">
|
||
<taxonomicName id="98C24D522705320EB1A6F911FE34F95F" authorityName="Boie" authorityYear="1827" box="[247,479,1711,1735]" class="Reptilia" family="Colubridae" genus="Lycodon" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="494" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC32705320EB1A6F911FEA6F95F" box="[247,333,1711,1735]" italics="true" pageId="7" pageNumber="494">Lycodon</emphasis>
|
||
cf.
|
||
<emphasis id="6DB6EAC32705320EB025F911FE34F95F" box="[372,479,1711,1735]" italics="true" pageId="7" pageNumber="494">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="3B534B202705320EB0A8F911FCE7F95F" author="Zhang, H. Y. & Wang, L. Y. & Hou, S. B. & Wu, Y. H. & Duan, Z. P. & Jin, Y. D. & Zuo, C. S. & Zhang, Y. S. & Wang, K. & Che, J." box="[505,780,1711,1735]" pageId="7" pageNumber="494" pagination="81 - 94" refId="ref24266" refString="Zhang, H. Y., Wang, L. Y., Hou, S. B., Wu, Y. H., Duan, Z. P., Jin, Y. D., Zuo, C. S., Zhang, Y. S., Wang, K. & Che, J. (2024) Provincial records of Lycodon septentrionalis and L. cf. subcinctus from Yunnan Province, China. Chinese Journal of Zoology, 59 (1), 81 - 94. [in Chinese] https: // doi. org / 10.13859 / j. cjz. 202423018" type="journal article" year="2024">
|
||
Zhang
|
||
<emphasis id="6DB6EAC32705320EB311F90EFD87F95F" box="[576,620,1711,1735]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2024: 81–94)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5F7D36D12705320EB1C6F970FC67F97E" blockId="7.[151,1437,1466,1828]" box="[151,908,1742,1766]" pageId="7" pageNumber="494">
|
||
“
|
||
<taxonomicName id="98C24D522705320EB1F3F970FE4EF97E" authorityName="Cope" authorityYear="1895" box="[162,421,1742,1766]" class="Reptilia" family="Colubridae" genus="Anoplophallus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="494" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="6DB6EAC32705320EB1F3F970FE4EF97E" box="[162,421,1742,1766]" italics="true" pageId="7" pageNumber="494">Anoplophallus maculatus</emphasis>
|
||
</taxonomicName>
|
||
” (not a validly described taxon)—
|
||
<bibRefCitation id="3B534B202705320EB254F970FC63F97E" author="Cope, E. D." box="[773,904,1742,1766]" pageId="7" pageNumber="494" pagination="186 - 219" refId="ref18952" refString="Cope, E. D. (1895). The classification of the Ophidia. Transactions of the American Philosophical Society, 18 (2), 186 - 219, pls. 14 - 33. https: // doi. org / 10.2307 / 1005387" type="journal article" year="1895">Cope (1895)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5F7D36D12705320EB1A6F952FCA7F89D" blockId="7.[151,1437,1466,1828]" box="[247,844,1772,1797]" pageId="7" pageNumber="494">
|
||
<taxonomicName id="98C24D522705320EB1A6F952FDC6F89C" authorityName="" (sensu Leviton" authorityYear="1955" box="[247,557,1772,1796]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="494" phylum="Chordata" rank="subSpecies" species="subcinctus" subSpecies="maculatus">
|
||
<emphasis id="6DB6EAC32705320EB1A6F952FDC6F89C" box="[247,557,1772,1796]" italics="true" pageId="7" pageNumber="494">Lycodon subcinctus maculatus</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="3B534B202705320EB316F952FCE7F89D" author="Leviton, A. E." box="[583,780,1772,1797]" pageId="7" pageNumber="494" pagination="195 - 203" refId="ref20767" refString="Leviton, A. E. (1955) Systematic notes on the Asian snake Lycodon subcinctus. The Philippine Journal of Science, 84 (2), 195 - 203." type="journal article" year="1955">Leviton (1955: 199</bibRefCitation>
|
||
, 201).
|
||
</paragraph>
|
||
<paragraph id="5F7D36D12705320EB1A6F8B5FB45F8BB" blockId="7.[151,1437,1466,1828]" box="[247,1198,1803,1828]" pageId="7" pageNumber="494">
|
||
<taxonomicName id="98C24D522705320EB1A6F8B5FE57F8BB" box="[247,444,1803,1827]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="494" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="6DB6EAC32705320EB1A6F8B5FE57F8BB" box="[247,444,1803,1827]" italics="true" pageId="7" pageNumber="494">Lycodon maculatus</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="3B534B202705320EB087F8B5FD96F8BB" author="Liu, S. & Hou, M. & Cai, B. & Li, S. & Zhang, Z. & Yu, R. & Rao, D. & Zhang, L." box="[470,637,1803,1827]" pageId="7" pageNumber="494" pagination="307 - 316" refId="ref21074" refString="Liu, S., Hou, M., Cai, B., Li, S., Zhang, Z., Yu, R., Rao, D. & Zhang, L. (2023) Taxonomic status of Lycodon subcinctus sensu lato in China (Serpentes, Colubridae). Herpetozoa, 36, 307 - 316. https: // doi. org / 10.3897 / herpetozoa. 36. e 114206" type="journal article" year="2023">
|
||
Liu
|
||
<emphasis id="6DB6EAC32705320EB0AEF8B2FDC7F8BB" box="[511,556,1803,1827]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2023)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="3B534B202705320EB3D9F8B2FCDEF8BB" author="Uetz, P. & Freed, P. & Aguilar, R. & Hosek, J." box="[648,821,1803,1827]" pageId="7" pageNumber="494" refId="ref23225" refString="Uetz, P., Freed, P., Aguilar, R. & Hosek, J. (2024) The Reptile Database. The Reptile Database. Available from: http: // www. reptile-database. org / (accessed 15 January 2024)" type="url" year="2024">
|
||
Uetz
|
||
<emphasis id="6DB6EAC32705320EB3EFF8B2FD00F8BB" box="[702,747,1803,1827]" italics="true" pageId="7" pageNumber="494">et al</emphasis>
|
||
. (2024
|
||
</bibRefCitation>
|
||
, page “
|
||
<taxonomicName id="98C24D522705320EB2D1F8B5FBAEF8BB" box="[896,1093,1803,1827]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="494" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="6DB6EAC32705320EB2D1F8B5FBAEF8BB" box="[896,1093,1803,1827]" italics="true" pageId="7" pageNumber="494">Lycodon maculatus</emphasis>
|
||
</taxonomicName>
|
||
”, in part).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="17D8655A27053202B1C6F8F5FBF0F924" lastPageId="11" lastPageNumber="498" pageId="7" pageNumber="494" type="materials_examined">
|
||
<paragraph id="5F7D36D12705320EB1C6F8F5FD5DF811" blockId="7.[151,1436,1866,1929]" pageId="7" pageNumber="494">
|
||
<materialsCitation id="EFAA3C8C2705320EB1C6F8F5FD5DF811" collectingDate="1990-06-25" collectionCode="MCZ" collectorName="S. J. Karsen" country="Hong Kong" location="Chek Lap Kok Island" pageId="7" pageNumber="494" specimenCode="MCZ R-175968" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="6DB6EAC32705320EB1C6F8F5FEEFF8FD" bold="true" box="[151,260,1867,1893]" pageId="7" pageNumber="494">
|
||
<typeStatus id="807988732705320EB1C6F8F5FEEFF8FD" box="[151,260,1867,1893]" pageId="7" pageNumber="494" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCode id="0F649EAA2705320EB044F8F5FE21F8FD" box="[277,458,1866,1893]" pageId="7" pageNumber="494">MCZ R-175968</specimenCode>
|
||
(
|
||
<specimenCount id="49C4FD582705320EB08AF8F5FDBEF8FD" box="[475,597,1867,1893]" pageId="7" pageNumber="494" type="adult">adult male</specimenCount>
|
||
) from
|
||
<location id="5A1D600A2705320EB3F4F8F5FC75F8FD" LSID="urn:lsid:plazi:treatment:D76B87C72705321CB1C6FB14FA6CFBC1:5A1D600A2705320EB3F4F8F5FC75F8FD" box="[677,926,1867,1893]" country="Hong Kong" name="Chek Lap Kok Island" pageId="7" pageNumber="494">Chek Lap Kok Island</location>
|
||
,
|
||
<collectingCountry id="27D576412705320EB2FAF8F5FA73F8FD" box="[939,1432,1867,1893]" name="Hong Kong" pageId="7" pageNumber="494">Hong Kong Special Administrative Region</collectingCountry>
|
||
,
|
||
<collectingCountry id="27D576412705320EB1C6F8D1FF31F811" box="[151,218,1903,1929]" name="China" pageId="7" pageNumber="494">China</collectingCountry>
|
||
collected on
|
||
<date id="2B7C10112705320EB021F8D0FDEEF811" box="[368,517,1902,1929]" pageId="7" pageNumber="494" value="1990-06-25">
|
||
<collectingDate id="3B38E9F92705320EB021F8D0FDEEF811" box="[368,517,1902,1929]" pageId="7" pageNumber="494" value="1990-06-25">25 June 1990</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="F23753072705320EB37EF8D1FD59F811" box="[559,690,1903,1929]" pageId="7" pageNumber="494">S.J. Karsen</collectorName>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="0BBD6659270A3201B1C6FA69FF02F93B" ID-DOI="http://doi.org/10.5281/zenodo.13916221" ID-Zenodo-Dep="13916221" httpUri="https://zenodo.org/record/13916221/files/figure.png" pageId="8" pageNumber="495" startId="8.[151,250,1495,1519]" targetBox="[151,1436,181,1471]" targetPageId="8" targetType="figure">
|
||
<paragraph id="5F7D36D1270A3201B1C6FA69FF02F93B" blockId="8.[151,1437,1495,1699]" pageId="8" pageNumber="495">
|
||
<emphasis id="6DB6EAC3270A3201B1C6FA69FEE5FA77" bold="true" box="[151,270,1495,1519]" pageId="8" pageNumber="495">FIGURE 1</emphasis>
|
||
. Colubrid snake species involved in the case of
|
||
<taxonomicName id="98C24D52270A3201B251FA69FB6AFA77" authority="Cope, 1895" authorityName="Cope" authorityYear="1895" box="[768,1153,1495,1519]" class="Reptilia" family="Colubridae" genus="Anoplophallus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="495" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="6DB6EAC3270A3201B251FA69FBEFFA77" box="[768,1028,1495,1519]" italics="true" pageId="8" pageNumber="495">Anoplophallus maculatus</emphasis>
|
||
<bibRefCitation id="3B534B20270A3201B55BFA69FB6AFA77" author="Cope, E. D." box="[1034,1153,1495,1519]" pageId="8" pageNumber="495" pagination="186 - 219" refId="ref18952" refString="Cope, E. D. (1895). The classification of the Ophidia. Transactions of the American Philosophical Society, 18 (2), 186 - 219, pls. 14 - 33. https: // doi. org / 10.2307 / 1005387" type="journal article" year="1895">Cope, 1895</bibRefCitation>
|
||
</taxonomicName>
|
||
.—USNM 7339, previously considered the holotype of “
|
||
<taxonomicName id="98C24D52270A3201B0ECFA45FD6FF98B" box="[445,644,1531,1555]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="495" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="6DB6EAC3270A3201B0ECFA45FD6FF98B" box="[445,644,1531,1555]" italics="true" pageId="8" pageNumber="495">Lycodon maculatus</emphasis>
|
||
</taxonomicName>
|
||
” reported by
|
||
<bibRefCitation id="3B534B20270A3201B244FA45FC2EF98B" author="Liu, S. & Hou, M. & Cai, B. & Li, S. & Zhang, Z. & Yu, R. & Rao, D. & Zhang, L." box="[789,965,1531,1555]" pageId="8" pageNumber="495" pagination="307 - 316" refId="ref21074" refString="Liu, S., Hou, M., Cai, B., Li, S., Zhang, Z., Yu, R., Rao, D. & Zhang, L. (2023) Taxonomic status of Lycodon subcinctus sensu lato in China (Serpentes, Colubridae). Herpetozoa, 36, 307 - 316. https: // doi. org / 10.3897 / herpetozoa. 36. e 114206" type="journal article" year="2023">
|
||
Liu
|
||
<emphasis id="6DB6EAC3270A3201B210FA42FC9DF98B" box="[833,886,1531,1555]" italics="true" pageId="8" pageNumber="495">et al.</emphasis>
|
||
(2023)
|
||
</bibRefCitation>
|
||
: A—General dorsal view; B—General ventral view; C—Dorsal view of the head; D—Lateral view of the head, left side; E—Live specimen of
|
||
<taxonomicName id="98C24D52270A3201B515F9A1FAFEF9AF" box="[1092,1301,1567,1591]" class="Reptilia" family="Colubridae" genus="Leptodeira" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="495" phylum="Chordata" rank="species" species="maculata">
|
||
<emphasis id="6DB6EAC3270A3201B515F9A1FAFEF9AF" box="[1092,1301,1567,1591]" italics="true" pageId="8" pageNumber="495">Leptodeira maculata</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="98C24D52270A3201B469F9A1FF14F9C3" authorityName="Hallowell" authorityYear="1861" class="Reptilia" family="Megalopidae" genus="Megalops" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="495" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="6DB6EAC3270A3201B469F9A1FF14F9C3" italics="true" pageId="8" pageNumber="495">Megalops maculatus</emphasis>
|
||
</taxonomicName>
|
||
) from Minatitlán, Colima, Mexico; F—Live specimen of
|
||
<taxonomicName id="98C24D52270A3201B207F9FDFBD7F9C3" box="[854,1084,1603,1627]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="495" phylum="Chordata" rank="species" species="subannulatus">
|
||
<emphasis id="6DB6EAC3270A3201B207F9FDFBD7F9C3" box="[854,1084,1603,1627]" italics="true" pageId="8" pageNumber="495">Lycodon subannulatus</emphasis>
|
||
</taxonomicName>
|
||
from Penang Island, Malaysia, of which
|
||
<taxonomicName id="98C24D52270A3201B18AF9D9FE10F9E7" authorityName="Günther" authorityYear="1864" box="[219,507,1639,1663]" genus="Nymphophidium" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="495" phylum="Chordata" rank="species" species="maculatum">
|
||
<emphasis id="6DB6EAC3270A3201B18AF9D9FE10F9E7" box="[219,507,1639,1663]" italics="true" pageId="8" pageNumber="495">Nymphophidium maculatum</emphasis>
|
||
</taxonomicName>
|
||
is a junior synonym. Images not to scale. Photos by T. Hsu (A–D), C. Grünwald (E), J. L. Lee (F).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5F7D36D1270A3200B196F948FE37FE11" blockId="8.[151,1436,1781,2024]" lastBlockId="9.[151,1437,150,393]" lastPageId="9" lastPageNumber="496" pageId="8" pageNumber="495">
|
||
<materialsCitation id="EFAA3C8C270A3201B196F948FC59F8AC" collectingDate="1990-06-29" collectionCode="MCZ" collectorName="S. J. Karsen" country="China" location="Pak Ngan Village" municipality="Mui Wo Town" pageId="8" pageNumber="495" specimenCode="MCZ R-175967" specimenCount="1" stateProvince="Lantau Island" typeStatus="paratype">
|
||
<emphasis id="6DB6EAC3270A3201B196F948FE70F897" bold="true" box="[199,411,1781,1808]" pageId="8" pageNumber="495">
|
||
<typeStatus id="80798873270A3201B196F948FED4F888" box="[199,319,1782,1808]" pageId="8" pageNumber="495" type="paratype">Paratypes</typeStatus>
|
||
(n = 8)
|
||
</emphasis>
|
||
:—
|
||
<emphasis id="6DB6EAC3270A3201B0EFF94BFDECF897" bold="true" box="[446,519,1781,1807]" pageId="8" pageNumber="495">
|
||
<collectingCountry id="27D57641270A3201B0EFF94BFDECF897" box="[446,519,1781,1807]" name="China" pageId="8" pageNumber="495">China</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<specimenCode id="0F649EAA270A3201B349F948FD26F888" box="[536,717,1781,1808]" pageId="8" pageNumber="495">MCZ R-175967</specimenCode>
|
||
(
|
||
<specimenCount id="49C4FD58270A3201B38FF94BFC84F897" box="[734,879,1781,1807]" pageId="8" pageNumber="495" type="adult">adult female</specimenCount>
|
||
) from
|
||
<location id="5A1D600A270A3201B2EEF94BFB67F897" LSID="urn:lsid:plazi:treatment:D76B87C72705321CB1C6FB14FA6CFBC1:5A1D600A270A3201B2EEF94BFB67F897" box="[959,1164,1781,1808]" country="China" municipality="Mui Wo Town" name="Pak Ngan Village" pageId="8" pageNumber="495" stateProvince="Lantau Island">Pak Ngan Village</location>
|
||
,
|
||
<collectingMunicipality id="BF19ACAB270A3201B5C8F94BFAAAF888" box="[1177,1345,1781,1808]" pageId="8" pageNumber="495">Mui Wo Town</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="9D06F833270A3201B41FF948FF35F8AB" pageId="8" pageNumber="495">Lantau Island</collectingRegion>
|
||
,
|
||
<collectingCountry id="27D57641270A3201B1BAF8A4FE9BF8AC" box="[235,368,1818,1844]" name="Hong Kong" pageId="8" pageNumber="495">Hong Kong</collectingCountry>
|
||
SAR. collected on
|
||
<date id="2B7C1011270A3201B301F8A4FD03F8AC" box="[592,744,1818,1844]" pageId="8" pageNumber="495" value="1990-06-29">
|
||
<collectingDate id="3B38E9F9270A3201B301F8A4FD03F8AC" box="[592,744,1818,1844]" pageId="8" pageNumber="495" value="1990-06-29">29 June 1990</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="F2375307270A3201B244F8A4FC7EF8AC" box="[789,917,1818,1844]" pageId="8" pageNumber="495">S.J. Karsen</collectorName>
|
||
—
|
||
</materialsCitation>
|
||
<materialsCitation id="EFAA3C8C270A3201B2E0F8A4FE76F838" collectingDate="2003-05-14" collectionCode="CAS" collectorName="Wogan G. O. U. & Wilkinson J. A. & Vindum J. V. & Win H. & Thin T" country="Myanmar" elevation="254" latitude="25.093166" location="Hepu Stream, Hepu Village" longLatPrecision="1" longitude="96.40019" municipality="Moenyin Township" pageId="8" pageNumber="495" specimenCode="CAS 235846" specimenCount="1" stateProvince="Kachin State">
|
||
<emphasis id="6DB6EAC3270A3201B2E0F8A4FBC2F8AC" bold="true" box="[945,1065,1818,1844]" pageId="8" pageNumber="495">
|
||
<collectingCountry id="27D57641270A3201B2E0F8A4FBC2F8AC" box="[945,1065,1818,1844]" name="Myanmar" pageId="8" pageNumber="495">Myanmar</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<specimenCode id="0F649EAA270A3201B568F8A4FB20F8AC" box="[1081,1227,1817,1844]" pageId="8" pageNumber="495">CAS 235846</specimenCode>
|
||
(
|
||
<specimenCount id="49C4FD58270A3201B58AF8A7FABDF8AB" box="[1243,1366,1817,1843]" pageId="8" pageNumber="495" type="adult">adult male</specimenCount>
|
||
) from
|
||
<location id="5A1D600A270A3201B1C6F880FE0FF8CF" LSID="urn:lsid:plazi:treatment:D76B87C72705321CB1C6FB14FA6CFBC1:5A1D600A270A3201B1C6F880FE0FF8CF" box="[151,484,1853,1880]" country="Myanmar" latitude="25.093166" longLatPrecision="1" longitude="96.40019" municipality="Moenyin Township" name="Hepu Stream, Hepu Village" pageId="8" pageNumber="495" stateProvince="Kachin State">Hepu Stream, Hepu Village</location>
|
||
, Indawgyi WS,
|
||
<collectingMunicipality id="BF19ACAB270A3201B3FDF883FC7BF8CF" box="[684,912,1853,1879]" pageId="8" pageNumber="495">Moenyin Township</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="9D06F833270A3201B2F3F883FBD7F8C0" box="[930,1084,1853,1880]" country="Myanmar" name="Kachin" pageId="8" pageNumber="495">Kachin State</collectingRegion>
|
||
, Myanmar (
|
||
<geoCoordinate id="3AF65016270A3201B581F883FA73F8C0" box="[1232,1432,1853,1880]" degrees="25.0931666667" direction="north" orientation="latitude" pageId="8" pageNumber="495" precision="1" value="25.093166">25.0931666667N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3AF65016270A3201B1C6F8DFFEBCF8E4" box="[151,343,1889,1916]" degrees="96.4001944444" direction="east" orientation="longitude" pageId="8" pageNumber="495" precision="1" value="96.40019">96.4001944444E</geoCoordinate>
|
||
; altitude
|
||
<quantity id="983A9B34270A3201B09BF8DFFDFDF8E4" box="[458,534,1889,1916]" metricMagnitude="2" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="495" unit="m" value="254.0">
|
||
<elevation id="D4EFD1E2270A3201B09BF8DFFDFDF8E4" box="[458,534,1889,1916]" metricMagnitude="2" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="495" unit="m" value="254.0">254 m</elevation>
|
||
</quantity>
|
||
asl.) collected on
|
||
<date id="2B7C1011270A3201B3A5F8DCFC78F8E4" box="[756,915,1890,1916]" pageId="8" pageNumber="495" value="2003-05-14">
|
||
<collectingDate id="3B38E9F9270A3201B3A5F8DCFC78F8E4" box="[756,915,1890,1916]" pageId="8" pageNumber="495" value="2003-05-14">14 May 2003</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="F2375307270A3201B297F8DCFB9AF8E4" box="[966,1137,1890,1916]" pageId="8" pageNumber="495">Wogan G.O.U.</collectorName>
|
||
,
|
||
<collectorName id="F2375307270A3201B5D5F8DCFAD9F8E4" box="[1156,1330,1890,1916]" pageId="8" pageNumber="495">Wilkinson J.A.</collectorName>
|
||
,
|
||
<collectorName id="F2375307270A3201B413F8DCFF28F838" pageId="8" pageNumber="495">Vindum J.V.</collectorName>
|
||
,
|
||
<collectorName id="F2375307270A3201B181F838FEC9F838" box="[208,290,1926,1952]" pageId="8" pageNumber="495">Win H.</collectorName>
|
||
&
|
||
<collectorName id="F2375307270A3201B019F838FE73F838" box="[328,408,1926,1952]" pageId="8" pageNumber="495">Thin T</collectorName>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="EFAA3C8C270A3201B0F4F838FC96F87F" collectingDate="2003-01-19" collectionCode="CAS" collectorName="Win H. & Tun H. & Lwin K. S. & Shein A. K. & Naing Z. M" elevation="45" latitude="13.867111" location="East of Thaye Chaung Town" longLatPrecision="1" longitude="98.28369" municipality="Thaye Chaung Township" pageId="8" pageNumber="495" specimenCode="CAS 229726" specimenCount="1" stateProvince="Dawei District">
|
||
<specimenCode id="0F649EAA270A3201B0F4F838FDD3F807" box="[421,568,1925,1952]" pageId="8" pageNumber="495">CAS 229726</specimenCode>
|
||
(
|
||
<specimenCount id="49C4FD58270A3201B319F838FD33F838" box="[584,728,1926,1952]" pageId="8" pageNumber="495" type="adult">adult female</specimenCount>
|
||
) from
|
||
<location id="5A1D600A270A3201B277F838FB85F838" LSID="urn:lsid:plazi:treatment:D76B87C72705321CB1C6FB14FA6CFBC1:5A1D600A270A3201B277F838FB85F838" box="[806,1134,1926,1952]" latitude="13.867111" longLatPrecision="1" longitude="98.28369" municipality="Thaye Chaung Township" name="East of Thaye Chaung Town" pageId="8" pageNumber="495" stateProvince="Dawei District">East of Thaye Chaung Town</location>
|
||
,
|
||
<collectingMunicipality id="BF19ACAB270A3201B52BF838FA73F838" box="[1146,1432,1926,1952]" pageId="8" pageNumber="495">Thaye Chaung Township</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="9D06F833270A3201B1C6F814FED5F85C" box="[151,318,1962,1988]" pageId="8" pageNumber="495">Dawei District</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F833270A3201B016F814FDF4F85C" box="[327,543,1962,1988]" country="Myanmar" name="Taninthayi" pageId="8" pageNumber="495">Tanintharyi Region</collectingRegion>
|
||
(
|
||
<geoCoordinate id="3AF65016270A3201B37CF817FD04F85B" box="[557,751,1961,1987]" degrees="13.8671111111" direction="north" orientation="latitude" pageId="8" pageNumber="495" precision="1" value="13.867111">13.8671111111N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3AF65016270A3201B3A8F817FC52F85C" box="[761,953,1961,1988]" degrees="98.2836944444" direction="east" orientation="longitude" pageId="8" pageNumber="495" precision="1" value="98.28369">98.2836944444E</geoCoordinate>
|
||
; altitude
|
||
<quantity id="983A9B34270A3201B570F817FBB1F85C" box="[1057,1114,1961,1988]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="495" unit="m" value="45.0">
|
||
<elevation id="D4EFD1E2270A3201B570F817FBB1F85C" box="[1057,1114,1961,1988]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="495" unit="m" value="45.0">45 m</elevation>
|
||
</quantity>
|
||
asl.) collected on
|
||
<date id="2B7C1011270A3201B472F814FF24F870" pageId="8" pageNumber="495" value="2003-01-19">
|
||
<collectingDate id="3B38E9F9270A3201B472F814FF24F870" pageId="8" pageNumber="495" value="2003-01-19">19 January 2003</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="F2375307270A3201B1AAF870FEBBF870" box="[251,336,1998,2024]" pageId="8" pageNumber="495">Win H.</collectorName>
|
||
,
|
||
<collectorName id="F2375307270A3201B00CF870FE45F870" box="[349,430,1998,2024]" pageId="8" pageNumber="495">Tun H.</collectorName>
|
||
,
|
||
<collectorName id="F2375307270A3201B0EDF870FDD9F870" box="[444,562,1998,2024]" pageId="8" pageNumber="495">Lwin K.S.</collectorName>
|
||
,
|
||
<collectorName id="F2375307270A3201B311F870FD56F870" box="[576,701,1998,2024]" pageId="8" pageNumber="495">Shein A.K.</collectorName>
|
||
&
|
||
<collectorName id="F2375307270A3201B3B2F870FC8AF870" box="[739,865,1998,2024]" pageId="8" pageNumber="495">Naing Z.M</collectorName>
|
||
<emphasis id="6DB6EAC3270A3201B230F873FC08F87F" bold="true" box="[865,995,1997,2023]" pageId="8" pageNumber="495">
|
||
—
|
||
<materialsCitation id="EFAA3C8C270A3200B22CF873FDDCFF4D" collectingDate="2020-10-05" collectionCode="DTU" collectorName="T. V. Nguyen" country="Vietnam" elevation="1000" lastPageId="9" lastPageNumber="496" latitude="11.467058" location="Gung Re Commune" longLatPrecision="1" longitude="108.069084" municipality="Di Linh District" pageId="8" pageNumber="495" specimenCode="DTU 552" specimenCount="2" specimenCount-adult="1" specimenCount-male="1" stateProvince="Lam Dong Province">
|
||
<collectingCountry id="27D57641270A3201B22CF873FC08F87F" box="[893,995,1997,2023]" name="Vietnam" pageId="8" pageNumber="495">Vietnam</collectingCountry>
|
||
.
|
||
<specimenCode id="0F649EAA270A3201B2A3F870FBB5F870" box="[1010,1118,1997,2024]" pageId="8" pageNumber="495">DTU 552</specimenCode>
|
||
(
|
||
<specimenCount id="49C4FD58270A3201B53FF873FB43F87F" box="[1134,1192,1997,2023]" pageId="8" pageNumber="495" type="adult">adult</specimenCount>
|
||
<specimenCount id="49C4FD58270A3201B5E1F873FB03F87F" box="[1200,1256,1997,2023]" pageId="8" pageNumber="495" type="male">male</specimenCount>
|
||
) from
|
||
<location id="5A1D600A270A3200B466F870FEE4FF29" LSID="urn:lsid:plazi:treatment:D76B87C72705321CB1C6FB14FA6CFBC1:5A1D600A270A3200B466F870FEE4FF29" country="Vietnam" lastPageId="9" lastPageNumber="496" latitude="11.467058" longLatPrecision="1" longitude="108.069084" municipality="Di Linh District" name="Gung Re Commune" pageId="8" pageNumber="495" stateProvince="Lam Dong Province">Gung Re Commune</location>
|
||
,
|
||
<collectingMunicipality id="BF19ACAB270B3200B04CFF29FE32FF29" box="[285,473,151,177]" pageId="9" pageNumber="496">Di Linh District</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="9D06F833270B3200B0B6FF29FD3AFF29" box="[487,721,151,177]" country="Vietnam" name="Lam Dong" pageId="9" pageNumber="496">Lam Dong Province</collectingRegion>
|
||
(
|
||
<geoCoordinate id="3AF65016270B3200B3B3FF28FC97FF29" box="[738,892,150,177]" degrees="11.467058" direction="north" orientation="latitude" pageId="9" pageNumber="496" precision="1" value="11.467058">11.467058°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3AF65016270B3200B2DBFF29FBC7FF29" box="[906,1068,151,177]" degrees="108.069084" direction="east" orientation="longitude" pageId="9" pageNumber="496" precision="1" value="108.069084">108.069084°E</geoCoordinate>
|
||
; altitude
|
||
<quantity id="983A9B34270B3200B5CDFF29FB1FFF29" box="[1180,1268,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="496" unit="m" value="1000.0">
|
||
<elevation id="D4EFD1E2270B3200B5CDFF29FB1FFF29" box="[1180,1268,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="496" unit="m" value="1000.0">1000 m</elevation>
|
||
</quantity>
|
||
asl.) collected on
|
||
<date id="2B7C1011270B3200B1ECFF04FE9BFF4D" box="[189,368,186,213]" pageId="9" pageNumber="496" value="2020-10-05">
|
||
<collectingDate id="3B38E9F9270B3200B1ECFF04FE9BFF4D" box="[189,368,186,213]" pageId="9" pageNumber="496" value="2020-10-05">5 October 2020</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="F2375307270B3200B0CEFF05FDD8FF4D" box="[415,563,187,213]" pageId="9" pageNumber="496">T.V. Nguyen</collectorName>
|
||
;
|
||
</materialsCitation>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
<materialsCitation id="EFAA3C8C270B3200B310FF05FAD3FF61" collectingDate="2021-07-15" collectionCode="DTU" collectorName="T. V. Nguyen" elevation="585" latitude="13.885446" location="Ia Pech Commune" longLatPrecision="1" longitude="107.82866" municipality="Ia Glai District" pageId="9" pageNumber="496" specimenCode="DTU 325" specimenCount="1" stateProvince="Gia Lai Province">
|
||
<specimenCode id="0F649EAA270B3200B310FF05FD45FF4D" box="[577,686,186,213]" pageId="9" pageNumber="496">DTU 325</specimenCode>
|
||
(
|
||
<specimenCount id="49C4FD58270B3200B3EEFF05FCBAFF4D" box="[703,849,187,213]" pageId="9" pageNumber="496" type="adult">adult female</specimenCount>
|
||
) from
|
||
<location id="5A1D600A270B3200B2F3FF05FB91FF4D" LSID="urn:lsid:plazi:treatment:D76B87C72705321CB1C6FB14FA6CFBC1:5A1D600A270B3200B2F3FF05FB91FF4D" box="[930,1146,187,213]" latitude="13.885446" longLatPrecision="1" longitude="107.82866" municipality="Ia Glai District" name="Ia Pech Commune" pageId="9" pageNumber="496" stateProvince="Gia Lai Province">Ia Pech Commune</location>
|
||
,
|
||
<collectingMunicipality id="BF19ACAB270B3200B5D6FF05FADCFF4D" box="[1159,1335,187,213]" pageId="9" pageNumber="496">Ia Glai District</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="9D06F833270B3200B414FF05FF10FF61" country="Vietnam" name="Gia Lai" pageId="9" pageNumber="496">Gia Lai Province</collectingRegion>
|
||
(
|
||
<geoCoordinate id="3AF65016270B3200B05AFF60FE4DFF61" box="[267,422,222,249]" degrees="13.885446" direction="north" orientation="latitude" pageId="9" pageNumber="496" precision="1" value="13.885446">13.885446°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3AF65016270B3200B0E2FF61FDBEFF61" box="[435,597,223,249]" degrees="107.828661" direction="east" orientation="longitude" pageId="9" pageNumber="496" precision="1" value="107.82866">107.828661°E</geoCoordinate>
|
||
; altitude
|
||
<quantity id="983A9B34270B3200B393FF60FCE0FF61" box="[706,779,222,249]" metricMagnitude="2" metricUnit="m" metricValue="5.85" pageId="9" pageNumber="496" unit="m" value="585.0">
|
||
<elevation id="D4EFD1E2270B3200B393FF60FCE0FF61" box="[706,779,222,249]" metricMagnitude="2" metricUnit="m" metricValue="5.85" pageId="9" pageNumber="496" unit="m" value="585.0">585 m</elevation>
|
||
</quantity>
|
||
asl.) collected on
|
||
<date id="2B7C1011270B3200B28EFF60FB9FFF61" box="[991,1140,222,249]" pageId="9" pageNumber="496" value="2021-07-15">
|
||
<collectingDate id="3B38E9F9270B3200B28EFF60FB9FFF61" box="[991,1140,222,249]" pageId="9" pageNumber="496" value="2021-07-15">15 July 2021</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="F2375307270B3200B5F0FF61FADFFF61" box="[1185,1332,223,249]" pageId="9" pageNumber="496">T.V. Nguyen</collectorName>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="EFAA3C8C270B3200B410FF61FC9FFE85" collectionCode="NHMW" location="Tam Dao NP" pageId="9" pageNumber="496" specimenCode="NHMW 39658" specimenCount="1" stateProvince="Vinh Phuc Province">
|
||
<specimenCode id="0F649EAA270B3200B410FF61FF0DFE85" pageId="9" pageNumber="496">NHMW 39658</specimenCode>
|
||
:1 (
|
||
<specimenCount id="49C4FD58270B3200B053FEBDFE7BFE85" box="[258,400,259,285]" pageId="9" pageNumber="496" type="adult">adult female</specimenCount>
|
||
) from
|
||
<location id="5A1D600A270B3200B08DFEBDFD85FE85" LSID="urn:lsid:plazi:treatment:D76B87C72705321CB1C6FB14FA6CFBC1:5A1D600A270B3200B08DFEBDFD85FE85" box="[476,622,259,285]" name="Tam Dao NP" pageId="9" pageNumber="496" stateProvince="Vinh Phuc Province">Tam Dao NP</location>
|
||
,
|
||
<collectingRegion id="9D06F833270B3200B329FEBDFCB3FE85" box="[632,856,259,285]" country="Vietnam" name="Vinh Phuc" pageId="9" pageNumber="496">Vinh Phuc Province</collectingRegion>
|
||
—
|
||
</materialsCitation>
|
||
<materialsCitation id="EFAA3C8C270B3200B225FEBCFE91FEFD" collectingDate="2018-05-13" collectionCode="RBINS" country="Cambodia" elevation="55" latitude="13.01917" location="Sambour District" longLatPrecision="1" longitude="105.92712" pageId="9" pageNumber="496" specimenCode="RBINS 2738, RBINS 18563" specimenCount="1" stateProvince="Kratie Province">
|
||
<emphasis id="6DB6EAC3270B3200B225FEBCFC1BFE84" bold="true" box="[884,1008,258,284]" pageId="9" pageNumber="496">
|
||
<collectingCountry id="27D57641270B3200B225FEBCFC1BFE84" box="[884,1008,258,284]" name="Cambodia" pageId="9" pageNumber="496">Cambodia</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<specimenCode id="0F649EAA270B3200B2AEFEBDFB7AFE85" box="[1023,1169,259,285]" pageId="9" pageNumber="496">RBINS 2738</specimenCode>
|
||
(
|
||
<specimenCount id="49C4FD58270B3200B5CEFEBDFAC4FE85" box="[1183,1327,259,285]" pageId="9" pageNumber="496" type="adult">adult female</specimenCount>
|
||
, formerly under
|
||
<specimenCode id="0F649EAA270B3200B1B1FE99FE69FED9" box="[224,386,294,321]" pageId="9" pageNumber="496">RBINS 18563</specimenCode>
|
||
) from
|
||
<location id="5A1D600A270B3200B083FE99FD7CFED9" LSID="urn:lsid:plazi:treatment:D76B87C72705321CB1C6FB14FA6CFBC1:5A1D600A270B3200B083FE99FD7CFED9" box="[466,663,295,321]" country="Cambodia" latitude="13.01917" longLatPrecision="1" longitude="105.92712" name="Sambour District" pageId="9" pageNumber="496" stateProvince="Kratie Province">Sambour District</location>
|
||
,
|
||
<collectingRegion id="9D06F833270B3200B3F5FE99FCBCFED9" box="[676,855,295,321]" country="Cambodia" name="Kratie" pageId="9" pageNumber="496">Kratie Province</collectingRegion>
|
||
(
|
||
<geoCoordinate id="3AF65016270B3200B236FE99FC03FED9" box="[871,1000,295,321]" degrees="13.01917" direction="north" orientation="latitude" pageId="9" pageNumber="496" precision="1" value="13.01917">13.01917N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3AF65016270B3200B2A4FE98FB96FED9" box="[1013,1149,294,321]" degrees="105.92712" direction="east" orientation="longitude" pageId="9" pageNumber="496" precision="1" value="105.92712">105.92712E</geoCoordinate>
|
||
; altitude
|
||
<quantity id="983A9B34270B3200B5BAFE98FACDFED9" box="[1259,1318,294,321]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="496" unit="m" value="55.0">
|
||
<elevation id="D4EFD1E2270B3200B5BAFE98FACDFED9" box="[1259,1318,294,321]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="496" unit="m" value="55.0">55 m</elevation>
|
||
</quantity>
|
||
) collected on
|
||
<date id="2B7C1011270B3200B1ECFEF5FEB2FEFD" box="[189,345,331,357]" pageId="9" pageNumber="496" value="2018-05-13">
|
||
<collectingDate id="3B38E9F9270B3200B1ECFEF5FEB2FEFD" box="[189,345,331,357]" pageId="9" pageNumber="496" value="2018-05-13">13 May 2018</collectingDate>
|
||
</date>
|
||
.—
|
||
</materialsCitation>
|
||
<materialsCitation id="EFAA3C8C270B3200B02BFEF4FE37FE11" collectingDate="1962" collectionCode="USNM" country="Thailand" latitude="18.7903" location="Chiang Mai Province" longLatPrecision="7" longitude="98.9972" pageId="9" pageNumber="496" specimenCode="USNM 164427" specimenCount="1" stateProvince="Chiang Mai Province">
|
||
<emphasis id="6DB6EAC3270B3200B02BFEF4FE0CFEFC" bold="true" box="[378,487,330,356]" pageId="9" pageNumber="496">
|
||
<collectingCountry id="27D57641270B3200B02BFEF4FE0CFEFC" box="[378,487,330,356]" name="Thailand" pageId="9" pageNumber="496">Thailand</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<specimenCode id="0F649EAA270B3200B0A6FEF5FD4EFEFD" box="[503,677,331,357]" pageId="9" pageNumber="496">USNM 164427</specimenCode>
|
||
(
|
||
<specimenCount id="49C4FD58270B3200B3E7FEF5FCDAFEFD" box="[694,817,331,357]" pageId="9" pageNumber="496" type="adult">adult male</specimenCount>
|
||
) from
|
||
<collectingRegion id="9D06F833270B3200B2D3FEF5FB92FEFD" box="[898,1145,331,357]" country="Thailand" name="Chiang Mai" pageId="9" pageNumber="496">Chiang Mai Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="27D57641270B3200B5D6FEF5FB00FEFD" box="[1159,1259,331,357]" name="Thailand" pageId="9" pageNumber="496">Thailand</collectingCountry>
|
||
(ca.
|
||
<geoCoordinate id="3AF65016270B3200B477FEF5FA73FEFD" box="[1318,1432,331,357]" degrees="18.7903" direction="north" orientation="latitude" pageId="9" pageNumber="496" precision="5" value="18.7903">18.7903N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3AF65016270B3200B1C6FED1FEE8FE11" box="[151,259,367,393]" degrees="98.9972" direction="east" orientation="longitude" pageId="9" pageNumber="496" precision="5" value="98.9972">98.9972E</geoCoordinate>
|
||
) collected in
|
||
<collectingDate id="3B38E9F9270B3200B0CDFED1FE3CFE11" box="[412,471,367,393]" pageId="9" pageNumber="496">1962</collectingDate>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="0BBD6659270B3200B1C6F977FBCBF8D6" ID-DOI="http://doi.org/10.5281/zenodo.13916223" ID-Zenodo-Dep="13916223" httpUri="https://zenodo.org/record/13916223/files/figure.png" pageId="9" pageNumber="496" startId="9.[151,250,1737,1761]" targetBox="[151,1436,417,1714]" targetPageId="9" targetType="figure">
|
||
<paragraph id="5F7D36D1270B3200B1C6F977FBCBF8D6" blockId="9.[151,1436,1737,1870]" pageId="9" pageNumber="496">
|
||
<emphasis id="6DB6EAC3270B3200B1C6F977FEE5F979" bold="true" box="[151,270,1737,1761]" pageId="9" pageNumber="496">FIGURE 2</emphasis>
|
||
. Photographs of
|
||
<taxonomicName id="98C24D52270B3200B0EDF977FD6FF979" authorityName="Boie" authorityYear="1827" box="[444,644,1737,1761]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="496" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC3270B3200B0EDF977FD6FF979" box="[444,644,1737,1761]" italics="true" pageId="9" pageNumber="496">Lycodon subcinctus</emphasis>
|
||
</taxonomicName>
|
||
in preservative. A, B—General dorsal and ventral views of MNHN 1884.120 (Syntype of
|
||
<taxonomicName id="98C24D52270B3200B044F953FE04F89D" authorityName="Sauvage" authorityYear="1884" box="[277,495,1773,1797]" class="Reptilia" family="Colubridae" genus="Elapsoides" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="496" phylum="Chordata" rank="species" species="annulatus">
|
||
<emphasis id="6DB6EAC3270B3200B044F953FE04F89D" box="[277,495,1773,1797]" italics="true" pageId="9" pageNumber="496">Elapsoides annulatus</emphasis>
|
||
</taxonomicName>
|
||
, subadult female); C, D—General dorsal and ventral views of MNHN 1884.121 (syntype of
|
||
<taxonomicName id="98C24D52270B3200B1C6F8AFFE84F8B1" authorityName="Sauvage" authorityYear="1884" box="[151,367,1809,1833]" class="Reptilia" family="Colubridae" genus="Elapsoides" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="496" phylum="Chordata" rank="species" species="annulatus">
|
||
<emphasis id="6DB6EAC3270B3200B1C6F8AFFE84F8B1" box="[151,367,1809,1833]" italics="true" pageId="9" pageNumber="496">Elapsoides annulatus</emphasis>
|
||
</taxonomicName>
|
||
, subadult female); E, F—General dorsal and ventral views of RBINS 2610 (neotype of
|
||
<taxonomicName id="98C24D52270B3200B585F8AFFA7DF8B1" authorityName="Nutphand" authorityYear="1986" box="[1236,1430,1809,1833]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="496" phylum="Chordata" rank="species" species="suratensis">
|
||
<emphasis id="6DB6EAC3270B3200B585F8AFFA7DF8B1" box="[1236,1430,1809,1833]" italics="true" pageId="9" pageNumber="496">Lycodon suratensis</emphasis>
|
||
</taxonomicName>
|
||
, adult female). Images not to scale. Photos by G. Vogel (A–D) and O.S.G. Pauwels (E–F).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5F7D36D1270B3203B196F827FC9BFE35" blockId="9.[151,1437,1945,2044]" lastBlockId="10.[151,1437,150,429]" lastPageId="10" lastPageNumber="497" pageId="9" pageNumber="496">
|
||
<emphasis id="6DB6EAC3270B3200B196F827FDE6F82B" bold="true" box="[199,525,1945,1971]" pageId="9" pageNumber="496">Additional material (n = 12)</emphasis>
|
||
:
|
||
<emphasis id="6DB6EAC3270B3200B34BF827FD88F82B" bold="true" box="[538,611,1945,1971]" pageId="9" pageNumber="496">
|
||
<collectingCountry id="27D57641270B3200B34BF827FD88F82B" box="[538,611,1945,1971]" name="China" pageId="9" pageNumber="496">China</collectingCountry>
|
||
</emphasis>
|
||
. MCZ R-176514–15, R-177133 (
|
||
<specimenCount id="49C4FD58270B3200B282F827FB92F82B" box="[979,1145,1945,1971]" count="3" pageId="9" pageNumber="496" type="juvenile">three juveniles</specimenCount>
|
||
, sex unknown) from Hong Kong SAR.—
|
||
<emphasis id="6DB6EAC3270B3200B066F803FE44F84F" bold="true" box="[311,431,1981,2007]" pageId="9" pageNumber="496">
|
||
<collectingCountry id="27D57641270B3200B066F803FE44F84F" box="[311,431,1981,2007]" name="Myanmar" pageId="9" pageNumber="496">Myanmar</collectingCountry>
|
||
</emphasis>
|
||
. CAS 245971 (
|
||
<specimenCount id="49C4FD58270B3200B30EF803FD06F84F" box="[607,749,1981,2007]" count="1" pageId="9" pageNumber="496" type="juvenile">one juvenile</specimenCount>
|
||
, sex unknow) collected from the
|
||
<collectingRegion id="9D06F833270B3200B537F803FB0CF84F" box="[1126,1255,1981,2007]" country="Myanmar" name="Taninthayi" pageId="9" pageNumber="496">Tanintharyi</collectingRegion>
|
||
Nature Reserve along a pipeline road near the vicinity of Khotama military camp, Yebyu Township, Dawei District,
|
||
<collectingRegion id="9D06F833270B3200B44DF85FFA76F863" box="[1308,1437,2017,2043]" country="Myanmar" name="Taninthayi" pageId="9" pageNumber="496">Tanintharyi</collectingRegion>
|
||
Region—
|
||
<emphasis id="6DB6EAC327083203B055FF28FE81FF28" bold="true" box="[260,362,150,176]" pageId="10" pageNumber="497">
|
||
<collectingCountry id="27D5764127083203B055FF28FE81FF28" box="[260,362,150,176]" name="Vietnam" pageId="10" pageNumber="497">Vietnam</collectingCountry>
|
||
</emphasis>
|
||
. SIFASV 98 (
|
||
<specimenCount id="49C4FD5827083203B352FF29FD80FF29" box="[515,619,151,177]" count="1" pageId="10" pageNumber="497" type="adult">one adult</specimenCount>
|
||
female, released) from Tan Thai Commune, Dai Tu District,
|
||
<collectingRegion id="9D06F83327083203B45AFF29FF15FF4D" country="Vietnam" name="Thai Nguyen" pageId="10" pageNumber="497">Thai Nguyen Province</collectingRegion>
|
||
; SIFASV 97 (
|
||
<specimenCount id="49C4FD5827083203B0C6FF05FDEBFF4D" box="[407,512,187,213]" count="1" pageId="10" pageNumber="497" type="adult">one adult</specimenCount>
|
||
female, released) from Ba Na-Nui Chua NR,
|
||
<collectingRegion id="9D06F83327083203B2ACFF05FB8AFF4D" box="[1021,1121,187,213]" country="Vietnam" name="Da Nang" pageId="10" pageNumber="497">Da Nang</collectingRegion>
|
||
City; SIFASV 99 (
|
||
<specimenCount id="49C4FD5827083203B465FF05FA77FF4D" box="[1332,1436,187,213]" count="1" pageId="10" pageNumber="497" type="adult">one adult</specimenCount>
|
||
female, released) from Pu Mat NP,
|
||
<collectingRegion id="9D06F83327083203B377FF61FD12FF61" box="[550,761,223,249]" country="Vietnam" name="Nghe An" pageId="10" pageNumber="497">Nghe An Province</collectingRegion>
|
||
; SIFASV 101 (
|
||
<specimenCount id="49C4FD5827083203B2F5FF61FBE5FF61" box="[932,1038,223,249]" count="1" pageId="10" pageNumber="497" type="adult">one adult</specimenCount>
|
||
female, released) from Pu Mat NP,
|
||
<collectingRegion id="9D06F83327083203B1C6FEBDFE82FE85" box="[151,361,259,285]" country="Vietnam" name="Nghe An" pageId="10" pageNumber="497">Nghe An Province</collectingRegion>
|
||
; SIFASV 102 (adult female, released) from Nui Chua NP,
|
||
<collectingRegion id="9D06F83327083203B2A2FEBDFB0DFE85" box="[1011,1254,259,285]" country="Vietnam" name="Ninh Thuan" pageId="10" pageNumber="497">Ninh Thuan Province</collectingRegion>
|
||
; USNM 166987 (
|
||
<specimenCount id="49C4FD5827083203B1CEFE99FEC4FED9" box="[159,303,295,321]" count="1" pageId="10" pageNumber="497" type="juvenile">one juvenile</specimenCount>
|
||
, sex unknown) from Bien Hoa City,
|
||
<collectingRegion id="9D06F83327083203B38CFE99FC56FED9" box="[733,957,295,321]" country="Vietnam" name="Dong Nai" pageId="10" pageNumber="497">Dong Nai Province</collectingRegion>
|
||
; SIFASV 96 (
|
||
<specimenCount id="49C4FD5827083203B531FE99FB1BFED9" box="[1120,1264,295,321]" count="1" pageId="10" pageNumber="497" type="juvenile">one juvenile</specimenCount>
|
||
, sex unknown, released) from Phu Quoc NP,
|
||
<collectingRegion id="9D06F83327083203B0B6FEF5FD38FEFD" box="[487,723,331,357]" country="Vietnam" name="Kien Giang" pageId="10" pageNumber="497">Kien Giang Province</collectingRegion>
|
||
—
|
||
<emphasis id="6DB6EAC327083203B3BEFEF4FCB7FEFC" bold="true" box="[751,860,330,356]" pageId="10" pageNumber="497">
|
||
<collectingCountry id="27D5764127083203B3BEFEF4FCB7FEFC" box="[751,860,330,356]" name="Thailand" pageId="10" pageNumber="497">Thailand</collectingCountry>
|
||
</emphasis>
|
||
. FMNH 180149 (
|
||
<specimenCount id="49C4FD5827083203B574FEF5FB5FFEFD" box="[1061,1204,331,357]" count="1" pageId="10" pageNumber="497" type="juvenile">one juvenile</specimenCount>
|
||
, male) from
|
||
<collectingRegion id="9D06F83327083203B415FEF5FE6DFE11" country="Thailand" name="Nakhon Ratchasima" pageId="10" pageNumber="497">Nakhon Ratchasima Province</collectingRegion>
|
||
collected on
|
||
<date id="2B7C101127083203B34FFED1FD46FE11" box="[542,685,367,393]" pageId="10" pageNumber="497" value="1969-04-06">6 April 1969</date>
|
||
by W. R. Heyer; RBINS 16990 (
|
||
<specimenCount id="49C4FD5827083203B575FED1FB58FE11" box="[1060,1203,367,393]" count="1" pageId="10" pageNumber="497" type="juvenile">one juvenile</specimenCount>
|
||
, sex unknown) from Sakaerat Biosphere Reserve Trail,
|
||
<collectingRegion id="9D06F83327083203B34AFE2DFC87FE35" box="[539,876,403,429]" country="Thailand" name="Nakhon Ratchasima" pageId="10" pageNumber="497">Nakhon Ratchasima Province</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<caption id="0BBD665927083203B1C6F940FBACF81A" ID-DOI="http://doi.org/10.5281/zenodo.13916225" ID-Zenodo-Dep="13916225" httpUri="https://zenodo.org/record/13916225/files/figure.png" pageId="10" pageNumber="497" startId="10.[151,250,1790,1814]" targetBox="[153,1423,463,1754]" targetPageId="10" targetType="figure">
|
||
<paragraph id="5F7D36D127083203B1C6F940FBACF81A" blockId="10.[151,1436,1790,1922]" pageId="10" pageNumber="497">
|
||
<emphasis id="6DB6EAC327083203B1C6F940FEE5F88E" bold="true" box="[151,270,1790,1814]" pageId="10" pageNumber="497">FIGURE 3</emphasis>
|
||
. Bayesian phylogenetic tree of
|
||
<taxonomicName id="98C24D5227083203B304F940FD40F88E" authorityName="Fitzinger" authorityYear="1826" box="[597,683,1790,1814]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="497" phylum="Chordata" rank="genus">
|
||
<emphasis id="6DB6EAC327083203B304F940FD40F88E" box="[597,683,1790,1814]" italics="true" pageId="10" pageNumber="497">Lycodon</emphasis>
|
||
</taxonomicName>
|
||
inferred from the mitochondrial cytb sequences. Numbers before slashes indicate Bayesian posterior probabilities and numbers after slashes indicate ML bootstrap supports. Nodes filled in with a black circle indicate strong support by both the ML and BI analyses, whereas nodes filled in with a white circle indicate strong support only in the BI analysis. Inset photo of
|
||
<taxonomicName id="98C24D5227083203B34AF8D4FCEEF81A" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[539,773,1898,1922]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="497" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC327083203B34AF8D4FCEEF81A" box="[539,773,1898,1922]" italics="true" pageId="10" pageNumber="497">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B827083203B25AF8D4FCB6F819" box="[779,861,1898,1922]" pageId="10" pageNumber="497" rank="species">
|
||
<emphasis id="6DB6EAC327083203B25AF8D4FCC8F81A" bold="true" box="[779,803,1898,1922]" pageId="10" pageNumber="497">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327083203B27EF8D4FCBDF81A" bold="true" box="[815,854,1898,1922]" pageId="10" pageNumber="497">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
taken by A.M. Bragin.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0BBD665927093202B1C6FC35FCE8FB88" ID-DOI="http://doi.org/10.5281/zenodo.13916227" ID-Zenodo-Dep="13916227" httpUri="https://zenodo.org/record/13916227/files/figure.png" pageId="11" pageNumber="498" startId="11.[151,250,907,931]" targetBox="[151,1436,188,896]" targetPageId="11" targetType="figure">
|
||
<paragraph id="5F7D36D127093202B1C6FC35FCE8FB88" blockId="11.[151,1436,907,1040]" pageId="11" pageNumber="498">
|
||
<emphasis id="6DB6EAC327093202B1C6FC35FEE4FC3C" bold="true" box="[151,271,907,932]" pageId="11" pageNumber="498">FIGURE 4</emphasis>
|
||
. A statistical summary of the
|
||
<taxonomicName id="98C24D5227093202B31BFC32FD39FC3C" box="[586,722,908,932]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="498" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC327093202B31BFC32FDB3FC3B" box="[586,600,908,931]" italics="true" pageId="11" pageNumber="498">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327093202B336FC32FD39FC3C" box="[615,722,908,932]" italics="true" pageId="11" pageNumber="498">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
species complex. (
|
||
<emphasis id="6DB6EAC327093202B2CBFC35FC47FC3B" bold="true" box="[922,940,907,931]" pageId="11" pageNumber="498">A</emphasis>
|
||
) principal components analysis (PCA) showing three discrete clusters corresponding to
|
||
<taxonomicName id="98C24D5227093202B37EFC0EFCF1FC50" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[559,794,944,968]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="498" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC327093202B37EFC0EFCF1FC50" box="[559,794,944,968]" italics="true" pageId="11" pageNumber="498">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B827093202B270FC0EFC9FFC5F" box="[801,884,944,968]" pageId="11" pageNumber="498" rank="species">
|
||
<emphasis id="6DB6EAC327093202B270FC0EFCD2FC50" bold="true" box="[801,825,944,968]" pageId="11" pageNumber="498">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327093202B217FC0EFC86FC50" bold="true" box="[838,877,944,968]" pageId="11" pageNumber="498">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="98C24D5227093202B2D3FC0EFC32FC50" box="[898,985,944,968]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="498" phylum="Chordata" rank="species" species="sealei">
|
||
<emphasis id="6DB6EAC327093202B2D3FC0EFC7BFC5F" box="[898,912,944,967]" italics="true" pageId="11" pageNumber="498">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327093202B2CFFC0EFC32FC50" box="[926,985,944,968]" italics="true" pageId="11" pageNumber="498">sealei</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="98C24D5227093202B543FC0EFB72FC50" box="[1042,1177,944,968]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="498" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC327093202B543FC0EFBCBFC5F" box="[1042,1056,944,967]" italics="true" pageId="11" pageNumber="498">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327093202B57FFC0EFB72FC50" box="[1070,1177,944,968]" italics="true" pageId="11" pageNumber="498">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
, respectively. (B–E) Box and whisker plots showing statistically significant differences in the number of ventrals (B), subcaudals (C), total body scales (D) and subcaudal ratio values (E) between the three species.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5F7D36D127093202B196FBDCFBF0F924" blockId="11.[151,1437,1122,2012]" pageId="11" pageNumber="498">
|
||
<emphasis id="6DB6EAC327093202B196FBDCFD1CFBE4" bold="true" box="[199,759,1122,1148]" pageId="11" pageNumber="498">Referred material from the literature (n = 24)</emphasis>
|
||
.—
|
||
<emphasis id="6DB6EAC327093202B24BFBDCFC6BFBE4" bold="true" box="[794,896,1122,1148]" pageId="11" pageNumber="498">
|
||
<collectingCountry id="27D5764127093202B24BFBDCFC6BFBE4" box="[794,896,1122,1148]" name="Vietnam" pageId="11" pageNumber="498">Vietnam</collectingCountry>
|
||
</emphasis>
|
||
. IEBR 4757 (
|
||
<specimenCount id="49C4FD5827093202B573FBDCFB65FBE4" box="[1058,1166,1122,1148]" count="1" pageId="11" pageNumber="498" type="adult">one adult</specimenCount>
|
||
female) from Na Hang NR,
|
||
<collectingRegion id="9D06F83327093202B19FFB38FE3CFB38" box="[206,471,1158,1184]" country="Vietnam" name="Tuyen Quang" pageId="11" pageNumber="498">Tuyen Quang Province</collectingRegion>
|
||
(see details in
|
||
<bibRefCitation id="3B534B2027093202B3DBFB38FCD9FB38" author="Le, D. T. & Luong, A. M. & Pham, C. T. & Pham, T. Q. & Nguyen, S. L. H. & Ziegler, T. & Nguyen, T. Q." box="[650,818,1158,1184]" pageId="11" pageNumber="498" pagination="201 - 219" refId="ref20602" refString="Le, D. T., Luong, A. M., Pham, C. T., Pham, T. Q., Nguyen, S. L. H., Ziegler, T. & Nguyen, T. Q. (2021) New records and an updated checklist of amphibians and snakes from Tuyen Quang Province, Vietnam. Bonn zoological Bulletin, 70 (1), 201 - 219. https: // doi. org / 10.20363 / BZB- 2021.70.1.201" type="journal article" year="2021">
|
||
Le
|
||
<emphasis id="6DB6EAC327093202B3E0FB39FD0EFB38" box="[689,741,1158,1184]" italics="true" pageId="11" pageNumber="498">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
); IEBR A.2010.42 (
|
||
<specimenCount id="49C4FD5827093202B548FB38FB6EFB38" box="[1049,1157,1158,1184]" count="1" pageId="11" pageNumber="498" type="adult">one adult</specimenCount>
|
||
male), ZFMK 91899 & IEBR A.2010.43 (
|
||
<specimenCount id="49C4FD5827093202B033FB14FE17FB5C" box="[354,508,1194,1220]" count="2" pageId="11" pageNumber="498" type="juvenile">two juveniles</specimenCount>
|
||
, sex unknown) from Cat Tien NP,
|
||
<collectingRegion id="9D06F83327093202B2D1FB14FBB2FB5C" box="[896,1113,1194,1220]" country="Vietnam" name="Dong Nai" pageId="11" pageNumber="498">Dong Nai Province</collectingRegion>
|
||
(see details in
|
||
<bibRefCitation id="3B534B2027093202B450FB14FF25FB70" author="Geissler, P. & Nguyen, T. Q. & Poyarkov, N. A. & Bohme, W." pageId="11" pageNumber="498" pagination="9 - 16" refId="ref19433" refString="Geissler, P., Nguyen, T. Q., Poyarkov, N. A. & Bohme, W. (2011) New records of snakes from Cat Tien National Park Dong Nai and Lam Dong provinces southern Vietnam. Bonn Zoological Bulletin, 60 (1), 9 - 16." type="journal article" year="2011">
|
||
Geissler
|
||
<emphasis id="6DB6EAC327093202B435FB15FA7EFB5C" box="[1380,1429,1194,1220]" italics="true" pageId="11" pageNumber="498">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
)—
|
||
<emphasis id="6DB6EAC327093202B1A2FB70FED7FB70" bold="true" box="[243,316,1230,1256]" pageId="11" pageNumber="498">
|
||
<collectingCountry id="27D5764127093202B1A2FB70FED7FB70" box="[243,316,1230,1256]" name="China" pageId="11" pageNumber="498">China</collectingCountry>
|
||
</emphasis>
|
||
. NHMUK 1937.2.1.4 (
|
||
<specimenCount id="49C4FD5827093202B36FFB70FD27FB70" box="[574,716,1230,1256]" count="1" pageId="11" pageNumber="498" type="juvenile">one juvenile</specimenCount>
|
||
, sex unknown) from Qionghai City,
|
||
<collectingRegion id="9D06F83327093202B50EFB70FAF1FB70" box="[1119,1306,1230,1256]" country="China" name="Hainan" pageId="11" pageNumber="498">Hainan Province</collectingRegion>
|
||
(see details in
|
||
<bibRefCitation id="3B534B2027093202B1E5FB4CFED1FA94" author="Smith, M. A." box="[180,314,1266,1292]" pageId="11" pageNumber="498" pagination="195 - 212" refId="ref22878" refString="Smith, M. A. (1923) On a collection of reptiles and batrachians from the island of Hainan. Journal of the Natural History Society of Siam, 6, 195 - 212." type="journal article" year="1923">Smith 1923</bibRefCitation>
|
||
); AMNH R-27755 (
|
||
<specimenCount id="49C4FD5827093202B34BFB4CFD68FA94" box="[538,643,1266,1292]" count="1" pageId="11" pageNumber="498" type="adult">one adult</specimenCount>
|
||
female) from Nada Town, Danzhou City,
|
||
<collectingRegion id="9D06F83327093202B50DFB4CFAF3FA94" box="[1116,1304,1266,1292]" country="China" name="Hainan" pageId="11" pageNumber="498">Hainan Province</collectingRegion>
|
||
(see details in
|
||
<bibRefCitation id="3B534B2027093202B1E2FAA9FEC6FAA9" author="Pope, C. H." box="[179,301,1302,1329]" pageId="11" pageNumber="498" refId="ref21916" refString="Pope, C. H. (1935) The reptiles of China. Turtles, crocodilians, snakes, lizards. Natural History of Central Asia. Vol. 10. American Museum of Natural History, New York, New York, 604 pp. https: // doi. org / 10.5962 / bhl. title. 12104" type="book" year="1935">Pope 1935</bibRefCitation>
|
||
); No. 58 (
|
||
<specimenCount id="49C4FD5827093202B0C9FAA8FDEBFAA8" box="[408,512,1302,1328]" count="1" pageId="11" pageNumber="498" type="adult">one adult</specimenCount>
|
||
male) & No. 10 (
|
||
<specimenCount id="49C4FD5827093202B3EEFAA8FCA7FAA8" box="[703,844,1302,1328]" count="1" pageId="11" pageNumber="498" type="juvenile">one juvenile</specimenCount>
|
||
, sex unknown) from
|
||
<collectingRegion id="9D06F83327093202B564FAA8FAEAFAA8" box="[1077,1281,1302,1328]" country="China" name="Guangxi" pageId="11" pageNumber="498">Guangxi Province</collectingRegion>
|
||
, No. 732 (
|
||
<specimenCount id="49C4FD5827093202B423FAA8FF24FACC" count="1" pageId="11" pageNumber="498" type="adult">one adult</specimenCount>
|
||
male) & No. 197, No. 655122, No. 524 (
|
||
<specimenCount id="49C4FD5827093202B3C7FA84FCE5FACC" box="[662,782,1338,1364]" count="3" pageId="11" pageNumber="498" type="adult">three adult</specimenCount>
|
||
females) from
|
||
<collectingRegion id="9D06F83327093202B2E9FA84FB9EFACC" box="[952,1141,1338,1364]" country="China" name="Hainan" pageId="11" pageNumber="498">Hainan Province</collectingRegion>
|
||
, No. 452 & No. 2701 (
|
||
<specimenCount id="49C4FD5827093202B420FA84FF14FAE0" count="2" pageId="11" pageNumber="498" type="juvenile">two juveniles</specimenCount>
|
||
, sex unknown) from
|
||
<collectingRegion id="9D06F83327093202B0BBFAE0FD70FAE0" box="[490,667,1374,1400]" country="China" name="Fujian" pageId="11" pageNumber="498">Fujian Province</collectingRegion>
|
||
(see details in
|
||
<bibRefCitation id="3B534B2027093202B211FAE0FC12FAE0" author="Zhao, E. M. & Huang, M. H. & Zong, Y. & Jiang, Y. M. & Huang, Q. Y. & Zhao, H. & Ma, J. F. & Zheng, J. & Huang, Z. J. & Wei, G. & Yang, D. Q. & Li, D. J." box="[832,1017,1374,1400]" pageId="11" pageNumber="498" refId="ref24466" refString="Zhao, E. M., Huang, M. H., Zong, Y., Jiang, Y. M., Huang, Q. Y., Zhao, H., Ma, J. F., Zheng, J., Huang, Z. J., Wei, G., Yang, D. Q. & Li, D. J. (1998) Fauna Sinica Reptilia. Vol. 3. Squamata: Serpentes. Science Press, Beijing, 522 pp. [in Chinese]" type="book" year="1998">
|
||
Zhao
|
||
<emphasis id="6DB6EAC327093202B2D1FAE1FC5BFAE0" box="[896,944,1374,1400]" italics="true" pageId="11" pageNumber="498">et al</emphasis>
|
||
. 1998
|
||
</bibRefCitation>
|
||
); KIZ 2023031 (
|
||
<specimenCount id="49C4FD5827093202B5E0FAE0FAF1FAE0" box="[1201,1306,1374,1400]" count="1" pageId="11" pageNumber="498" type="adult">one adult</specimenCount>
|
||
male) from Mengla County, Xishuangbanna Dai Autonomous Prefecture,
|
||
<collectingRegion id="9D06F83327093202B214FA3CFBEDFA04" box="[837,1030,1410,1436]" country="China" name="Yunnan" pageId="11" pageNumber="498">Yunnan Province</collectingRegion>
|
||
, KIZ 2023034 (
|
||
<specimenCount id="49C4FD5827093202B5E2FA3CFAF0FA04" box="[1203,1307,1410,1436]" count="1" pageId="11" pageNumber="498" type="adult">one adult</specimenCount>
|
||
male) from Jinghong County, Xishuangbanna Dai Autonomous Prefecture,
|
||
<collectingRegion id="9D06F83327093202B207FA19FBFCFA58" box="[854,1047,1446,1473]" country="China" name="Yunnan" pageId="11" pageNumber="498">Yunnan Province</collectingRegion>
|
||
, KIZ 2009061301 (
|
||
<specimenCount id="49C4FD5827093202B5BEFA18FABCFA58" box="[1263,1367,1446,1472]" count="1" pageId="11" pageNumber="498" type="adult">one adult</specimenCount>
|
||
male) from Tian Town, Hechi City,
|
||
<collectingRegion id="9D06F83327093202B0B5FA74FD59FA7C" box="[484,690,1482,1508]" country="China" name="Guangxi" pageId="11" pageNumber="498">Guangxi Province</collectingRegion>
|
||
, KIZ 20190902 (
|
||
<specimenCount id="49C4FD5827093202B224FA74FC35FA7C" box="[885,990,1482,1508]" count="1" pageId="11" pageNumber="498" type="adult">one adult</specimenCount>
|
||
female) from Huangjiang Town, Hechi City,
|
||
<collectingRegion id="9D06F83327093202B184FA50FE4FF990" box="[213,420,1518,1544]" country="China" name="Guangxi" pageId="11" pageNumber="498">Guangxi Province</collectingRegion>
|
||
, KIZ 2023032 (
|
||
<specimenCount id="49C4FD5827093202B30BFA50FD2FF990" box="[602,708,1518,1544]" count="1" pageId="11" pageNumber="498" type="adult">one adult</specimenCount>
|
||
male) from Luohu Town, Shenzhen City,
|
||
<collectingRegion id="9D06F83327093202B5F6FA50FA73F990" box="[1191,1432,1518,1544]" country="China" name="Guangdong" pageId="11" pageNumber="498">Guangdong Province</collectingRegion>
|
||
, KIZ 2023029 (
|
||
<specimenCount id="49C4FD5827093202B06EF9ACFE42F9B4" box="[319,425,1554,1580]" count="1" pageId="11" pageNumber="498" type="adult">one adult</specimenCount>
|
||
female) from Huangpu Town, Guangzhou City,
|
||
<collectingRegion id="9D06F83327093202B29FF9ACFB55F9B4" box="[974,1214,1554,1580]" country="China" name="Guangdong" pageId="11" pageNumber="498">Guangdong Province</collectingRegion>
|
||
, KIZ 2023044 (
|
||
<specimenCount id="49C4FD5827093202B423F9ACFF1FF9C8" count="1" pageId="11" pageNumber="498" type="juvenile">one juvenile</specimenCount>
|
||
, sex unknown) from Xiegang Town, Dongguan City,
|
||
<collectingRegion id="9D06F83327093202B202F988FBAFF9C8" box="[851,1092,1590,1616]" country="China" name="Guangdong" pageId="11" pageNumber="498">Guangdong Province</collectingRegion>
|
||
, CIB 9820 & CIB 78124 (
|
||
<specimenCount id="49C4FD5827093202B420F988FF24F9EC" count="2" pageId="11" pageNumber="498" type="adult">two adult</specimenCount>
|
||
females) from Sanya City,
|
||
<collectingRegion id="9D06F83327093202B359F9E4FD2CF9EC" box="[520,711,1626,1652]" country="China" name="Hainan" pageId="11" pageNumber="498">Hainan Province</collectingRegion>
|
||
, KIZ 2023030 (adult female) from Hanjiang Town, Putian City,
|
||
<collectingRegion id="9D06F83327093202B1C6F9C0FEA0F900" box="[151,331,1662,1688]" country="China" name="Fujian" pageId="11" pageNumber="498">Fujian Province</collectingRegion>
|
||
(see details in
|
||
<bibRefCitation id="3B534B2027093202B0A6F9C0FD4FF900" author="Liu, S. & Hou, M. & Cai, B. & Li, S. & Zhang, Z. & Yu, R. & Rao, D. & Zhang, L." box="[503,676,1662,1688]" pageId="11" pageNumber="498" pagination="307 - 316" refId="ref21074" refString="Liu, S., Hou, M., Cai, B., Li, S., Zhang, Z., Yu, R., Rao, D. & Zhang, L. (2023) Taxonomic status of Lycodon subcinctus sensu lato in China (Serpentes, Colubridae). Herpetozoa, 36, 307 - 316. https: // doi. org / 10.3897 / herpetozoa. 36. e 114206" type="journal article" year="2023">
|
||
Liu
|
||
<emphasis id="6DB6EAC327093202B377F9C1FDB3F900" box="[550,600,1662,1688]" italics="true" pageId="11" pageNumber="498">et al</emphasis>
|
||
. 2023
|
||
</bibRefCitation>
|
||
); KIZ 039729 (adult female) from Mengla County, Xishuangbanna Dai Autonomous Prefecture,
|
||
<collectingRegion id="9D06F83327093202B08CF91CFD76F924" box="[477,669,1698,1724]" country="China" name="Yunnan" pageId="11" pageNumber="498">Yunnan Province</collectingRegion>
|
||
(see details in
|
||
<bibRefCitation id="3B534B2027093202B214F91CFBE4F924" author="Zhang, H. Y. & Wang, L. Y. & Hou, S. B. & Wu, Y. H. & Duan, Z. P. & Jin, Y. D. & Zuo, C. S. & Zhang, Y. S. & Wang, K. & Che, J." box="[837,1039,1698,1724]" pageId="11" pageNumber="498" pagination="81 - 94" refId="ref24266" refString="Zhang, H. Y., Wang, L. Y., Hou, S. B., Wu, Y. H., Duan, Z. P., Jin, Y. D., Zuo, C. S., Zhang, Y. S., Wang, K. & Che, J. (2024) Provincial records of Lycodon septentrionalis and L. cf. subcinctus from Yunnan Province, China. Chinese Journal of Zoology, 59 (1), 81 - 94. [in Chinese] https: // doi. org / 10.13859 / j. cjz. 202423018" type="journal article" year="2024">
|
||
Zhang
|
||
<emphasis id="6DB6EAC327093202B2C2F91DFC2EF924" box="[915,965,1698,1724]" italics="true" pageId="11" pageNumber="498">et al</emphasis>
|
||
. 2024
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="17D8655A27093202B196F978FE02F80C" pageId="11" pageNumber="498" type="etymology">
|
||
<paragraph id="5F7D36D127093202B196F978FE02F80C" blockId="11.[151,1437,1122,2012]" pageId="11" pageNumber="498">
|
||
<emphasis id="6DB6EAC327093202B196F978FEA3F978" bold="true" box="[199,328,1734,1760]" pageId="11" pageNumber="498">Etymology</emphasis>
|
||
. The species
|
||
<emphasis id="6DB6EAC327093202B0BDF979FDDCF978" box="[492,567,1735,1760]" italics="true" pageId="11" pageNumber="498">nomen</emphasis>
|
||
is composed of the prefix
|
||
<emphasis id="6DB6EAC327093202B22BF979FC49F978" box="[890,930,1735,1760]" italics="true" pageId="11" pageNumber="498">neo</emphasis>
|
||
-, drawn from the classical Greek adjective νἐος (α, ν) (
|
||
<emphasis id="6DB6EAC327093202B070F955FEBFF89C" box="[289,340,1771,1796]" italics="true" pageId="11" pageNumber="498">neos</emphasis>
|
||
,
|
||
<emphasis id="6DB6EAC327093202B035F955FE67F89C" box="[356,396,1771,1796]" italics="true" pageId="11" pageNumber="498">nea</emphasis>
|
||
,
|
||
<emphasis id="6DB6EAC327093202B0CDF955FE39F89C" box="[412,466,1771,1796]" italics="true" pageId="11" pageNumber="498">neon</emphasis>
|
||
), meaning “new”, and the Latin adjective
|
||
<taxonomicName id="98C24D5227093202B2E8F954FBC7F89C" authorityName="Hallowell" authorityYear="1861" box="[953,1068,1770,1796]" class="Reptilia" family="Megalopidae" genus="Megalops" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="498" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="6DB6EAC327093202B2E8F954FBC7F89C" box="[953,1068,1770,1796]" italics="true" pageId="11" pageNumber="498">maculatus</emphasis>
|
||
</taxonomicName>
|
||
(-
|
||
<emphasis id="6DB6EAC327093202B516F955FBBEF89C" box="[1095,1109,1771,1796]" italics="true" pageId="11" pageNumber="498">a</emphasis>
|
||
, -
|
||
<emphasis id="6DB6EAC327093202B53FF955FB7BF89C" box="[1134,1168,1771,1796]" italics="true" pageId="11" pageNumber="498">um</emphasis>
|
||
), meaning “spotted” or “blotched”, by allusion to the dorsal pattern of this species. We selected this species
|
||
<emphasis id="6DB6EAC327093202B521F8B1FB50F8B0" box="[1136,1211,1807,1832]" italics="true" pageId="11" pageNumber="498">nomen</emphasis>
|
||
, “the new
|
||
<taxonomicName id="98C24D5227093202B46CF8B0FEE1F8D4" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="498" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="6DB6EAC327093202B46CF8B0FEE1F8D4" italics="true" pageId="11" pageNumber="498">Lycodon maculatus</emphasis>
|
||
</taxonomicName>
|
||
”, because it describes both the long use of the combination in the literature following
|
||
<bibRefCitation id="3B534B2027093202B5ABF88CFA77F8D5" author="Cope, E. D." box="[1274,1436,1842,1869]" pageId="11" pageNumber="498" pagination="186 - 219" refId="ref18952" refString="Cope, E. D. (1895). The classification of the Ophidia. Transactions of the American Philosophical Society, 18 (2), 186 - 219, pls. 14 - 33. https: // doi. org / 10.2307 / 1005387" type="journal article" year="1895">Cope’s (1895)</bibRefCitation>
|
||
misidentification, and our own correction in the present paper, from “
|
||
<taxonomicName id="98C24D5227093202B2C8F8E8FABEF8E9" authority="" sensu Cope (1895)" authorityName="" sensu Cope" authorityYear="1895" box="[921,1365,1878,1905]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="498" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="6DB6EAC327093202B2C8F8E8FB99F8E8" box="[921,1138,1878,1904]" italics="true" pageId="11" pageNumber="498">Lycodon maculatus</emphasis>
|
||
” sensu
|
||
<bibRefCitation id="3B534B2027093202B598F8E8FABEF8E9" author="Cope, E. D." box="[1225,1365,1878,1905]" pageId="11" pageNumber="498" pagination="186 - 219" refId="ref18952" refString="Cope, E. D. (1895). The classification of the Ophidia. Transactions of the American Philosophical Society, 18 (2), 186 - 219, pls. 14 - 33. https: // doi. org / 10.2307 / 1005387" type="journal article" year="1895">Cope (1895)</bibRefCitation>
|
||
</taxonomicName>
|
||
to our own definition of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="17D8655A27093205B196F820FD85FE6C" lastPageId="12" lastPageNumber="499" pageId="11" pageNumber="498" type="diagnosis">
|
||
<paragraph id="5F7D36D127093205B196F820FD85FE6C" blockId="11.[151,1437,1122,2012]" lastBlockId="12.[151,1437,150,2038]" lastPageId="12" lastPageNumber="499" pageId="11" pageNumber="498">
|
||
<emphasis id="6DB6EAC327093202B196F820FED1F820" bold="true" box="[199,314,1950,1976]" pageId="11" pageNumber="498">Diagnosis</emphasis>
|
||
. A species of the genus
|
||
<taxonomicName id="98C24D5227093202B30CF820FD56F820" authorityName="Fitzinger" authorityYear="1826" box="[605,701,1950,1976]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="498" phylum="Chordata" rank="genus">
|
||
<emphasis id="6DB6EAC327093202B30CF820FD56F820" box="[605,701,1950,1976]" italics="true" pageId="11" pageNumber="498">Lycodon</emphasis>
|
||
</taxonomicName>
|
||
characterized by the following characters: body size moderate, maximum total length up to
|
||
<quantity id="983A9B3427093202B08AF87CFDD7F844" box="[475,572,1986,2012]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="498" unit="mm" value="900.0">900 mm</quantity>
|
||
, slender; no preocular, both loreal and prefrontal entering orbit; dorsal scale rows 17-17-15, with the outermost rows smooth and the remaining rows keeled or feebly keeled; ventral scales 187– 208; subcaudals 68–84, paired; supralabials 8, with the 3
|
||
<superScript id="A8B79B99270E3205B248FF04FCCCFF50" attach="none" box="[793,807,186,200]" fontSize="6" pageId="12" pageNumber="499">rd</superScript>
|
||
–5
|
||
<superScript id="A8B79B99270E3205B212FF04FCBBFF50" attach="left" box="[835,848,186,200]" fontSize="6" pageId="12" pageNumber="499">th</superScript>
|
||
(rarely 3
|
||
<superScript id="A8B79B99270E3205B2E6FF04FC2EFF50" attach="none" box="[951,965,186,200]" fontSize="6" pageId="12" pageNumber="499">rd</superScript>
|
||
–6
|
||
<superScript id="A8B79B99270E3205B2B0FF04FC05FF50" attach="none" box="[993,1006,186,200]" fontSize="6" pageId="12" pageNumber="499">th</superScript>
|
||
) contacting the orbit; postoculars 2/2; temporals usually 1+2; cloacal plate divided; dorsum black or black gray, dark brown or gray brown posteriorly; 5–10 white cross-bands on the body in adults (often less in larger specimens), widely separated and relatively narrow (about 4–7 vertebral dorsal scales long); in adults, dorsal bands darken along the anterior edges of each dorsal scale, producing a speckled, reticulated or maculated pattern; large adult specimens lack body and tail bands along the posterior portion of the dorsum but maintain at least a few whitish bands anteriorly; in juveniles, 19–27 plain white body and tail bands; venter plain, cream or light gray, occasionally a mid-dorsal line present along the underside of the tail (data from
|
||
<bibRefCitation id="3B534B20270E3205B0ACFE09FD68FE49" author="Smith, M. A." box="[509,643,439,465]" pageId="12" pageNumber="499" pagination="195 - 212" refId="ref22878" refString="Smith, M. A. (1923) On a collection of reptiles and batrachians from the island of Hainan. Journal of the Natural History Society of Siam, 6, 195 - 212." type="journal article" year="1923">Smith 1923</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B20270E3205B3DEFE09FCE1FE49" author="Pope, C. H." box="[655,778,438,465]" pageId="12" pageNumber="499" refId="ref21916" refString="Pope, C. H. (1935) The reptiles of China. Turtles, crocodilians, snakes, lizards. Natural History of Central Asia. Vol. 10. American Museum of Natural History, New York, New York, 604 pp. https: // doi. org / 10.5962 / bhl. title. 12104" type="book" year="1935">Pope 1935</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B20270E3205B247FE09FC39FE49" author="Zhao, E. M. & Huang, M. H. & Zong, Y. & Jiang, Y. M. & Huang, Q. Y. & Zhao, H. & Ma, J. F. & Zheng, J. & Huang, Z. J. & Wei, G. & Yang, D. Q. & Li, D. J." box="[790,978,439,465]" pageId="12" pageNumber="499" refId="ref24466" refString="Zhao, E. M., Huang, M. H., Zong, Y., Jiang, Y. M., Huang, Q. Y., Zhao, H., Ma, J. F., Zheng, J., Huang, Z. J., Wei, G., Yang, D. Q. & Li, D. J. (1998) Fauna Sinica Reptilia. Vol. 3. Squamata: Serpentes. Science Press, Beijing, 522 pp. [in Chinese]" type="book" year="1998">
|
||
Zhao
|
||
<emphasis id="6DB6EAC3270E3205B206FE09FC62FE49" box="[855,905,439,465]" italics="true" pageId="12" pageNumber="499">et al</emphasis>
|
||
. 1998
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B20270E3205B28EFE09FB57FE49" author="Geissler, P. & Nguyen, T. Q. & Poyarkov, N. A. & Bohme, W." box="[991,1212,439,465]" pageId="12" pageNumber="499" pagination="9 - 16" refId="ref19433" refString="Geissler, P., Nguyen, T. Q., Poyarkov, N. A. & Bohme, W. (2011) New records of snakes from Cat Tien National Park Dong Nai and Lam Dong provinces southern Vietnam. Bonn Zoological Bulletin, 60 (1), 9 - 16." type="journal article" year="2011">
|
||
Geissler
|
||
<emphasis id="6DB6EAC3270E3205B513FE09FB9FFE49" box="[1090,1140,439,465]" italics="true" pageId="12" pageNumber="499">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B20270E3205B598FE09FA82FE49" author="Le, D. T. & Luong, A. M. & Pham, C. T. & Pham, T. Q. & Nguyen, S. L. H. & Ziegler, T. & Nguyen, T. Q." box="[1225,1385,439,465]" pageId="12" pageNumber="499" pagination="201 - 219" refId="ref20602" refString="Le, D. T., Luong, A. M., Pham, C. T., Pham, T. Q., Nguyen, S. L. H., Ziegler, T. & Nguyen, T. Q. (2021) New records and an updated checklist of amphibians and snakes from Tuyen Quang Province, Vietnam. Bonn zoological Bulletin, 70 (1), 201 - 219. https: // doi. org / 10.20363 / BZB- 2021.70.1.201" type="journal article" year="2021">
|
||
Le
|
||
<emphasis id="6DB6EAC3270E3205B5BFFE09FACBFE49" box="[1262,1312,439,465]" italics="true" pageId="12" pageNumber="499">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B20270E3205B424FE09FEFAFE6C" author="Liu, S. & Hou, M. & Cai, B. & Li, S. & Zhang, Z. & Yu, R. & Rao, D. & Zhang, L." pageId="12" pageNumber="499" pagination="307 - 316" refId="ref21074" refString="Liu, S., Hou, M., Cai, B., Li, S., Zhang, Z., Yu, R., Rao, D. & Zhang, L. (2023) Taxonomic status of Lycodon subcinctus sensu lato in China (Serpentes, Colubridae). Herpetozoa, 36, 307 - 316. https: // doi. org / 10.3897 / herpetozoa. 36. e 114206" type="journal article" year="2023">
|
||
Liu
|
||
<emphasis id="6DB6EAC3270E3205B1C6FE65FF23FE6C" box="[151,200,474,500]" italics="true" pageId="12" pageNumber="499">et al</emphasis>
|
||
. 2023
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B20270E3205B04CFE64FE0DFE6C" author="Zhang, H. Y. & Wang, L. Y. & Hou, S. B. & Wu, Y. H. & Duan, Z. P. & Jin, Y. D. & Zuo, C. S. & Zhang, Y. S. & Wang, K. & Che, J." box="[285,486,474,501]" pageId="12" pageNumber="499" pagination="81 - 94" refId="ref24266" refString="Zhang, H. Y., Wang, L. Y., Hou, S. B., Wu, Y. H., Duan, Z. P., Jin, Y. D., Zuo, C. S., Zhang, Y. S., Wang, K. & Che, J. (2024) Provincial records of Lycodon septentrionalis and L. cf. subcinctus from Yunnan Province, China. Chinese Journal of Zoology, 59 (1), 81 - 94. [in Chinese] https: // doi. org / 10.13859 / j. cjz. 202423018" type="journal article" year="2024">
|
||
Zhang
|
||
<emphasis id="6DB6EAC3270E3205B03AFE65FE76FE6C" box="[363,413,474,500]" italics="true" pageId="12" pageNumber="499">et al</emphasis>
|
||
. 2024
|
||
</bibRefCitation>
|
||
; this study).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="17D8655A270E3218B196FE40FCB8F9E8" lastPageId="17" lastPageNumber="504" pageId="12" pageNumber="499" type="description">
|
||
<paragraph id="5F7D36D1270E3205B196FE40FB10FD53" blockId="12.[151,1437,150,2038]" pageId="12" pageNumber="499">
|
||
<emphasis id="6DB6EAC3270E3205B196FE40FDA2FD80" bold="true" box="[199,585,510,536]" pageId="12" pageNumber="499">
|
||
Description of the
|
||
<typeStatus id="80798873270E3205B0F8FE40FDFBFD80" box="[425,528,510,536]" pageId="12" pageNumber="499" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="C7F92A54270E3205B371FE40FDA2FD80" box="[544,585,510,536]" captionStart="FIGURE 5" captionStartId="13.[151,250,1495,1519]" captionTargetBox="[151,1436,181,1471]" captionTargetId="figure-18@13.[151,1436,181,1471]" captionTargetPageId="13" captionText="FIGURE 5. Lycodon neomaculatus sp. nov. in preservative—MCZ R-175968 (holotype, adult male) A—General dorsal view; B—General ventral view; C—Lateral view of the head, right side; D—Lateral view of the head, left side; E—Dorsal view of the head; F—Ventral view of the tail with the everted hemipenes. Images not to scale. Photos by M. Gage and J. Martinez." figureDoi="http://doi.org/10.5281/zenodo.13916229" httpUri="https://zenodo.org/record/13916229/files/figure.png" pageId="12" pageNumber="499">Fig</figureCitation>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC3270E3205B309FE40FD9BFD80" bold="true" box="[600,624,510,536]" pageId="12" pageNumber="499">5)</emphasis>
|
||
. Specimen MCZ R-
|
||
<quantity id="983A9B34270E3205B206FE40FC21FD80" box="[855,970,510,537]" metricMagnitude="3" metricUnit="m" metricValue="4.469587199999999" pageId="12" pageNumber="499" unit="in" value="175968.0">175968 in</quantity>
|
||
excellent condition, no incisions present along ventral surface, both hemipenes everted, but right organ more fully prepared than left. Body slender; tail long, gradually tapering to a blunt and enlarged terminal scute; head oblong shaped, longer than wide (HeadW/HeadL 0.57), slightly flattened, moderately distinct from the neck; snout elongate, projecting over the lower jaw, rounded in dorsal profile, truncate and slightly depressed in lateral profile with a weak canthus rostralis; nostrils large, positioned dorso-laterally, round in shape; eyes medium with an elliptical and slightly vertical pupil.
|
||
</paragraph>
|
||
<paragraph id="5F7D36D1270E3205B196FD6BFB07FC8B" blockId="12.[151,1437,150,2038]" pageId="12" pageNumber="499">
|
||
<emphasis id="6DB6EAC3270E3205B196FD6BFE82FD76" box="[199,361,725,750]" italics="true" pageId="12" pageNumber="499">Measurements</emphasis>
|
||
. SVL
|
||
<quantity id="983A9B34270E3205B0E7FD6AFDFCFD77" box="[438,535,724,751]" metricMagnitude="-1" metricUnit="m" metricValue="4.56" pageId="12" pageNumber="499" unit="mm" value="456.0">456 mm</quantity>
|
||
, TaL
|
||
<quantity id="983A9B34270E3205B309FD6BFD52FD77" box="[600,697,725,751]" metricMagnitude="-1" metricUnit="m" metricValue="1.29" pageId="12" pageNumber="499" unit="mm" value="129.0">129 mm</quantity>
|
||
, TL
|
||
<quantity id="983A9B34270E3205B3A1FD6AFCA5FD77" box="[752,846,724,751]" metricMagnitude="-1" metricUnit="m" metricValue="5.85" pageId="12" pageNumber="499" unit="mm" value="585.0">585 mm</quantity>
|
||
(ratio TaL/TL 0.221), HeadL
|
||
<quantity id="983A9B34270E3205B5F7FD6AFAE5FD77" box="[1190,1294,724,751]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="12" pageNumber="499" unit="mm" value="16.2">16.2 mm</quantity>
|
||
, HeadW
|
||
<quantity id="983A9B34270E3205B428FD6BFF2EFC8B" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="12" pageNumber="499" unit="mm" value="9.3">9.3 mm</quantity>
|
||
, SnL
|
||
<quantity id="983A9B34270E3205B054FD46FEB6FC8B" box="[261,349,760,787]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="12" pageNumber="499" unit="mm" value="5.2">5.2 mm</quantity>
|
||
, SnW
|
||
<quantity id="983A9B34270E3205B0F6FD46FE14FC8B" box="[423,511,760,787]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="12" pageNumber="499" unit="mm" value="4.1">4.1 mm</quantity>
|
||
, IOD
|
||
<quantity id="983A9B34270E3205B315FD46FD77FC8B" box="[580,668,760,787]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="12" pageNumber="499" unit="mm" value="5.3">5.3 mm</quantity>
|
||
, EyeD
|
||
<quantity id="983A9B34270E3205B3BFFD46FCADFC8B" box="[750,838,760,787]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="499" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, FrontalL
|
||
<quantity id="983A9B34270E3205B2EBFD46FBF9FC8B" box="[954,1042,760,787]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="12" pageNumber="499" unit="mm" value="3.8">3.8 mm</quantity>
|
||
, FrontalW
|
||
<quantity id="983A9B34270E3205B5DEFD46FB03FC8B" box="[1167,1256,760,787]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="12" pageNumber="499" unit="mm" value="3.6">3.6 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5F7D36D1270E3205B196FCA2FC7CFC39" blockId="12.[151,1437,150,2038]" pageId="12" pageNumber="499">
|
||
<emphasis id="6DB6EAC3270E3205B196FCA2FE84FCAE" box="[199,367,796,822]" italics="true" pageId="12" pageNumber="499">Body scalation</emphasis>
|
||
. Dorsal scale rows 17-17-15; dorsal scale rows smooth anteriorly, remaining scale rows feebly keeled at midbody and posteriorly, except for the outermost row, which is always smooth; scales on the vertebral row not enlarged; no apical pits; 192 ventral scales, laterally angulate; 72 subcaudals, paired and laterally angulate; total body scales 265; subcaudal ratio 27.17%; cloacal plate divided.
|
||
</paragraph>
|
||
<paragraph id="5F7D36D1270E3205B196FC15FF06F9D1" blockId="12.[151,1437,150,2038]" pageId="12" pageNumber="499">
|
||
<emphasis id="6DB6EAC3270E3205B196FC15FE9BFC5D" box="[199,368,939,965]" italics="true" pageId="12" pageNumber="499">Head scalation</emphasis>
|
||
. Rostral wider than high, barely visible from above, posterior suture not projecting into internasal region, forming a straight angle when viewed in dorsal profile; nasal 1.5 times longer than high, partially divided by a small suture above the nostril; posterior half of nasal subpentagonal, slightly larger than anterior nasal; nasal surrounded by the first two supralabials, rostral, internasal, prefrontal and loreal; internasals paired, anterior sutures slightly concave in dorsal profile, in contact with rostral, nasal, and prefrontal; each internasal 1.7 times wider than long; prefrontals paired, approximately 2.2 times longer than internasals, subrectangular, each scale 1.6 times shorter than frontal; each prefrontal 1.1 times wider than long, in contact with internasal, nasal, loreal, eye, and frontal; supraoculars paired, subrectangular shaped, 1.5 times longer than wide, in contact with prefrontal, posterior suture of each scale 1.9 times wider than anterior suture; frontal small, hexagonal, shield shaped, slightly longer than wide (ratio FrontalL/FrontalW 1.05), tapering posteriorly; anterior suture of frontal projecting past eye sockets in dorsal profile; angle formed by the posterior vertex of frontal slightly acute; parietals paired, suture 1.2 times longer than wide and 1.1 times longer than frontal, entire length of each parietal 1.7 times longer than wide; loreal 1/1, subrectangular, 1.4 times longer than high, in contact with eye; preocular absent; subocular absent; postoculars 2/2, uppermost scale approximately 1.5 times larger than lowermost; temporals 1+2; supralabials 8/8, first and second in contact with nasal, second and third in contact with loreal, third and fifth in contact with eye, sixth supralabial largest; infralabials 9/9, first pair in broad contact, first to fifth in contact with the anterior pair of chin shields; mental subtriangular in ventral profile, 2.1 times wider than long; anterior chin shields 1.4 times longer than posterior shields; posterior chin shields in narrow contact medially, separated by skin tissue along the mental groove.
|
||
</paragraph>
|
||
<paragraph id="5F7D36D1270E3205B196F9ECFE1FF86D" blockId="12.[151,1437,150,2038]" pageId="12" pageNumber="499">
|
||
<emphasis id="6DB6EAC3270E3205B196F9ECFE1DF9F4" box="[199,502,1618,1644]" italics="true" pageId="12" pageNumber="499">Coloration in preservation</emphasis>
|
||
. After 34 years of preservation, dorsal surface of body black anteriorly, gradually transitioning to dark brown by midbody and pale brown posteriorly, lighter along flanks; seven white body bands present, the first six with a conspicuous dark brown or black spot on the anterior end of each dorsal scale, forming a speckled pattern across every band, and the seventh band more subdued with dark brown pigment engulfing all of the dorsal scales; remaining posterior portion of the dorsum plain; tail tip cream. Dorsal surface of head dark brown from the anterior half of the frontal and supraocular scales to the snout, including the rostral, nasal, loreal and first two supralabials; remaining dorsal portion of head covered with pale brown marbling stretching from the posterior half of the supraoculars to the end of the parietals, curving in a slightly concave manner across the occipital region; within the marbling a small dark-brown spot present medially along the parietal suture; eye black, pupil white; remaining six supralabials cream; mental, first pair of infralabials and anterior edge of anterior chin shields dark brown; remaining underside of head white, immaculate. Ventral surface of head, body, tail white, and immaculate. Coloration in life not recorded.
|
||
</paragraph>
|
||
<caption id="0BBD6659270F3204B1C6FA69FAA0F9AF" ID-DOI="http://doi.org/10.5281/zenodo.13916229" ID-Zenodo-Dep="13916229" httpUri="https://zenodo.org/record/13916229/files/figure.png" pageId="13" pageNumber="500" startId="13.[151,250,1495,1519]" targetBox="[151,1436,181,1471]" targetPageId="13" targetType="figure">
|
||
<paragraph id="5F7D36D1270F3204B1C6FA69FAA0F9AF" blockId="13.[151,1436,1495,1592]" pageId="13" pageNumber="500">
|
||
<emphasis id="6DB6EAC3270F3204B1C6FA69FEE6FA77" bold="true" box="[151,269,1495,1519]" pageId="13" pageNumber="500">FIGURE 5</emphasis>
|
||
.
|
||
<taxonomicName id="98C24D52270F3204B048FA69FDE8FA77" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[281,515,1495,1519]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="500" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC3270F3204B048FA69FDE8FA77" box="[281,515,1495,1519]" italics="true" pageId="13" pageNumber="500">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B8270F3204B358FA69FDB0FA77" box="[521,603,1495,1519]" pageId="13" pageNumber="500" rank="species">
|
||
<emphasis id="6DB6EAC3270F3204B358FA69FDCAFA77" bold="true" box="[521,545,1495,1519]" pageId="13" pageNumber="500">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC3270F3204B37CFA69FDBFFA77" bold="true" box="[557,596,1495,1519]" pageId="13" pageNumber="500">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
in preservative—MCZ R-175968 (holotype, adult male) A—General dorsal view; B—General ventral view; C—Lateral view of the head, right side; D—Lateral view of the head, left side; E—Dorsal view of the head; F—Ventral view of the tail with the everted hemipenes. Images not to scale. Photos by M. Gage and J. Martinez.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5F7D36D1270F3204B196F937FE27F87F" blockId="13.[151,1437,1673,2023]" pageId="13" pageNumber="500">
|
||
<emphasis id="6DB6EAC3270F3204B196F937FDBCF93B" bold="true" box="[199,599,1673,1700]" pageId="13" pageNumber="500">General description and variation</emphasis>
|
||
(
|
||
<tableCitation id="1240036A270F3204B332F937FD5CF93C" box="[611,695,1673,1700]" captionStart="TABLE 1" captionStartId="16.[152,244,151,175]" captionTargetPageId="16" captionText="TABLE 1. Comparison of morphological characters of Lycodon neomaculatus sp. nov. with L. subcinctus and L. sealei. Diagnostic differences between the new species and L. subcinctus + L. sealei are marked in bold." httpUri="http://table.plazi.org/id/0BBD665927123219B1C9FF29FB48FF4B" pageId="13" pageNumber="500" tableUuid="0BBD665927123219B1C9FF29FB48FF4B">Table 1</tableCitation>
|
||
; Appendix
|
||
<tableCitation id="1240036A270F3204B265F937FC7DF93C" box="[820,918,1673,1700]" captionStart="TABLE 1" captionStartId="16.[152,244,151,175]" captionTargetPageId="16" captionText="TABLE 1. Comparison of morphological characters of Lycodon neomaculatus sp. nov. with L. subcinctus and L. sealei. Diagnostic differences between the new species and L. subcinctus + L. sealei are marked in bold." httpUri="http://table.plazi.org/id/0BBD665927123219B1C9FF29FB48FF4B" pageId="13" pageNumber="500" tableUuid="0BBD665927123219B1C9FF29FB48FF4B">Table S1</tableCitation>
|
||
;
|
||
<figureCitation id="C7F92A54270F3204B2F0F937FC07F93C" box="[929,1004,1673,1700]" captionStart="FIGURE 1" captionStartId="8.[151,250,1495,1519]" captionTargetBox="[151,1436,181,1471]" captionTargetId="figure-29@8.[151,1436,181,1471]" captionTargetPageId="8" captionText="FIGURE 1. Colubrid snake species involved in the case of Anoplophallus maculatus Cope, 1895.—USNM 7339, previously considered the holotype of “Lycodon maculatus” reported by Liu et al. (2023): A—General dorsal view; B—General ventral view; C—Dorsalview ofthe head;D—Lateralview of thehead,leftside; E—Livespecimen of Leptodeira maculata (= Megalops maculatus) from Minatitlán, Colima, Mexico; F—Live specimen of Lycodon subannulatus from Penang Island, Malaysia, of which Nymphophidium maculatum is a junior synonym. Images not to scale. Photos by T. Hsu (A–D), C. Grünwald (E), J. L. Lee (F)." figureDoi="http://doi.org/10.5281/zenodo.13916221" httpUri="https://zenodo.org/record/13916221/files/figure.png" pageId="13" pageNumber="500">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="C7F92A54270F3204B2A9F937FBE3F93C" box="[1016,1032,1673,1700]" captionStart="FIGURE 5" captionStartId="13.[151,250,1495,1519]" captionTargetBox="[151,1436,181,1471]" captionTargetId="figure-18@13.[151,1436,181,1471]" captionTargetPageId="13" captionText="FIGURE 5. Lycodon neomaculatus sp. nov. in preservative—MCZ R-175968 (holotype, adult male) A—General dorsal view; B—General ventral view; C—Lateral view of the head, right side; D—Lateral view of the head, left side; E—Dorsal view of the head; F—Ventral view of the tail with the everted hemipenes. Images not to scale. Photos by M. Gage and J. Martinez." figureDoi="http://doi.org/10.5281/zenodo.13916229" httpUri="https://zenodo.org/record/13916229/files/figure.png" pageId="13" pageNumber="500">5</figureCitation>
|
||
,
|
||
<figureCitation id="C7F92A54270F3204B543F937FBC9F93B" box="[1042,1058,1673,1699]" captionStart="FIGURE 6" captionStartId="14.[151,250,1495,1519]" captionTargetBox="[151,1436,181,1471]" captionTargetId="figure-29@14.[151,1436,181,1471]" captionTargetPageId="14" captionText="FIGURE 6. Lycodon neomaculatus sp. nov. in preservative—specimen CAS 235846 (paratype, adult male) A—General dorsal view; B—General ventral view; C—dorsal view of the head; D—Ventral view of the head; E—Lateral view of the head, right side; F—Ventral view of the tail with the everted hemipenes. Photos by J.L. Lee." figureDoi="http://doi.org/10.5281/zenodo.13916233" httpUri="https://zenodo.org/record/13916233/files/figure.png" pageId="13" pageNumber="500">6</figureCitation>
|
||
,
|
||
<figureCitation id="C7F92A54270F3204B57DF934FB84F93C" box="[1068,1135,1674,1700]" captionStart="FIGURE 7" captionStartId="15.[151,250,1323,1347]" captionTargetBox="[397,1191,669,1299]" captionTargetId="figure-260@15.[397,1191,669,1299]" captionTargetPageId="15" captionText="FIGURE 7. Differences in dorsal body band coloration in preservative between Lycodon neomaculatus sp. nov. (A, B) and L. subcinctus (C, D). A—FMNH 180149 from Nakhon Ratchasima, Thailand; B—CAS 229729 from Tanintharyi, Myanmar; C—FMNH 229855 from Selangor, Malaysia; D—FMNH 183772 from Selangor, Malaysia. Images not to scale. Photos by J.L. Lee." figureDoi="http://doi.org/10.5281/zenodo.13916235" httpUri="https://zenodo.org/record/13916235/files/figure.png" pageId="13" pageNumber="500">7A, B</figureCitation>
|
||
,
|
||
<figureCitation id="C7F92A54270F3204B528F934FB62F93C" box="[1145,1161,1674,1700]" captionStart="FIGURE 8" captionStartId="17.[151,250,1084,1108]" captionTargetBox="[396,1191,181,1060]" captionTargetId="figure-18@17.[396,1191,181,1060]" captionTargetPageId="17" captionText="FIGURE 8. Lycodon neomaculatus sp. nov. in life (not collected, sex not determined): A, B—Hong Kong, China (adult, subadult; respectively); C—Qiongzhong, Hainan, China (adult); D—Xishuangbanna, Yunnan, China; E—Pu Mat NP, Nghe An, Vietnam; F—Ma Da NR, Dong Nai, Vietnam (adult); G—Cat Tien NP, Dong Nai, Vietnam (subadult); H—Loc Bac, Lam Dong, Vietnam (adult); I—Phuoc Binh NP, Ninh Thuan, Vietnam (adult); J—Hinhurp, Vientiane, Laos (subadult); K—Kompong Seila, Seihanuk, Cambodia (adult); L—Chai Prakarn, Chiang Mai, Thailand (subadult); M—Mae Yom NP, Phrae, Thailand (adult); N—Wang Nam Khiao, Nakhon Ratchasima, Thailand (adult); O—Kaeng Krachan NP, Phetchaburi, Thailand (subadult). Photos by:A. Tomaszek (A, B); C.X. Liao (C); Z.H. Ma (D); P.S. Hamilton (E); H.X.N. Nguyen (F); A.M. Bragin (G, I); N.A. Poyarkov (H); T. Calame (J); T. Samnang (K); A. Kaosung (L); R. Grassby-Lewis (M, N); A. Pierce (O)." figureDoi="http://doi.org/10.5281/zenodo.13916237" httpUri="https://zenodo.org/record/13916237/files/figure.png" pageId="13" pageNumber="500">8</figureCitation>
|
||
). Based on
|
||
<specimenCount id="49C4FD58270F3204B456F934FA76F93B" box="[1287,1437,1673,1700]" count="47" pageId="13" pageNumber="500" type="generic">47 specimens</specimenCount>
|
||
(
|
||
<specimenCount id="49C4FD58270F3204B1F1F910FEE3F95F" box="[160,264,1709,1736]" count="12" pageId="13" pageNumber="500" type="male">12 males</specimenCount>
|
||
,
|
||
<specimenCount id="49C4FD58270F3204B045F910FE79F95F" box="[276,402,1709,1736]" count="22" pageId="13" pageNumber="500" type="female">22 females</specimenCount>
|
||
, and
|
||
<specimenCount id="49C4FD58270F3204B09FF910FDB0F95F" box="[462,603,1709,1736]" count="13" pageId="13" pageNumber="500" type="juvenile">13 juveniles</specimenCount>
|
||
). The maximum length is
|
||
<quantity id="983A9B34270F3204B2D2F910FC0BF950" box="[899,992,1710,1736]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="500" unit="mm" value="900.0">900 mm</quantity>
|
||
(SVL
|
||
<quantity id="983A9B34270F3204B57DF910FB66F950" box="[1068,1165,1710,1736]" metricMagnitude="-1" metricUnit="m" metricValue="7.4" pageId="13" pageNumber="500" unit="mm" value="740.0">740 mm</quantity>
|
||
; TaL
|
||
<quantity id="983A9B34270F3204B59DF913FAC6F950" box="[1228,1325,1709,1736]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="500" unit="mm" value="160.0">160 mm</quantity>
|
||
; based on an adult female specimen [no. 197] reported in
|
||
<bibRefCitation id="3B534B20270F3204B3E5F96FFC46F974" author="Zhao, E. M. & Huang, M. H. & Zong, Y. & Jiang, Y. M. & Huang, Q. Y. & Zhao, H. & Ma, J. F. & Zheng, J. & Huang, Z. J. & Wei, G. & Yang, D. Q. & Li, D. J." box="[692,941,1745,1772]" pageId="13" pageNumber="500" refId="ref24466" refString="Zhao, E. M., Huang, M. H., Zong, Y., Jiang, Y. M., Huang, Q. Y., Zhao, H., Ma, J. F., Zheng, J., Huang, Z. J., Wei, G., Yang, D. Q. & Li, D. J. (1998) Fauna Sinica Reptilia. Vol. 3. Squamata: Serpentes. Science Press, Beijing, 522 pp. [in Chinese]" type="book" year="1998">
|
||
Zhao
|
||
<emphasis id="6DB6EAC3270F3204B3A4F96CFCC3F973" box="[757,808,1745,1771]" italics="true" pageId="13" pageNumber="500">et al</emphasis>
|
||
. 1998: 184
|
||
</bibRefCitation>
|
||
). The maximum length in males is
|
||
<quantity id="983A9B34270F3204B46FF96CFA76F974" box="[1342,1437,1746,1772]" metricMagnitude="-1" metricUnit="m" metricValue="8.1" pageId="13" pageNumber="500" unit="mm" value="810.0">810 mm</quantity>
|
||
(SVL
|
||
<quantity id="983A9B34270F3204B18AF94BFED2F888" box="[219,313,1781,1808]" metricMagnitude="-1" metricUnit="m" metricValue="6.51" pageId="13" pageNumber="500" unit="mm" value="651.0">651 mm</quantity>
|
||
, TaL
|
||
<quantity id="983A9B34270F3204B024F94BFE3FF888" box="[373,468,1781,1808]" metricMagnitude="-1" metricUnit="m" metricValue="1.59" pageId="13" pageNumber="500" unit="mm" value="159.0">159 mm</quantity>
|
||
; specimen IEBR A.2010.42;
|
||
<bibRefCitation id="3B534B20270F3204B246F94BFBF1F888" author="Geissler, P. & Nguyen, T. Q. & Poyarkov, N. A. & Bohme, W." box="[791,1050,1781,1808]" pageId="13" pageNumber="500" pagination="9 - 16" refId="ref19433" refString="Geissler, P., Nguyen, T. Q., Poyarkov, N. A. & Bohme, W. (2011) New records of snakes from Cat Tien National Park Dong Nai and Lam Dong provinces southern Vietnam. Bonn Zoological Bulletin, 60 (1), 9 - 16." type="journal article" year="2011">
|
||
Geissler
|
||
<emphasis id="6DB6EAC3270F3204B228F948FC41F897" box="[889,938,1781,1807]" italics="true" pageId="13" pageNumber="500">et al</emphasis>
|
||
. 2011: 13
|
||
</bibRefCitation>
|
||
); ratio TaL/TL
|
||
<geoCoordinate id="3AF65016270F3204B595F948FAE8F888" box="[1220,1283,1782,1808]" degrees="0.178" direction="north" orientation="latitude" pageId="13" pageNumber="500" precision="55" value="0.178">0.178</geoCoordinate>
|
||
<geoCoordinate id="3AF65016270F3204B452F948FABBF888" box="[1283,1360,1782,1808]" degrees="0.221" direction="west" orientation="longitude" pageId="13" pageNumber="500" precision="55" value="-0.221">–0.221</geoCoordinate>
|
||
(mean = 0.201 ± 0.012, n = 42), without sexual dimorphism. Most specimens greatly resemble the
|
||
<typeStatus id="80798873270F3204B5C0F8A7FB1CF8AB" box="[1169,1271,1817,1843]" pageId="13" pageNumber="500" type="holotype">holotype</typeStatus>
|
||
. Body slender; tail resembling the
|
||
<typeStatus id="80798873270F3204B03EF883FE3FF8CF" box="[367,468,1853,1879]" pageId="13" pageNumber="500" type="holotype">holotype</typeStatus>
|
||
, gradually tapering to a blunt and enlarged terminal scute; head oblong shaped, longer than wide, subrectangular, widest posteriorly and slightly flattened, distinct from neck; snout elongate, rounded dorsally, truncate and slightly depressed laterally, weak canthus rostralis; nostrils always pointed dorsolaterally, large, oval shaped, dividing the nasal scale anteriorly and occasionally posteriorly as well; eyes medium with an elliptical and vertical pupil.
|
||
</paragraph>
|
||
<caption id="0BBD6659270C3207B1C6FA69FC21F9AF" ID-DOI="http://doi.org/10.5281/zenodo.13916233" ID-Zenodo-Dep="13916233" httpUri="https://zenodo.org/record/13916233/files/figure.png" pageId="14" pageNumber="501" startId="14.[151,250,1495,1519]" targetBox="[151,1436,181,1471]" targetPageId="14" targetType="figure">
|
||
<paragraph id="5F7D36D1270C3207B1C6FA69FC21F9AF" blockId="14.[151,1436,1495,1591]" pageId="14" pageNumber="501">
|
||
<emphasis id="6DB6EAC3270C3207B1C6FA69FEE6FA77" bold="true" box="[151,269,1495,1519]" pageId="14" pageNumber="501">FIGURE 6</emphasis>
|
||
.
|
||
<taxonomicName id="98C24D52270C3207B049FA69FDE9FA77" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[280,514,1495,1519]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="501" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC3270C3207B049FA69FDE9FA77" box="[280,514,1495,1519]" italics="true" pageId="14" pageNumber="501">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6DB6EAC3270C3207B356FA69FDBCFA77" bold="true" box="[519,599,1495,1519]" pageId="14" pageNumber="501">
|
||
<taxonomicNameLabel id="768557B8270C3207B356FA69FDBCFA77" box="[519,599,1495,1519]" pageId="14" pageNumber="501" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in preservative—specimen CAS 235846 (paratype, adult male) A—General dorsal view; B—General ventral view; C—dorsal view of the head; D—Ventral view of the head; E—Lateral view of the head, right side; F—Ventral view of the tail with the everted hemipenes. Photos by J.L. Lee.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5F7D36D1270C3207B196F937FE6CF8CF" blockId="14.[151,1437,1673,2024]" pageId="14" pageNumber="501">
|
||
<emphasis id="6DB6EAC3270C3207B196F937FE84F93B" box="[199,367,1673,1699]" italics="true" pageId="14" pageNumber="501">Body scalation</emphasis>
|
||
. Dorsal scales in 17-17-15 rows; all dorsal scale rows smooth anteriorly, first to fifth outermost rows smooth at midbody and posteriorly, remaining scales keeled; scales of the outermost dorsal scale row slightly enlarged; ventrals 187–208 (mean = 198.98 ± 4.90; n = 57), without sexual dimorphism, laterally angulate; subcaudals 68–84 (mean = 75.07±4.65, n = 30), without sexual dimorphism, all paired, laterally angulate; total body scales 260–290 (mean = 274.36 ± 7.46, n = 47); subcaudal ratio 25.09–30.21% (mean = 27.48 ± 1.26, n = 42); cloacal plate divided.
|
||
</paragraph>
|
||
<paragraph id="5F7D36D1270C3206B196F8DFFCEBFD1D" blockId="14.[151,1437,1673,2024]" lastBlockId="15.[151,1437,151,645]" lastPageId="15" lastPageNumber="502" pageId="14" pageNumber="501">
|
||
<emphasis id="6DB6EAC3270C3207B196F8DFFE9CF8E3" box="[199,375,1889,1915]" italics="true" pageId="14" pageNumber="501">Head scalation</emphasis>
|
||
. Arrangement of cephalic scales complete, including two internasals, two prefrontals, two supraoculars, one frontal, and two parietals, with all except the frontal arranged in pairs. Rostral wider than high, visible from above, posterior suture not projecting into internasal region or separating the internasals only a short distance medially; angle formed by the posterior suture of the rostral scale and the internasals straight or broadly obtuse angled; nasal partially divided or completely divided into two distinct scales by a suture above or below the nostril; anterior half subrectangular, posterior half pentagonal and distinctly larger; internasals subrectangular, not narrowing anteriorly, wider than long, separate from loreal; prefrontals subrectangular, larger than internasals, in broad contact with loreal, in contact with orbit; supraoculars subrectangular, posterior end of each scale slightly wider than anterior end; frontal hexagonal or pentagonal, shield shaped; angle formed by the posterior vertex of frontal scale acute angled; anterior edge of frontal scale positioned in front of eye sockets; parietals paired, each scale and suture larger and longer than frontal; loreal longer than high, subrectangular, in broad contact with nasal and prefrontal; loreal in contact with orbit; supralabials normally 8/8, 1
|
||
<superScript id="A8B79B99270D3206B29DFE2DFC3CFE39" attach="none" box="[972,983,403,417]" fontSize="6" pageId="15" pageNumber="502">st</superScript>
|
||
–2
|
||
<superScript id="A8B79B99270D3206B2A2FE2CFBE8FE38" attach="left" box="[1011,1027,402,416]" fontSize="6" pageId="15" pageNumber="502">nd</superScript>
|
||
supralabials in contact with nasals, 2
|
||
<superScript id="A8B79B99270D3206B1F4FE08FF5EFE5C" attach="none" box="[165,181,438,452]" fontSize="6" pageId="15" pageNumber="502">nd</superScript>
|
||
–3
|
||
<superScript id="A8B79B99270D3206B183FE08FF0BFE5C" attach="left" box="[210,224,438,452]" fontSize="6" pageId="15" pageNumber="502">rd</superScript>
|
||
supralabials in contact with loreal, 3
|
||
<superScript id="A8B79B99270D3206B3C5FE08FD49FE5C" attach="none" box="[660,674,438,452]" fontSize="6" pageId="15" pageNumber="502">rd</superScript>
|
||
–5
|
||
<superScript id="A8B79B99270D3206B3EFFE08FD20FE5C" attach="left" box="[702,715,438,452]" fontSize="6" pageId="15" pageNumber="502">th</superScript>
|
||
(45/
|
||
<specimenCount id="49C4FD58270D3206B253FE08FC4BFE49" box="[770,928,438,465]" count="49" pageId="15" pageNumber="502" type="generic">49 specimens</specimenCount>
|
||
) or rarely 3
|
||
<superScript id="A8B79B99270D3206B561FE08FBD5FE5C" attach="none" box="[1072,1086,438,452]" fontSize="6" pageId="15" pageNumber="502">rd</superScript>
|
||
–6
|
||
<superScript id="A8B79B99270D3206B508FE08FB8DFE5C" attach="left" box="[1113,1126,438,452]" fontSize="6" pageId="15" pageNumber="502">th</superScript>
|
||
(4/
|
||
<specimenCount id="49C4FD58270D3206B5C1FE09FAC6FE49" box="[1168,1325,439,465]" count="49" pageId="15" pageNumber="502" type="generic">49 specimens</specimenCount>
|
||
) entering orbit, 6
|
||
<superScript id="A8B79B99270D3206B1B9FE64FF1EFE70" attach="left" box="[232,245,474,488]" fontSize="6" pageId="15" pageNumber="502">th</superScript>
|
||
supralabial largest; preocular absent; postoculars two, uppermost scalelargest; temporals 1+2 (rarely 1+1); infralabials 9/9 (42/
|
||
<specimenCount id="49C4FD58270D3206B028FE41FDF3FD81" box="[377,536,511,537]" count="49" pageId="15" pageNumber="502" type="generic">49 specimens</specimenCount>
|
||
), rarely 8/8 (7/
|
||
<specimenCount id="49C4FD58270D3206B392FE41FC88FD81" box="[707,867,511,537]" count="49" pageId="15" pageNumber="502" type="generic">49 specimens</specimenCount>
|
||
), 2
|
||
<superScript id="A8B79B99270D3206B2D7FE40FC7DFD94" attach="none" box="[902,918,510,524]" fontSize="6" pageId="15" pageNumber="502">nd</superScript>
|
||
–3
|
||
<superScript id="A8B79B99270D3206B2E2FE40FC2AFD94" attach="left" box="[947,961,510,524]" fontSize="6" pageId="15" pageNumber="502">rd</superScript>
|
||
infralabials small, first pair larger and in contact with each other; 1
|
||
<superScript id="A8B79B99270D3206B096FD9DFE39FDA9" attach="none" box="[455,466,547,561]" fontSize="6" pageId="15" pageNumber="502">st</superScript>
|
||
–4
|
||
<superScript id="A8B79B99270D3206B0BFFD9CFE10FDA8" attach="left" box="[494,507,546,560]" fontSize="6" pageId="15" pageNumber="502">th</superScript>
|
||
(40/
|
||
<specimenCount id="49C4FD58270D3206B37FFD9DFD20FDA5" box="[558,715,547,573]" count="49" pageId="15" pageNumber="502" type="generic">49 specimens</specimenCount>
|
||
) or rarely 1
|
||
<superScript id="A8B79B99270D3206B208FD9DFC8FFDA9" attach="none" box="[857,868,547,561]" fontSize="6" pageId="15" pageNumber="502">st</superScript>
|
||
–5
|
||
<superScript id="A8B79B99270D3206B2D1FD9CFC66FDA8" attach="left" box="[896,909,546,560]" fontSize="6" pageId="15" pageNumber="502">th</superScript>
|
||
(9/
|
||
<specimenCount id="49C4FD58270D3206B2E7FD9DFBB8FDA5" box="[950,1107,547,573]" count="49" pageId="15" pageNumber="502" type="generic">49 specimens</specimenCount>
|
||
) infralabials in contact with anterior chin shields, 5
|
||
<superScript id="A8B79B99270D3206B0C8FDF8FE4DFDCC" attach="left" box="[409,422,582,596]" fontSize="6" pageId="15" pageNumber="502">th</superScript>
|
||
and 6
|
||
<superScript id="A8B79B99270D3206B0B4FDF8FE19FDCC" attach="left" box="[485,498,582,596]" fontSize="6" pageId="15" pageNumber="502">th</superScript>
|
||
infralabials greatly enlarged; mental triangular, small; anterior chin shields as long as or slightly longer than posterior chin shields, paired.
|
||
</paragraph>
|
||
<caption id="0BBD6659270D3206B1C6FA95FF2FFA36" ID-DOI="http://doi.org/10.5281/zenodo.13916235" ID-Zenodo-Dep="13916235" httpUri="https://zenodo.org/record/13916235/files/figure.png" pageId="15" pageNumber="502" startId="15.[151,250,1323,1347]" targetBox="[397,1191,669,1299]" targetPageId="15" targetType="figure">
|
||
<paragraph id="5F7D36D1270D3206B1C6FA95FF2FFA36" blockId="15.[151,1437,1323,1454]" pageId="15" pageNumber="502">
|
||
<emphasis id="6DB6EAC3270D3206B1C6FA95FEE4FADB" bold="true" box="[151,271,1323,1347]" pageId="15" pageNumber="502">FIGURE 7</emphasis>
|
||
. Differences in dorsal body band coloration in preservative between
|
||
<taxonomicName id="98C24D52270D3206B28FFA95FB22FADB" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[990,1225,1323,1347]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="502" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC3270D3206B28FFA95FB22FADB" box="[990,1225,1323,1347]" italics="true" pageId="15" pageNumber="502">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6DB6EAC3270D3206B580FA95FACFFADB" bold="true" box="[1233,1316,1323,1347]" pageId="15" pageNumber="502">
|
||
<taxonomicNameLabel id="768557B8270D3206B580FA95FACFFADB" box="[1233,1316,1323,1347]" pageId="15" pageNumber="502" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A, B) and
|
||
<taxonomicName id="98C24D52270D3206B1C6FAEEFEF5FAFF" box="[151,286,1359,1383]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="502" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC3270D3206B1C6FAEEFF4EFAFF" box="[151,165,1360,1383]" italics="true" pageId="15" pageNumber="502">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC3270D3206B1E2FAF1FEF5FAFF" box="[179,286,1359,1383]" italics="true" pageId="15" pageNumber="502">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
(C, D). A—FMNH 180149 from Nakhon Ratchasima, Thailand; B—CAS 229729 from Tanintharyi, Myanmar; C—FMNH 229855 from Selangor, Malaysia; D—FMNH 183772 from Selangor, Malaysia. Images not to scale. Photos by J.L. Lee.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5F7D36D1270D3206B196F9BAFAA6F911" blockId="15.[151,1437,1539,1889]" pageId="15" pageNumber="502">
|
||
<emphasis id="6DB6EAC3270D3206B196F9BAFEDAF985" box="[199,305,1540,1565]" italics="true" pageId="15" pageNumber="502">Dentition</emphasis>
|
||
. Based on
|
||
<specimenCount id="49C4FD58270D3206B0E2F9BDFD88F985" box="[435,611,1539,1565]" count="4" pageId="15" pageNumber="502" type="generic">four specimens</specimenCount>
|
||
, namely CAS 229726 (
|
||
<typeStatus id="80798873270D3206B23FF9BAFC3FF986" box="[878,980,1540,1566]" pageId="15" pageNumber="502" type="paratype">paratype</typeStatus>
|
||
), CAS 235846 (
|
||
<typeStatus id="80798873270D3206B5DFF9BAFB1FF986" box="[1166,1268,1540,1566]" pageId="15" pageNumber="502" type="paratype">paratype</typeStatus>
|
||
), CAS 245971 (
|
||
<typeStatus id="80798873270D3206B1CEF996FEEEF9DA" box="[159,261,1576,1602]" pageId="15" pageNumber="502" type="paratype">paratype</typeStatus>
|
||
), and FMNH 180149 (additional voucher specimen). Total of 12–14 maxillary teeth on each upper jaw, with 8–9 small anterior teeth + a wide diastema, longer than the largest teeth + 5–6 moderately enlarged posterior teeth, without gap, last one very large. The maxilla is strongly arched and distinctly bent inwards anteriorly.
|
||
</paragraph>
|
||
<paragraph id="5F7D36D1270D3206B196F92DFBC6F8F9" blockId="15.[151,1437,1539,1889]" pageId="15" pageNumber="502">
|
||
<emphasis id="6DB6EAC3270D3206B196F92DFE37F935" box="[199,476,1683,1709]" italics="true" pageId="15" pageNumber="502">Hemipenial morphology</emphasis>
|
||
. Based on
|
||
<specimenCount id="49C4FD58270D3206B335F92DFCF7F935" box="[612,796,1683,1709]" count="3" pageId="15" pageNumber="502" type="generic">three specimens</specimenCount>
|
||
(MCZ R-175968 (
|
||
<typeStatus id="80798873270D3206B2A8F92DFBB5F935" box="[1017,1118,1683,1709]" pageId="15" pageNumber="502" type="holotype">holotype</typeStatus>
|
||
) see
|
||
<figureCitation id="C7F92A54270D3206B5F1F92DFB1CF936" box="[1184,1271,1683,1710]" captionStart="FIGURE 5" captionStartId="13.[151,250,1495,1519]" captionTargetBox="[151,1436,181,1471]" captionTargetId="figure-18@13.[151,1436,181,1471]" captionTargetPageId="13" captionText="FIGURE 5. Lycodon neomaculatus sp. nov. in preservative—MCZ R-175968 (holotype, adult male) A—General dorsal view; B—General ventral view; C—Lateral view of the head, right side; D—Lateral view of the head, left side; E—Dorsal view of the head; F—Ventral view of the tail with the everted hemipenes. Images not to scale. Photos by M. Gage and J. Martinez." figureDoi="http://doi.org/10.5281/zenodo.13916229" httpUri="https://zenodo.org/record/13916229/files/figure.png" pageId="15" pageNumber="502">Fig. 5F</figureCitation>
|
||
; CAS 235846 (
|
||
<typeStatus id="80798873270D3206B1CEF906FEE8F94A" box="[159,259,1720,1746]" pageId="15" pageNumber="502" type="paratype">paratype</typeStatus>
|
||
) see
|
||
<figureCitation id="C7F92A54270D3206B06EF909FE79F949" box="[319,402,1719,1745]" captionStart="FIGURE 6" captionStartId="14.[151,250,1495,1519]" captionTargetBox="[151,1436,181,1471]" captionTargetId="figure-29@14.[151,1436,181,1471]" captionTargetPageId="14" captionText="FIGURE 6. Lycodon neomaculatus sp. nov. in preservative—specimen CAS 235846 (paratype, adult male) A—General dorsal view; B—General ventral view; C—dorsal view of the head; D—Ventral view of the head; E—Lateral view of the head, right side; F—Ventral view of the tail with the everted hemipenes. Photos by J.L. Lee." figureDoi="http://doi.org/10.5281/zenodo.13916233" httpUri="https://zenodo.org/record/13916233/files/figure.png" pageId="15" pageNumber="502">Fig. 6F</figureCitation>
|
||
, and USMN 7339 fide
|
||
<bibRefCitation id="3B534B20270D3206B3CDF906FCF7F94A" author="Cope, E. D." box="[668,796,1719,1746]" pageId="15" pageNumber="502" pagination="186 - 219" refId="ref18952" refString="Cope, E. D. (1895). The classification of the Ophidia. Transactions of the American Philosophical Society, 18 (2), 186 - 219, pls. 14 - 33. https: // doi. org / 10.2307 / 1005387" type="journal article" year="1895">Cope 1895</bibRefCitation>
|
||
). The hemipenis is elongate and unilobed. The proximal half of the hemipenis is poorly ornamented with approximately three or four rows of irregularly arranged spines present in asulcate and sulcate profile. The distal half is calyculate, encircled by small, longitudinally arranged flounces each with a single dense row of small spinose calyces. The apical region of the hemipenis is entirely nude. The
|
||
<emphasis id="6DB6EAC3270D3206B19BF8F9FE75F8F9" box="[202,414,1863,1889]" italics="true" pageId="15" pageNumber="502">sulcus spermaticus</emphasis>
|
||
is simple and arranged centripetally, sulcus lips indistinct.
|
||
</paragraph>
|
||
<caption id="0BBD665927123219B1C9FF29FB48FF4B" ID-Table-UUID="0BBD665927123219B1C9FF29FB48FF4B" httpUri="http://table.plazi.org/id/0BBD665927123219B1C9FF29FB48FF4B" pageId="16" pageNumber="503" startId="16.[152,244,151,175]" targetBox="[163,1425,227,1164]" targetIsTable="true" targetPageId="16" targetType="table">
|
||
<paragraph id="5F7D36D127123219B1C9FF29FA76FF37" blockId="16.[151,1437,151,211]" box="[152,1437,151,176]" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B1C9FF29FEE3FF37" bold="true" box="[152,264,151,175]" pageId="16" pageNumber="503">TABLE 1</emphasis>
|
||
. Comparison of morphological characters of
|
||
<taxonomicName id="98C24D5227123219B3BDFF29FC09FF37" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[748,994,151,175]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="503" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC327123219B3BDFF29FC09FF37" box="[748,994,151,175]" italics="true" pageId="16" pageNumber="503">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B827123219B2B9FF26FBD4FF37" box="[1000,1087,152,176]" pageId="16" pageNumber="503" rank="species">
|
||
<emphasis id="6DB6EAC327123219B2B9FF26FBEAFF28" bold="true" box="[1000,1025,152,176]" pageId="16" pageNumber="503">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B55EFF26FBD3FF28" bold="true" box="[1039,1080,152,176]" pageId="16" pageNumber="503">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
with
|
||
<taxonomicName id="98C24D5227123219B52AFF26FAE3FF37" box="[1147,1288,151,175]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="503" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC327123219B52AFF26FB61FF37" box="[1147,1162,152,175]" italics="true" pageId="16" pageNumber="503">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B5C6FF29FAE3FF37" box="[1175,1288,151,175]" italics="true" pageId="16" pageNumber="503">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="98C24D5227123219B46AFF26FA7EFF37" box="[1339,1429,151,175]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="503" phylum="Chordata" rank="species" species="sealei">
|
||
<emphasis id="6DB6EAC327123219B46AFF26FAA1FF37" box="[1339,1354,152,175]" italics="true" pageId="16" pageNumber="503">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B406FF29FA7EFF37" box="[1367,1429,151,175]" italics="true" pageId="16" pageNumber="503">sealei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5F7D36D127123219B1C6FF05FB48FF4B" blockId="16.[151,1437,151,211]" box="[151,1187,187,211]" pageId="16" pageNumber="503">
|
||
Diagnostic differences between the new species and
|
||
<taxonomicName id="98C24D5227123219B398FF02FCBDFF4B" box="[713,854,187,211]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="503" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC327123219B398FF02FD33FF4B" box="[713,728,188,211]" italics="true" pageId="16" pageNumber="503">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B3B4FF05FCBDFF4B" box="[741,854,187,211]" italics="true" pageId="16" pageNumber="503">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="98C24D5227123219B223FF02FC27FF4B" box="[882,972,187,211]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="503" phylum="Chordata" rank="species" species="sealei">
|
||
<emphasis id="6DB6EAC327123219B223FF02FC6AFF4B" box="[882,897,188,211]" italics="true" pageId="16" pageNumber="503">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B2DFFF05FC27FF4B" box="[910,972,187,211]" italics="true" pageId="16" pageNumber="503">sealei</emphasis>
|
||
</taxonomicName>
|
||
are marked in bold.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5F7D36D127123219B1F2FF5DFAC3FBF6" pageId="16" pageNumber="503">
|
||
<table id="2DC2C4712712CDF6B1F2FF5DFA7AFB14" box="[163,1425,227,1164]" gridcols="4" gridrows="12" pageId="16" pageNumber="503">
|
||
<tr id="E1F234932712CDF6B1F2FF5DFA7AFF64" box="[163,1425,227,252]" gridrow="0" pageId="16" pageNumber="503">
|
||
<th id="A2235DEF2712CDF6B1F2FF5DFEDAFF64" box="[163,305,227,252]" gridcol="0" gridrow="0" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B1F2FF5DFF19FF63" bold="true" box="[163,242,227,251]" pageId="16" pageNumber="503">Species</emphasis>
|
||
</th>
|
||
<th id="A2235DEF2712CDF6B01BFF5DFD70FF64" box="[330,667,227,252]" gridcol="1" gridrow="0" pageId="16" pageNumber="503">
|
||
<taxonomicName id="98C24D5227123219B03EFF5AFDF6FF63" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[367,541,227,251]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="503" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC327123219B03EFF5AFE95FF63" bold="true" box="[367,382,228,251]" italics="true" pageId="16" pageNumber="503">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B0DAFF5DFDF6FF63" bold="true" box="[395,541,227,251]" italics="true" pageId="16" pageNumber="503">neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B827123219B375FF5AFD9DFF63" box="[548,630,228,252]" pageId="16" pageNumber="503" rank="species">
|
||
<emphasis id="6DB6EAC327123219B375FF5AFDD7FF64" bold="true" box="[548,572,228,252]" pageId="16" pageNumber="503">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B318FF5AFD9BFF64" bold="true" box="[585,624,228,252]" pageId="16" pageNumber="503">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</th>
|
||
<th id="A2235DEF2712CDF6B3E7FF5DFBA9FF64" box="[694,1090,227,252]" gridcol="2" gridrow="0" pageId="16" pageNumber="503">
|
||
<taxonomicName id="98C24D5227123219B267FF5AFC2AFF63" box="[822,961,227,251]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="503" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC327123219B267FF5AFCAEFF63" bold="true" box="[822,837,228,251]" italics="true" pageId="16" pageNumber="503">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B203FF5DFC2AFF63" bold="true" box="[850,961,227,251]" italics="true" pageId="16" pageNumber="503">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="A2235DEF2712CDF6B50DFF5DFA7AFF64" box="[1116,1425,227,252]" gridcol="3" gridrow="0" pageId="16" pageNumber="503">
|
||
<taxonomicName id="98C24D5227123219B59AFF5AFAC9FF63" box="[1227,1314,227,251]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="503" phylum="Chordata" rank="species" species="sealei">
|
||
<emphasis id="6DB6EAC327123219B59AFF5AFB31FF63" bold="true" box="[1227,1242,228,251]" italics="true" pageId="16" pageNumber="503">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B5B6FF5DFAC9FF63" bold="true" box="[1255,1314,227,251]" italics="true" pageId="16" pageNumber="503">sealei</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="E1F234932712CDF6B1F2FEB5FA7AFED9" box="[163,1425,267,321]" gridrow="1" pageId="16" pageNumber="503">
|
||
<th id="A2235DEF2712CDF6B1F2FEB5FEDAFED9" box="[163,305,267,321]" gridcol="0" gridrow="1" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B1F2FEB2FF1CFEBB" bold="true" box="[163,247,267,292]" pageId="16" pageNumber="503">TaL/TL</emphasis>
|
||
</th>
|
||
<td id="A2235DEF2712CDF6B01BFEB5FD70FED9" box="[330,667,267,321]" gridcol="1" gridrow="1" pageId="16" pageNumber="503">0.178–0.221 (0.201±0.012, n=42)</td>
|
||
<td id="A2235DEF2712CDF6B3E7FEB5FBA9FED9" box="[694,1090,267,321]" gridcol="2" gridrow="1" pageId="16" pageNumber="503">0.168–0.204 (0.187±0.009, n=52)</td>
|
||
<td id="A2235DEF2712CDF6B50DFEB5FA7AFED9" box="[1116,1425,267,321]" gridcol="3" gridrow="1" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B5E6FEB2FB2FFEBC" bold="true" box="[1207,1220,268,292]" pageId="16" pageNumber="503">0</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B59BFEB2FAE2FEBC" bold="true" box="[1226,1289,268,292]" pageId="16" pageNumber="503">154–0</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B441FEB2FB49FED9" bold="true" pageId="16" pageNumber="503">184 (0</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B5F9FE97FB02FED9" bold="true" box="[1192,1257,297,321]" pageId="16" pageNumber="503">168±0</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B5BEFE97FAFEFED9" bold="true" box="[1263,1301,297,321]" pageId="16" pageNumber="503">009</emphasis>
|
||
,
|
||
<emphasis id="6DB6EAC327123219B473FE97FA8BFED9" bold="true" box="[1314,1376,297,321]" pageId="16" pageNumber="503">n=18)</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="E1F234932712CDF6B1F2FEEFFA7AFE1F" box="[163,1425,337,391]" gridrow="2" pageId="16" pageNumber="503">
|
||
<th id="A2235DEF2712CDF6B1F2FEEFFEDAFE1F" box="[163,305,337,391]" gridcol="0" gridrow="2" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B1F2FEEFFF33FEF1" bold="true" box="[163,216,337,361]" pageId="16" pageNumber="503">VEN</emphasis>
|
||
</th>
|
||
<td id="A2235DEF2712CDF6B01BFEEFFD70FE1F" box="[330,667,337,391]" gridcol="1" gridrow="2" pageId="16" pageNumber="503">187–208 (198.98±4.90, n=57)</td>
|
||
<td id="A2235DEF2712CDF6B3E7FEEFFBA9FE1F" box="[694,1090,337,391]" gridcol="2" gridrow="2" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B21EFEEFFCABFE1F" bold="true" pageId="16" pageNumber="503">201–224 (212</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B216FED1FC90FE1F" bold="true" box="[839,891,367,391]" pageId="16" pageNumber="503">59±5</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B2D0FED1FC71FE1F" bold="true" box="[897,922,367,391]" pageId="16" pageNumber="503">50</emphasis>
|
||
,
|
||
<emphasis id="6DB6EAC327123219B2F6FED1FC0EFE1F" bold="true" box="[935,997,367,391]" pageId="16" pageNumber="503">n=47)</emphasis>
|
||
</td>
|
||
<td id="A2235DEF2712CDF6B50DFEEFFA7AFE1F" box="[1116,1425,337,391]" gridcol="3" gridrow="2" pageId="16" pageNumber="503">198–211 (204.68±4.24, n=19)</td>
|
||
</tr>
|
||
<tr id="E1F234932712CDF6B1F2FE28FA7AFE55" box="[163,1425,406,461]" gridrow="3" pageId="16" pageNumber="503">
|
||
<th id="A2235DEF2712CDF6B1F2FE28FEDAFE55" box="[163,305,406,461]" gridcol="0" gridrow="3" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B1F2FE28FF28FE36" bold="true" box="[163,195,406,430]" pageId="16" pageNumber="503">SC</emphasis>
|
||
</th>
|
||
<td id="A2235DEF2712CDF6B01BFE28FD70FE55" box="[330,667,406,461]" gridcol="1" gridrow="3" pageId="16" pageNumber="503">68–84 (75.07±4.65, n=30)</td>
|
||
<td id="A2235DEF2712CDF6B3E7FE28FBA9FE55" box="[694,1090,406,461]" gridcol="2" gridrow="3" pageId="16" pageNumber="503">68–87 (77.37±4.93, n=49)</td>
|
||
<td id="A2235DEF2712CDF6B50DFE28FA7AFE55" box="[1116,1425,406,461]" gridcol="3" gridrow="3" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B586FE29FB5EFE54" bold="true" pageId="16" pageNumber="503">59–69 (63</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B5EAFE0AFB04FE54" bold="true" box="[1211,1263,436,460]" pageId="16" pageNumber="503">58±5</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B5A7FE0AFAE4FE54" bold="true" box="[1270,1295,436,460]" pageId="16" pageNumber="503">35</emphasis>
|
||
,
|
||
<emphasis id="6DB6EAC327123219B44DFE0AFAB1FE54" bold="true" box="[1308,1370,436,460]" pageId="16" pageNumber="503">n=19)</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="E1F234932712CDF6B1F2FE62FA7AFD8A" box="[163,1425,476,530]" gridrow="4" pageId="16" pageNumber="503">
|
||
<th id="A2235DEF2712CDF6B1F2FE62FEDAFD8A" box="[163,305,476,530]" gridcol="0" gridrow="4" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B1F2FE62FF12FE6C" bold="true" box="[163,249,476,500]" pageId="16" pageNumber="503">TOTAL</emphasis>
|
||
</th>
|
||
<td id="A2235DEF2712CDF6B01BFE62FD70FD8A" box="[330,667,476,530]" gridcol="1" gridrow="4" pageId="16" pageNumber="503">260–290 (274.36±7.46, n=42)</td>
|
||
<td id="A2235DEF2712CDF6B3E7FE62FBA9FD8A" box="[694,1090,476,530]" gridcol="2" gridrow="4" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B21EFE63FCABFD8A" bold="true" pageId="16" pageNumber="503">274–310 (289</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B216FE44FC91FD8A" bold="true" box="[839,890,506,530]" pageId="16" pageNumber="503">81±7</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B2D0FE44FC71FD8A" bold="true" box="[897,922,506,530]" pageId="16" pageNumber="503">29</emphasis>
|
||
,
|
||
<emphasis id="6DB6EAC327123219B2F6FE44FC0EFD8A" bold="true" box="[935,997,506,530]" pageId="16" pageNumber="503">n=45)</emphasis>
|
||
</td>
|
||
<td id="A2235DEF2712CDF6B50DFE62FA7AFD8A" box="[1116,1425,476,530]" gridcol="3" gridrow="4" pageId="16" pageNumber="503">261–274 (268.56 ± 5.32, n=19)</td>
|
||
</tr>
|
||
<tr id="E1F234932712CDF6B1F2FD9CFA7AFDC0" box="[163,1425,546,600]" gridrow="5" pageId="16" pageNumber="503">
|
||
<th id="A2235DEF2712CDF6B1F2FD9CFEDAFDC0" box="[163,305,546,600]" gridcol="0" gridrow="5" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B1F2FD9CFF3DFDA2" bold="true" box="[163,214,546,570]" pageId="16" pageNumber="503">SCR</emphasis>
|
||
</th>
|
||
<td id="A2235DEF2712CDF6B01BFD9CFD70FDC0" box="[330,667,546,600]" gridcol="1" gridrow="5" pageId="16" pageNumber="503">25.09–30.21 (27.48±1.26, n=42)</td>
|
||
<td id="A2235DEF2712CDF6B3E7FD9CFBA9FDC0" box="[694,1090,546,600]" gridcol="2" gridrow="5" pageId="16" pageNumber="503">24.37–29.19 (26.71±1.18, n=48)</td>
|
||
<td id="A2235DEF2712CDF6B50DFD9CFA7AFDC0" box="[1116,1425,546,600]" gridcol="3" gridrow="5" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B5E6FD9CFB3BFDA2" bold="true" box="[1207,1232,546,570]" pageId="16" pageNumber="503">22</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B586FD9CFAFCFDA2" bold="true" box="[1239,1303,546,570]" pageId="16" pageNumber="503">61–25</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B44CFD9CFB5EFDCF" bold="true" pageId="16" pageNumber="503">27 (23</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B5EAFD81FB04FDCF" bold="true" box="[1211,1263,575,599]" pageId="16" pageNumber="503">70±0</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327123219B5A7FD81FAE4FDCF" bold="true" box="[1270,1295,575,599]" pageId="16" pageNumber="503">08</emphasis>
|
||
,
|
||
<emphasis id="6DB6EAC327123219B44DFD81FAB1FDCF" bold="true" box="[1308,1370,575,599]" pageId="16" pageNumber="503">n=19)</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="E1F234932712CDF6B1F2FDD6FA7AFD05" box="[163,1425,616,669]" gridrow="6" pageId="16" pageNumber="503">
|
||
<th id="A2235DEF2712CDF6B1F2FDD6FEDAFD05" box="[163,305,616,669]" gridcol="0" gridrow="6" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B1F2FDD6FEF8FD04" bold="true" pageId="16" pageNumber="503">BB+TB (juveniles)</emphasis>
|
||
</th>
|
||
<td id="A2235DEF2712CDF6B01BFDD6FD70FD05" box="[330,667,616,669]" gridcol="1" gridrow="6" pageId="16" pageNumber="503">19–27 (22.78±3.11, n=9)</td>
|
||
<td id="A2235DEF2712CDF6B3E7FDD6FBA9FD05" box="[694,1090,616,669]" gridcol="2" gridrow="6" pageId="16" pageNumber="503">12–33 (20.75±6.13, n=24)</td>
|
||
<td id="A2235DEF2712CDF6B50DFDD6FA7AFD05" box="[1116,1425,616,669]" gridcol="3" gridrow="6" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B58CFDD6FAB8FD05" bold="true" pageId="16" pageNumber="503">8–12 (10.50±1.52, n=6)</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="E1F234932712CDF6B1F2FD12FA7AFD5E" box="[163,1425,684,710]" gridrow="7" pageId="16" pageNumber="503">
|
||
<th id="A2235DEF2712CDF6B1F2FD12FEDAFD5E" box="[163,305,684,710]" gridcol="0" gridrow="7" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B1F2FD13FEF0FD5D" bold="true" box="[163,283,685,709]" pageId="16" pageNumber="503">Largest SL</emphasis>
|
||
</th>
|
||
<td id="A2235DEF2712CDF6B01BFD12FD70FD5E" box="[330,667,684,710]" gridcol="1" gridrow="7" pageId="16" pageNumber="503">6th (n= 25)</td>
|
||
<td id="A2235DEF2712CDF6B3E7FD12FBA9FD5E" box="[694,1090,684,710]" gridcol="2" gridrow="7" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B213FD13FCA4FD5D" bold="true" box="[834,847,685,709]" pageId="16" pageNumber="503">7</emphasis>
|
||
<emphasis id="6DB6EAC327123219B21EFD12FCB7FD22" bold="true" box="[847,860,684,698]" pageId="16" pageNumber="503">th</emphasis>
|
||
<emphasis id="6DB6EAC327123219B233FD13FC5EFD5D" bold="true" box="[866,949,685,709]" pageId="16" pageNumber="503">(n = 54)</emphasis>
|
||
</td>
|
||
<td id="A2235DEF2712CDF6B50DFD12FA7AFD5E" box="[1116,1425,684,710]" gridcol="3" gridrow="7" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B5ECFD13FB21FD5D" bold="true" box="[1213,1226,685,709]" pageId="16" pageNumber="503">7</emphasis>
|
||
<emphasis id="6DB6EAC327123219B598FD12FB3DFD22" bold="true" box="[1225,1238,684,698]" pageId="16" pageNumber="503">th</emphasis>
|
||
<emphasis id="6DB6EAC327123219B58CFD13FADAFD5D" bold="true" box="[1245,1329,685,709]" pageId="16" pageNumber="503">(n = 19)</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="E1F234932712CDF6B1F2FD6BFA7AFD76" box="[163,1425,725,750]" gridrow="8" pageId="16" pageNumber="503">
|
||
<th id="A2235DEF2712CDF6B1F2FD6BFEDAFD76" box="[163,305,725,750]" gridcol="0" gridrow="8" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B1F2FD6BFEF3FD75" bold="true" box="[163,280,725,749]" pageId="16" pageNumber="503">Body color</emphasis>
|
||
</th>
|
||
<td id="A2235DEF2712CDF6B01BFD6BFD70FD76" box="[330,667,725,750]" gridcol="1" gridrow="8" pageId="16" pageNumber="503">black or black-gray</td>
|
||
<td id="A2235DEF2712CDF6B3E7FD6BFBA9FD76" box="[694,1090,725,750]" gridcol="2" gridrow="8" pageId="16" pageNumber="503">black</td>
|
||
<td id="A2235DEF2712CDF6B50DFD6BFA7AFD76" box="[1116,1425,725,750]" gridcol="3" gridrow="8" pageId="16" pageNumber="503">black</td>
|
||
</tr>
|
||
<tr id="E1F234932712CDF6B1F2FD43FA7AFCC9" box="[163,1425,765,849]" gridrow="9" pageId="16" pageNumber="503">
|
||
<th id="A2235DEF2712CDF6B1F2FD43FEDAFCC9" box="[163,305,765,849]" gridcol="0" gridrow="9" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B1F2FD43FF0FFCAB" bold="true" pageId="16" pageNumber="503">Color of bands</emphasis>
|
||
</th>
|
||
<td id="A2235DEF2712CDF6B01BFD43FD70FCC9" box="[330,667,765,849]" gridcol="1" gridrow="9" pageId="16" pageNumber="503">cream or white, relatively wide in juveniles, speckled and reticulated with gray in adults</td>
|
||
<td id="A2235DEF2712CDF6B3E7FD43FBA9FCC9" box="[694,1090,765,849]" gridcol="2" gridrow="9" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B397FD43FBA9FCAB" bold="true" pageId="16" pageNumber="503">white, width variable in juveniles, smeared with black or gray in adults</emphasis>
|
||
</td>
|
||
<td id="A2235DEF2712CDF6B50DFD43FA7AFCC9" box="[1116,1425,765,849]" gridcol="3" gridrow="9" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B5D2FD43FA9AFCC8" bold="true" pageId="16" pageNumber="503">white, width broad in juveniles, smeared with black or gray in adults</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="E1F234932712CDF6B1F2FCDEFA7AFC76" box="[163,1425,864,1006]" gridrow="10" pageId="16" pageNumber="503">
|
||
<th id="A2235DEF2712CDF6B1F2FCDEFEDAFC76" box="[163,305,864,1006]" gridcol="0" gridrow="10" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B1F2FCDEFEDAFCE0" bold="true" box="[163,305,864,888]" pageId="16" pageNumber="503">Distributions</emphasis>
|
||
</th>
|
||
<td id="A2235DEF2712CDF6B01BFCDEFD70FC76" box="[330,667,864,1006]" gridcol="1" gridrow="10" pageId="16" pageNumber="503">China, Myanmar, Vietnam, Laos, Cambodia, Thailand (except southern peninsular)</td>
|
||
<td id="A2235DEF2712CDF6B3E7FCDEFBA9FC76" box="[694,1090,864,1006]" gridcol="2" gridrow="10" pageId="16" pageNumber="503">Thailand (peninsular), Malaysia (peninsular), Singapore, Indonesia (Sumatra, Java, Bali, Nusa Tenggar, Alor, Flores, Sumba, Sumba), East Timor, India (Nicobar Islands?)</td>
|
||
<td id="A2235DEF2712CDF6B50DFCDEFA7AFC76" box="[1116,1425,864,1006]" gridcol="3" gridrow="10" pageId="16" pageNumber="503">Philippines (Palawan, Mindanao?), Malaysia (Sabah, Sarawak), Indonesia (West & Central Kalimantan), Brunei Darussalam</td>
|
||
</tr>
|
||
<tr id="E1F234932712CDF6B1F2FC40FA7AFB14" box="[163,1425,1022,1164]" gridrow="11" pageId="16" pageNumber="503">
|
||
<th id="A2235DEF2712CDF6B1F2FC40FEDAFB14" box="[163,305,1022,1164]" gridcol="0" gridrow="11" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B1F2FC40FEF2FB8E" bold="true" box="[163,281,1022,1046]" pageId="16" pageNumber="503">References</emphasis>
|
||
</th>
|
||
<td id="A2235DEF2712CDF6B01BFC40FD70FB14" box="[330,667,1022,1164]" gridcol="1" gridrow="11" pageId="16" pageNumber="503">
|
||
<bibRefCitation id="3B534B2027123219B036FC40FE1AFB8F" author="Smith, M. A." box="[359,497,1022,1047]" pageId="16" pageNumber="503" pagination="195 - 212" refId="ref22878" refString="Smith, M. A. (1923) On a collection of reptiles and batrachians from the island of Hainan. Journal of the Natural History Society of Siam, 6, 195 - 212." type="journal article" year="1923">Smith (1923)</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B2027123219B0AAFC41FD90FB8E" author="Pope, C. H." box="[507,635,1022,1046]" pageId="16" pageNumber="503" refId="ref21916" refString="Pope, C. H. (1935) The reptiles of China. Turtles, crocodilians, snakes, lizards. Natural History of Central Asia. Vol. 10. American Museum of Natural History, New York, New York, 604 pp. https: // doi. org / 10.5962 / bhl. title. 12104" type="book" year="1935">Pope (1935)</bibRefCitation>
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||
;
|
||
<bibRefCitation id="3B534B2027123219B01AFBA2FDEEFBAC" author="Zhao, E. M. & Huang, M. H. & Zong, Y. & Jiang, Y. M. & Huang, Q. Y. & Zhao, H. & Ma, J. F. & Zheng, J. & Huang, Z. J. & Wei, G. & Yang, D. Q. & Li, D. J." box="[331,517,1052,1076]" pageId="16" pageNumber="503" refId="ref24466" refString="Zhao, E. M., Huang, M. H., Zong, Y., Jiang, Y. M., Huang, Q. Y., Zhao, H., Ma, J. F., Zheng, J., Huang, Z. J., Wei, G., Yang, D. Q. & Li, D. J. (1998) Fauna Sinica Reptilia. Vol. 3. Squamata: Serpentes. Science Press, Beijing, 522 pp. [in Chinese]" type="book" year="1998">
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||
Zhao
|
||
<emphasis id="6DB6EAC327123219B0D4FBA3FE59FBAC" box="[389,434,1052,1076]" italics="true" pageId="16" pageNumber="503">et al</emphasis>
|
||
. (1998)
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B2027123219B35EFBA2FE75FBC9" author="Geissler, P. & Nguyen, T. Q. & Poyarkov, N. A. & Bohme, W." pageId="16" pageNumber="503" pagination="9 - 16" refId="ref19433" refString="Geissler, P., Nguyen, T. Q., Poyarkov, N. A. & Bohme, W. (2011) New records of snakes from Cat Tien National Park Dong Nai and Lam Dong provinces southern Vietnam. Bonn Zoological Bulletin, 60 (1), 9 - 16." type="journal article" year="2011">
|
||
Geissler
|
||
<emphasis id="6DB6EAC327123219B339FBA3FD7EFBAC" box="[616,661,1052,1076]" italics="true" pageId="16" pageNumber="503">et al</emphasis>
|
||
. (2011)
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B2027123219B0F6FB84FDA3FBC9" author="Le, D. T. & Luong, A. M. & Pham, C. T. & Pham, T. Q. & Nguyen, S. L. H. & Ziegler, T. & Nguyen, T. Q." box="[423,584,1081,1105]" pageId="16" pageNumber="503" pagination="201 - 219" refId="ref20602" refString="Le, D. T., Luong, A. M., Pham, C. T., Pham, T. Q., Nguyen, S. L. H., Ziegler, T. & Nguyen, T. Q. (2021) New records and an updated checklist of amphibians and snakes from Tuyen Quang Province, Vietnam. Bonn zoological Bulletin, 70 (1), 201 - 219. https: // doi. org / 10.20363 / BZB- 2021.70.1.201" type="journal article" year="2021">
|
||
Le
|
||
<emphasis id="6DB6EAC327123219B098FB84FE1EFBC9" box="[457,501,1081,1105]" italics="true" pageId="16" pageNumber="503">et al</emphasis>
|
||
. (2021)
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B2027123219B303FB87FE50FBF7" author="Liu, S. & Hou, M. & Cai, B. & Li, S. & Zhang, Z. & Yu, R. & Rao, D. & Zhang, L." pageId="16" pageNumber="503" pagination="307 - 316" refId="ref21074" refString="Liu, S., Hou, M., Cai, B., Li, S., Zhang, Z., Yu, R., Rao, D. & Zhang, L. (2023) Taxonomic status of Lycodon subcinctus sensu lato in China (Serpentes, Colubridae). Herpetozoa, 36, 307 - 316. https: // doi. org / 10.3897 / herpetozoa. 36. e 114206" type="journal article" year="2023">
|
||
Liu
|
||
<emphasis id="6DB6EAC327123219B32DFB84FE82FBF6" italics="true" pageId="16" pageNumber="503">et al</emphasis>
|
||
. (2023)
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B2027123219B097FBE8FD67FBF7" author="Zhang, H. Y. & Wang, L. Y. & Hou, S. B. & Wu, Y. H. & Duan, Z. P. & Jin, Y. D. & Zuo, C. S. & Zhang, Y. S. & Wang, K. & Che, J." box="[454,652,1110,1135]" pageId="16" pageNumber="503" pagination="81 - 94" refId="ref24266" refString="Zhang, H. Y., Wang, L. Y., Hou, S. B., Wu, Y. H., Duan, Z. P., Jin, Y. D., Zuo, C. S., Zhang, Y. S., Wang, K. & Che, J. (2024) Provincial records of Lycodon septentrionalis and L. cf. subcinctus from Yunnan Province, China. Chinese Journal of Zoology, 59 (1), 81 - 94. [in Chinese] https: // doi. org / 10.13859 / j. cjz. 202423018" type="journal article" year="2024">
|
||
Zhang
|
||
<emphasis id="6DB6EAC327123219B35CFBE9FDD1FBF6" box="[525,570,1110,1134]" italics="true" pageId="16" pageNumber="503">et al</emphasis>
|
||
. (2024)
|
||
</bibRefCitation>
|
||
; this study
|
||
</td>
|
||
<td id="A2235DEF2712CDF6B3E7FC40FBA9FB14" box="[694,1090,1022,1164]" gridcol="2" gridrow="11" pageId="16" pageNumber="503">
|
||
<bibRefCitation id="3B534B2027123219B39AFC41FC85FB8F" author="Sauvage, H. E." box="[715,878,1023,1047]" pageId="16" pageNumber="503" pagination="142 - 147" refId="ref22733" refString="Sauvage, H. E. (1884) Sur quelques reptiles de la collection du Museum d'histoire naturelle. Bulletin de la Societe Philomathique de Paris, 7 (1883 - 1884), 142 - 147." type="journal article" year="1884">Sauvage (1884)</bibRefCitation>
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||
;
|
||
<bibRefCitation id="3B534B2027123219B229FC40FBC2FB8E" author="Nutphand, W." box="[888,1065,1022,1046]" pageId="16" pageNumber="503" pagination="153 - 159" refId="ref21551" refString="Nutphand, W. (1986) Ngoo plang chanuan. Wolf snakes. Thai Zoological Center, Bangkok, 1 (9), 153 - 159. [in Thai]" type="journal article" year="1986">Nutphand (1986)</bibRefCitation>
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,
|
||
<bibRefCitation id="3B534B2027123219B39FFBA2FC43FBAC" author="Pauwels, O. S. G. & David, P." box="[718,936,1052,1076]" pageId="16" pageNumber="503" pagination="114 - 120" refId="ref21857" refString="Pauwels, O. S. G., David, P. & Chan-ard, T. (2006) Dryocalamus davisonii tungsongensis Nutphand 1986 and Lycodon suratensis Nutphand 1986 (Serpentes: Colubridae): translation of their original description and taxonomic status. Hamadryad, 30 (1 - 2), 114 - 120." type="book chapter" year="2006">
|
||
Pauwels
|
||
<emphasis id="6DB6EAC327123219B278FBA3FCBEFBAC" box="[809,853,1052,1076]" italics="true" pageId="16" pageNumber="503">et al</emphasis>
|
||
. (2006)
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B2027123219B2E2FBA2FC83FBC9" author="David, P. & Teynie, A. & Vogel, G." pageId="16" pageNumber="503" refId="ref19064" refString="David, P., Teynie, A. & Vogel, G. (2023) The Snakes of Laos. Edition Chimaira, Frankfurt am Main, 960 pp." type="book" year="2023">
|
||
David
|
||
<emphasis id="6DB6EAC327123219B2A6FBA3FBCFFBAC" box="[1015,1060,1052,1076]" italics="true" pageId="16" pageNumber="503">et al</emphasis>
|
||
. (2023)
|
||
</bibRefCitation>
|
||
; this study
|
||
</td>
|
||
<td id="A2235DEF2712CDF6B50DFC40FA7AFB14" box="[1116,1425,1022,1164]" gridcol="3" gridrow="11" pageId="16" pageNumber="503">
|
||
<bibRefCitation id="3B534B2027123219B524FC40FAC7FB8F" author="Boulenger, G. A." box="[1141,1324,1022,1047]" pageId="16" pageNumber="503" refId="ref18624" refString="Boulenger, G. A. (1893) Catalogue of the snakes in the British Museum (Natural History), Vol. I. Containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae, and Colubridae Aglyphae, part. Order of the Trustees, British Museum (Natural History), London, xiii + 448 pp., 28 pls. https: // doi. org / 10.5962 / bhl. title. 8316" type="book" year="1893">Boulenger (1893)</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B2027123219B467FC40FAF3FBAC" author="Taylor, E. H." pageId="16" pageNumber="503" refId="ref23151" refString="Taylor, E. H. (1922) The snakes of the Philippine Islands. Philippine Bureau of Science Publications, Manila, 312 pp. https: // doi. org / 10.5962 / bhl. title. 55346" type="book" year="1922">Taylor (1922)</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B2027123219B50DFB87FB13FBC9" author="Leviton, A. E." box="[1116,1272,1081,1105]" pageId="16" pageNumber="503" pagination="195 - 203" refId="ref20767" refString="Leviton, A. E. (1955) Systematic notes on the Asian snake Lycodon subcinctus. The Philippine Journal of Science, 84 (2), 195 - 203." type="journal article" year="1955">Leviton (1955)</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B2027123219B453FB84FA66FBC9" author="Lanza, B." box="[1282,1421,1081,1105]" pageId="16" pageNumber="503" pagination="89 - 104" refId="ref20480" refString="Lanza, B. (1999) A new species of Lycodon from the Philippines, with a key to the genus (Reptilia Serpentes Colubridae). Tropical Zoology, 12 (1), 89 - 104. https: // doi. org / 10.1080 / 03946975.1999.10539380" type="journal article" year="1999">Lanza (1999)</bibRefCitation>
|
||
; this study
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="5F7D36D127123219B196FB5CFAD5F8F8" blockId="16.[151,1437,1250,1888]" pageId="16" pageNumber="503">
|
||
<emphasis id="6DB6EAC327123219B196FB5CFEA9FB64" box="[199,322,1250,1276]" italics="true" pageId="16" pageNumber="503">Coloration</emphasis>
|
||
(
|
||
<figureCitation id="C7F92A5427123219B005FB5CFE48FB64" box="[340,419,1250,1276]" captionStart="FIGURE 1" captionStartId="8.[151,250,1495,1519]" captionTargetBox="[151,1436,181,1471]" captionTargetId="figure-29@8.[151,1436,181,1471]" captionTargetPageId="8" captionText="FIGURE 1. Colubrid snake species involved in the case of Anoplophallus maculatus Cope, 1895.—USNM 7339, previously considered the holotype of “Lycodon maculatus” reported by Liu et al. (2023): A—General dorsal view; B—General ventral view; C—Dorsalview ofthe head;D—Lateralview of thehead,leftside; E—Livespecimen of Leptodeira maculata (= Megalops maculatus) from Minatitlán, Colima, Mexico; F—Live specimen of Lycodon subannulatus from Penang Island, Malaysia, of which Nymphophidium maculatum is a junior synonym. Images not to scale. Photos by T. Hsu (A–D), C. Grünwald (E), J. L. Lee (F)." figureDoi="http://doi.org/10.5281/zenodo.13916221" httpUri="https://zenodo.org/record/13916221/files/figure.png" pageId="16" pageNumber="503">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="C7F92A5427123219B0E5FB5CFE2FFB65" box="[436,452,1250,1277]" captionStart="FIGURE 5" captionStartId="13.[151,250,1495,1519]" captionTargetBox="[151,1436,181,1471]" captionTargetId="figure-18@13.[151,1436,181,1471]" captionTargetPageId="13" captionText="FIGURE 5. Lycodon neomaculatus sp. nov. in preservative—MCZ R-175968 (holotype, adult male) A—General dorsal view; B—General ventral view; C—Lateral view of the head, right side; D—Lateral view of the head, left side; E—Dorsal view of the head; F—Ventral view of the tail with the everted hemipenes. Images not to scale. Photos by M. Gage and J. Martinez." figureDoi="http://doi.org/10.5281/zenodo.13916229" httpUri="https://zenodo.org/record/13916229/files/figure.png" pageId="16" pageNumber="503">5</figureCitation>
|
||
,
|
||
<figureCitation id="C7F92A5427123219B083FB5CFE09FB64" box="[466,482,1250,1276]" captionStart="FIGURE 6" captionStartId="14.[151,250,1495,1519]" captionTargetBox="[151,1436,181,1471]" captionTargetId="figure-29@14.[151,1436,181,1471]" captionTargetPageId="14" captionText="FIGURE 6. Lycodon neomaculatus sp. nov. in preservative—specimen CAS 235846 (paratype, adult male) A—General dorsal view; B—General ventral view; C—dorsal view of the head; D—Ventral view of the head; E—Lateral view of the head, right side; F—Ventral view of the tail with the everted hemipenes. Photos by J.L. Lee." figureDoi="http://doi.org/10.5281/zenodo.13916233" httpUri="https://zenodo.org/record/13916233/files/figure.png" pageId="16" pageNumber="503">6</figureCitation>
|
||
,
|
||
<figureCitation id="C7F92A5427123219B0A1FB5CFDDDFB64" box="[496,566,1250,1276]" captionStart="FIGURE 7" captionStartId="15.[151,250,1323,1347]" captionTargetBox="[397,1191,669,1299]" captionTargetId="figure-260@15.[397,1191,669,1299]" captionTargetPageId="15" captionText="FIGURE 7. Differences in dorsal body band coloration in preservative between Lycodon neomaculatus sp. nov. (A, B) and L. subcinctus (C, D). A—FMNH 180149 from Nakhon Ratchasima, Thailand; B—CAS 229729 from Tanintharyi, Myanmar; C—FMNH 229855 from Selangor, Malaysia; D—FMNH 183772 from Selangor, Malaysia. Images not to scale. Photos by J.L. Lee." figureDoi="http://doi.org/10.5281/zenodo.13916235" httpUri="https://zenodo.org/record/13916235/files/figure.png" pageId="16" pageNumber="503">7A–B</figureCitation>
|
||
,
|
||
<figureCitation id="C7F92A5427123219B315FB5CFDBFFB64" box="[580,596,1250,1276]" captionStart="FIGURE 8" captionStartId="17.[151,250,1084,1108]" captionTargetBox="[396,1191,181,1060]" captionTargetId="figure-18@17.[396,1191,181,1060]" captionTargetPageId="17" captionText="FIGURE 8. Lycodon neomaculatus sp. nov. in life (not collected, sex not determined): A, B—Hong Kong, China (adult, subadult; respectively); C—Qiongzhong, Hainan, China (adult); D—Xishuangbanna, Yunnan, China; E—Pu Mat NP, Nghe An, Vietnam; F—Ma Da NR, Dong Nai, Vietnam (adult); G—Cat Tien NP, Dong Nai, Vietnam (subadult); H—Loc Bac, Lam Dong, Vietnam (adult); I—Phuoc Binh NP, Ninh Thuan, Vietnam (adult); J—Hinhurp, Vientiane, Laos (subadult); K—Kompong Seila, Seihanuk, Cambodia (adult); L—Chai Prakarn, Chiang Mai, Thailand (subadult); M—Mae Yom NP, Phrae, Thailand (adult); N—Wang Nam Khiao, Nakhon Ratchasima, Thailand (adult); O—Kaeng Krachan NP, Phetchaburi, Thailand (subadult). Photos by:A. Tomaszek (A, B); C.X. Liao (C); Z.H. Ma (D); P.S. Hamilton (E); H.X.N. Nguyen (F); A.M. Bragin (G, I); N.A. Poyarkov (H); T. Calame (J); T. Samnang (K); A. Kaosung (L); R. Grassby-Lewis (M, N); A. Pierce (O)." figureDoi="http://doi.org/10.5281/zenodo.13916237" httpUri="https://zenodo.org/record/13916237/files/figure.png" pageId="16" pageNumber="503">8</figureCitation>
|
||
). Dorsal surface black or black gray anteriorly, gradually transitioning to gray brown or pale brown at midbody and posteriorly; lighter brown along flanks; 5–10 white or cream colored body bands in adults, widely separated, about 4–7 dorsal scales long on the vertebral row, slightly expanded ventrolaterally, 7–10 dorsal scales long above the ventral plates; the first body band starts at the level of the 15
|
||
<superScript id="A8B79B9927123219B40AFAF0FA8DFAC4" attach="none" box="[1371,1382,1358,1372]" fontSize="6" pageId="16" pageNumber="503">st</superScript>
|
||
–20
|
||
<superScript id="A8B79B9927123219B4C1FAF3FA76FAC3" attach="right" box="[1424,1437,1357,1371]" fontSize="6" pageId="16" pageNumber="503">th</superScript>
|
||
ventral scale, and the first two to four cross-bands are separated from one another by 10–16 vertebral dorsal scales; in adults, the dorsal bands become progressively more heavily speckled with dark brown along the anterior portion of each dorsal scale until the band becomes completely subdued with dark pigment and appears indistinct relative to the rest of the dorsal surface; the posteriormost bands along the body and tail darken first, and frequently only 4–7 of the anteriormost pale bands are visible in large specimens; remaining posterior portion of the dorsum is plain in adults; tail tip is cream. Dorsal surface of the head blackish-brown or black; in some specimens the posterior half of the head beyond the frontal scale is slightly marbled with shades of brown, but in larger individuals the marbling is restricted to the posterior half of the parietals and the adjacent occipital scales; snout dark brown, with dark pigment normally invading the first two supralabials and ventral surface around the mental scale; supralabials cream or white, especially in their lower half; chin and throat plain white, sometimes pale yellow, with occasional dark brown spots on the first few infralabials; eye brown in life, pupil black. Venter white, cream or pale yellow, with a dark brown, diffuse spot of the anterior edge of each ventral scale; progressively, the ventral scales become more strongly speckled or tinted with brown at midline; in some specimens the entire ventral surface is plain cream or white. The underside of the tail is plain white but in some specimens is dark grey with white or pale yellow marbling.
|
||
</paragraph>
|
||
<caption id="0BBD665927133218B1C6FB82FBBDFAC8" ID-DOI="http://doi.org/10.5281/zenodo.13916237" ID-Zenodo-Dep="13916237" httpUri="https://zenodo.org/record/13916237/files/figure.png" pageId="17" pageNumber="504" startId="17.[151,250,1084,1108]" targetBox="[396,1191,181,1060]" targetPageId="17" targetType="figure">
|
||
<paragraph id="5F7D36D127133218B1C6FB82FBBDFAC8" blockId="17.[151,1436,1084,1361]" pageId="17" pageNumber="504">
|
||
<emphasis id="6DB6EAC327133218B1C6FB82FEFBFBCC" bold="true" box="[151,272,1084,1108]" pageId="17" pageNumber="504">FIGURE 8</emphasis>
|
||
.
|
||
<taxonomicName id="98C24D5227133218B071FB82FDE5FBCC" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[288,526,1084,1108]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="504" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC327133218B071FB82FDE5FBCC" box="[288,526,1084,1108]" italics="true" pageId="17" pageNumber="504">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B827133218B346FB82FD86FBCC" box="[535,621,1084,1108]" pageId="17" pageNumber="504" rank="species">
|
||
<emphasis id="6DB6EAC327133218B346FB82FDC4FBCC" bold="true" box="[535,559,1084,1108]" pageId="17" pageNumber="504">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327133218B36EFB82FD8DFBCC" bold="true" box="[575,614,1084,1108]" pageId="17" pageNumber="504">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
in life (not collected, sex not determined): A, B—Hong Kong, China (adult, subadult; respectively); C—Qiongzhong, Hainan, China (adult); D—Xishuangbanna, Yunnan, China; E—Pu Mat NP, Nghe An, Vietnam; F—Ma Da NR, Dong Nai, Vietnam (adult); G—Cat Tien NP, Dong Nai, Vietnam (subadult); H—Loc Bac, Lam Dong, Vietnam (adult); I—Phuoc Binh NP, Ninh Thuan, Vietnam (adult); J—Hinhurp, Vientiane, Laos (subadult); K—Kompong Seila, Seihanuk, Cambodia (adult); L—Chai Prakarn, Chiang Mai, Thailand (subadult); M—Mae Yom NP, Phrae, Thailand (adult); N—Wang Nam Khiao, Nakhon Ratchasima, Thailand (adult); O—Kaeng Krachan NP, Phetchaburi, Thailand (subadult). Photos by: A. Tomaszek (A, B); C.X. Liao (C); Z.H. Ma (D); P.S. Hamilton (E); H.X.N. Nguyen (F); A.M. Bragin (G, I); N.A. Poyarkov (H); T. Calame (J); T. Samnang (K); A. Kaosung (L); R. Grassby-Lewis (M, N); A. Pierce (O).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5F7D36D127133218B196FA1CFCB8F9E8" blockId="17.[151,1437,1442,2009]" pageId="17" pageNumber="504">Juvenile specimens are more strongly patterned than adults and are usually black dorsally with 11–19 white, uniform bands on the body, generally visible up to the vent, and 7–12 bands on the tail. The head is largely as black along the dorsal surface and on the sides of the snout, however the posterior portion of the head has a white facial collar present across the entirety of the parietals, the posterior half of the frontal, most of the temporals and the remaining occipital region and nape; supralabials white or cream, occasionally with dark gray mottling near the snout and eyes. The ventral surface is uniform cream or white.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="17D8655A2713321AB196F9C4FBBCFA3A" lastPageId="19" lastPageNumber="506" pageId="17" pageNumber="504" type="discussion">
|
||
<paragraph id="5F7D36D127133218B196F9C4FC0DF899" blockId="17.[151,1437,1442,2009]" pageId="17" pageNumber="504">
|
||
<emphasis id="6DB6EAC327133218B196F9C4FE8DF90C" bold="true" box="[199,358,1658,1684]" pageId="17" pageNumber="504">Comparisons</emphasis>
|
||
.
|
||
<taxonomicName id="98C24D5227133218B029F9C4FD94F90C" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[376,639,1658,1684]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="504" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC327133218B029F9C4FD94F90C" box="[376,639,1658,1684]" italics="true" pageId="17" pageNumber="504">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B827133218B3D8F9C5FD03F90D" box="[649,744,1659,1685]" pageId="17" pageNumber="504" rank="species">
|
||
<emphasis id="6DB6EAC327133218B3D8F9C5FD48F90D" bold="true" box="[649,675,1659,1685]" pageId="17" pageNumber="504">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327133218B3E7F9C5FD09F90D" bold="true" box="[694,738,1659,1685]" pageId="17" pageNumber="504">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
can be easily distinguished from all other known
|
||
<taxonomicName id="98C24D5227133218B46CF9C4FF03F920" pageId="17" pageNumber="504">
|
||
<emphasis id="6DB6EAC327133218B46CF9C4FA76F90C" box="[1341,1437,1658,1684]" italics="true" pageId="17" pageNumber="504">Lycodon</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(except
|
||
<taxonomicName id="98C24D5227133218B01AF921FE0BF920" box="[331,480,1694,1721]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="504" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC327133218B01AF921FEB0F920" box="[331,347,1695,1720]" italics="true" pageId="17" pageNumber="504">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327133218B03BF920FE0BF920" box="[362,480,1694,1720]" italics="true" pageId="17" pageNumber="504">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="98C24D5227133218B0A1F921FDBBF920" box="[496,592,1694,1721]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="504" phylum="Chordata" rank="species" species="sealei">
|
||
<emphasis id="6DB6EAC327133218B0A1F921FDEBF920" box="[496,512,1695,1720]" italics="true" pageId="17" pageNumber="504">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327133218B35EF920FDBBF920" box="[527,592,1694,1720]" italics="true" pageId="17" pageNumber="504">sealei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="98C24D5227133218B30EF921FB88F920" authority="Wostl, Hamidy, Kurniawan & Smith" authorityName="Wostl, Hamidy, Kurniawan & Smith" box="[607,1123,1694,1721]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="504" phylum="Chordata" rank="species" species="sidiki">
|
||
<emphasis id="6DB6EAC327133218B30EF921FD84F920" box="[607,623,1695,1720]" italics="true" pageId="17" pageNumber="504">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327133218B32FF920FD50F920" box="[638,699,1694,1720]" italics="true" pageId="17" pageNumber="504">sidiki</emphasis>
|
||
Wostl, Hamidy, Kurniawan & Smith
|
||
</taxonomicName>
|
||
) by the lack of a preocular, and by having both the loreal and prefrontal scales enter the orbit. We summarize the main characters separating
|
||
<taxonomicName id="98C24D5227133218B1C6F958FE71F898" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[151,410,1766,1792]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="504" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC327133218B1C6F958FE71F898" box="[151,410,1766,1792]" italics="true" pageId="17" pageNumber="504">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B827133218B0F1F959FE10F899" box="[416,507,1767,1793]" pageId="17" pageNumber="504" rank="species">
|
||
<emphasis id="6DB6EAC327133218B0F1F959FE51F899" bold="true" box="[416,442,1767,1793]" pageId="17" pageNumber="504">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327133218B098F959FE1EF899" bold="true" box="[457,501,1767,1793]" pageId="17" pageNumber="504">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from
|
||
<taxonomicName id="98C24D5227133218B311F959FD38F898" box="[576,723,1766,1793]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="504" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC327133218B311F959FDBBF898" box="[576,592,1767,1792]" italics="true" pageId="17" pageNumber="504">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327133218B30CF958FD38F898" box="[605,723,1766,1792]" italics="true" pageId="17" pageNumber="504">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="98C24D5227133218B25BF959FC82F898" box="[778,873,1766,1793]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="504" phylum="Chordata" rank="species" species="sealei">
|
||
<emphasis id="6DB6EAC327133218B25BF959FCF1F898" box="[778,794,1767,1792]" italics="true" pageId="17" pageNumber="504">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327133218B279F958FC82F898" box="[808,873,1766,1792]" italics="true" pageId="17" pageNumber="504">sealei</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<tableCitation id="1240036A27133218B2DDF958FC0BF899" box="[908,992,1766,1793]" captionStart="TABLE 1" captionStartId="16.[152,244,151,175]" captionTargetPageId="16" captionText="TABLE 1. Comparison of morphological characters of Lycodon neomaculatus sp. nov. with L. subcinctus and L. sealei. Diagnostic differences between the new species and L. subcinctus + L. sealei are marked in bold." httpUri="http://table.plazi.org/id/0BBD665927123219B1C9FF29FB48FF4B" pageId="17" pageNumber="504" tableUuid="0BBD665927123219B1C9FF29FB48FF4B">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5F7D36D12713321BB196F8B4FD94FF61" blockId="17.[151,1437,1442,2009]" lastBlockId="18.[151,1436,151,249]" lastPageId="18" lastPageNumber="505" pageId="17" pageNumber="504">
|
||
<taxonomicName id="98C24D5227133218B196F8B4FE20F8BC" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[199,459,1802,1828]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="504" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC327133218B196F8B4FE20F8BC" box="[199,459,1802,1828]" italics="true" pageId="17" pageNumber="504">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B827133218B083F8B5FDC5F8BD" box="[466,558,1803,1829]" pageId="17" pageNumber="504" rank="species">
|
||
<emphasis id="6DB6EAC327133218B083F8B5FE07F8BD" bold="true" box="[466,492,1803,1829]" pageId="17" pageNumber="504">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327133218B0AAF8B5FDCCF8BD" bold="true" box="[507,551,1803,1829]" pageId="17" pageNumber="504">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="98C24D5227133218B395F8B5FCB2F8BC" box="[708,857,1802,1829]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="504" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC327133218B395F8B5FD3FF8BC" box="[708,724,1803,1828]" italics="true" pageId="17" pageNumber="504">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC327133218B3B2F8B4FCB2F8BC" box="[739,857,1802,1828]" italics="true" pageId="17" pageNumber="504">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
by having: a lower number of ventrals (
|
||
<collectingRegion id="9D06F83327133218B470F8B5FAB7F8BD" box="[1313,1372,1803,1829]" country="Austria" name="Wien" pageId="17" pageNumber="504">VEN</collectingRegion>
|
||
187– 208 [mean = 198.98 ± 4.90] vs. 201–224 [mean = 212.59 ± 5.50]), a lower number of total body scales (TOTAL 260–290 [mean = 274.36 ± 7.46] vs. 274–311 [mean = 289.81 ± 7.29]), relative size of the largest supralabial scale (sixth supralabial largest vs. seventh supralabial largest), and by the pattern of pigmentation on the white bands in adults (white bands darkening along the anterior edge of each dorsal scale, creating a speckled, reticulate or ‘maculate’ patterning vs. white bands darkening along the entire dorsal scale, pattern more smeared, patched or ‘piebald’ in appearance; see
|
||
<figureCitation id="C7F92A542710321BB083FF29FDCBFF29" box="[466,544,151,177]" captionStart="FIGURE 7" captionStartId="15.[151,250,1323,1347]" captionTargetBox="[397,1191,669,1299]" captionTargetId="figure-260@15.[397,1191,669,1299]" captionTargetPageId="15" captionText="FIGURE 7. Differences in dorsal body band coloration in preservative between Lycodon neomaculatus sp. nov. (A, B) and L. subcinctus (C, D). A—FMNH 180149 from Nakhon Ratchasima, Thailand; B—CAS 229729 from Tanintharyi, Myanmar; C—FMNH 229855 from Selangor, Malaysia; D—FMNH 183772 from Selangor, Malaysia. Images not to scale. Photos by J.L. Lee." figureDoi="http://doi.org/10.5281/zenodo.13916235" httpUri="https://zenodo.org/record/13916235/files/figure.png" pageId="18" pageNumber="505">Figs. 7</figureCitation>
|
||
,
|
||
<figureCitation id="C7F92A542710321BB37AFF29FDD0FF29" box="[555,571,151,177]" captionStart="FIGURE 9" captionStartId="18.[151,250,1435,1459]" captionTargetBox="[151,1436,273,1412]" captionTargetId="figure-73@18.[151,1436,273,1412]" captionTargetPageId="18" captionText="FIGURE 9. Lycodon subcinctus in life (not collected, sex not determined): A—Kathu, Phuket, Thailand (adult); B—Hala Bala WS, Narathiwat, Thailand (subadult); C—Khao Phanom Bencha NP, Krabi, Thailand (subadult); D—Khao Sok NP, Surat Thani, Thailand (adult); E—Baling, Kedah, Malaysia (adult); F—Fraser’s Hill, Pahang, Malaysia (subadult); G—West Malaysia (adult); H—Upper Seletar Reservoir, Singapore (adult); I—Padang, West Sumatra, Indonesia (adult); J—Bekasi, Java, Indonesia (adult); K—Jembrana, Bali, Indonesia (adult); L—Alor, East Nusa Tenggara, Indonesia (subadult). Photos by: A. Tomaszek (A, K); R. Grassby-Lewis (B); R. Jaihan (C, D), P. Pawangkhanant (E, I); G. Vogel (G), R. Kittrell (F); R. Tann (H); A.R. Mudzakir (J); C. Trainor (L)." figureDoi="http://doi.org/10.5281/zenodo.13916239" httpUri="https://zenodo.org/record/13916239/files/figure.png" pageId="18" pageNumber="505">9</figureCitation>
|
||
). In addition, some adult
|
||
<taxonomicName id="98C24D522710321BB204FF29FC03FF29" box="[853,1000,151,177]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="505" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC32710321BB204FF29FC03FF29" box="[853,1000,151,177]" italics="true" pageId="18" pageNumber="505">L. subcinctus</emphasis>
|
||
</taxonomicName>
|
||
(and
|
||
<taxonomicName id="98C24D522710321BB577FF29FB6FFF29" box="[1062,1156,151,177]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="505" phylum="Chordata" rank="species" species="sealei">
|
||
<emphasis id="6DB6EAC32710321BB577FF29FB6FFF29" box="[1062,1156,151,177]" italics="true" pageId="18" pageNumber="505">L. sealei</emphasis>
|
||
</taxonomicName>
|
||
) specimens may become completely melanistic, whereas all adult
|
||
<taxonomicName id="98C24D522710321BB30FFF05FCFCFF4D" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[606,791,187,213]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="505" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC32710321BB30FFF05FCFCFF4D" box="[606,791,187,213]" italics="true" pageId="18" pageNumber="505">L. neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6DB6EAC32710321BB24CFF05FC9DFF4D" bold="true" box="[797,886,187,213]" pageId="18" pageNumber="505">
|
||
<taxonomicNameLabel id="768557B82710321BB24CFF05FC9DFF4D" box="[797,886,187,213]" pageId="18" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
we have examined retain light pigment along the dorsal bands and are never fully melanistic.
|
||
</paragraph>
|
||
<caption id="0BBD66592710321BB1C6FA25FE1FF913" ID-DOI="http://doi.org/10.5281/zenodo.13916239" ID-Zenodo-Dep="13916239" httpUri="https://zenodo.org/record/13916239/files/figure.png" pageId="18" pageNumber="505" startId="18.[151,250,1435,1459]" targetBox="[151,1436,273,1412]" targetPageId="18" targetType="figure">
|
||
<paragraph id="5F7D36D12710321BB1C6FA25FE1FF913" blockId="18.[151,1437,1435,1675]" pageId="18" pageNumber="505">
|
||
<emphasis id="6DB6EAC32710321BB1C6FA25FEFBFA2B" bold="true" box="[151,272,1435,1459]" pageId="18" pageNumber="505">FIGURE 9</emphasis>
|
||
.
|
||
<taxonomicName id="98C24D522710321BB071FA25FE00FA2B" box="[288,491,1435,1459]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="505" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC32710321BB071FA25FE00FA2B" box="[288,491,1435,1459]" italics="true" pageId="18" pageNumber="505">Lycodon subcinctus</emphasis>
|
||
</taxonomicName>
|
||
in life (not collected, sex not determined): A—Kathu, Phuket, Thailand (adult); B—Hala Bala WS, Narathiwat, Thailand (subadult); C—Khao Phanom Bencha NP, Krabi, Thailand (subadult); D—Khao Sok NP, Surat Thani, Thailand (adult); E—Baling, Kedah, Malaysia (adult); F—Fraser’s Hill, Pahang, Malaysia (subadult); G—West Malaysia (adult); H—Upper Seletar Reservoir, Singapore (adult); I—Padang, West Sumatra, Indonesia (adult); J—Bekasi, Java, Indonesia (adult); K—Jembrana, Bali, Indonesia (adult); L—Alor, East Nusa Tenggara, Indonesia (subadult). Photos by: A. Tomaszek (A, K); R. Grassby-Lewis (B); R. Jaihan (C, D), P. Pawangkhanant (E, I); G. Vogel (G), R. Kittrell (F); R. Tann (H); A.R. Mudzakir (J); C. Trainor (L).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5F7D36D12710321BB196F963FDAAF86C" blockId="18.[151,1437,1757,2036]" pageId="18" pageNumber="505">
|
||
<taxonomicName id="98C24D522710321BB196F963FE20F96F" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[199,459,1757,1783]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="505" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC32710321BB196F963FE20F96F" box="[199,459,1757,1783]" italics="true" pageId="18" pageNumber="505">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82710321BB082F960FDDBF960" box="[467,560,1758,1784]" pageId="18" pageNumber="505" rank="species">
|
||
<emphasis id="6DB6EAC32710321BB082F960FE06F960" bold="true" box="[467,493,1758,1784]" pageId="18" pageNumber="505">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32710321BB0ACF960FDC2F960" bold="true" box="[509,553,1758,1784]" pageId="18" pageNumber="505">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="98C24D522710321BB398F960FCC2F96F" box="[713,809,1757,1784]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="505" phylum="Chordata" rank="species" species="sealei">
|
||
<emphasis id="6DB6EAC32710321BB398F960FD32F96F" box="[713,729,1758,1783]" italics="true" pageId="18" pageNumber="505">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32710321BB3B9F963FCC2F96F" box="[744,809,1757,1783]" italics="true" pageId="18" pageNumber="505">sealei</emphasis>
|
||
</taxonomicName>
|
||
by having a higher relative tail length (TaL/TL
|
||
<geoCoordinate id="3AF650162710321BB41EF960FA65F960" box="[1359,1422,1758,1784]" degrees="0.178" direction="north" orientation="latitude" pageId="18" pageNumber="505" precision="55" value="0.178">0.178</geoCoordinate>
|
||
<geoCoordinate id="3AF650162710321BB4DFF960FF3DF884" degrees="0.221" direction="west" orientation="longitude" pageId="18" pageNumber="505" precision="55" value="-0.221">– 0.221</geoCoordinate>
|
||
[mean = 0.201 ± 0.012] vs.
|
||
<geoCoordinate id="3AF650162710321BB342F8BFFDB9F884" box="[531,594,1793,1820]" degrees="0.154" direction="north" orientation="latitude" pageId="18" pageNumber="505" precision="55" value="0.154">0.154</geoCoordinate>
|
||
<geoCoordinate id="3AF650162710321BB303F8BFFD74F884" box="[594,671,1793,1820]" degrees="0.184" direction="west" orientation="longitude" pageId="18" pageNumber="505" precision="55" value="-0.184">–0.184</geoCoordinate>
|
||
[mean = 0.168 ± 0.009]), a higher number of subcaudals (SC 68–84 [mean = 75.07 ± 4.65] vs. 59–69 [mean = 63.58 ± 5.53]), a higher subcaudal ratio (SCR 25.09–30.21 [mean = 27.48 ± 1.26] vs. 22.61–25.27 [mean = 23.70 ± 0.08]), relative size of the largest supralabial scale (sixth supralabial largest vs. seventh supralabial largest), and a higher number of dorsal body bands in juvenile specimens (BB+TB 19–27 vs. 8–12; see
|
||
<figureCitation id="C7F92A542710321BB028F82FFE20F834" box="[377,459,1937,1964]" captionStart="FIGURE 10" captionStartId="19.[151,250,1057,1081]" captionTargetBox="[151,1436,181,1033]" captionTargetId="figure-18@19.[151,1436,181,1033]" captionTargetPageId="19" captionText="FIGURE 10. Lycodon sealei in life (not collected, sex not determined): A—Puerto Princesca, Palawan, Philippines (adult); B—Taytay, Palawan, Philippines (subadult); C—Balabac, Palawan, Philippines (adult); D—Ranau, Sabah, Malaysia (subadult); E—Marudi, Sarawak, Malaysia (subadult); F—Sandakan, Sabah, Malaysia (subadult); G—Kuching, Sarawak, Malaysia (adult); H—Petra Jaya, Kuching, Sarawak, Malaysia (adult); I—West Kotawaringin, Central Kalimantan, Indonesia (adult). Photos by: R. Grassby-Lewis (A); A. Sonlee (B); C. Supsup (C); R. Monday (D); C.C. Lee (E, H); S. Lee (F); L. Platania (G); H. Kurniawan (I)." figureDoi="http://doi.org/10.5281/zenodo.13916241" httpUri="https://zenodo.org/record/13916241/files/figure.png" pageId="18" pageNumber="505">Fig. 10</figureCitation>
|
||
). The same color pattern characteristics that differentiate
|
||
<taxonomicName id="98C24D522710321BB516F82CFAEBF833" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[1095,1280,1937,1964]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="505" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC32710321BB516F82CFBBCF833" box="[1095,1111,1938,1963]" italics="true" pageId="18" pageNumber="505">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32710321BB535F82FFAEBF833" box="[1124,1280,1937,1963]" italics="true" pageId="18" pageNumber="505">neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82710321BB457F82CFA8BF834" box="[1286,1376,1938,1964]" pageId="18" pageNumber="505" rank="species">
|
||
<emphasis id="6DB6EAC32710321BB457F82CFACBF834" bold="true" box="[1286,1312,1938,1964]" pageId="18" pageNumber="505">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32710321BB47CF82CFAB2F834" bold="true" box="[1325,1369,1938,1964]" pageId="18" pageNumber="505">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from
|
||
<taxonomicName id="98C24D522710321BB1C6F808FEC1F857" box="[151,298,1973,2000]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="505" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC32710321BB1C6F808FF4CF857" box="[151,167,1974,1999]" italics="true" pageId="18" pageNumber="505">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32710321BB1E5F80BFEC1F857" box="[180,298,1973,1999]" italics="true" pageId="18" pageNumber="505">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82710321BB060F808FE9AF848" box="[305,369,1974,2000]" pageId="18" pageNumber="505" sensu="stricto">
|
||
<emphasis id="6DB6EAC32710321BB060F808FED7F848" bold="true" box="[305,316,1974,2000]" pageId="18" pageNumber="505">s</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32710321BB018F808FE81F848" bold="true" box="[329,362,1974,2000]" pageId="18" pageNumber="505">str</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
also apply to
|
||
<taxonomicName id="98C24D522710321BB35AF808FD82F857" box="[523,617,1973,2000]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="505" phylum="Chordata" rank="species" species="sealei">
|
||
<emphasis id="6DB6EAC32710321BB35AF808FDF0F857" box="[523,539,1974,1999]" italics="true" pageId="18" pageNumber="505">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32710321BB379F80BFD82F857" box="[552,617,1973,1999]" italics="true" pageId="18" pageNumber="505">sealei</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, the geographic ranges of both species are separated by that of
|
||
<taxonomicName id="98C24D522710321BB1C6F864FEC0F86B" box="[151,299,2009,2036]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="505" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC32710321BB1C6F864FF4CF86B" box="[151,167,2010,2035]" italics="true" pageId="18" pageNumber="505">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32710321BB1E4F867FEC0F86B" box="[181,299,2009,2035]" italics="true" pageId="18" pageNumber="505">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82710321BB063F864FE98F86C" box="[306,371,2010,2036]" pageId="18" pageNumber="505" sensu="stricto">
|
||
<emphasis id="6DB6EAC32710321BB063F864FED6F86C" bold="true" box="[306,317,2010,2036]" pageId="18" pageNumber="505">s</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32710321BB01AF864FE87F86C" bold="true" box="[331,364,2010,2036]" pageId="18" pageNumber="505">str</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(see
|
||
<figureCitation id="C7F92A542710321BB0FFF867FDE2F86C" box="[430,521,2009,2036]" captionStart="FIGURE 11" captionStartId="20.[151,250,1043,1067]" captionTargetBox="[205,1382,185,1016]" captionTargetId="figure-29@20.[201,1386,181,1019]" captionTargetPageId="20" captionText="FIGURE 11. Distribution ranges of Lycodon sealei, L. subcinctus, and L. cf. subcinctus (Nicobar Island). Notes: numbers indicate different localities where the species have been recorded (see Appendix Table S5 for the details of localities)." figureDoi="http://doi.org/10.5281/zenodo.13916243" httpUri="https://zenodo.org/record/13916243/files/figure.png" pageId="18" pageNumber="505">Figs. 11</figureCitation>
|
||
,
|
||
<figureCitation id="C7F92A542710321BB344F864FDDFF86C" box="[533,564,2010,2036]" captionStart="FIGURE 12" captionStartId="20.[151,250,1980,2004]" captionTargetBox="[211,1376,1130,1954]" captionTargetId="figure-71@20.[207,1380,1127,1957]" captionTargetPageId="20" captionText="FIGURE 12. Distribution ranges of Lycodon neomaculatus sp. nov. Numbers indicate various localities where the species have been recorded (see Appendix Table S6 for the details of localities)." figureDoi="http://doi.org/10.5281/zenodo.13916245" httpUri="https://zenodo.org/record/13916245/files/figure.png" pageId="18" pageNumber="505">12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="0BBD66592711321AB1C6FB9FFF5CFB76" ID-DOI="http://doi.org/10.5281/zenodo.13916241" ID-Zenodo-Dep="13916241" httpUri="https://zenodo.org/record/13916241/files/figure.png" pageId="19" pageNumber="506" startId="19.[151,250,1057,1081]" targetBox="[151,1436,181,1033]" targetPageId="19" targetType="figure">
|
||
<paragraph id="5F7D36D12711321AB1C6FB9FFF5CFB76" blockId="19.[151,1437,1057,1262]" pageId="19" pageNumber="506">
|
||
<emphasis id="6DB6EAC32711321AB1C6FB9FFEF0FBA1" bold="true" box="[151,283,1057,1081]" pageId="19" pageNumber="506">FIGURE 10</emphasis>
|
||
.
|
||
<taxonomicName id="98C24D522711321AB07BFB9FFE2FFBA1" box="[298,452,1057,1081]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="506" phylum="Chordata" rank="species" species="sealei">
|
||
<emphasis id="6DB6EAC32711321AB07BFB9FFE2FFBA1" box="[298,452,1057,1081]" italics="true" pageId="19" pageNumber="506">Lycodon sealei</emphasis>
|
||
</taxonomicName>
|
||
in life (not collected, sex not determined): A—Puerto Princesca, Palawan, Philippines (adult); B—Taytay, Palawan, Philippines (subadult); C—Balabac, Palawan, Philippines (adult); D—Ranau, Sabah, Malaysia (subadult); E—Marudi, Sarawak, Malaysia (subadult); F—Sandakan, Sabah, Malaysia (subadult); G—Kuching, Sarawak, Malaysia (adult); H—Petra Jaya, Kuching, Sarawak, Malaysia (adult); I—West Kotawaringin, Central Kalimantan, Indonesia (adult). Photos by: R. Grassby-Lewis (A); A. Sonlee (B); C. Supsup (C); R. Monday (D); C.C. Lee (E, H); S. Lee (F); L. Platania (G); H. Kurniawan (I).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5F7D36D12711321AB196FAFEFBBCFA3A" blockId="19.[151,1436,1344,1946]" pageId="19" pageNumber="506">
|
||
In addition,
|
||
<taxonomicName id="98C24D522711321AB004FAFEFDFAFAC2" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[341,529,1344,1370]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="506" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC32711321AB004FAFEFE8EFAC1" box="[341,357,1344,1369]" italics="true" pageId="19" pageNumber="506">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32711321AB024FAFEFDFAFAC2" box="[373,529,1344,1370]" italics="true" pageId="19" pageNumber="506">neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82711321AB34BFAFEFD93FAC2" box="[538,632,1344,1370]" pageId="19" pageNumber="506" rank="species">
|
||
<emphasis id="6DB6EAC32711321AB34BFAFEFDDFFAC2" bold="true" box="[538,564,1344,1370]" pageId="19" pageNumber="506">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32711321AB314FAFEFD9AFAC2" bold="true" box="[581,625,1344,1370]" pageId="19" pageNumber="506">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="98C24D522711321AB245FAFEFC9AFAC2" box="[788,881,1344,1370]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="506" phylum="Chordata" rank="species" species="sidiki">
|
||
<emphasis id="6DB6EAC32711321AB245FAFEFCCFFAC1" box="[788,804,1344,1369]" italics="true" pageId="19" pageNumber="506">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32711321AB265FAFEFC9AFAC2" box="[820,881,1344,1370]" italics="true" pageId="19" pageNumber="506">sidiki</emphasis>
|
||
</taxonomicName>
|
||
by having the prefrontal enter the orbit (vs. not entering); temporals 1+2 (vs. 2+3); cloacal plate divided (vs. undivided); body in adults with pale and wide body bands (vs. dark and narrow crossbars) (see Wostl
|
||
<emphasis id="6DB6EAC32711321AB393FA36FD11FA3A" box="[706,762,1416,1442]" italics="true" pageId="19" pageNumber="506">et al.</emphasis>
|
||
2017 and
|
||
<bibRefCitation id="3B534B202711321AB23EFA36FBA0FA3A" author="Nguyen, T. V. & David, P. & Vogel, G." box="[879,1099,1416,1442]" pageId="19" pageNumber="506" pagination="563 - 581" refId="ref21621" refString="Nguyen, T. V., David, P. & Vogel, G. (2024) Expanded description & range extension of Lycodon sidiki Wostl, Hamidy, Kurniawan & Smith, 2017 (Squamata: Colubridae: Lycodon). Zootaxa, 5492 (4), 563 - 581. https: // doi. org / 10.11646 / zootaxa. 5492.4.5" type="journal article" year="2024">
|
||
Nguyen
|
||
<emphasis id="6DB6EAC32711321AB29EFA36FBECFA3A" box="[975,1031,1416,1442]" italics="true" pageId="19" pageNumber="506">et al.</emphasis>
|
||
2024
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="17D8655A2711321AB196FA15FB9AF802" pageId="19" pageNumber="506" type="distribution">
|
||
<paragraph id="5F7D36D12711321AB196FA15FB9AF802" blockId="19.[151,1436,1344,1946]" pageId="19" pageNumber="506">
|
||
<emphasis id="6DB6EAC32711321AB196FA15FE54FA5D" bold="true" box="[199,447,1451,1477]" pageId="19" pageNumber="506">
|
||
Distribution (
|
||
<figureCitation id="C7F92A542711321AB03AFA15FE54FA5D" box="[363,447,1451,1477]" captionStart="FIGURE 12" captionStartId="20.[151,250,1980,2004]" captionTargetBox="[211,1376,1130,1954]" captionTargetId="figure-71@20.[207,1380,1127,1957]" captionTargetPageId="20" captionText="FIGURE 12. Distribution ranges of Lycodon neomaculatus sp. nov. Numbers indicate various localities where the species have been recorded (see Appendix Table S6 for the details of localities)." figureDoi="http://doi.org/10.5281/zenodo.13916245" httpUri="https://zenodo.org/record/13916245/files/figure.png" pageId="19" pageNumber="506">Fig. 12</figureCitation>
|
||
</emphasis>
|
||
). Based on our definition of
|
||
<taxonomicName id="98C24D522711321AB240FA12FBFEFA5E" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[785,1045,1452,1478]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="506" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC32711321AB240FA12FBFEFA5E" box="[785,1045,1452,1478]" italics="true" pageId="19" pageNumber="506">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82711321AB54FFA12FB97FA5E" box="[1054,1148,1452,1478]" pageId="19" pageNumber="506" rank="species">
|
||
<emphasis id="6DB6EAC32711321AB54FFA12FBD3FA5E" bold="true" box="[1054,1080,1452,1478]" pageId="19" pageNumber="506">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32711321AB518FA12FB9EFA5E" bold="true" box="[1097,1141,1452,1478]" pageId="19" pageNumber="506">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
we establish its range as follows: China (southern), Myanmar (northern and peninsular), Vietnam (throughout), Laos (throughout), Cambodia (throughout), and Thailand (except peninsular). According to our data and an examination of records from the literature,
|
||
<taxonomicName id="98C24D522711321AB058F9A6FE29F9AA" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[265,450,1560,1586]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="506" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC32711321AB058F9A6FEF2F9A9" box="[265,281,1560,1585]" italics="true" pageId="19" pageNumber="506">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32711321AB077F9A6FE29F9AA" box="[294,450,1560,1586]" italics="true" pageId="19" pageNumber="506">neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82711321AB099F9A6FDC9F9AA" box="[456,546,1560,1586]" pageId="19" pageNumber="506" rank="species">
|
||
<emphasis id="6DB6EAC32711321AB099F9A6FE09F9AA" bold="true" box="[456,482,1560,1586]" pageId="19" pageNumber="506">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32711321AB0BEF9A6FDF0F9AA" bold="true" box="[495,539,1560,1586]" pageId="19" pageNumber="506">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
inhabits the following provinces/states:—
|
||
<emphasis id="6DB6EAC32711321AB2A8F9A9FBA9F9A9" bold="true" box="[1017,1090,1559,1585]" pageId="19" pageNumber="506">
|
||
<collectingCountry id="27D576412711321AB2A8F9A9FBA9F9A9" box="[1017,1090,1559,1585]" name="China" pageId="19" pageNumber="506">China</collectingCountry>
|
||
</emphasis>
|
||
, south-east: Hong Kong SAR.;
|
||
<collectingRegion id="9D06F8332711321AB1C6F982FF09F9CE" box="[151,226,1596,1622]" country="China" name="Fujian" pageId="19" pageNumber="506">Fujian</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB1BAF982FE9AF9CE" box="[235,369,1596,1622]" country="China" name="Guangdong" pageId="19" pageNumber="506">Guangdong</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB02AF982FE2DF9CE" box="[379,454,1596,1622]" country="China" name="Fujian" pageId="19" pageNumber="506">Fujian</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB09EF982FDD9F9CE" box="[463,562,1596,1622]" country="China" name="Guangxi" pageId="19" pageNumber="506">Guangxi</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB36AF982FD7FF9CE" box="[571,660,1596,1622]" country="China" name="Yunnan" pageId="19" pageNumber="506">Yunnan</collectingRegion>
|
||
, and
|
||
<collectingRegion id="9D06F8332711321AB39AF982FCF7F9CE" box="[715,796,1596,1622]" country="China" name="Hainan" pageId="19" pageNumber="506">Hainan</collectingRegion>
|
||
provinces;—
|
||
<emphasis id="6DB6EAC32711321AB2E3F982FBC1F9CE" bold="true" box="[946,1066,1596,1622]" pageId="19" pageNumber="506">
|
||
<collectingCountry id="27D576412711321AB2E3F982FBC1F9CE" box="[946,1066,1596,1622]" name="Myanmar" pageId="19" pageNumber="506">Myanmar</collectingCountry>
|
||
</emphasis>
|
||
: northern and peninsular:
|
||
<collectingRegion id="9D06F8332711321AB41AF982FF39F9E2" country="Myanmar" name="Kachin" pageId="19" pageNumber="506">Kachin State</collectingRegion>
|
||
;
|
||
<collectingRegion id="9D06F8332711321AB18DF9DEFEA6F9E2" box="[220,333,1632,1658]" country="Myanmar" name="Mandalay" pageId="19" pageNumber="506">Mandalay</collectingRegion>
|
||
, and
|
||
<collectingRegion id="9D06F8332711321AB0D4F9DEFDEDF9E2" box="[389,518,1632,1658]" country="Myanmar" name="Taninthayi" pageId="19" pageNumber="506">Tanintharyi</collectingRegion>
|
||
Regions);—
|
||
<emphasis id="6DB6EAC32711321AB3C5F9E1FD11F9E1" bold="true" box="[660,762,1631,1657]" pageId="19" pageNumber="506">
|
||
<collectingCountry id="27D576412711321AB3C5F9E1FD11F9E1" box="[660,762,1631,1657]" name="Vietnam" pageId="19" pageNumber="506">Vietnam</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectingRegion id="9D06F8332711321AB256F9DEFCBCF9E2" box="[775,855,1632,1658]" country="Vietnam" name="Ha Noi" pageId="19" pageNumber="506">Ha Noi</collectingRegion>
|
||
capital;
|
||
<collectingRegion id="9D06F8332711321AB2E5F9DEFBF3F9E2" box="[948,1048,1632,1658]" country="Vietnam" name="Da Nang" pageId="19" pageNumber="506">Da Nang</collectingRegion>
|
||
City;
|
||
<collectingRegion id="9D06F8332711321AB50BF9DEFB57F9E2" box="[1114,1212,1632,1658]" country="Vietnam" name="Bac Kan" pageId="19" pageNumber="506">Bac Kan</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB597F9DEFAB4F9E2" box="[1222,1375,1632,1658]" country="Vietnam" name="Tuyen Quang" pageId="19" pageNumber="506">Tuyen Quang</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB438F9DEFF19F906" country="Vietnam" name="Thai Nguyen" pageId="19" pageNumber="506">Thai Nguyen</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB1AEF93AFE92F906" box="[255,377,1668,1694]" country="Vietnam" name="Vinh Phuc" pageId="19" pageNumber="506">Vinh Phuc</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB0D7F93AFE19F906" box="[390,498,1668,1694]" country="Vietnam" name="Lai Chau" pageId="19" pageNumber="506">Lai Chau</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB0AFF93AFDB8F906" box="[510,595,1668,1694]" country="Vietnam" name="Son La" pageId="19" pageNumber="506">Son La</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB30EF93AFD0AF906" box="[607,737,1668,1694]" country="Vietnam" name="Thanh Hoa" pageId="19" pageNumber="506">Thanh Hoa</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB3BFF93AFC83F906" box="[750,872,1668,1694]" country="Vietnam" name="Ninh Binh" pageId="19" pageNumber="506">Ninh Binh</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB224F93AFC35F906" box="[885,990,1668,1694]" country="Vietnam" name="Nghe An" pageId="19" pageNumber="506">Nghe An</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB2BBF93AFB8AF906" box="[1002,1121,1668,1694]" country="Vietnam" name="Quang Tri" pageId="19" pageNumber="506">Quang Tri</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB53FF93AFAC0F906" box="[1134,1323,1668,1694]" country="Vietnam" name="Thua Thien-Hue" pageId="19" pageNumber="506">Thua Thien-Hue</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB469F93AFA73F906" box="[1336,1432,1668,1694]" country="Vietnam" name="Ha Tinh" pageId="19" pageNumber="506">Ha Tinh</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB1C6F916FECAF95A" box="[151,289,1704,1730]" country="Vietnam" name="Quang Nam" pageId="19" pageNumber="506">Quang Nam</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB07DF916FE73F95A" box="[300,408,1704,1730]" country="Vietnam" name="Kon Tum" pageId="19" pageNumber="506">Kon Tum</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB0F5F916FDC5F95A" box="[420,558,1704,1730]" country="Vietnam" name="Quang Ngai" pageId="19" pageNumber="506">Quang Ngai</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB369F916FD7BF95A" box="[568,656,1704,1730]" country="Vietnam" name="Gia Lai" pageId="19" pageNumber="506">Gia Lai</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB3CBF916FD15F95A" box="[666,766,1704,1730]" country="Vietnam" name="Dak Lak" pageId="19" pageNumber="506">Dak Lak</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB258F916FC83F95A" box="[777,872,1704,1730]" country="Vietnam" name="Phu Yen" pageId="19" pageNumber="506">Phu Yen</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB222F916FC1CF95A" box="[883,1015,1704,1730]" country="Vietnam" name="Khanh Hoa" pageId="19" pageNumber="506">Khanh Hoa</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB553F916FB66F95A" box="[1026,1165,1704,1730]" country="Vietnam" name="Ninh Thuan" pageId="19" pageNumber="506">Ninh Thuan</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB5C9F916FAFEF95A" box="[1176,1301,1704,1730]" country="Vietnam" name="Lam Dong" pageId="19" pageNumber="506">Lam Dong</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB471F916FA7EF95A" box="[1312,1429,1704,1730]" country="Vietnam" name="Dak Nong" pageId="19" pageNumber="506">Dak Nong</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB1C6F972FEF4F97E" box="[151,287,1740,1766]" country="Vietnam" name="Binh Phuoc" pageId="19" pageNumber="506">Binh Phuoc</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB07DF972FE75F97E" box="[300,414,1740,1766]" country="Vietnam" name="Dong Nai" pageId="19" pageNumber="506">Dong Nai</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB0FBF972FD98F97E" box="[426,627,1740,1766]" country="Vietnam" name="Ba Ria-Vung Tau" pageId="19" pageNumber="506">Ba Ria-Vung Tau</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB3D1F972FD06F97E" box="[640,749,1740,1766]" country="Vietnam" name="Tay Ninh" pageId="19" pageNumber="506">Tay Ninh</collectingRegion>
|
||
, and
|
||
<collectingRegion id="9D06F8332711321AB27BF972FC46F97E" box="[810,941,1740,1766]" country="Vietnam" name="Kien Giang" pageId="19" pageNumber="506">Kien Giang</collectingRegion>
|
||
provinces;—
|
||
<emphasis id="6DB6EAC32711321AB516F972FB6AF97E" bold="true" box="[1095,1153,1740,1766]" pageId="19" pageNumber="506">
|
||
<collectingCountry id="27D576412711321AB516F972FB6AF97E" box="[1095,1153,1740,1766]" name="Laos" pageId="19" pageNumber="506">Laos</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectingRegion id="9D06F8332711321AB5C0F972FB36F97E" box="[1169,1245,1740,1766]" country="Laos" name="Bokeo" pageId="19" pageNumber="506">Bokeo</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB5B8F972FA73F97E" box="[1257,1432,1740,1766]" country="Laos" name="Louangphabang" pageId="19" pageNumber="506">Luang Prabang</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB1C6F94EFEEEF892" box="[151,261,1776,1802]" country="Laos" name="Viangchan" pageId="19" pageNumber="506">Vientiane</collectingRegion>
|
||
, Bolikhamxay,
|
||
<collectingRegion id="9D06F8332711321AB0E4F94EFDBEF892" box="[437,597,1776,1802]" country="Laos" name="Khammouan" pageId="19" pageNumber="506">Khammouane</collectingRegion>
|
||
, and
|
||
<collectingRegion id="9D06F8332711321AB3C0F94EFCFFF892" box="[657,788,1776,1802]" country="Laos" name="Champasak" pageId="19" pageNumber="506">Champasak</collectingRegion>
|
||
provinces;—
|
||
<emphasis id="6DB6EAC32711321AB2FDF951FBC3F891" bold="true" box="[940,1064,1775,1801]" pageId="19" pageNumber="506">
|
||
<collectingCountry id="27D576412711321AB2FDF951FBC3F891" box="[940,1064,1775,1801]" name="Cambodia" pageId="19" pageNumber="506">Cambodia</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectingRegion id="9D06F8332711321AB569F94EFB5EF892" box="[1080,1205,1776,1802]" country="Cambodia" name="Siem Reap" pageId="19" pageNumber="506">Siem Reap</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB590F94EFAE1F892" box="[1217,1290,1776,1802]" country="Cambodia" name="Kratie" pageId="19" pageNumber="506">Kratie</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB447F94EFA72F892" box="[1302,1433,1776,1802]" country="Cambodia" name="Mondolkiri" pageId="19" pageNumber="506">Mondulkiri</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB1C6F8AAFEE6F8B6" box="[151,269,1812,1838]" country="Cambodia" name="Koh Kong" pageId="19" pageNumber="506">Koh Kong</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB046F8AAFE5EF8B6" box="[279,437,1812,1838]" country="Cambodia" name="Sihanoukville" pageId="19" pageNumber="506">Sihanoukville</collectingRegion>
|
||
, and
|
||
<collectingRegion id="9D06F8332711321AB0BDF8AAFDF0F8B6" box="[492,539,1812,1838]" country="Cambodia" name="Kep" pageId="19" pageNumber="506">Kep</collectingRegion>
|
||
provinces;—
|
||
<emphasis id="6DB6EAC32711321AB3E1F8ADFCF6F8B5" bold="true" box="[688,797,1811,1837]" pageId="19" pageNumber="506">
|
||
<collectingCountry id="27D576412711321AB3E1F8ADFCF6F8B5" box="[688,797,1811,1837]" name="Thailand" pageId="19" pageNumber="506">Thailand</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectingRegion id="9D06F8332711321AB27BF8AAFC41F8B6" box="[810,938,1812,1838]" country="Thailand" name="Chiang Rai" pageId="19" pageNumber="506">Chiang Rai</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB2E2F8AAFBD2F8B6" box="[947,1081,1812,1838]" country="Thailand" name="Chiang Mai" pageId="19" pageNumber="506">Chiang Mai</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB513F8AAFB6EF8B6" box="[1090,1157,1812,1838]" country="Thailand" name="Phrae" pageId="19" pageNumber="506">Phrae</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB5DFF8AAFAFEF8B6" box="[1166,1301,1812,1838]" country="Thailand" name="Phitsanulok" pageId="19" pageNumber="506">Phitsanulok</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB44EF8AAFABEF8B6" box="[1311,1365,1812,1838]" country="Thailand" name="Loei" pageId="19" pageNumber="506">Loei</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB40FF8AAFF3BF8CA" country="Thailand" name="Nong Khai" pageId="19" pageNumber="506">Nong Khai</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB18CF886FE8DF8CA" box="[221,358,1848,1874]" country="Thailand" name="Udon Thani" pageId="19" pageNumber="506">Udon Thani</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB022F886FE20F8CA" box="[371,459,1848,1874]" country="Thailand" name="Kalasin" pageId="19" pageNumber="506">Kalasin</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB086F886FD55F8CA" box="[471,702,1848,1874]" country="Thailand" name="Nakhon Ratchasima" pageId="19" pageNumber="506">Nakhon Ratchasima</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB39BF886FC93F8CA" box="[714,888,1848,1874]" country="Thailand" name="Nakhon Nayok" pageId="19" pageNumber="506">Nakhon Nayok</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB2D5F886FC0DF8CA" box="[900,998,1848,1874]" country="Thailand" name="Sa Kaeo" pageId="19" pageNumber="506">Sa Kaeo</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB2A3F886FB73F8CA" box="[1010,1176,1848,1874]" country="Thailand" name="Chachoengsao" pageId="19" pageNumber="506">Chachoengsao</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB5F4F886FB14F8CA" box="[1189,1279,1848,1874]" country="Thailand" name="Rayong" pageId="19" pageNumber="506">Rayong</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB45AF886FA73F8CA" box="[1291,1432,1848,1874]" country="Thailand" name="Chanthaburi" pageId="19" pageNumber="506">Chanthaburi</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB1C6F8E2FEC9F8EE" box="[151,290,1884,1910]" country="Thailand" name="Uthai Thani" pageId="19" pageNumber="506">Uthai Thani</collectingRegion>
|
||
,
|
||
<collectingRegion id="9D06F8332711321AB061F8E2FE52F8EE" box="[304,441,1884,1910]" country="Thailand" name="Phetchaburi" pageId="19" pageNumber="506">Phetchaburi</collectingRegion>
|
||
, and
|
||
<collectingRegion id="9D06F8332711321AB0A9F8E2FD01F8EE" box="[504,746,1884,1910]" country="Thailand" name="Prachuap Khiri Khan" pageId="19" pageNumber="506">Prachuap Khiri Khan</collectingRegion>
|
||
provinces (based on
|
||
<bibRefCitation id="3B534B202711321AB2B0F8E2FBB5F8EE" author="Pope, C. H." box="[993,1118,1883,1910]" pageId="19" pageNumber="506" refId="ref21916" refString="Pope, C. H. (1935) The reptiles of China. Turtles, crocodilians, snakes, lizards. Natural History of Central Asia. Vol. 10. American Museum of Natural History, New York, New York, 604 pp. https: // doi. org / 10.5962 / bhl. title. 12104" type="book" year="1935">Pope 1935</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B202711321AB53DF8E2FAC6F8EE" author="Zhao, E. M. & Huang, M. H. & Zong, Y. & Jiang, Y. M. & Huang, Q. Y. & Zhao, H. & Ma, J. F. & Zheng, J. & Huang, Z. J. & Wei, G. & Yang, D. Q. & Li, D. J." box="[1132,1325,1884,1910]" pageId="19" pageNumber="506" refId="ref24466" refString="Zhao, E. M., Huang, M. H., Zong, Y., Jiang, Y. M., Huang, Q. Y., Zhao, H., Ma, J. F., Zheng, J., Huang, Z. J., Wei, G., Yang, D. Q. & Li, D. J. (1998) Fauna Sinica Reptilia. Vol. 3. Squamata: Serpentes. Science Press, Beijing, 522 pp. [in Chinese]" type="book" year="1998">
|
||
Zhao
|
||
<emphasis id="6DB6EAC32711321AB5FFF8E2FB09F8EE" box="[1198,1250,1884,1910]" italics="true" pageId="19" pageNumber="506">et al</emphasis>
|
||
. 1998
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B202711321AB46AF8E2FF39F802" author="Le, D. T. & Luong, A. M. & Pham, C. T. & Pham, T. Q. & Nguyen, S. L. H. & Ziegler, T. & Nguyen, T. Q." pageId="19" pageNumber="506" pagination="201 - 219" refId="ref20602" refString="Le, D. T., Luong, A. M., Pham, C. T., Pham, T. Q., Nguyen, S. L. H., Ziegler, T. & Nguyen, T. Q. (2021) New records and an updated checklist of amphibians and snakes from Tuyen Quang Province, Vietnam. Bonn zoological Bulletin, 70 (1), 201 - 219. https: // doi. org / 10.20363 / BZB- 2021.70.1.201" type="journal article" year="2021">
|
||
Le
|
||
<emphasis id="6DB6EAC32711321AB433F8E2FA7EF8EE" box="[1378,1429,1884,1910]" italics="true" pageId="19" pageNumber="506">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B202711321AB18FF83EFE48F802" author="David, P. & Teynie, A. & Vogel, G." box="[222,419,1920,1946]" pageId="19" pageNumber="506" refId="ref19064" refString="David, P., Teynie, A. & Vogel, G. (2023) The Snakes of Laos. Edition Chimaira, Frankfurt am Main, 960 pp." type="book" year="2023">
|
||
David
|
||
<emphasis id="6DB6EAC32711321AB078F83EFEB0F802" box="[297,347,1920,1946]" italics="true" pageId="19" pageNumber="506">et al</emphasis>
|
||
. 2023
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B202711321AB0FEF83EFDBCF802" author="Liu, S. & Hou, M. & Cai, B. & Li, S. & Zhang, Z. & Yu, R. & Rao, D. & Zhang, L." box="[431,599,1920,1946]" pageId="19" pageNumber="506" pagination="307 - 316" refId="ref21074" refString="Liu, S., Hou, M., Cai, B., Li, S., Zhang, Z., Yu, R., Rao, D. & Zhang, L. (2023) Taxonomic status of Lycodon subcinctus sensu lato in China (Serpentes, Colubridae). Herpetozoa, 36, 307 - 316. https: // doi. org / 10.3897 / herpetozoa. 36. e 114206" type="journal article" year="2023">
|
||
Liu
|
||
<emphasis id="6DB6EAC32711321AB08CF83EFDE5F802" box="[477,526,1920,1946]" italics="true" pageId="19" pageNumber="506">et al</emphasis>
|
||
. 2023
|
||
</bibRefCitation>
|
||
; examined specimens;records from iNaturalist).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="0BBD66592716321DB1C6FBADFAABFBD7" ID-DOI="http://doi.org/10.5281/zenodo.13916243" ID-Zenodo-Dep="13916243" httpUri="https://zenodo.org/record/13916243/files/figure.png" pageId="20" pageNumber="507" startId="20.[151,250,1043,1067]" targetBox="[205,1382,185,1016]" targetPageId="20" targetType="figure">
|
||
<paragraph id="5F7D36D12716321DB1C6FBADFAABFBD7" blockId="20.[151,1436,1043,1103]" pageId="20" pageNumber="507">
|
||
<emphasis id="6DB6EAC32716321DB1C6FBADFEC9FBB3" bold="true" box="[151,290,1043,1067]" pageId="20" pageNumber="507">FIGURE 11.</emphasis>
|
||
Distribution ranges of
|
||
<taxonomicName id="98C24D522716321DB348FBADFD5FFBB3" box="[537,692,1043,1067]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="507" phylum="Chordata" rank="species" species="sealei">
|
||
<emphasis id="6DB6EAC32716321DB348FBADFD5FFBB3" box="[537,692,1043,1067]" italics="true" pageId="20" pageNumber="507">Lycodon sealei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="98C24D522716321DB395FBAAFCA7FBB3" box="[708,844,1043,1067]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="507" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC32716321DB395FBAAFD39FBB3" box="[708,722,1044,1067]" italics="true" pageId="20" pageNumber="507">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32716321DB3B0FBADFCA7FBB3" box="[737,844,1043,1067]" italics="true" pageId="20" pageNumber="507">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="98C24D522716321DB2DBFBAAFBDDFBB3" box="[906,1078,1043,1067]" class="Reptilia" family="Colubridae" genus="Lycodon" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="507" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC32716321DB2DBFBAAFC73FBB3" box="[906,920,1044,1067]" italics="true" pageId="20" pageNumber="507">L</emphasis>
|
||
. cf.
|
||
<emphasis id="6DB6EAC32716321DB29AFBADFBDDFBB3" box="[971,1078,1043,1067]" italics="true" pageId="20" pageNumber="507">subcinctus</emphasis>
|
||
</taxonomicName>
|
||
(Nicobar Island). Notes: numbers indicate different localities where the species have been recorded (see Appendix Table S5 for the details of localities).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0BBD66592716321DB1C6F802FCD0F861" ID-DOI="http://doi.org/10.5281/zenodo.13916245" ID-Zenodo-Dep="13916245" httpUri="https://zenodo.org/record/13916245/files/figure.png" pageId="20" pageNumber="507" startId="20.[151,250,1980,2004]" targetBox="[211,1376,1130,1954]" targetPageId="20" targetType="figure">
|
||
<paragraph id="5F7D36D12716321DB1C6F802FCD0F861" blockId="20.[151,1436,1980,2041]" pageId="20" pageNumber="507">
|
||
<emphasis id="6DB6EAC32716321DB1C6F802FEF4F84D" bold="true" box="[151,287,1980,2005]" pageId="20" pageNumber="507">FIGURE 12.</emphasis>
|
||
Distribution ranges of
|
||
<taxonomicName id="98C24D522716321DB356F803FD1BF84D" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[519,752,1981,2005]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="507" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC32716321DB356F803FD1BF84D" box="[519,752,1981,2005]" italics="true" pageId="20" pageNumber="507">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82716321DB3A4F803FCADF84C" box="[757,838,1981,2005]" pageId="20" pageNumber="507" rank="species">
|
||
<emphasis id="6DB6EAC32716321DB3A4F803FCE6F84D" bold="true" box="[757,781,1981,2005]" pageId="20" pageNumber="507">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32716321DB248F803FCABF84D" bold="true" box="[793,832,1981,2005]" pageId="20" pageNumber="507">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Numbers indicate various localities where the species have been recorded (see Appendix Table S6 for the details of localities).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="17D8655A2717321CB196FF29FD98FDF9" pageId="21" pageNumber="508" type="biology_ecology">
|
||
<paragraph id="5F7D36D12717321CB196FF29FD98FDF9" blockId="21.[151,1437,150,1113]" pageId="21" pageNumber="508">
|
||
<emphasis id="6DB6EAC32717321CB196FF29FE6DFF28" bold="true" box="[199,390,150,177]" pageId="21" pageNumber="508">Natural history</emphasis>
|
||
.
|
||
<taxonomicName id="98C24D522717321CB0CAFF29FD4FFF29" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[411,676,151,177]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC32717321CB0CAFF29FD4FFF29" box="[411,676,151,177]" italics="true" pageId="21" pageNumber="508">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82717321CB3E3FF29FCFFFF29" box="[690,788,151,177]" pageId="21" pageNumber="508" rank="species">
|
||
<emphasis id="6DB6EAC32717321CB3E3FF29FD27FF29" bold="true" box="[690,716,151,177]" pageId="21" pageNumber="508">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32717321CB3B0FF29FCE6FF29" bold="true" box="[737,781,151,177]" pageId="21" pageNumber="508">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
inhabits regions covered with primary and secondary rainforests, tropical evergreen forests and adjacent lowlands between
|
||
<quantity id="983A9B342717321CB2F9FF04FBC3FF4D" box="[936,1064,186,213]" metricMagnitude="2" metricUnit="m" metricValue="4.75" metricValueMax="8.0" metricValueMin="1.5" pageId="21" pageNumber="508" unit="m" value="475.0" valueMax="800.0" valueMin="150.0">150–800 m</quantity>
|
||
asl. (
|
||
<bibRefCitation id="3B534B202717321CB534FF05FB0AFF4D" author="Pope, C. H." box="[1125,1249,186,213]" pageId="21" pageNumber="508" refId="ref21916" refString="Pope, C. H. (1935) The reptiles of China. Turtles, crocodilians, snakes, lizards. Natural History of Central Asia. Vol. 10. American Museum of Natural History, New York, New York, 604 pp. https: // doi. org / 10.5962 / bhl. title. 12104" type="book" year="1935">Pope 1935</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B202717321CB5BFFF05FA7CFF4D" author="Liu, S. & Hou, M. & Cai, B. & Li, S. & Zhang, Z. & Yu, R. & Rao, D. & Zhang, L." box="[1262,1431,187,213]" pageId="21" pageNumber="508" pagination="307 - 316" refId="ref21074" refString="Liu, S., Hou, M., Cai, B., Li, S., Zhang, Z., Yu, R., Rao, D. & Zhang, L. (2023) Taxonomic status of Lycodon subcinctus sensu lato in China (Serpentes, Colubridae). Herpetozoa, 36, 307 - 316. https: // doi. org / 10.3897 / herpetozoa. 36. e 114206" type="journal article" year="2023">
|
||
Liu
|
||
<emphasis id="6DB6EAC32717321CB44DFF05FAA5FF4D" box="[1308,1358,187,213]" italics="true" pageId="21" pageNumber="508">et al</emphasis>
|
||
. 2023
|
||
</bibRefCitation>
|
||
; this study). This species may also be found in disturbed moist forests, thickets, tangled vegetation, and near human habitats such as shrublands, plantations, rice fields, abandoned structures, edges of cultivated areas, and villages (
|
||
<bibRefCitation id="3B534B202717321CB1CEFE99FEF6FED9" author="Zhao, E. M." box="[159,285,295,321]" pageId="21" pageNumber="508" refId="ref24423" refString="Zhao, E. M. (2006) Snakes of China. Vols. I - II. Anhui Science and Technology Publishing House, Hefei, 390 + 287 pp., 194 col. pls. [in Chinese]" type="book" year="2006">Zhao 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B202717321CB079FE99FE06FED9" author="David, P. & Teynie, A. & Vogel, G." box="[296,493,295,321]" pageId="21" pageNumber="508" refId="ref19064" refString="David, P., Teynie, A. & Vogel, G. (2023) The Snakes of Laos. Edition Chimaira, Frankfurt am Main, 960 pp." type="book" year="2023">
|
||
David
|
||
<emphasis id="6DB6EAC32717321CB022FE99FE4FFED9" box="[371,420,295,321]" italics="true" pageId="21" pageNumber="508">et al</emphasis>
|
||
. 2023
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B202717321CB0A8FE99FD4AFED9" author="Liu, S. & Hou, M. & Cai, B. & Li, S. & Zhang, Z. & Yu, R. & Rao, D. & Zhang, L." box="[505,673,295,321]" pageId="21" pageNumber="508" pagination="307 - 316" refId="ref21074" refString="Liu, S., Hou, M., Cai, B., Li, S., Zhang, Z., Yu, R., Rao, D. & Zhang, L. (2023) Taxonomic status of Lycodon subcinctus sensu lato in China (Serpentes, Colubridae). Herpetozoa, 36, 307 - 316. https: // doi. org / 10.3897 / herpetozoa. 36. e 114206" type="journal article" year="2023">
|
||
Liu
|
||
<emphasis id="6DB6EAC32717321CB376FE99FDB3FED9" box="[551,600,295,321]" italics="true" pageId="21" pageNumber="508">et al</emphasis>
|
||
. 2023
|
||
</bibRefCitation>
|
||
; this study). This species is chiefly nocturnal (active between 1930– 2330 hrs) but can also be active in the early morning. It is mainly terrestrial and is frequently found crawling on the forest floor or climbing in the forest understory. During the day, it retreats under stones, stumps, pieces of wood, and vegetation. This species can also be arboreal, sometimes ascending to the crowns of large trees, such as rubber and coffee trees (T.V. Nguyen, pers. obs.). There have been no reports of its reproduction, but like other
|
||
<taxonomicName id="98C24D522717321CB455FE09FA8FFE49" authorityName="Fitzinger" authorityYear="1826" box="[1284,1380,439,465]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="genus">
|
||
<emphasis id="6DB6EAC32717321CB455FE09FA8FFE49" box="[1284,1380,439,465]" italics="true" pageId="21" pageNumber="508">Lycodon</emphasis>
|
||
</taxonomicName>
|
||
, it is presumably oviparous.
|
||
<taxonomicName id="98C24D522717321CB0CEFE65FD49FE6D" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[415,674,475,501]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC32717321CB0CEFE65FD49FE6D" box="[415,674,475,501]" italics="true" pageId="21" pageNumber="508">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82717321CB3F8FE65FCEEFE6D" box="[681,773,475,501]" pageId="21" pageNumber="508" rank="species">
|
||
<emphasis id="6DB6EAC32717321CB3F8FE65FD28FE6D" bold="true" box="[681,707,475,501]" pageId="21" pageNumber="508">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32717321CB383FE65FD15FE6D" bold="true" box="[722,766,475,501]" pageId="21" pageNumber="508">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is known to eat amphibians and lizards specifically skinks like
|
||
<taxonomicName id="98C24D522717321CB196FE41FDFBFD81" authority="(Kuhl)" baseAuthorityName="Kuhl" baseAuthorityYear="1820" box="[199,528,511,537]" class="Reptilia" family="Scincidae" genus="Eutropis" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="multifasciata">
|
||
<emphasis id="6DB6EAC32717321CB196FE41FE54FD81" box="[199,447,511,537]" italics="true" pageId="21" pageNumber="508">Eutropis multifasciata</emphasis>
|
||
(Kuhl)
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="3B534B202717321CB31BFE41FD21FD81" author="Pope, C. H." box="[586,714,510,537]" pageId="21" pageNumber="508" refId="ref21916" refString="Pope, C. H. (1935) The reptiles of China. Turtles, crocodilians, snakes, lizards. Natural History of Central Asia. Vol. 10. American Museum of Natural History, New York, New York, 604 pp. https: // doi. org / 10.5962 / bhl. title. 12104" type="book" year="1935">Pope, 1935</bibRefCitation>
|
||
),
|
||
<taxonomicName id="98C24D522717321CB38CFE41FBDBFD81" authority="Gray" authorityName="Gray" authorityYear="1845" box="[733,1072,511,537]" class="Reptilia" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="chinensis">
|
||
<emphasis id="6DB6EAC32717321CB38CFE41FC19FD81" box="[733,1010,511,537]" italics="true" pageId="21" pageNumber="508">Ateuchosaurus chinensis</emphasis>
|
||
Gray
|
||
</taxonomicName>
|
||
(see https://www.inaturalist.org/ observations/14752734),
|
||
<taxonomicName id="98C24D522717321CB0E2FD9DFCDDFDA5" authority="Gunther" authorityName="Gunther" box="[435,822,547,573]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="microlepis">
|
||
<emphasis id="6DB6EAC32717321CB0E2FD9DFD3FFDA5" box="[435,724,547,573]" italics="true" pageId="21" pageNumber="508">Tropidophorus microlepis</emphasis>
|
||
Günther
|
||
</taxonomicName>
|
||
(https://www.inaturalist.org/observations/214567395), and possibly small snakes and small birds.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="17D8655A2717321CB196FDD4FA6CFBC1" pageId="21" pageNumber="508" type="conservation">
|
||
<paragraph id="5F7D36D12717321CB196FDD4FA6CFBC1" blockId="21.[151,1437,150,1113]" pageId="21" pageNumber="508">
|
||
<emphasis id="6DB6EAC32717321CB196FDD4FE5CFD1D" bold="true" box="[199,439,618,645]" pageId="21" pageNumber="508">Conservation status</emphasis>
|
||
. According to the IUCN Red List, both
|
||
<taxonomicName id="98C24D522717321CB2DFFDD5FB86FD1D" box="[910,1133,619,645]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="6DB6EAC32717321CB2DFFDD5FB86FD1D" box="[910,1133,619,645]" italics="true" pageId="21" pageNumber="508">Lycodon subcinctus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="98C24D522717321CB5F9FDD5FAE2FD1D" box="[1192,1289,619,645]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="sealei">
|
||
<emphasis id="6DB6EAC32717321CB5F9FDD5FB53FD1C" box="[1192,1208,619,644]" italics="true" pageId="21" pageNumber="508">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32717321CB599FDD5FAE2FD1D" box="[1224,1289,619,645]" italics="true" pageId="21" pageNumber="508">sealei</emphasis>
|
||
</taxonomicName>
|
||
are listed as Species of Least Concern (LC) due to their wide geographic distributions and ecological adaptability, presence in several large, protected areas, and slight adaptability to habitat degradation (
|
||
<bibRefCitation id="3B534B202717321CB57BFD0DFAC7FD55" author="Kim, P. & Schoppe, S." box="[1066,1324,691,717]" pageId="21" pageNumber="508" refId="ref20425" refString="Kim, P. & Schoppe, S. (2023) Lycodon sealei. The IUCN Red List of Threatened Species, 2023, e. T 186135286 A 186135381. https: // doi. org / 10.2305 / IUCN. UK. 20231. RLTS. T 186135286 A 186135381. en" type="journal volume" year="2023">Kim & Schoppe 2023</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3B534B202717321CB46AFD0DFF11FD69" author="Lilley, R. & Nguyen, T. Q. & Iskandar, D. & Grismer, L. & Luu, V. & Reilly, S. & Sumontha, M." pageId="21" pageNumber="508" refId="ref20990" refString="Lilley, R., Nguyen, T. Q., Iskandar, D., Grismer, L., Luu, V., Reilly, S. & Sumontha, M. (2023) Lycodon subcinctus. The IUCN Red List of Threatened Species, 2023, e. T 186136203 A 186136744. https: // doi. org / 10.2305 / IUCN. UK. 2023 - 1. RLTS. T 186136203 A 186136744. en" type="journal volume" year="2023">
|
||
Lilley
|
||
<emphasis id="6DB6EAC32717321CB4D9FD0DFF46FD69" italics="true" pageId="21" pageNumber="508">et al</emphasis>
|
||
. 2023
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="98C24D522717321CB05EFD69FDF8FD69" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[271,531,727,753]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC32717321CB05EFD69FDF8FD69" box="[271,531,727,753]" italics="true" pageId="21" pageNumber="508">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82717321CB34DFD69FD92FD69" box="[540,633,727,753]" pageId="21" pageNumber="508" rank="species">
|
||
<emphasis id="6DB6EAC32717321CB34DFD69FDDDFD69" bold="true" box="[540,566,727,753]" pageId="21" pageNumber="508">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32717321CB317FD69FD99FD69" bold="true" box="[582,626,727,753]" pageId="21" pageNumber="508">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is also distributed across in
|
||
<collectingCountry id="27D576412717321CB292FD69FBEDFD69" box="[963,1030,727,753]" name="China" pageId="21" pageNumber="508">China</collectingCountry>
|
||
and Indochina and inhabits several national parks and other protected areas but does not appear to be abundant wherever it occurs. No major threats are known for this species; however, some local populations could be threatened by deforestation and other forms of habitat loss. Many species of wolf snakes, including
|
||
<taxonomicName id="98C24D522717321CB3B3FCFDFBF0FCC5" authority="Blanford" authorityName="Blanford" box="[738,1051,835,861]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="davisonii">
|
||
<emphasis id="6DB6EAC32717321CB3B3FCFDFC44FCC5" box="[738,943,835,861]" italics="true" pageId="21" pageNumber="508">Lycodon davisonii</emphasis>
|
||
Blanford
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="98C24D522717321CB577FCFDFAEFFCC5" authority="Gunther" authorityName="Gunther" authorityYear="1864" box="[1062,1284,835,861]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="laoensis">
|
||
<emphasis id="6DB6EAC32717321CB577FCFDFBDDFCC4" box="[1062,1078,835,860]" italics="true" pageId="21" pageNumber="508">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32717321CB512FCFDFB74FCC5" box="[1091,1183,835,861]" italics="true" pageId="21" pageNumber="508">laoensis</emphasis>
|
||
Günther
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="98C24D522717321CB46CFCFDFED8FC19" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC32717321CB46CFCFDFED8FC19" italics="true" pageId="21" pageNumber="508">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82717321CB068FCD9FE78FC19" box="[313,403,871,897]" pageId="21" pageNumber="508" rank="species">
|
||
<emphasis id="6DB6EAC32717321CB068FCD9FEB8FC19" bold="true" box="[313,339,871,897]" pageId="21" pageNumber="508">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32717321CB031FCD9FE67FC19" bold="true" box="[352,396,871,897]" pageId="21" pageNumber="508">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
resemble venomous elapid snakes, such as kraits in the genus
|
||
<taxonomicName id="98C24D522717321CB515FCD9FB5AFC18" box="[1092,1201,871,896]" class="Reptilia" family="Elapidae" genus="Bungarus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="genus">
|
||
<emphasis id="6DB6EAC32717321CB515FCD9FB5AFC18" box="[1092,1201,871,896]" italics="true" pageId="21" pageNumber="508">Bungarus</emphasis>
|
||
</taxonomicName>
|
||
(including
|
||
<taxonomicName id="98C24D522717321CB47EFCD9FE6BFC3D" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Reptilia" family="Elapidae" genus="Bungarus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="candidus">
|
||
<emphasis id="6DB6EAC32717321CB47EFCD9FF17FC3D" italics="true" pageId="21" pageNumber="508">Bungarus candidus</emphasis>
|
||
(Linnaeus)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="98C24D522717321CB0DAFC35FD90FC3D" authority="Blyth" authorityName="Blyth" authorityYear="1861" box="[395,635,907,933]" class="Reptilia" family="Elapidae" genus="Bungarus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="multicinctus">
|
||
<emphasis id="6DB6EAC32717321CB0DAFC35FE77FC3C" box="[395,412,907,932]" italics="true" pageId="21" pageNumber="508">B</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32717321CB0FBFC35FDD8FC3D" box="[426,563,907,933]" italics="true" pageId="21" pageNumber="508">multicinctus</emphasis>
|
||
Blyth
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="98C24D522717321CB3D7FC35FA92FC3D" authority="Aksornneam, Rujirawan, Yodthong, Sung & Aowphol" authorityName="Aksornneam, Rujirawan, Yodthong, Sung & Aowphol" authorityYear="2024" box="[646,1401,907,933]" class="Reptilia" family="Elapidae" genus="Bungarus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="sagittatus">
|
||
<emphasis id="6DB6EAC32717321CB3D7FC35FD7CFC3C" box="[646,663,907,932]" italics="true" pageId="21" pageNumber="508">B</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32717321CB3F4FC35FCF9FC3C" box="[677,786,907,932]" italics="true" pageId="21" pageNumber="508">sagittatus</emphasis>
|
||
Aksornneam, Rujirawan, Yodthong, Sung & Aowphol
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="98C24D522717321CB4D5FC35FDBBFC51" authority="Chen, Shi, Vogel, Ding & Shi" authorityName="Chen, Shi, Vogel, Ding & Shi" authorityYear="2021" class="Reptilia" family="Elapidae" genus="Bungarus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="suzhenae">
|
||
<emphasis id="6DB6EAC32717321CB4D5FC35FF15FC51" italics="true" pageId="21" pageNumber="508">B. suzhenae</emphasis>
|
||
Chen, Shi, Vogel, Ding & Shi
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="98C24D522717321CB3DBFC11FC68FC51" authority="Pope" authorityName="Pope" authorityYear="1928" box="[650,899,943,969]" class="Reptilia" family="Elapidae" genus="Bungarus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="wanghaotingi">
|
||
<emphasis id="6DB6EAC32717321CB3DBFC11FD70FC50" box="[650,667,943,968]" italics="true" pageId="21" pageNumber="508">B</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32717321CB3F8FC11FCA8FC51" box="[681,835,943,969]" italics="true" pageId="21" pageNumber="508">wanghaotingi</emphasis>
|
||
Pope
|
||
</taxonomicName>
|
||
) and often live in areas where humans co-occur. Consequently, wolf snakes, although non-venomous, are occasionally the victims of misidentifications and are needlessly killed.
|
||
<taxonomicName id="98C24D522717321CB031FC49FDF2FB89" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[352,537,1015,1041]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC32717321CB031FC49FE9BFB88" box="[352,368,1015,1040]" italics="true" pageId="21" pageNumber="508">L</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32717321CB02CFC49FDF2FB89" box="[381,537,1015,1041]" italics="true" pageId="21" pageNumber="508">neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82717321CB34EFC49FD91FB89" box="[543,634,1015,1041]" pageId="21" pageNumber="508" rank="species">
|
||
<emphasis id="6DB6EAC32717321CB34EFC49FDD2FB89" bold="true" box="[543,569,1015,1041]" pageId="21" pageNumber="508">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32717321CB319FC49FD9FFB89" bold="true" box="[584,628,1015,1041]" pageId="21" pageNumber="508">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is not protected by national or international regulations and is not listed in CITES Appendices. Given this information, we suggest
|
||
<taxonomicName id="98C24D522717321CB268FBA5FBD6FBAD" authorityName="Nguyen & Lee & Pauwels & Kennedy-Gold & Poyarkov & David & Vogel" authorityYear="2024" box="[825,1085,1051,1077]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="508" phylum="Chordata" rank="species" species="neomaculatus" status="sp. nov.">
|
||
<emphasis id="6DB6EAC32717321CB268FBA5FBD6FBAD" box="[825,1085,1051,1077]" italics="true" pageId="21" pageNumber="508">Lycodon neomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="768557B82717321CB517FBA5FB48FBAD" box="[1094,1187,1051,1077]" pageId="21" pageNumber="508" rank="species">
|
||
<emphasis id="6DB6EAC32717321CB517FBA5FB8BFBAD" bold="true" box="[1094,1120,1051,1077]" pageId="21" pageNumber="508">sp</emphasis>
|
||
.
|
||
<emphasis id="6DB6EAC32717321CB521FBA5FB77FBAD" bold="true" box="[1136,1180,1051,1077]" pageId="21" pageNumber="508">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
should be considered Least Concern (LC) following the IUCN’s Red List categories (
|
||
<bibRefCitation id="3B534B202717321CB231FB81FA97FBC1" author="IUCN Standards & Petitions Committee" box="[864,1404,1087,1113]" pageId="21" pageNumber="508" refId="ref19856" refString="IUCN Standards and Petitions Committee (2024) Guidelines for using the IUCN red list categories and criteria. Ver. 16. Prepared by the standards and petitions committee. The IUCN Red List of Threatened Species. Available from: https: // www. iucnredlist. org / documents / RedList - Guidelines. pdf (accessed 1 January 2024)" type="journal volume" year="2024">IUCN Standards and Petitions Committee 2024</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |